treatments-xml/data/03/B1/DB/03B1DB4C8B3D4E0B4FCDFAA5384EFC1D.xml

<document id="C2C7D514F649EEA7071D28D93CA5E76E" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B3D4E0B4FCDFAA5384EFC1D" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria conoidea Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, comb. nov." docType="treatment" docVersion="15" lastPageNumber="19" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="17" updateTime="1716321918122" updateUser="carolina">
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<mods:title id="15812CD9424AF27FC8CF88EDAB23C8FD">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:namePart id="095D37E4F68F556E731A0AA86A8174B4">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="D29CECDD5A6D2CFC0B1FC0B83FFE288E">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="6A4D383A1F2C0A21ABA405BBEFE9C81E" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="4E9412B13CD5A564A65664E3CAF9C2D3">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="257138479757E56018509935D15E0DA4">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="BBA9F25564E8AA2E17728578249AE79A">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="60E2A74EACE260333E24CE5F2E06C3B4">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:date id="582041DA5C4AB8BD3F9F3A76DECB7AF8">2019</mods:date>
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<treatment id="03B1DB4C8B3D4E0B4FCDFAA5384EFC1D" ID-GBIF-Taxon="157239579" LSID="urn:lsid:plazi:treatment:03B1DB4C8B3D4E0B4FCDFAA5384EFC1D" httpUri="http://treatment.plazi.org/id/03B1DB4C8B3D4E0B4FCDFAA5384EFC1D" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C30239D18B3D4E094FCDFAA53831FAAF" box="[504,1084,1369,1396]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BA76A5A8B3D4E094FCDFAA53831FAAF" blockId="16.[504,1084,1369,1432]" box="[504,1084,1369,1396]" pageId="16" pageNumber="17">
<heading id="D0EFDD368B3D4E094FCDFAA53831FAAF" box="[504,1084,1369,1396]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="4C1811D98B3D4E094FCDFAA53FBEFAAF" ID-CoL="7RTH8" authority="(Leschke, 1914)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[504,947,1369,1396]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="B96CB6488B3D4E094FCDFAA53EE0FAAF" bold="true" box="[504,749,1369,1396]" italics="true" pageId="16" pageNumber="17">Landouria conoidea</emphasis>
(
<bibRefCitation id="EF8917AB8B3D4E094CC9FAA63FA6FAAF" author="Leschke" box="[764,939,1370,1396]" firstAuthor="Leschke" pageId="16" pageNumber="17" pagination="205 - 284" refId="ref33668" refString="Leschke M. 1914. Zur Molluskenfauna von Java und Celebes. Mitteilungen aus dem naturhistorischen Museum in Hamburg 31: 205 - 284, 1 pl." type="journal article" year="1914">Leschke, 1914</bibRefCitation>
)
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<taxonomicNameLabel id="A25F0B338B3D4E094D8FFAA63831FAAF" box="[954,1084,1370,1396]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C30239D18B3D4E094CBFFA813FA7FA4C" box="[650,938,1405,1432]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BA76A5A8B3D4E094CBFFA813FA7FA4C" blockId="16.[504,1084,1369,1432]" box="[650,938,1405,1432]" pageId="16" pageNumber="17">
<figureCitation id="132376DF8B3D4E094CBFFA813ED9FA4C" box="[650,724,1405,1431]" captionStart="Figs 2–5" captionStartId="11.[189,241,1860,1886]" captionTargetBox="[231,1357,265,1834]" captionTargetId="figure@11.[231,1357,265,1834]" captionTargetPageId="11" captionText="Figs 2–5. Shells of species of Landouria from Java. 2. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype of Helix winteriana, ZMA 376193a). 3. L. winteriana, West Java, Cibodas (syntype of Plectotropis kraepelini Leschke, 1914, ZMH 98416). 4. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 5. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak (neotype of Helix rotatoria, MZB 19205). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152223/files/figure.png" pageId="16" pageNumber="17">Figs 4</figureCitation>
, 
<figureCitation id="132376DF8B3D4E094CD4FA813EFFFA4C" box="[737,754,1405,1431]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="16" pageNumber="17">6</figureCitation>
, 
<figureCitation id="132376DF8B3D4E094CCBFA813F13FA43" box="[766,798,1405,1432]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="16" pageNumber="17">53</figureCitation>
; 
<tableCitation id="C69A5FE18B3D4E094D1FFA813FA7FA4C" box="[810,938,1405,1431]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="16" pageNumber="17" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
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<subSubSection id="C30239D18B3D4E094E88FA3A3880F9F2" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BA76A5A8B3D4E094E88FA3A3FDAFA3A" blockId="16.[189,983,1478,1505]" box="[189,983,1478,1505]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB084D748B3D4E094E88FA3A3FDAFA3A" box="[189,983,1478,1505]" pageId="16" pageNumber="17">
<taxonomicName id="4C1811D98B3D4E094E88FA3A3EA2FA3B" ID-CoL="7SRF7" authority="Leschke, 1914: 212" authorityName="Leschke" authorityPageNumber="212" authorityYear="1914" box="[189,687,1478,1504]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3D4E094E88FA3A3DB2FA3B" box="[189,447,1478,1504]" italics="true" pageId="16" pageNumber="17">Plectotropis conoidea</emphasis>
<treatmentCitation id="0AB94C4B8B3D4E094FF0FA3A3EA2FA3B" author="Leschke M." box="[453,687,1478,1504]" page="212" pageId="16" pageNumber="17" year="1914">
<bibRefCitation id="EF8917AB8B3D4E094FF0FA3A3EA2FA3B" author="Leschke M." box="[453,687,1478,1504]" firstAuthor="Leschke" pageId="16" pageNumber="17" pagination="205 - 284" refId="ref33668" refString="Leschke M. 1914. Zur Molluskenfauna von Java und Celebes. Mitteilungen aus dem naturhistorischen Museum in Hamburg 31: 205 - 284, 1 pl." type="journal article" year="1914">Leschke, 1914: 212</bibRefCitation>
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</taxonomicName>
, figs 8–9 (“Buitenzorg”).
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</paragraph>
<paragraph id="8BA76A5A8B3D4E094E88F9F33880F9F2" blockId="16.[189,1165,1551,1578]" box="[189,1165,1551,1578]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB084D748B3D4E094E88F9F33880F9F2" box="[189,1165,1551,1578]" pageId="16" pageNumber="17">
<taxonomicName id="4C1811D98B3D4E094E88F9F33F3AF9F2" ID-CoL="3S6S5" authority="- van Benthem Jutting 1950: 461" authorityName="- van Benthem Jutting" authorityPageNumber="461" authorityYear="1950" box="[189,823,1551,1578]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B3D4E094E88F9F33DA4F9F2" box="[189,425,1551,1577]" italics="true" pageId="16" pageNumber="17">Landouria rotatoria</emphasis>
– 
<treatmentCitation id="0AB94C4B8B3D4E094FF3F9F33F3AF9F2" author="van Benthem Jutting W. S. S." box="[454,823,1551,1578]" page="461" pageId="16" pageNumber="17" year="1950">
<bibRefCitation id="EF8917AB8B3D4E094FF3F9F33F3AF9F2" author="van Benthem Jutting W. S. S." box="[454,823,1551,1578]" firstAuthor="van Benthem Jutting" pageId="16" pageNumber="17" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting 1950: 461</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(in part). (not 
<bibRefCitation id="EF8917AB8B3D4E094DD6F9F33888F9F2" author="Pfeiffer" box="[995,1157,1551,1577]" firstAuthor="Pfeiffer" pageId="16" pageNumber="17" refId="ref34385" refString="Pfeiffer L. 1842. Symbolae ad Historiam Heliceorum. Sectio alterna. T. Fischer, Cassel [Kassel, Germany]. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
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</paragraph>
</subSubSection>
<subSubSection id="C30239D18B3D4E094E88F9A43D55F91F" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BA76A5A8B3D4E094E88F9A43D3BF9A9" blockId="16.[189,310,1624,1650]" box="[189,310,1624,1650]" pageId="16" pageNumber="17">
<heading id="D0EFDD368B3D4E094E88F9A43D3BF9A9" bold="true" box="[189,310,1624,1650]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B96CB6488B3D4E094E88F9A43D3BF9A9" bold="true" box="[189,310,1624,1650]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B3D4E094E88F97A3D55F91F" blockId="16.[189,1399,1670,1732]" pageId="16" pageNumber="17">
<taxonomicName id="4C1811D98B3D4E094E88F97A3DA1F97B" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[189,428,1670,1696]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3D4E094E88F97A3DA1F97B" box="[189,428,1670,1696]" italics="true" pageId="16" pageNumber="17">Landouria conoidea</emphasis>
</taxonomicName>
is characterized by a small, conical, sharply keeled shell with tubercles all over the shell surface.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B3D4E094E88F90E384FF810" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BA76A5A8B3D4E094E88F90E3DA7F8D7" blockId="16.[189,426,1778,1804]" box="[189,426,1778,1804]" pageId="16" pageNumber="17">
<heading id="D0EFDD368B3D4E094E88F90E3DA7F8D7" bold="true" box="[189,426,1778,1804]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B96CB6488B3D4E094E88F90E3DA7F8D7" bold="true" box="[189,426,1778,1804]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7060078B3D4E094EDFF8DD3822F884" ID-GBIF-Occurrence="2243137473" collectionCode="ZMH" country="Indonesia" latitude="-6.6" location="Bogor" longLatPrecision="1304" longitude="106.8" pageId="16" pageNumber="17" specimenCode="98144" specimenCount="8" stateProvince="West Java" typeStatus="syntype">
<paragraph id="8BA76A5A8B3D4E094EDFF8DD3D57F8E0" blockId="16.[189,1076,1825,1887]" box="[234,346,1825,1851]" pageId="16" pageNumber="17">
<typeStatus id="54A3D4F88B3D4E094EDFF8DD3D57F8E0" box="[234,346,1825,1851]" pageId="16" pageNumber="17" type="syntype">
<emphasis id="B96CB6488B3D4E094EDFF8DD3D57F8E0" bold="true" box="[234,346,1825,1851]" pageId="16" pageNumber="17">Syntypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B3D4E094E88F8B93839F884" blockId="16.[189,1076,1825,1887]" box="[189,1076,1859,1887]" pageId="16" pageNumber="17">
<collectingCountry id="F30F2ACA8B3D4E094E88F8B93D56F884" box="[189,347,1861,1887]" name="Indonesia" pageId="16" pageNumber="17">INDONESIA</collectingCountry>
• 
<specimenCount id="9D1EA1D38B3D4E094F41F8B93DC4F884" box="[372,457,1861,1887]" count="8" pageId="16" pageNumber="17" type="generic">8 spec.</specimenCount>
; 
<collectingRegion id="49DCA4B88B3D4E094FE1F8B93E40F884" box="[468,589,1861,1887]" country="Indonesia" name="Jawa Barat" pageId="16" pageNumber="17">West Java</collectingRegion>
, 
<location id="8EC73C818B3D4E094C6DF8B93EA8F884" LSID="urn:lsid:plazi:treatment:03B1DB4C8B3D4E0B4FCDFAA5384EFC1D:8EC73C818B3D4E094C6DF8B93EA8F884" box="[600,677,1861,1887]" country="Indonesia" latitude="-6.6" longLatPrecision="1304" longitude="106.8" name="Bogor" pageId="16" pageNumber="17" stateProvince="West Java">Bogor</location>
; 
<geoCoordinate id="EE2C0C9D8B3D4E094C84F8BF3F04F884" box="[689,777,1859,1887]" degrees="6" direction="south" minutes="36" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-6.6">6°36′ S</geoCoordinate>
, 
<geoCoordinate id="EE2C0C9D8B3D4E094D23F8BF3F83F884" box="[790,910,1859,1887]" degrees="106" direction="east" minutes="48" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="106.8">106°48′ E</geoCoordinate>
; 
<collectionCode id="ED09F29F8B3D4E094DAFF8B93FD6F884" box="[922,987,1861,1887]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="16" pageNumber="17" type="Museum">ZMH</collectionCode>
<specimenCode id="DBBEC2218B3D4E094DD7F8B93822F884" box="[994,1071,1861,1887]" pageId="16" pageNumber="17">98144</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BA76A5A8B3D4E094EDFF8713DABF87C" blockId="16.[189,1090,1933,1995]" box="[234,422,1933,1959]" pageId="16" pageNumber="17">
<emphasis id="B96CB6488B3D4E094EDFF8713DABF87C" bold="true" box="[234,422,1933,1959]" pageId="16" pageNumber="17">Other material</emphasis>
</paragraph>
<paragraph id="8BA76A5A8B3D4E094E88F84D384FF810" blockId="16.[189,1090,1933,1995]" box="[189,1090,1967,1995]" pageId="16" pageNumber="17">
<materialsCitation id="3B7060078B3D4E094E88F84D3833F810" ID-GBIF-Occurrence="2243137355" box="[189,1086,1967,1995]" collectionCode="ZMH" country="Indonesia" latitude="-6.6" location="Bogor" longLatPrecision="1304" longitude="106.8" pageId="16" pageNumber="17" specimenCode="98145" specimenCount="10" stateProvince="West Java">
<collectingCountry id="F30F2ACA8B3D4E094E88F84D3D56F810" box="[189,347,1969,1995]" name="Indonesia" pageId="16" pageNumber="17">INDONESIA</collectingCountry>
• 
<specimenCount id="9D1EA1D38B3D4E094F41F84D3DD5F810" box="[372,472,1969,1995]" count="10" pageId="16" pageNumber="17" type="generic">10 spec.</specimenCount>
; 
<collectingRegion id="49DCA4B88B3D4E094FD6F84D3E56F810" box="[483,603,1969,1995]" country="Indonesia" name="Jawa Barat" pageId="16" pageNumber="17">West Java</collectingRegion>
, 
<location id="8EC73C818B3D4E094C52F84D3EB9F810" LSID="urn:lsid:plazi:treatment:03B1DB4C8B3D4E0B4FCDFAA5384EFC1D:8EC73C818B3D4E094C52F84D3EB9F810" box="[615,692,1969,1995]" country="Indonesia" latitude="-6.6" longLatPrecision="1304" longitude="106.8" name="Bogor" pageId="16" pageNumber="17" stateProvince="West Java">Bogor</location>
; 
<geoCoordinate id="EE2C0C9D8B3D4E094CF5F8533F1AF810" box="[704,791,1967,1995]" degrees="6" direction="south" minutes="36" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-6.6">6°36′ S</geoCoordinate>
, 
<geoCoordinate id="EE2C0C9D8B3D4E094D11F8533F91F810" box="[804,924,1967,1995]" degrees="106" direction="east" minutes="48" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="106.8">106°48′ E</geoCoordinate>
; 
<collectionCode id="ED09F29F8B3D4E094D9CF84D3FE7F810" box="[937,1002,1969,1995]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="16" pageNumber="17" type="Museum">ZMH</collectionCode>
<specimenCode id="DBBEC2218B3D4E094DC4F84C3833F810" box="[1009,1086,1968,1995]" pageId="16" pageNumber="17">98145</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B3C4E0B4E88FEF73DB6FE76" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BA76A5A8B3C4E084E88FEF73D43FEFE" blockId="17.[189,334,267,293]" box="[189,334,267,293]" pageId="17" pageNumber="18">
<heading id="D0EFDD368B3C4E084E88FEF73D43FEFE" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B96CB6488B3C4E084E88FEF73D43FEFE" bold="true" box="[189,334,267,293]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B3C4E0B4E88FEC63E1AFEB7" blockId="17.[189,1398,313,446]" lastBlockId="18.[189,1398,267,364]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
SHELL (
<figureCitation id="132376DF8B3C4E084F24FEC53D53FE8F" box="[273,350,313,340]" captionStart="Figs 2–5" captionStartId="11.[189,241,1860,1886]" captionTargetBox="[231,1357,265,1834]" captionTargetId="figure@11.[231,1357,265,1834]" captionTargetPageId="11" captionText="Figs 2–5. Shells of species of Landouria from Java. 2. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype of Helix winteriana, ZMA 376193a). 3. L. winteriana, West Java, Cibodas (syntype of Plectotropis kraepelini Leschke, 1914, ZMH 98416). 4. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 5. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak (neotype of Helix rotatoria, MZB 19205). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152223/files/figure.png" pageId="17" pageNumber="18">Figs 4</figureCitation>
, 
<figureCitation id="132376DF8B3C4E084F5FFEC53D76FE88" box="[362,379,313,339]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="17" pageNumber="18">6</figureCitation>
; 
<tableCitation id="C69A5FE18B3C4E084FB2FEC53E05FE8F" box="[391,520,313,340]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="17" pageNumber="18" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Conical, with 4.75–5 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side without distinct incised spiral lines, umbilical side with indistinct spiral lines; with tubercles all over shell surface that carry scaly processes or hairs mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled at beginning; aperture almost circular; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 30–43% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<caption id="DF673AD28B3C4E084E88F907391CF833" httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,241,1787,1813]" targetBox="[189,1399,501,1756]" targetPageId="17">
<paragraph id="8BA76A5A8B3C4E084E88F907391CF833" blockId="17.[189,1399,1787,2024]" pageId="17" pageNumber="18">
<emphasis id="B96CB6488B3C4E084E88F9073D37F8CE" bold="true" box="[189,314,1787,1813]" pageId="17" pageNumber="18">Figs 6–11.</emphasis>
Shell sculpture of species of 
<taxonomicName id="4C1811D98B3C4E084CA9F9073F15F8CE" authorityName="Godwin-Austen" authorityYear="1918" box="[668,792,1787,1813]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B3C4E084CA9F9073F15F8CE" box="[668,792,1787,1813]" italics="true" pageId="17" pageNumber="18">Landouria</emphasis>
</taxonomicName>
from 
<collectingRegion id="49DCA4B88B3C4E084D57F9073F9BF8CE" box="[866,918,1787,1813]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">Java</collectingRegion>
(umbilical side of shell in 6; apical side in 7–11). 
<emphasis id="B96CB6488B3C4E084F04F8E23D4DF8E3" bold="true" box="[305,320,1822,1848]" pageId="17" pageNumber="18">6</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084F65F8E33EA7F8E3" authority="(Leschke, 1914)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[336,682,1822,1848]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3C4E084F65F8E33DD0F8E3" box="[336,477,1822,1848]" italics="true" pageId="17" pageNumber="18">L. conoidea</emphasis>
(
<bibRefCitation id="EF8917AB8B3C4E084FDBF8E23EADF8E3" author="Leschke" box="[494,672,1822,1848]" firstAuthor="Leschke" pageId="17" pageNumber="18" pagination="205 - 284" refId="ref33668" refString="Leschke M. 1914. Zur Molluskenfauna von Java und Celebes. Mitteilungen aus dem naturhistorischen Museum in Hamburg 31: 205 - 284, 1 pl." type="journal article" year="1914">Leschke, 1914</bibRefCitation>
)
</taxonomicName>
, 
<collectingRegion id="49DCA4B88B3C4E084C82F8E23F3CF8E3" box="[695,817,1822,1848]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="18">West Java</collectingRegion>
, Bogor (
<typeStatus id="54A3D4F88B3C4E084DAFF8E23FF7F8E3" box="[922,1018,1822,1848]" pageId="17" pageNumber="18" type="syntype">syntype</typeStatus>
, ZMH 98144). 
<emphasis id="B96CB6488B3C4E084A80F8E238C9F8E3" bold="true" box="[1205,1220,1822,1848]" pageId="17" pageNumber="18">7</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084AE0F8E3397AF8E3" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1237,1399,1822,1848]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
<emphasis id="B96CB6488B3C4E084AE0F8E3397AF8E3" box="[1237,1399,1822,1848]" italics="true" pageId="17" pageNumber="18">L. tholiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3C4E084E88F8BE3D28F887" box="[189,293,1858,1884]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B3C4E084F0EF8BD3DC9F880" box="[315,452,1857,1883]" country="Indonesia" name="Yogyakarta" pageId="17" pageNumber="18">Yogyakarta</collectingRegion>
, Gunung Kidul, Tepus, near Poktunggal Beach (
<typeStatus id="54A3D4F88B3C4E084A7DF8BD38BEF880" box="[1096,1203,1857,1883]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
, MZB 17999). 
<emphasis id="B96CB6488B3C4E084E88F8983CC1F8A5" bold="true" box="[189,204,1892,1918]" pageId="17" pageNumber="18">8</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084ED4F8993D81F8A5" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[225,396,1892,1918]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="madurensis" status="sp. nov.">
<emphasis id="B96CB6488B3C4E084ED4F8993D81F8A5" box="[225,396,1892,1918]" italics="true" pageId="17" pageNumber="18">L. madurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3C4E084FACF8993DF0F8A4" box="[409,509,1893,1919]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B3C4E084C3AF8993E8BF8A4" box="[527,646,1893,1919]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">East Java</collectingRegion>
, Madura Island, Sampang, Nepa Monkey lowland rainforest (
<typeStatus id="54A3D4F88B3C4E084EF0F8743D22F879" box="[197,303,1928,1954]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, MZB 19209). 
<emphasis id="B96CB6488B3C4E084FDEF87B3DF7F87A" bold="true" box="[491,506,1927,1953]" pageId="17" pageNumber="18">9</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084C3EF8743ECDF87A" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[523,704,1928,1953]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="pacitanensis" status="sp. nov.">
<emphasis id="B96CB6488B3C4E084C3EF8743ECDF87A" box="[523,704,1928,1953]" italics="true" pageId="17" pageNumber="18">L. pacitanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3C4E084CFFF8743F27F879" box="[714,810,1928,1954]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B3C4E084D0DF8743FA1F879" box="[824,940,1928,1954]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">East Java</collectingRegion>
, Pacitan, Srau Beach (
<typeStatus id="54A3D4F88B3C4E084A88F874392AF879" box="[1213,1319,1928,1954]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, ZMH 148510). 
<emphasis id="B96CB6488B3C4E084F04F8573D43F81E" bold="true" box="[305,334,1963,1989]" pageId="17" pageNumber="18">10</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084F54F8503F40F81D" authority="van Benthem Jutting, 1950" authorityName="van Benthem Jutting" authorityYear="1950" box="[353,845,1963,1990]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="B96CB6488B3C4E084F54F8503DF5F81E" box="[353,504,1963,1989]" italics="true" pageId="17" pageNumber="18">L. monticola</emphasis>
<bibRefCitation id="EF8917AB8B3C4E084C31F8573F40F81D" author="van Benthem Jutting" box="[516,845,1963,1990]" firstAuthor="van Benthem Jutting" pageId="17" pageNumber="18" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting, 1950</bibRefCitation>
</taxonomicName>
, 
<collectingRegion id="49DCA4B88B3C4E084D69F8573FD5F81E" box="[860,984,1963,1989]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="18">West Java</collectingRegion>
, Gunung Cikuray (
<typeStatus id="54A3D4F88B3C4E084AF6F8573920F81E" box="[1219,1325,1963,1989]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, SMF 8880). 
<emphasis id="B96CB6488B3C4E084F25F8323D27F833" bold="true" box="[272,298,1998,2024]" pageId="17" pageNumber="18">11</emphasis>
. 
<taxonomicName id="4C1811D98B3C4E084F0FF8333EA8F833" authority="(Mousson, 1848)" authorityName="Mousson" authorityYear="1848" baseAuthorityName="Mousson" baseAuthorityYear="1848" box="[314,677,1998,2024]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="smimensis">
<emphasis id="B96CB6488B3C4E084F0FF8333DDFF833" box="[314,466,1999,2024]" italics="true" pageId="17" pageNumber="18">L. smimensis</emphasis>
(
<bibRefCitation id="EF8917AB8B3C4E084FD4F8323E96F833" author="Mousson" box="[481,667,1998,2024]" firstAuthor="Mousson" pageId="17" pageNumber="18" pagination="264 - 273" refId="ref33832" refString="Mousson A. 1848. Ueber die Land- und Susswassermollusken von Java. Mittheilungen der naturforschenden Gesellschaft in Zurich 1: 264 - 273." type="journal article" year="1848">Mousson, 1848</bibRefCitation>
)
</taxonomicName>
, 
<collectingRegion id="49DCA4B88B3C4E084C85F8323F2CF833" box="[688,801,1998,2024]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">East Java</collectingRegion>
, Tosari (ZMA 390881). Scale bars: 
<quantity id="4CE0C7BF8B3C4E084AF0F8323901F833" box="[1221,1292,1998,2024]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="8BA76A5A8B3F4E0B4E88FE6F3DB6FE76" blockId="18.[189,443,403,430]" box="[189,443,403,430]" pageId="18" pageNumber="19">GENITALIA. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30239D18B3F4E0B4E88FE293D81FC8D" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BA76A5A8B3F4E0B4E88FE293D22FE34" blockId="18.[189,303,469,495]" box="[189,303,469,495]" pageId="18" pageNumber="19">
<heading id="D0EFDD368B3F4E0B4E88FE293D22FE34" bold="true" box="[189,303,469,495]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B96CB6488B3F4E0B4E88FE293D22FE34" bold="true" box="[189,303,469,495]" pageId="18" pageNumber="19">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B3F4E0B4E88FDFF3E10FD53" blockId="18.[189,1399,515,648]" pageId="18" pageNumber="19">
<taxonomicName id="4C1811D98B3F4E0B4E88FDFF3DA3FDC6" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[189,430,515,541]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3F4E0B4E88FDFF3DA3FDC6" box="[189,430,515,541]" italics="true" pageId="18" pageNumber="19">Landouria conoidea</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C1811D98B3F4E0B4C7BFDF83EE3FDC6" authorityName="- van Benthem Jutting" authorityYear="1950" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[590,750,516,541]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="winteriana">
<emphasis id="B96CB6488B3F4E0B4C7BFDF83EE3FDC6" box="[590,750,516,541]" italics="true" pageId="18" pageNumber="19">L. winteriana</emphasis>
</taxonomicName>
, which occurs in the same region in 
<collectingRegion id="49DCA4B88B3F4E0B4A9EFDF83928FDC5" box="[1195,1317,516,542]" country="Indonesia" name="Jawa Barat" pageId="18" pageNumber="19">West Java</collectingRegion>
, in the smaller shell diameter (
<quantity id="4CE0C7BF8B3F4E0B4FE9FDDA3E62FD9A" box="[476,623,550,577]" metricMagnitude="-3" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.7" metricValueMin="7.6" pageId="18" pageNumber="19" unit="mm" value="8.149999999999999" valueMax="8.7" valueMin="7.6">7.6–8.7 mm</quantity>
vs 
<quantity id="4CE0C7BF8B3F4E0B4C95FDDA3F4DFD9A" box="[672,832,550,577]" metricMagnitude="-2" metricUnit="m" metricValue="1.1149999999999998" metricValueMax="1.26" metricValueMin="0.97" pageId="18" pageNumber="19" unit="mm" value="11.149999999999999" valueMax="12.6" valueMin="9.7">9.7–12.6 mm</quantity>
in 
<taxonomicName id="4C1811D98B3F4E0B4D5BFDDB381DFD9B" authorityName="- van Benthem Jutting" authorityYear="1950" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[878,1040,551,576]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="winteriana">
<emphasis id="B96CB6488B3F4E0B4D5BFDDB381DFD9B" box="[878,1040,551,576]" italics="true" pageId="18" pageNumber="19">L. winteriana</emphasis>
</taxonomicName>
), the higher shell form (D/H 1.48–1.54 vs 
<quantity id="4CE0C7BF8B3F4E0B4F6BFDB53DF9FDB8" box="[350,500,585,612]" metricMagnitude="-2" metricUnit="m" metricValue="4.5974" metricValueMax="5.2578" metricValueMin="3.937" pageId="18" pageNumber="19" unit="in" value="1.81" valueMax="2.07" valueMin="1.55">1.55–2.07 in</quantity>
<taxonomicName id="4C1811D98B3F4E0B4FC9FDB63E91FDB8" authorityName="- van Benthem Jutting" authorityYear="1950" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[508,668,586,611]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="winteriana">
<emphasis id="B96CB6488B3F4E0B4FC9FDB63E91FDB8" box="[508,668,586,611]" italics="true" pageId="18" pageNumber="19">L. winteriana</emphasis>
</taxonomicName>
), the more pronounced keel and the presence of tubercles all over the shell surface (
<figureCitation id="132376DF8B3F4E0B4FF2FD913E02FD53" box="[455,527,621,648]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BA76A5A8B3F4E0B4E88FD523D81FC8D" blockId="18.[189,1399,686,854]" pageId="18" pageNumber="19">
With regard to the conical shell form, 
<taxonomicName id="4C1811D98B3F4E0B4CB3FD533F1FFD13" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[646,786,686,712]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3F4E0B4CB3FD533F1FFD13" box="[646,786,686,712]" italics="true" pageId="18" pageNumber="19">L. conoidea</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C1811D98B3F4E0B4DACFD53380AFD13" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[921,1031,687,712]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B3F4E0B4DACFD53380AFD13" box="[921,1031,687,712]" italics="true" pageId="18" pageNumber="19">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3F4E0B4A25FD533861FD12" box="[1040,1132,687,713]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
(Fig. 25), from which it differs in the less elevated (D/H 1.48–1.54 vs 
<quantity id="4CE0C7BF8B3F4E0B4CC2FD2D3F86FD30" box="[759,907,721,747]" metricMagnitude="-2" metricUnit="m" metricValue="3.2766" metricValueMax="3.4798" metricValueMin="3.0734" pageId="18" pageNumber="19" unit="in" value="1.29" valueMax="1.37" valueMin="1.21">1.21–1.37 in</quantity>
<taxonomicName id="4C1811D98B3F4E0B4DA6FD2E380CFD30" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[915,1025,722,747]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B3F4E0B4DA6FD2E380CFD30" box="[915,1025,722,747]" italics="true" pageId="18" pageNumber="19">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3F4E0B4A3FFD2E3865FD37" box="[1034,1128,722,748]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
), smaller shell (D 
<quantity id="4CE0C7BF8B3F4E0B4B76FD2D3D16FCD4" metricMagnitude="-3" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.7" metricValueMin="7.6" pageId="18" pageNumber="19" unit="mm" value="8.149999999999999" valueMax="8.7" valueMin="7.6">7.6– 8.7 mm</quantity>
vs 11.6–14.0 mm in 
<taxonomicName id="4C1811D98B3F4E0B4C10FD093E9BFCD5" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[549,662,757,782]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B3F4E0B4C10FD093E9BFCD5" box="[549,662,757,782]" italics="true" pageId="18" pageNumber="19">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3F4E0B4C94FD093F0CFCD4" box="[673,769,757,783]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
) with fewer whorls (4.75–5 vs 
<quantity id="4CE0C7BF8B3F4E0B4AB2FD0838F7FCD5" box="[1159,1274,756,783]" metricMagnitude="-1" metricUnit="m" metricValue="1.68275" metricValueMax="1.778" metricValueMin="1.5875" pageId="18" pageNumber="19" unit="in" value="6.625" valueMax="7.0" valueMin="6.25">6.25–7 in</quantity>
<taxonomicName id="4C1811D98B3F4E0B4B30FD09397BFCD5" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1285,1398,757,782]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B3F4E0B4B30FD09397BFCD5" box="[1285,1398,757,782]" italics="true" pageId="18" pageNumber="19">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3F4E0B4E88FCE43D11FCE9" box="[189,284,792,818]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
), the more pronounced keel and the shell sculpture consisting of tubercles vs longish scales in 
<taxonomicName id="4C1811D98B3F4E0B4E88FCC03D27FC8E" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,298,828,853]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B3F4E0B4E88FCC03D27FC8E" box="[189,298,828,853]" italics="true" pageId="18" pageNumber="19">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B3F4E0B4F07FCC03D81FC8D" box="[306,396,828,854]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B3F4E0B4E88FC80384EFC1D" box="[189,1091,892,966]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BA76A5A8B3F4E0B4E88FC803D5BFC4D" blockId="18.[189,342,892,918]" box="[189,342,892,918]" pageId="18" pageNumber="19">
<heading id="D0EFDD368B3F4E0B4E88FC803D5BFC4D" bold="true" box="[189,342,892,918]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B96CB6488B3F4E0B4E88FC803D5BFC4D" bold="true" box="[189,342,892,918]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B3F4E0B4E88FC57384EFC1D" blockId="18.[189,1091,939,966]" box="[189,1091,939,966]" pageId="18" pageNumber="19">
<taxonomicName id="4C1811D98B3F4E0B4E88FC573DA1FC1E" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[189,428,939,965]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B3F4E0B4E88FC573DA1FC1E" box="[189,428,939,965]" italics="true" pageId="18" pageNumber="19">Landouria conoidea</emphasis>
</taxonomicName>
is so far known only from Bogor in 
<collectingRegion id="49DCA4B88B3F4E0B4D6CFC573FC3FC1E" box="[857,974,939,965]" country="Indonesia" name="Jawa Barat" pageId="18" pageNumber="19">West Java</collectingRegion>
(Fig. 53).
</paragraph>
</subSubSection>
</treatment>
</document>