190 lines
32 KiB
XML
190 lines
32 KiB
XML
<document id="AE85838BF35B3200717046DBA460B1CC" ID-CLB-Dataset="22144" ID-DOI="10.5852/ejt.2017.309" ID-GBIF-Dataset="4e7bcd4d-1cdb-45cf-b2c3-54da2466f171" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589468946001" checkinUser="carolina" docAuthor="Herbert, David G." docDate="2017" docId="039F87A56745FF9E4FC4FC2EFD72F812" docLanguage="en" docName="ejt-2017-309.pdf.imf" docOrigin="European Journal of Taxonomy 309" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sheldonia Herbert, 2017, s.l." docType="key" docVersion="6" lastPageNumber="3" masterDocId="FFA6FFDD6747FF9C4F79FFC5FFA2FFF9" masterDocTitle="A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="3" updateTime="1698839690494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B900D2A8302041E75F31F668CCA4BC82">A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)</mods:title>
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<mods:roleTerm id="1B990B9638AE9DE8984A2430F94538B1">Author</mods:roleTerm>
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<mods:namePart id="9382608BC0E59D9C876FCBB4874ECCD3">Herbert, David G.</mods:namePart>
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<mods:title id="E9CC64448A7474FD0DBEB4DAF947C0E5">European Journal of Taxonomy</mods:title>
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<mods:part id="A3D96F211E495DE9DC37CCBF08A06661">
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<mods:date id="4230435342411131ECFB5C901F1BD77C">2017</mods:date>
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<mods:number id="169B702B893FBC80F46FBCDF96B8FD80">2017-04-10</mods:number>
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<mods:number id="0FF815BCF92BD0FB1D9FD33F8EF97F11">309</mods:number>
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<treatment id="039F87A56745FF9E4FC4FC2EFD72F812" ID-DOI="http://doi.org/10.5281/zenodo.3846852" ID-GBIF-Taxon="164151032" ID-Zenodo-Dep="3846852" LSID="urn:lsid:plazi:treatment:039F87A56745FF9E4FC4FC2EFD72F812" httpUri="http://treatment.plazi.org/id/039F87A56745FF9E4FC4FC2EFD72F812" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="C32C65386745FF9E4FC4FC2EFDCFFBFD" box="[189,621,1002,1029]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B8936B36745FF9E4FC4FC2EFDCFFBFD" blockId="2.[189,621,1002,1029]" box="[189,621,1002,1029]" pageId="2" pageNumber="3">
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<heading id="D0C181DF6745FF9E4FC4FC2EFDCFFBFD" bold="true" box="[189,621,1002,1029]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="B942EAA16745FF9E4FC4FC2EFDE6FBFC" bold="true" box="[189,580,1002,1029]" pageId="2" pageNumber="3">
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Key to genera within
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<taxonomicName id="4C364D306745FF9E4EB3FC2FFDE6FBFC" ID-CoL="7PGPT" authorityName="Ancey" authorityYear="1887" box="[458,580,1002,1029]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" sensu="lato">
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<emphasis id="B942EAA16745FF9E4EB3FC2FFDE6FBFC" box="[458,580,1002,1029]" italics="true" pageId="2" pageNumber="3">Sheldonia</emphasis>
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</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A27157DA6745FF9E4D32FC2FFDCFFBFD" box="[587,621,1002,1029]" pageId="2" pageNumber="3" sensu="lato">
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<emphasis id="B942EAA16745FF9E4D32FC2FFDC5FBFC" bold="true" box="[587,615,1002,1029]" pageId="2" pageNumber="3">s.l</emphasis>
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.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386745FF9E4FC4FBDFFAD4F955" pageId="2" pageNumber="3" type="key">
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<key id="FFA761826745FF9E4FC4FBDFFAD4F955" pageId="2" pageNumber="3">
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<keyStep id="30C22E166745FF9E4FC4FBDFFAD5FB85" pageId="2" pageNumber="3">
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<paragraph id="8B8936B36745FF9E4FC4FBDFFAD4FBA1" blockId="2.[189,1399,1050,1148]" pageId="2" pageNumber="3">
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<keyLead id="30C795866745FF9E4FC4FBDFFAD4FBA1" pageId="2" pageNumber="3">
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1. Epiphallic caecum arising close to penis apex and adjacent to insertion of penial retractor muscle (
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<figureCitation id="130D2A366745FF9E4F8AFBFBFEF3FBA1" box="[243,337,1086,1112]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
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) ……………………………………………………………………………………………2
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</keyLead>
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</paragraph>
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<paragraph id="8B8936B36745FF9E4FB8FBA7FAD5FB85" blockId="2.[189,1399,1050,1148]" box="[193,1399,1122,1148]" pageId="2" pageNumber="3">
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<keyLead id="30C795866745FF9E4FB8FBA7FAD5FB85" box="[193,1399,1122,1148]" pageId="2" pageNumber="3">
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– Epiphallic caecum arising nearer to mid-point of epiphallus (
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<figureCitation id="130D2A366745FF9E4CCFFBA7FC55FB85" box="[950,1015,1122,1148]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
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B–C) …………………………3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C22E166745FF9E4FC4FB61FAD5FAB6" pageId="2" pageNumber="3">
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<paragraph id="8B8936B36745FF9E4FC4FB61FAD5FB1B" blockId="2.[189,1399,1188,1359]" pageId="2" pageNumber="3">
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<keyLead id="30C795866745FF9E4FC4FB61FAD5FB1B" pageId="2" pageNumber="3">
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2. Flagellum short and stout (
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<figureCitation id="130D2A366745FF9E4D4EFB61FD3AFB47" box="[567,664,1188,1214]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1D</figureCitation>
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); epiphallus short; vas deferens thick, long and convoluted; spermatophore tail plumose (
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<figureCitation id="130D2A366745FF9E4D4FFB0DFD29FB1B" box="[566,651,1224,1250]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1E</figureCitation>
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);protoconch spirally punctate ………
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<taxonomicName id="4C364D306745FF9E4B3BFB0DFAD5FB1B" ID-CoL="7PF62" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[1090,1399,1224,1251]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAA16745FF9E4B3BFB0DFB71FB1A" bold="true" box="[1090,1235,1224,1251]" italics="true" pageId="2" pageNumber="3">Ptilototheca</emphasis>
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Herbert, 2016
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B8936B36745FF9E4FB8FB28FAD5FAB6" blockId="2.[189,1399,1188,1359]" pageId="2" pageNumber="3">
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<keyLead id="30C795866745FF9E4FB8FB28FAD5FAB6" pageId="2" pageNumber="3">
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– Flagellum longer, divided into distinct f1 and f2 components (
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<figureCitation id="130D2A366745FF9E4CAEFB29FB95FAFE" box="[983,1079,1260,1287]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
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); vas deferens slender and relatively short, not convoluted; spermatophore tail with branched spines (
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<figureCitation id="130D2A366745FF9E4B31FAD5FB3CFAD2" box="[1096,1182,1296,1323]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1F</figureCitation>
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), but not plumose; protoconch smooth (rarely spirally lirate), not punctate. ………
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<taxonomicName id="4C364D306745FF9E4CB7FAF1FAD5FAB6" ID-CoL="7P6H6" authority="Godwin-Austen, 1912" authorityName="Godwin-Austen" authorityYear="1912" box="[974,1399,1332,1359]" class="Gastropoda" family="Urocyclidae" genus="Kerkophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAA16745FF9E4CB7FAF1FBCEFAB6" bold="true" box="[974,1132,1332,1359]" italics="true" pageId="2" pageNumber="3">Kerkophorus</emphasis>
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Godwin-Austen, 1912
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C22E166745FF9E4FC4FAB2FAD5FA20" pageId="2" pageNumber="3">
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<paragraph id="8B8936B36745FF9E4FC4FAB2FAD5FA4C" blockId="2.[189,1399,1399,1498]" pageId="2" pageNumber="3">
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<keyLead id="30C795866745FF9E4FC4FAB2FAD5FA4C" pageId="2" pageNumber="3">
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3. Epiphallic caecum small and globular (
|
||
<figureCitation id="130D2A366745FF9E4DC3FAB2FCB7FA68" box="[698,789,1399,1425]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1B</figureCitation>
|
||
); spermatophore tail bifid (
|
||
<figureCitation id="130D2A366745FF9E4B20FAB2FB16FA68" box="[1113,1204,1399,1425]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1G</figureCitation>
|
||
) ……………… …………………………………………………………………
|
||
<taxonomicName id="4C364D306745FF9E4CB1FA5EFAD5FA4C" ID-CoL="7P8WJ" authority="Godwin-Austen, 1912" authorityName="Godwin-Austen" authorityYear="1912" box="[968,1399,1435,1462]" class="Gastropoda" family="Urocyclidae" genus="Microkerkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAA16745FF9E4CB1FA5EFBC1FA4F" bold="true" box="[968,1123,1435,1462]" italics="true" pageId="2" pageNumber="3">Microkerkus</emphasis>
|
||
Godwin-Austen, 1912
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8936B36745FF9E4FB8FA05FAD5FA20" blockId="2.[189,1399,1399,1498]" box="[193,1399,1471,1498]" pageId="2" pageNumber="3">
|
||
<keyLead id="30C795866745FF9E4FB8FA05FAD5FA20" box="[193,1399,1471,1498]" pageId="2" pageNumber="3">
|
||
– Epiphallic caecum large, cylindrical and usually curved (
|
||
<figureCitation id="130D2A366745FF9E4CF0FA7AFC47FA20" box="[905,997,1471,1497]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1C</figureCitation>
|
||
); spermatophore tail not bifid …4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C22E166745FF9E4FC4F9C7FAD4F955" pageId="2" pageNumber="3">
|
||
<paragraph id="8B8936B36745FF9E4FC4F9C7FAD5F99D" blockId="2.[189,1399,1537,1708]" pageId="2" pageNumber="3">
|
||
<keyLead id="30C795866745FF9E4FC4F9C7FAD5F99D" pageId="2" pageNumber="3">
|
||
4. Protoconch spirally punctate or with numerous, close-set, incised spiral microstriae; nonspinose distal portion of spermatophore tail long, the subterminal region with short, stout projections on both sides (
|
||
<figureCitation id="130D2A366745FF9E4D54F98CFD2FF99D" box="[557,653,1609,1636]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1H</figureCitation>
|
||
, inset) ……………………………
|
||
<emphasis id="B942EAA16745FF9E4B52F98CFB74F99A" box="[1067,1238,1609,1636]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C364D306745FF9E4B52F98CFB07F99D" ID-CoL="7PGPT" authorityName="Ancey" authorityYear="1887" box="[1067,1189,1609,1636]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" sensu="stricto">
|
||
<emphasis id="B942EAA16745FF9E4B52F98CFB07F99D" bold="true" box="[1067,1189,1609,1636]" italics="true" pageId="2" pageNumber="3">Sheldonia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157DA6745FF9E4BC9F98FFB74F99A" box="[1200,1238,1610,1635]" pageId="2" pageNumber="3" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<bibRefCitation id="EFA74B426745FF9E4BA6F98FFAD5F99D" author="Ancey C. F." box="[1247,1399,1610,1636]" pageId="2" pageNumber="3" pagination="53 - 54" refId="ref23111" refString="Ancey C. F. 1887. Descriptions of new genera or subgenera of Helicidae. Conchologist's Exchange 1 (9 - 10): 53 - 54." type="journal article" year="1887">Ancey, 1887</bibRefCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8936B36745FF9E4FB8F9ABFAD4F955" blockId="2.[189,1399,1537,1708]" pageId="2" pageNumber="3">
|
||
<keyLead id="30C795866745FF9E4FB8F9ABFAD4F955" pageId="2" pageNumber="3">
|
||
– Protoconch essentially smooth; non-spinose distal portion of spermatophore tail short, lacking lateral projections (
|
||
<figureCitation id="130D2A366745FF9E4EAEF957FD8EF955" box="[471,556,1682,1708]" captionStart="Fig" captionStartId="3.[188,231,1647,1673]" captionTargetBox="[208,1396,574,1598]" captionTargetId="figure@3.[189,1399,552,1601]" captionTargetPageId="3" captionText="Fig. 1. Illustrations of characters mentioned in key to genera within Sheldonia s.l. A. Kerkophorus vittarubra sp. nov., epiphallus, caecum and flagellum (paratype, NMSA V4725/T4062). B. Microkerkus burnupi (Godwin-Austen, 1914), epiphallus and caecum (NMSA W3660). C. Sheldonia fingolandensis sp. nov., epiphallus, caecum and flagellum (paratype, NMSA W4169/T3385). D. Ptilototheca soutpansbergensis Herbert, 2016, epiphallus, caecum and flagellum (paratype, NMSA P0304/T4060). E. Ptilototheca soutpansbergensis, spermatophore (paratype, NMSA W2259/T4067). F. Kerkophorus poeppigii (Pfeiffer, 1846), spermatophore (NMSA W4911). G. Microkerkus arnotti (Benson, 1864), spermatophore (NMSA W6082). H. Sheldonia capsula (Benson, 1864), spermatophore, arrow indicates short, stout projections on both sides of the subterminal region of the tail, enlarged in inset (NMSA W4686). I. Selatodryas luteosoma gen. et sp. nov., spermatophore, arrow indicates non-spinose distal portion of tail (paratype, NMSA W9691/T3874)." figureDoi="http://doi.org/10.5281/zenodo.3827191" httpUri="https://zenodo.org/record/3827191/files/figure.png" pageId="2" pageNumber="3">Fig. 1I</figureCitation>
|
||
, arrow) …………………………………………
|
||
<taxonomicName id="4C364D306745FF9E4B14F954FB5BF955" ID-CoL="7PGHK" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[1133,1273,1681,1708]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="B942EAA16745FF9E4B14F954FB5BF955" bold="true" box="[1133,1273,1681,1708]" italics="true" pageId="2" pageNumber="3">Selatodryas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157DA6745FF9E4A7CF957FAD4F955" box="[1285,1398,1682,1708]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="C32C65386745FF9E4FC4F911FD72F812" pageId="2" pageNumber="3" type="discussion">
|
||
<paragraph id="8B8936B36745FF9E4FC4F911FD72F812" blockId="2.[189,1399,1748,2027]" pageId="2" pageNumber="3">
|
||
This key must be considered provisional and is provided as a guide to the identification of the existing supra-specific taxa within
|
||
<emphasis id="B942EAA16745FF9E4E81F93DFD3BF8EB" box="[504,665,1784,1810]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C364D306745FF9E4E81F93DFDCFF8EB" authorityName="Ancey" authorityYear="1887" box="[504,621,1784,1810]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" sensu="lato">Sheldonia</taxonomicName>
|
||
<taxonomicNameLabel id="A27157DA6745FF9E4D0EF93DFD3BF8EB" box="[631,665,1784,1810]" pageId="2" pageNumber="3" sensu="lato">s.l.</taxonomicNameLabel>
|
||
</emphasis>
|
||
However, there remain taxa within
|
||
<taxonomicName id="4C364D306745FF9E4B32F93DFB62F8EB" authorityName="Ancey" authorityYear="1887" box="[1099,1216,1784,1810]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" sensu="lato">
|
||
<emphasis id="B942EAA16745FF9E4B32F93DFB62F8EB" box="[1099,1216,1784,1810]" italics="true" pageId="2" pageNumber="3">Sheldonia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157DA6745FF9E4BB0F93DFB49F8EA" box="[1225,1259,1784,1811]" pageId="2" pageNumber="3" sensu="lato">
|
||
<emphasis id="B942EAA16745FF9E4BB0F93DFB46F8EB" box="[1225,1252,1784,1810]" italics="true" pageId="2" pageNumber="3">s.l</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
that do not fit well into any of these taxa (e.g.,
|
||
<taxonomicName id="4C364D306745FF9E4D24F8D9FBB3F8CF" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[605,1041,1820,1846]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="monsmaripi">
|
||
<emphasis id="B942EAA16745FF9E4D24F8D9FCC4F8CF" box="[605,870,1820,1846]" italics="true" pageId="2" pageNumber="3">Sheldonia monsmaripi</emphasis>
|
||
Herbert, 2016
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D306745FF9E4B33F8D8FF5FF8A2" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="wolkbergensis">
|
||
<emphasis id="B942EAA16745FF9E4B33F8D8FAACF8CF" box="[1098,1294,1820,1847]" italics="true" pageId="2" pageNumber="3">S. wolkbergensis</emphasis>
|
||
Herbert, 2016
|
||
</taxonomicName>
|
||
). Establishing the true relationships of these enigmatic species will require additional insights from molecular data. Furthermore, even within these genera, there is significant variation and future research may reveal that this encompasses additional lineages worthy of recognition as distinct genera, particularly within
|
||
<taxonomicName id="4C364D306745FF9E4ED8F868FD98F83E" authorityName="Godwin-Austen" authorityYear="1912" box="[417,570,1965,1991]" class="Gastropoda" family="Urocyclidae" genus="Kerkophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAA16745FF9E4ED8F868FD98F83E" box="[417,570,1965,1991]" italics="true" pageId="2" pageNumber="3">Kerkophorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D306745FF9E4D0EF868FCA8F83E" authorityName="Godwin-Austen" authorityYear="1912" box="[631,778,1965,1991]" class="Gastropoda" family="Urocyclidae" genus="Microkerkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAA16745FF9E4D0EF868FCA8F83E" box="[631,778,1965,1991]" italics="true" pageId="2" pageNumber="3">Microkerkus</emphasis>
|
||
</taxonomicName>
|
||
. Construction of a key should thus be viewed as an iterative process, for which this is a first step.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |