228 lines
35 KiB
XML
228 lines
35 KiB
XML
<document id="65495EB3D21C0DCF2BB98812E99A45D2" ID-DOI="10.11646/zootaxa.3955.1.6" ID-GBIF-Dataset="af56c0dd-3bd2-48e9-8062-4c49ddcc7d66" ID-ISSN="1175-5326" ID-Zenodo-Dep="241004" ID-ZooBank="A6799083-28D0-470A-A958-ACEBC7B1AC07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049276364" checkinUser="plazi" docAuthor="Kubecek, Vaclav, Bray, Timothy C. & Bocak, Ladislav" docDate="2015" docId="039F8791FFB8FF8AE4A8FAA6FD88FBFE" docLanguage="en" docName="zt03955p122.pdf" docOrigin="Zootaxa 3955 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ochinoeus Kubecek, Bray & Bocak, 2015, gen. nov." docType="treatment" docVersion="8" lastPageNumber="117" masterDocId="FFA6FFE9FFBBFF8EE43FFFD9FFCEFD69" masterDocTitle="Molecular phylogeny of Metanoeina net-winged beetles identifies Ochinoeus, a new genus from China and Laos (Coleoptera: Lycidae)" masterLastPageNumber="122" masterPageNumber="113" pageNumber="116" updateTime="1698578172603" updateUser="plazi">
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<mods:titleInfo id="722DEACB647F5273A0371D54519BF289">
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<mods:title id="992E53132B913DD56F69AAD55D32EE17">Molecular phylogeny of Metanoeina net-winged beetles identifies Ochinoeus, a new genus from China and Laos (Coleoptera: Lycidae)</mods:title>
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<mods:namePart id="FF3562CE4B37C50D9B3A88C2DAD94B11">Kubecek, Vaclav</mods:namePart>
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<mods:namePart id="6E81C5D497F681AE003BB3A32BF01701">Bray, Timothy C.</mods:namePart>
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<mods:namePart id="EF6374F93E7DB227652B27C8D3610494">Bocak, Ladislav</mods:namePart>
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<mods:title id="BD4D99492082081FA9B0EB10105DBC93">Zootaxa</mods:title>
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<mods:date id="07CF9CB866D9EE12B24AE614B7D0FAE5">2015</mods:date>
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<mods:number id="F4BAD4A94DBFC6EA28E3FDA9ACEC3173">3955</mods:number>
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<mods:identifier id="2CA60EE02A1E5D465C0177355B12B1C1" type="DOI">10.11646/zootaxa.3955.1.6</mods:identifier>
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<treatment id="039F8791FFB8FF8AE4A8FAA6FD88FBFE" ID-DOI="http://doi.org/10.5281/zenodo.6111807" ID-GBIF-Taxon="119611470" ID-Zenodo-Dep="6111807" LSID="urn:lsid:plazi:treatment:039F8791FFB8FF8AE4A8FAA6FD88FBFE" httpUri="http://treatment.plazi.org/id/039F8791FFB8FF8AE4A8FAA6FD88FBFE" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116">
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<subSubSection id="C32C650CFFB8FF8DE4A8FAA6FE40F8F3" box="[151,398,1407,1434]" pageId="3" pageNumber="116" type="nomenclature">
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<paragraph id="8B893687FFB8FF8DE4A8FAA6FE40F8F3" blockId="3.[151,398,1407,1434]" box="[151,398,1407,1434]" pageId="3" pageNumber="116">
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<heading id="D0C181EBFFB8FF8DE4A8FAA6FE40F8F3" bold="true" box="[151,398,1407,1434]" fontSize="11" level="1" pageId="3" pageNumber="116" reason="1">
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<emphasis id="B942EA95FFB8FF8DE4A8FAA6FE40F8F3" bold="true" box="[151,398,1407,1434]" pageId="3" pageNumber="116">
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<taxonomicName id="4C364D04FFB8FF8DE4A8FAA6FED7F8F0" box="[151,281,1407,1433]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B942EA95FFB8FF8DE4A8FAA6FED7F8F0" bold="true" box="[151,281,1407,1433]" italics="true" pageId="3" pageNumber="116">Ochinoeus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157EEFFB8FF8DE51FFA59FE40F8F3" box="[288,398,1408,1434]" pageId="3" pageNumber="116" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C650CFFB8FF8DE4A8FA1EFDB4F8B5" box="[151,634,1478,1500]" pageId="3" pageNumber="116" type="materials_examined">
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<paragraph id="8B893687FFB8FF8DE4A8FA1EFDB4F8B5" blockId="3.[151,634,1478,1500]" box="[151,634,1478,1500]" pageId="3" pageNumber="116">
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<typeStatus id="548D8825FFB8FF8DE4A8FA1EFF07F8B5" box="[151,201,1479,1500]" pageId="3" pageNumber="116">Type</typeStatus>
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species.
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<taxonomicName id="4C364D04FFB8FF8DE51AFA1FFDEDF8B5" box="[293,547,1478,1500]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="huaphanensis" status="sp. nov.">
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<emphasis id="B942EA95FFB8FF8DE51AFA1FFDEDF8B5" box="[293,547,1478,1500]" italics="true" pageId="3" pageNumber="116">Ochinoeus huaphanensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157EEFFB8FF8DE616FA1EFDB4F8B5" box="[553,634,1479,1500]" pageId="3" pageNumber="116" rank="species">
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<emphasis id="B942EA95FFB8FF8DE616FA1EFDB4F8B5" bold="true" box="[553,634,1479,1500]" pageId="3" pageNumber="116">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C650CFFB8FF8DE4A8F9D0FB2CFA0F" pageId="3" pageNumber="116" type="diagnosis">
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<paragraph id="8B893687FFB8FF8DE4A8F9D0FB2CFA0F" blockId="3.[151,1437,1545,2038]" pageId="3" pageNumber="116">
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<emphasis id="B942EA95FFB8FF8DE4A8F9D0FEDEFB4B" bold="true" box="[151,272,1545,1570]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
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<taxonomicName id="4C364D04FFB8FF8DE525F9D0FE5FFB48" box="[282,401,1545,1569]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B942EA95FFB8FF8DE525F9D0FE5FFB48" box="[282,401,1545,1569]" italics="true" pageId="3" pageNumber="116">Ochinoeus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C364D04FFB8FF8DE613F9D0FD03FB48" box="[556,717,1545,1569]" class="Insecta" family="Lycidae" genus="Matsudanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EA95FFB8FF8DE613F9D0FD03FB48" box="[556,717,1545,1569]" italics="true" pageId="3" pageNumber="116">Matsudanoeus</emphasis>
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</taxonomicName>
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in the structure of male and female genitalia:
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<taxonomicName id="4C364D04FFB8FF8DE0D8F9D0FA90FB48" box="[1255,1374,1545,1569]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B942EA95FFB8FF8DE0D8F9D0FA90FB48" box="[1255,1374,1545,1569]" italics="true" pageId="3" pageNumber="116">Ochinoeus</emphasis>
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</taxonomicName>
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have phallus widened apically, the sclerotized thorns are inconspicuous, short and similar in length (
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<figureCitation id="130D2A02FFB8FF8DE0FFF9F4FA89FB2F" box="[1216,1351,1581,1606]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="3" pageNumber="116">Figs. 25–32</figureCitation>
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) unlike
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<taxonomicName id="4C364D04FFB8FF8DE4A8F988FEF7FB00" box="[151,313,1617,1641]" class="Insecta" family="Lycidae" genus="Matsudanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EA95FFB8FF8DE4A8F988FEF7FB00" box="[151,313,1617,1641]" italics="true" pageId="3" pageNumber="116">Matsudanoeus</emphasis>
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</taxonomicName>
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which have asymmetrical, long and well-sclerotized thorns in the internal sac (
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<figureCitation id="130D2A02FFB8FF8DE13EF988FA5FFB03" box="[1281,1425,1617,1642]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="3" pageNumber="116">Figs. 23–24</figureCitation>
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). Ovipositors have short, closely attached valvifers in the only slightly divergent position, aligned with the axis of the coxites (
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<figureCitation id="130D2A02FFB8FF8DE4CAF940FE86FBDB" box="[245,328,1689,1714]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="3" pageNumber="116">Fig. 33</figureCitation>
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), differing from the almost perpendicular position of the coxites in
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<emphasis id="B942EA95FFB8FF8DE004F940FB2AFBD8" box="[1083,1252,1689,1713]" italics="true" pageId="3" pageNumber="116">
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<taxonomicName id="4C364D04FFB8FF8DE004F940FB2EFBD8" box="[1083,1248,1689,1713]" class="Insecta" family="Lycidae" genus="Matsudanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">Matsudanoeus</taxonomicName>
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.
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</emphasis>
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Both genera are similar in superficial appearance and differ only in the irregular elytral secondary costae, which are inconspicuous to missing in the middle part of the elytron in
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<taxonomicName id="4C364D04FFB8FF8DE6F6F938FCF1FB90" box="[713,831,1761,1785]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B942EA95FFB8FF8DE6F6F938FCF1FB90" box="[713,831,1761,1785]" italics="true" pageId="3" pageNumber="116">Ochinoeus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130D2A02FFB8FF8DE768F938FBD6FB93" box="[855,1048,1761,1786]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="3" pageNumber="116">Figs. 18, 20–22</figureCitation>
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) and regular, well-developed in
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<taxonomicName id="4C364D04FFB8FF8DE4A8F8DCFEF7FA74" box="[151,313,1797,1821]" class="Insecta" family="Lycidae" genus="Matsudanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EA95FFB8FF8DE4A8F8DCFEF7FA74" box="[151,313,1797,1821]" italics="true" pageId="3" pageNumber="116">Matsudanoeus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130D2A02FFB8FF8DE578F8DCFE54FA77" box="[327,410,1797,1822]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="3" pageNumber="116">Fig. 19</figureCitation>
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). All
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<taxonomicName id="4C364D04FFB8FF8DE5E8F8DFFD9EFA74" box="[471,592,1798,1821]" class="Insecta" family="Lycidae" genus="Metanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EA95FFB8FF8DE5E8F8DFFD9EFA74" box="[471,592,1798,1821]" italics="true" pageId="3" pageNumber="116">Metanoeus</emphasis>
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</taxonomicName>
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differ in structure of pronotal carinae (
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<figureCitation id="130D2A02FFB8FF8DE7C2F8DCFB82FA77" box="[1021,1100,1797,1822]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="3" pageNumber="116">Fig. 11</figureCitation>
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), presenting very small elytral cells and regular secondary and transverse costae.
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<taxonomicName id="4C364D04FFB8FF8DE6F2F8F0FCB8FA28" box="[717,886,1833,1857]" class="Insecta" family="Lycidae" genus="Xylometanoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EA95FFB8FF8DE6F2F8F0FCB8FA28" box="[717,886,1833,1857]" italics="true" pageId="3" pageNumber="116">Xylometanoeus</emphasis>
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</taxonomicName>
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have only primary costae present (
|
||
<bibRefCitation id="EFA74B76FFB8FF8DE13EF8F0FF38FA0F" author="Sklenarova" pageId="3" pageNumber="116" refString="Sklenarova, K., Kubecek, V. & Bocak, L. (2014) Subtribal classification of Metriorrhynchini (Coleoptera: Lycidae): an integrative approach using molecular phylogeny and morphology of adults and larvae. Arthropod Systematics and Phylogeny, 72, 37 - 54." type="journal article" year="2014">
|
||
Sklenarova
|
||
<emphasis id="B942EA95FFB8FF8DE1B7F8F3FF7AFA0C" italics="true" pageId="3" pageNumber="116">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). The superficially similar
|
||
<taxonomicName id="4C364D04FFB8FF8DE620F897FDB3FA0C" box="[543,637,1870,1893]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EA95FFB8FF8DE620F897FDB3FA0C" box="[543,637,1870,1893]" italics="true" pageId="3" pageNumber="116">Cautires</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130D2A02FFB8FF8DE6B3F894FD12FA0F" box="[652,732,1869,1894]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="3" pageNumber="116">Fig. 10</figureCitation>
|
||
) have lanceolate phallus and slender valvifers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C650CFFB8FF8AE4F8F8A8FDDFFB6E" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116" type="description">
|
||
<paragraph id="8B893687FFB8FF8DE4F8F8A8FD4FFA9F" blockId="3.[151,1437,1545,2038]" pageId="3" pageNumber="116">
|
||
<emphasis id="B942EA95FFB8FF8DE4F8F8A8FE97FAE3" bold="true" box="[199,345,1905,1930]" pageId="3" pageNumber="116">Description.</emphasis>
|
||
Adult. Body
|
||
<quantity id="4CCE9B62FFB8FF8DE63FF8ABFD49FAE0" box="[512,647,1906,1930]" metricMagnitude="-2" metricUnit="m" metricValue="1.045" metricValueMax="1.2" metricValueMin="0.89" pageId="3" pageNumber="116" unit="mm" value="10.45" valueMax="12.0" valueMin="8.9">8.9–12 mm</quantity>
|
||
, dorso-ventrally flattened, moderately slender, body length 3.9–4.6 times width at humeri, elytra slightly widened backward (
|
||
<figureCitation id="130D2A02FFB8FF8DE704F84CFC6FFAC7" box="[827,929,1941,1966]" captionStart="FIGURES 2 – 4. 2" captionStartId="4.[151,264,741,764]" captionTargetBox="[151,1431,193,717]" captionTargetId="figure@4.[151,1436,193,721]" captionTargetPageId="4" captionText="FIGURES 2 – 4. 2 — The distribution of Ochinoeus gen. nov. General appearance. 3 — Ochinoeus huaphanensis sp. nov., 4 — O. hainanensis sp. nov. Scale 2 mm." httpUri="https://zenodo.org/record/241006/files/figure.png" pageId="3" pageNumber="116">Fig. 3–4</figureCitation>
|
||
). Body dark brown to black, pronotum and elytra often brightly coloured, ochreous, brown to dark brown, scutellum brown to light brown. Head, antennae and legs brown, covered with short pubescence.
|
||
</paragraph>
|
||
<caption id="DF49660FFFBFFF8AE4A8FD3CFE22FE73" httpUri="https://zenodo.org/record/241006/files/figure.png" pageId="4" pageNumber="117" targetBox="[151,1431,193,717]" targetPageId="4">
|
||
<paragraph id="8B893687FFBFFF8AE4A8FD3CFE22FE73" blockId="4.[151,1436,741,794]" pageId="4" pageNumber="117">
|
||
<emphasis id="B942EA95FFBFFF8AE4A8FD3CFEF4FF92" bold="true" box="[151,314,741,764]" pageId="4" pageNumber="117">FIGURES 2–4.</emphasis>
|
||
2—The distribution of
|
||
<taxonomicName id="4C364D04FFBFFF8AE615FD3FFD58FF95" box="[554,662,742,764]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B942EA95FFBFFF8AE615FD3FFD58FF95" box="[554,662,742,764]" italics="true" pageId="4" pageNumber="117">Ochinoeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157EEFFBFFF8AE6A3FD3EFD34FF95" box="[668,762,743,764]" pageId="4" pageNumber="117" rank="genus">
|
||
<emphasis id="B942EA95FFBFFF8AE6A3FD3EFD34FF95" bold="true" box="[668,762,743,764]" pageId="4" pageNumber="117">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
General appearance. 3—
|
||
<taxonomicName id="4C364D04FFBFFF8AE7C3FD3FFB37FF95" box="[1020,1273,742,764]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="huaphanensis" status="sp. nov.">
|
||
<emphasis id="B942EA95FFBFFF8AE7C3FD3FFB37FF95" box="[1020,1273,742,764]" italics="true" pageId="4" pageNumber="117">Ochinoeus huaphanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157EEFFBFFF8AE13FFD3EFA9EFF95" box="[1280,1360,743,764]" pageId="4" pageNumber="117" rank="species">
|
||
<emphasis id="B942EA95FFBFFF8AE13FFD3EFA9EFF95" bold="true" box="[1280,1360,743,764]" pageId="4" pageNumber="117">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 4—
|
||
<taxonomicName id="4C364D04FFBFFF8AE1BCFD3EFEDEFE73" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
|
||
<emphasis id="B942EA95FFBFFF8AE1BCFD3EFEDEFE73" italics="true" pageId="4" pageNumber="117">O. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157EEFFBFFF8AE529FCDCFEA9FE73" box="[278,359,773,794]" pageId="4" pageNumber="117" rank="species">
|
||
<emphasis id="B942EA95FFBFFF8AE529FCDCFEA9FE73" bold="true" box="[278,359,773,794]" pageId="4" pageNumber="117">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Scale 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B893687FFBFFF8AE4F8FC9BFA95F95A" blockId="4.[151,1437,834,1687]" pageId="4" pageNumber="117">
|
||
Head small, slightly hypognathous, partly retracted in pronotum. Eyes large, hemispherically prominent, interocular distance 1.00–1.26 times maximum eye diameter in males and 1.22–1.23 times larger in females. Mandibles slender, strongly curved. Maxillary palpi 4-segmented, basal palpomere 1 short, triangular, palpomere 2 elongate, 1.67 times longer than palpomere 1, palpomere 3 very short, apical palpomere robust, almost parallelsided, obliquely cut at apex. Labial palpi 3-segmented, basal palpomere robust, palpomere 2 short, square, apical palpomere robust, widest apically. Antennae 11-segmented, finely pubescent, scapus pear-shaped, pedicel short, rectangular, partly hidden in scapus; antennomeres 3–10 flabellate in males, serrate in females (
|
||
<figureCitation id="130D2A02FFBFFF8AE0FAFBC3FA80F95A" box="[1221,1358,1050,1075]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="4" pageNumber="117">Figs. 14–17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B893687FFBFFF8AE4F8FBE6FDADF83A" blockId="4.[151,1437,834,1687]" pageId="4" pageNumber="117">
|
||
Pronotum trapezoidal, widely rounded frontally, with projected posterior angles (
|
||
<figureCitation id="130D2A02FFBFFF8AE066FBE7FB0AF93E" box="[1113,1220,1086,1111]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="4" pageNumber="117">Figs. 6–9</figureCitation>
|
||
) pronotum 1.3–1.5 times wider at posterior angles than long at midline. Pronotum with seven areolae, but postero-lateral carinae sometimes inconspicuous and only five clear areolae present in disc of pronotum as result; lateral margins of pronotum only slightly elevated. Scutellum well-developed, wide at base, emarginate apically, with two prominent processes (
|
||
<figureCitation id="130D2A02FFBFFF8AE52EFB17FEADF98E" box="[273,355,1230,1255]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="4" pageNumber="117">Fig. 12</figureCitation>
|
||
). Elytra slender, 3.7–4.0 times longer than wide at humeri in males, 3.5 times longer than wide at humeri in females, flat, slightly widened posteriorly; four primary costae developed in entire length, secondary costae much weaker, irregular to missing especially in middle part of elytra (
|
||
<figureCitation id="130D2A02FFBFFF8AE03AFACFFB73F846" box="[1029,1213,1302,1327]" captionStart="FIGURES 5 – 22" captionStartId="5.[151,264,1490,1513]" captionTargetBox="[153,1432,197,1468]" captionTargetId="figure@5.[151,1436,193,1469]" captionTargetPageId="5" captionText="FIGURES 5 – 22. Pronotum. 5 — Matsudanoeus yuasai (Nakane), 6 — Ochinoeus xunyanbaensis sp. nov., 7 — O. hainanensis sp. nov., 8 — O. habashanensis sp. nov., 9 — O. huaphanensis sp. nov., 10 — Cautires sp., 11 — Metanoeus sp. Scutellum. 12 — Ochinoeus huaphanensis sp. nov. Antenna. 13 — Matsudanoeus yuasai, 14 — Ochinoeus xunyanbaensis sp. nov., 15 — O. hainanensis, sp. nov., 16 — O. habashanensis sp. nov., and 17 — O. huaphanensis sp. nov. Middle part of elytra. 18 — O. huaphanensis sp. nov., 19 — Matsudanoeus yuasai, 20 — O. xunyanbaensis sp. nov., 21 — O. hainanensis sp. nov., 22 – O. habashanensis sp. nov. Figs. 18 – 22 a, b. Details located as shown. Scales: 5 – 12 0.5 mm, 13 – 22 1 mm, 18 – 22 a, b 0.5 mm." httpUri="https://zenodo.org/record/241007/files/figure.png" pageId="4" pageNumber="117">Figs. 18, 20–22</figureCitation>
|
||
); bottoms of elytral cells smooth, with very short microsetae.
|
||
</paragraph>
|
||
<paragraph id="8B893687FFBFFF8AE4F8FA87FDDFFB6E" blockId="4.[151,1437,834,1687]" pageId="4" pageNumber="117">
|
||
Male genitalia without parameres, phallobase circular, with extensive ventral membrane ventrally, phallus club-shaped, widest anteriorly, with widely rounded or slender apex (
|
||
<figureCitation id="130D2A02FFBFFF8AE7D4FA5BFBB2F8F2" box="[1003,1148,1410,1435]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="4" pageNumber="117">Figs. 25–32</figureCitation>
|
||
), internal sac with two inconspicuous thorns apically (
|
||
<figureCitation id="130D2A02FFBFFF8AE63EFA7FFD5EF8D6" box="[513,656,1446,1471]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="4" pageNumber="117">Figs. 25–32</figureCitation>
|
||
). Female genitalia short and wide, with slender coxites and short, closely attached valvifers, valvifers always shorter than coxites. Vagina wide, membranous, lateral glands with basally sclerotized ducts (
|
||
<figureCitation id="130D2A02FFBFFF8AE587FA37FDC7FB6E" box="[440,521,1518,1543]" captionStart="FIGURES 23 – 33" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1427,628,1844]" captionTargetId="figure@6.[151,1436,628,1849]" captionTargetPageId="6" captionText="FIGURES 23 – 33. Genitalia. Male genitalia (ventral and lateral views). 23 – 24 Matsudanoeus yuasai (Nakane), 25 – 26 Ochinoeus habashanensis sp. nov., 27 – 28 O. huaphanensis sp. nov., 29 – 30 O. xunyanbaensis sp. nov., 31 – 32 O. hainanensis sp. nov.. Female genitalia. 33 — O. huaphanensis sp. nov. Scale 0.5 mm." httpUri="https://zenodo.org/record/241008/files/figure.png" pageId="4" pageNumber="117">Fig. 33</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C650CFFBFFF8AE4F8F9CBFCB2FB27" pageId="4" pageNumber="117" type="distribution">
|
||
<paragraph id="8B893687FFBFFF8AE4F8F9CBFCB2FB27" blockId="4.[151,1437,834,1687]" pageId="4" pageNumber="117">
|
||
<emphasis id="B942EA95FFBFFF8AE4F8F9CBFEAEFB42" bold="true" box="[199,352,1554,1579]" pageId="4" pageNumber="117">Distribution.</emphasis>
|
||
Four known species were recorded from various regions of
|
||
<collectingCountry id="F3217617FFBFFF8AE01DF9CBFBABFB42" box="[1058,1125,1554,1579]" name="China" pageId="4" pageNumber="117">China</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3217617FFBFFF8AE09EF9CAFB19FB43" box="[1185,1239,1555,1578]" name="Laos" pageId="4" pageNumber="117">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="130D2A02FFBFFF8AE0D5F9CBFAFFFB42" box="[1258,1329,1554,1579]" captionStart="FIGURES 2 – 4. 2" captionStartId="4.[151,264,741,764]" captionTargetBox="[151,1431,193,717]" captionTargetId="figure@4.[151,1436,193,721]" captionTargetPageId="4" captionText="FIGURES 2 – 4. 2 — The distribution of Ochinoeus gen. nov. General appearance. 3 — Ochinoeus huaphanensis sp. nov., 4 — O. hainanensis sp. nov. Scale 2 mm." httpUri="https://zenodo.org/record/241006/files/figure.png" pageId="4" pageNumber="117">Fig. 2</figureCitation>
|
||
), namely Yunnan, Shanxi, Hainan and Hua Phan province in northern
|
||
<collectingCountry id="F3217617FFBFFF8AE701F9EEFCB6FB27" box="[830,888,1591,1614]" name="Laos" pageId="4" pageNumber="117">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C650CFFBFFF8AE4F8F983FD88FBFE" pageId="4" pageNumber="117" type="etymology">
|
||
<paragraph id="8B893687FFBFFF8AE4F8F983FD88FBFE" blockId="4.[151,1437,834,1687]" pageId="4" pageNumber="117">
|
||
<emphasis id="B942EA95FFBFFF8AE4F8F983FE80FB1A" bold="true" box="[199,334,1626,1651]" pageId="4" pageNumber="117">Etymology.</emphasis>
|
||
The generic name
|
||
<taxonomicName id="4C364D04FFBFFF8AE60FF983FD69FB1B" box="[560,679,1626,1650]" class="Insecta" family="Lycidae" genus="Ochinoeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B942EA95FFBFFF8AE60FF983FD69FB1B" box="[560,679,1626,1650]" italics="true" pageId="4" pageNumber="117">Ochinoeus</emphasis>
|
||
</taxonomicName>
|
||
is a patronym in honour of Teruo Ochi, Kobe,
|
||
<collectingCountry id="F3217617FFBFFF8AE0EBF982FAD6FB1B" box="[1236,1304,1627,1650]" name="Japan" pageId="4" pageNumber="117">Japan</collectingCountry>
|
||
. The genus name is the noun of masculine gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |