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<document id="A23859E5A1F1DE9F8F5E7BF6327235E5" ID-CLB-Dataset="55761" ID-DOI="10.18590/euscorpius.2019.vol2019.iss277.1" ID-GBIF-Dataset="e9986ae3-7799-4a66-afc1-b1e1ecd827d8" ID-ISSN="1536-9307" ID-Zenodo-Dep="6511455" ID-ZooBank="64451868-1FB3-4D20-B88D-FBF7304EB775" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638261050631" checkinUser="felipe" docAuthor="Lowe, Graeme, Kovařík, František, Stockmann, Mark &amp; Šťáhlavský, František" docDate="2019" docId="D81E87F7512BFF8E5D94D991FD8BFEA0" docLanguage="en" docName="Euscorpius.277.1-30.pdf" docOrigin="Euscorpius 277" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Trypanothacus barnesi Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský 2019, sp. n." docType="treatment" docUuid="B0598EB1-8878-4F09-A850-E36129A16ED0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="21" masterDocId="2427FF8F512DFF995E5DDA5DFF8BFFC1" masterDocTitle="Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="5" updateTime="1699471307476" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4544EA23FE4178DD7F486A93B86705AC">Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="D7A72E52BC413AACAA41532E3A3BC96E">Kovařík, František</mods:namePart>
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<mods:namePart id="13F74628D081F19F095F377929F032F0">Stockmann, Mark</mods:namePart>
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<taxonomicName id="97B74D62512BFF9F5D94D991FB6BFC26" ID-CoL="7CXQH" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[969,1248,972,999]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF3512BFF9F5D94D991FB6BFC26" bold="true" box="[969,1248,972,999]" italics="true" pageId="6" pageNumber="5">Trypanothacus barnesi</emphasis>
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(
<figureCitation id="C88C2A64512BFF9F5D1BD9B3FC30FBC7" box="[838,955,1006,1030]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–10" captionStart-2="Figures 11–12" captionStart-3="Figures 13–14" captionStart-4="Figures 15–22" captionStart-5="Figures 23–28" captionStart-6="Figures 29–46" captionStartId-0="3.[113,190,1967,1989]" captionStartId-1="5.[113,190,1457,1479]" captionStartId-2="7.[116,193,1657,1679]" captionStartId-3="9.[113,190,1589,1611]" captionStartId-4="10.[113,190,1958,1980]" captionStartId-5="11.[116,193,1958,1980]" captionStartId-6="12.[115,192,1871,1893]" captionTargetBox-0="[132,1458,266,1903]" captionTargetBox-1="[113,1474,223,1433]" captionTargetBox-2="[116,1473,234,1632]" captionTargetBox-3="[113,1474,223,1572]" captionTargetBox-4="[120,1471,258,1929]" captionTargetBox-5="[113,1474,234,1927]" captionTargetBox-6="[131,1463,242,1849]" captionTargetId-0="figure-7@3.[113,1474,253,1912]" captionTargetId-1="figure-291@5.[113,1474,223,1449]" captionTargetId-2="figure-143@7.[114,1475,234,1632]" captionTargetId-3="figure-7@9.[113,1474,223,1574]" captionTargetId-4="figure-11@10.[113,1474,234,1934]" captionTargetId-5="figure-69@11.[111,1474,234,1930]" captionTargetId-6="figure-164@12.[113,1471,234,1854]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." captionText-2="Figures 11–12: Trypanothacus barnesi gen. et sp. n. Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm." captionText-3="Figures 13–14: Trypanothacus barnesi gen. et sp. n. Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm." captionText-4="Figures 15–22: Trypanothacus barnesi gen. et sp. n. Figures 15, 20–22. Male holotype, metasoma V and telson lateral view (15), metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Figures. 16–19. Female paratype from type locality, metasoma V and telson lateral view (16), metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Scale bars: 4 mm (17–19, 20–21)." captionText-5="Figures 23–28: Trypanothacus barnesi gen. et sp. n. Figures 23–25. Male paratype from Thumrait (VII.1997), metasoma and telson dorsal (23), lateral (24) and ventral (25) views. Figures 26–28. Female paratype from Yalooni (XII.1988), metasoma and telson dorsal (26), lateral (27) and ventral (28) views. UV fluorescence. Scale bar: 4 mm." captionText-6="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi-0="http://doi.org/10.5281/zenodo.6511465" figureDoi-1="http://doi.org/10.5281/zenodo.6511467" figureDoi-2="http://doi.org/10.5281/zenodo.6511473" figureDoi-3="http://doi.org/10.5281/zenodo.6511477" figureDoi-4="http://doi.org/10.5281/zenodo.6511479" figureDoi-5="http://doi.org/10.5281/zenodo.6511481" figureDoi-6="http://doi.org/10.5281/zenodo.6511485" httpUri-0="https://zenodo.org/record/6511465/files/figure.png" httpUri-1="https://zenodo.org/record/6511467/files/figure.png" httpUri-2="https://zenodo.org/record/6511473/files/figure.png" httpUri-3="https://zenodo.org/record/6511477/files/figure.png" httpUri-4="https://zenodo.org/record/6511479/files/figure.png" httpUri-5="https://zenodo.org/record/6511481/files/figure.png" httpUri-6="https://zenodo.org/record/6511485/files/figure.png" pageId="6" pageNumber="5">Figs. 1–42</figureCitation>
, 
<figureCitation id="C88C2A64512BFF9F5D98D9B3FB81FBC7" box="[965,1034,1006,1030]" captionStart-0="Figures 47–58" captionStart-1="Figures 59–70" captionStart-2="Figures 71–77" captionStartId-0="13.[113,190,1959,1981]" captionStartId-1="14.[116,193,1907,1929]" captionStartId-2="15.[113,190,1918,1940]" captionTargetBox-0="[113,1474,234,1930]" captionTargetBox-1="[124,1469,237,1889]" captionTargetBox-2="[113,1460,257,1882]" captionTargetId-0="figure-7@13.[113,1474,234,1932]" captionTargetId-1="figure-11@14.[113,1469,232,1890]" captionTargetId-2="figure-8@15.[110,1474,236,1899]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figures 47–58: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of male paratype from Thumrait (VII.1997), chela dorsal (47), external (48), ventral (49) and internal (50) views. Patella dorsal (51), external (52), ventral (53) and internal (54) views. Femur and trochanter dorsal (55), external (56), ventral (57) and internal (58) views. UV fluorescence. Scale bar: 2 mm." captionText-1="Figures 59–70: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of female paratypes, chela dorsal (59), external (60), ventral (61) and internal (62) views. Patella dorsal (63), external (64), ventral (65) and internal (66) views. Femur and trochanter dorsal (67), external (68), ventral (69) and internal (70) views. Dorsal views (59, 63, 67): paratype female from Thumrait (XI.1989) (with normal dorsointernal carina on femur). External (60, 64, 68), ventral (61, 65, 69) and internal (62, 66, 70) views: paratype female from Yalooni (XII.1988) (with malformed dorsointernal carina on femur). UV fluorescence. Scale bars: 2 mm (59, 63, 67), 2 mm (60–62, 64–66, 69–70)." captionText-2="Figures 71–77: Trypanothacus barnesi gen. et sp. n. Figures 71–73. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. Figures 74–77. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77)." figureDoi-0="http://doi.org/10.5281/zenodo.6511487" figureDoi-1="http://doi.org/10.5281/zenodo.6511489" figureDoi-2="http://doi.org/10.5281/zenodo.6511493" httpUri-0="https://zenodo.org/record/6511487/files/figure.png" httpUri-1="https://zenodo.org/record/6511489/files/figure.png" httpUri-2="https://zenodo.org/record/6511493/files/figure.png" pageId="6" pageNumber="5">47–77</figureCitation>
, 
<figureCitation id="C88C2A64512BFF9F5A48D9B3FBD0FBC7" box="[1045,1115,1006,1030]" captionStart-0="Figures 84–85" captionStart-1="Figures 86–89" captionStartId-0="17.[116,193,1988,2010]" captionStartId-1="18.[113,190,1969,1991]" captionTargetBox-0="[113,1474,248,1949]" captionTargetBox-1="[114,1474,236,1939]" captionTargetId-0="figure-35@17.[113,1474,248,1949]" captionTargetId-1="figure-12@18.[114,1474,234,1939]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figures 84–85: Trypanothacus barnesi gen. et sp. n., in vivo habitus, male holotype (84) and female paratype from type locality (85)." captionText-1="Figures 86–89: Trypanothacus barnesi gen. et sp. n.. Figures 86–87. Male (86) and female (87) from Thumrait, photos by J. N. Barnes. Figures 88–89. Type locality, Oman, S of Schelim, 18.041070°N 55.612262°E." figureDoi-0="http://doi.org/10.5281/zenodo.6511498" figureDoi-1="http://doi.org/10.5281/zenodo.6511503" httpUri-0="https://zenodo.org/record/6511498/files/figure.png" httpUri-1="https://zenodo.org/record/6511503/files/figure.png" pageId="6" pageNumber="5">84–87</figureCitation>
, 
<figureCitation id="C88C2A64512BFF9F5A38D9B3FB4EFBC7" box="[1125,1221,1006,1030]" captionStart="Figures 107–110" captionStartId="25.[113,189,1408,1430]" captionTargetBox="[121,1439,242,1379]" captionTargetId="figure-494@25.[113,1474,234,1386]" captionTargetPageId="25" captionText="Figures 107–110: Morphometric scaling relations of the telson of fossorial ‘Buthus group’ scorpions. Figure 107. Double logarithmic scatterplot of telson vesicle length/ depth (L/ D) ratio vs. aculeus length normalized to vesicle length (aculeus L/ vesicle L), for large pelophiles/ semipsammophiles (4 genera, 53 species) (Buthus, Odontobuthus, Pantobuthus: orange circles; Trypanothacus gen. n.: blue circles) and large psammophiles/ ultrapsammophiles (6 genera, 29 species) (Buthacus: cyan circles; Vachoniolus: dark green circles; Apistobuthus, Buthiscus, Liobuthus, and Plesiobuthus: yellow circles). Dark line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.57). Upper right inset: diagram of measurements estimated from inverted telson lateral profiles, with dorsal surface of vesicle approximately leveled; SA = ventral profile length of aculeus from base point to tip, LV = length of vesicle from anterior limit or inflection point to aculeus base point; HV = vesicle depth or height (distance between dorsal and ventral tangent planes). Base point was placed at local minimum of inverted ventral profile curve. SA was calculated by cubic spline fit (&gt;10 anchor points). For a minority of broken tips, SA was estimated by intersecting extrapolated dorsal and ventral profile curves (&lt;5%). Figure 108. Scatterplot of fossorial buthids in Figure 107, superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression fit to Order Scorpiones data (including data in Figure 107) (Pearson correlation coefficient, R = –0.62).Also plotted are data from small ‘Buthus group’ psammophiles/ ultrapsammophiles (5 genera, 10 species: Compsobuthus arabicus, Lanzatus somalilandus, Neobuthus kutcheri, Picobuthus dundoni, P. wahibaensis, Anomalobuthus krivochatskyi, A. pavlovskyi, A. rickmersi, A. lowei, A. talebii) (dark yellow circles). Figure 109. Double logarithmic scatterplot of carapace length (L) vs. aculeus length normalized to vesicle length (i.e., aculeus L/ vesicle L) for psammophile/ ultrapsammophile members of the ‘Buthus group’, including large species in Figure 107, and small species in Figure 108 (dark yellow circles). Blue line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.74). Figure 110. Scatterplot of psammophile buthid data in Figure 109 (all in one color, blue circles) superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression to Order Scorpiones data (including data in Figure 109) (Pearson correlation coefficient, R = –0.01). Also plotted are data from bothriurid genus Brachistosternus (43 species) (magenta triangles), and corresponding linear least squares regression fit (R = 0.39). All plots in Figures 107-110 include data from both males and females. Data were assembled by mining taxonomic literature or directly measuring specimens. Included are 1854 species or subspecies (1331 ♂, 1201 ♀) representing all known extant genera (1841 species or subspecies) plus 7 fossil genera (13 species)." figureDoi="http://doi.org/10.5281/zenodo.6511515" httpUri="https://zenodo.org/record/6511515/files/figure.png" pageId="6" pageNumber="5">107–108</figureCitation>
, 
<figureCitation id="C88C2A64512BFF9F5A8DD9B3FAA6FBC7" box="[1232,1325,1006,1030]" captionStart-0="Figures 111–113" captionStart-1="Figures 114–116" captionStart-2="Figures 117–118" captionStartId-0="26.[113,190,671,693]" captionStartId-1="27.[113,190,1990,2012]" captionStartId-2="28.[113,190,1984,2006]" captionTargetBox-0="[113,1474,235,649]" captionTargetBox-1="[113,1474,234,1972]" captionTargetBox-2="[113,1474,236,1962]" captionTargetId-0="figure-786@26.[113,1474,234,659]" captionTargetId-1="figure-7@27.[113,1474,234,1972]" captionTargetId-2="figure-11@28.[113,1474,236,1964]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionText-0="Figures 111–113: Trypanothacus barnesi gen. et sp. n., mitotic metaphase (111), postpachytene (112) and ideogram (113) (y axis - % of the chromosome length of the haploid set) of male holotype. Scale bar = 5 μm." captionText-1="Figures 114–116: Trypanothacus barnesi gen. et sp. n., promenade à deux (courtship dance) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." captionText-2="Figures 117–118: Trypanothacus barnesi gen. et sp. n., cheliceral massage (117) and spermatophore deposition (118) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi-0="http://doi.org/10.5281/zenodo.6511517" figureDoi-1="http://doi.org/10.5281/zenodo.6511519" figureDoi-2="http://doi.org/10.5281/zenodo.6511521" httpUri-0="https://zenodo.org/record/6511517/files/figure.png" httpUri-1="https://zenodo.org/record/6511519/files/figure.png" httpUri-2="https://zenodo.org/record/6511521/files/figure.png" pageId="6" pageNumber="5">111–118</figureCitation>
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<tableCitation id="1D35035A512BFF9F5B6AD9B3FA27FBC7" box="[1335,1452,1006,1030]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="8.[112,168,987,1008]" captionStartId-1="19.[113,169,1225,1246]" captionTargetId-0="graphics-540@8.[155,1419,222,960]" captionTargetId-1="graphics-432@19.[117,1474,234,1210]" captionTargetPageId-0="8" captionTargetPageId-1="19" captionText-0="Table 1. Comparative measurements of adults of Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 2. Variation in morphometric ratios of adult Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it" httpUri-0="http://table.plazi.org/id/04C866695125FF915E2DD986FD87FBCF" httpUri-1="http://table.plazi.org/id/04C86669513EFF8A5E2CDE94FD7EFB3D" pageId="6" pageNumber="5" tableUuid-0="04C866695125FF915E2DD986FD87FBCF" tableUuid-1="04C86669513EFF8A5E2CDE94FD7EFB3D">Tables 1–2</tableCitation>
) 
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<collectingCountry id="28A07671512BFF9F5AE1DE33FA88FB47" box="[1212,1283,1134,1158]" name="Oman" pageId="6" pageNumber="5">Oman</collectingCountry>
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<collectionCode id="36A6AE24512BFF9F5D6DDE93FBD9FB27" box="[816,1106,1230,1254]" pageId="6" pageNumber="5">TYPE MATERIAL EXAMINED</collectionCode>
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<collectingCountry id="28A07671512BFF9F5A3DDE93FB2CFB27" box="[1120,1191,1230,1254]" name="Oman" pageId="6" pageNumber="5">Oman</collectingCountry>
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), 
<collectingMunicipality id="B06CAC9B512BFF9F5B35DE93FA35FB27" box="[1384,1470,1230,1254]" pageId="6" pageNumber="5">Yalooni</collectingMunicipality>
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<date id="24091021512BFF9F5AE0DEB3FAABFAC7" box="[1213,1312,1262,1286]" pageId="6" pageNumber="5" value="1988-12">
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<collectorName id="FD425337512BFF9F5D6DDF53FC08FAE7" box="[816,899,1294,1318]" pageId="6" pageNumber="5">Yalooni</collectorName>
185, 
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<materialsCitation id="E0DF3CBC512BFF9F5A14DF53FC5AFAA4" ID-GBIF-Occurrence="3758760310" collectingDate="1989-11-10" collectionCode="NHMB" collectorName="I. A. J. Brown" country="Oman" latitude="17.65" location="Thamarit" longLatPrecision="1277" longitude="54.033333" municipality="Thamarit" pageId="6" pageNumber="5" specimenCode="ONHM 1416" specimenCount="1" specimenCount-female="1" stateProvince="Dhofar" typeStatus="paratype">
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<collectingMunicipality id="B06CAC9B512BFF9F5B07DF53FA34FAE7" box="[1370,1471,1294,1318]" country="Oman" latitude="17.65" longLatPrecision="1277" longitude="54.033333" name="Thamarit" pageId="6" pageNumber="5" stateProvince="Zufar">Thamarit</collectingMunicipality>
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<date id="24091021512BFF9F5DA6DF73FBFCFA87" box="[1019,1143,1326,1350]" pageId="6" pageNumber="5" value="1989-11-10">
<collectingDate id="344DE9C9512BFF9F5DA6DF73FBFCFA87" box="[1019,1143,1326,1350]" pageId="6" pageNumber="5" value="1989-11-10">10.XI.1989</collectingDate>
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<collectorName id="FD425337512BFF9F5AE7DF73FAEAFA84" box="[1210,1377,1326,1350]" pageId="6" pageNumber="5">I. A. J. Brown</collectorName>
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<locationDeviation id="41BA2FC3512BFF9F5B59DF13FAB3FA47" pageId="6" pageNumber="5">wadi area SE of Thumrait in slightly raised sedimentary plateau</locationDeviation>
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<collectedFrom id="E3AB751C512BFF9F5B18DF33FBEBFA67" pageId="6" pageNumber="5">under sheet wood, and in aircraft shelter</collectedFrom>
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<geoCoordinate id="35835026512BFF9F5A97DFD3FAAAFA67" box="[1226,1313,1422,1446]" degrees="53" direction="east" minutes="59" orientation="longitude" pageId="6" pageNumber="5" precision="925" value="53.983334">53°59'E</geoCoordinate>
, 
<date id="24091021512BFF9F5B71DFD3FA1BFA67" box="[1324,1424,1422,1446]" pageId="6" pageNumber="5" value="1997-07">
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<collectorName id="FD425337512BFF9F5D6DDFF3FC38FA04" box="[816,947,1454,1477]" pageId="6" pageNumber="5">J. N. Barnes</collectorName>
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<collectionCode id="36A6AE24512BFF9F5A44DFF3FBEBFA07" box="[1049,1120,1454,1478]" pageId="6" pageNumber="5">GLPC</collectionCode>
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<collectionCode id="36A6AE24512BFF9F5A34DFF3FB35FA07" box="[1129,1214,1454,1478]" pageId="6" pageNumber="5">ONHM</collectionCode>
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<materialsCitation id="E0DF3CBC512BFF9F5A9ADFF3FB64F9C4" ID-GBIF-Occurrence="3758760303" collectedFrom="in aircraft shelter" collectingDate="1998-07-23" collectionCode="NHMB" collectorName="J. N. Barnes" country="Oman" latitude="17.7" location="Thumrait" longLatPrecision="1277" longitude="53.816666" municipality="Thumrait" pageId="6" pageNumber="5" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1" typeStatus="paratype">
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), 
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airfield, 
<collectedFrom id="E3AB751C512BFF9F5A51DF93FB56FA27" box="[1036,1245,1486,1510]" pageId="6" pageNumber="5">in aircraft shelter</collectedFrom>
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<geoCoordinate id="35835026512BFF9F5AA9DF93FAC4FA27" box="[1268,1359,1486,1510]" degrees="17" direction="north" minutes="42" orientation="latitude" pageId="6" pageNumber="5" precision="925" value="17.7">17°42'N</geoCoordinate>
, 
<geoCoordinate id="35835026512BFF9F5B3BDF93FA36FA27" box="[1382,1469,1486,1510]" degrees="53" direction="east" minutes="49" orientation="longitude" pageId="6" pageNumber="5" precision="925" value="53.816666">53°49'E</geoCoordinate>
, 
<collectingDate id="344DE9C9512BFF9F5D6DDFB3FC3EF9C7" box="[816,949,1518,1542]" pageId="6" pageNumber="5">
<date id="24091021512BFF9F5D6DDFB3FCDBF9C7" box="[816,848,1518,1542]" pageId="6" pageNumber="5" value="1998-07-23">23.</date>
VII.1998
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, leg. 
<collectorName id="FD425337512BFF9F5DA4DFB3FB03F9C4" box="[1017,1160,1518,1541]" pageId="6" pageNumber="5">J. N. Barnes</collectorName>
, 
<collectionCode id="36A6AE24512BFF9F5AC5DFB3FB60F9C4" box="[1176,1259,1518,1541]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="6" pageNumber="5">NHMB</collectionCode>
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3♂ (holotype No. 1121 and 
<specimenCode id="00119E9A512BFF9F5DC0DC53FC21F9E7" box="[925,938,1550,1574]" pageId="6" pageNumber="5">2</specimenCode>
<typeStatus id="8F0C8843512BFF9F5DEEDC53FB92F9E7" box="[947,1049,1550,1574]" pageId="6" pageNumber="5" type="paratype">paratypes</typeStatus>
<specimenCode id="00119E9A512BFF9F5A7EDC53FB07F9E7" box="[1059,1164,1550,1574]" pageId="6" pageNumber="5">No. 1594</specimenCode>
), 
<locationDeviation id="41BA2FC3512BFF9F5AFCDC53FAADF9E7" box="[1185,1318,1550,1574]" location="Shalim" pageId="6" pageNumber="5">S of Shalim</locationDeviation>
, 
<geoCoordinate id="35835026512BFF9F5B6EDC53FA49F9E7" box="[1331,1474,1550,1574]" degrees="18.041070" direction="north" orientation="latitude" pageId="6" pageNumber="5" precision="1" value="18.04107">18.041070°N</geoCoordinate>
<geoCoordinate id="35835026512BFF9F5D6DDC73FC4BF987" box="[816,960,1582,1606]" degrees="55.612262" direction="east" orientation="longitude" pageId="6" pageNumber="5" precision="1" value="55.612263">55.612262°E</geoCoordinate>
, 
<date id="24091021512BFF9F5D89DC73FBC9F987" box="[980,1090,1582,1606]" pageId="6" pageNumber="5" value="2016-09-06">
<collectingDate id="344DE9C9512BFF9F5D89DC73FBC9F987" box="[980,1090,1582,1606]" pageId="6" pageNumber="5" value="2016-09-06">6.IX.2016</collectingDate>
</date>
, 21:00–23:00 h, full moon, UV detection, 
<collectionCode id="36A6AE24512BFF9F5DC3DC13FC6AF9A7" box="[926,993,1614,1638]" pageId="6" pageNumber="5">FKCP</collectionCode>
, 
<collectionCode id="36A6AE24512BFF9F5DB7DC13FBA5F9A7" box="[1002,1070,1614,1638]" pageId="6" pageNumber="5">GLPC</collectionCode>
(hemispermatophore);
</materialsCitation>
<materialsCitation id="E0DF3CBC512BFF9F5B7ADC13FABDF967" ID-GBIF-Occurrence="3758760308" collectingDate="2017-10-27" collectingDateMax="2017-10-28" collectingDateMin="2017-10-27" collectionCode="FKCP" collectorName="M. Stockmann" country="Oman" latitude="18.04107" location="S of Shalim" longLatPrecision="1" longitude="55.612263" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="1" stateProvince="Dhofar Province" typeStatus="paratype">
<specimenCount id="46B1FD68512BFF9F5B7ADC13FAC3F9A7" box="[1319,1352,1614,1638]" pageId="6" pageNumber="5" type="female">1♀</specimenCount>
(
<typeStatus id="8F0C8843512BFF9F5B08DC13FA3DF9A7" box="[1365,1462,1614,1638]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
), same locality as holotype, 
<date id="24091021512BFF9F5A1ADC33FB69F947" box="[1095,1250,1646,1670]" pageId="6" pageNumber="5" value="2017-10-27" valueMax="2017-10-28" valueMin="2017-10-27">
<collectingDate id="344DE9C9512BFF9F5A1ADC33FB69F947" box="[1095,1250,1646,1670]" pageId="6" pageNumber="5" value="2017-10-27" valueMax="2017-10-28" valueMin="2017-10-27">27–28.X.2017</collectingDate>
</date>
, 23:00–01:00 h, 62% moon, UV detection, leg. 
<collectorName id="FD425337512BFF9F5A19DCD3FB6FF967" box="[1092,1252,1678,1702]" pageId="6" pageNumber="5">M. Stockmann</collectorName>
, 
<collectionCode id="36A6AE24512BFF9F5AB2DCD3FAB9F967" box="[1263,1330,1678,1702]" pageId="6" pageNumber="5">FKCP</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="500836E1512BFF9F5D6DDCF3FB82F8A4" blockId="6.[816,1475,1230,1894]" pageId="6" pageNumber="5">
OTHERMATERIALEXAMINED. 
<materialsCitation id="E0DF3CBC512BFF9F5A0EDCF3FAAAF8C7" ID-GBIF-Occurrence="3758760301" collectedFrom="under rock" collectingDate="1980-03-22" collectionCode="MNHN" collectorName="J. N. Barnes &amp; Barnes" country="Oman" elevation="450" latitude="17.666666" location="Dhofar" longLatPrecision="1277" longitude="54.033333" pageId="6" pageNumber="5" specimenCode="RS8787" specimenCount="1" specimenCount-juvenile="1" stateProvince="Dhofar">
<emphasis id="62C3EAF3512BFF9F5A0EDCF3FB11F907" bold="true" box="[1107,1178,1710,1734]" pageId="6" pageNumber="5">
<collectingCountry id="28A07671512BFF9F5A0EDCF3FB11F907" box="[1107,1178,1710,1734]" name="Oman" pageId="6" pageNumber="5">Oman</collectingCountry>
</emphasis>
: 
<specimenCount id="46B1FD68512BFF9F5AFEDCF3FB58F907" box="[1187,1235,1710,1734]" pageId="6" pageNumber="5" type="juvenile">1juv</specimenCount>
, 
<locationDeviation id="41BA2FC3512BFF9F5A85DCF3FAE0F907" box="[1240,1387,1710,1734]" pageId="6" pageNumber="5">nearThamarit</locationDeviation>
, 
<collectingRegion id="9273F803512BFF9F5B2CDCF3FA35F907" box="[1393,1470,1710,1734]" country="Oman" name="Zufar" pageId="6" pageNumber="5">Dhofar</collectingRegion>
, 
<collectedFrom id="E3AB751C512BFF9F5D6DDC93FC2DF927" box="[816,934,1742,1766]" pageId="6" pageNumber="5">under rock</collectedFrom>
, 
<elevation id="DB9AD1D2512BFF9F5DECDC93FBA5F927" box="[945,1070,1742,1766]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="5" unit="m" value="450.0">
<quantity id="974F9B04512BFF9F5DECDC93FC7EF924" box="[945,1013,1742,1766]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="5" unit="m" value="450.0">450 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="35835026512BFF9F5A66DC93FB18F927" box="[1083,1171,1742,1766]" degrees="17" direction="north" minutes="40" orientation="latitude" pageId="6" pageNumber="5" precision="925" value="17.666666">17°40'N</geoCoordinate>
<geoCoordinate id="35835026512BFF9F5AC7DC93FB79F927" box="[1178,1266,1742,1766]" degrees="54" direction="east" minutes="02" orientation="longitude" pageId="6" pageNumber="5" precision="925" value="54.033333">54°02'E</geoCoordinate>
, 
<collectingDate id="344DE9C9512BFF9F5AA0DC93FAFCF927" box="[1277,1399,1742,1766]" pageId="6" pageNumber="5">
<date id="24091021512BFF9F5AA0DC93FAB0F927" box="[1277,1339,1742,1766]" pageId="6" pageNumber="5" value="1980-03-22">22.III</date>
.1980
</collectingDate>
, leg. 
<collectorName id="FD425337512BFF9F5BECDC93FC14F8C4" pageId="6" pageNumber="5">J. N. Barnes</collectorName>
, Sc74, 
<collectorName id="FD425337512BFF9F5DB2DCB3FBB2F8C4" box="[1007,1081,1774,1797]" pageId="6" pageNumber="5">Barnes</collectorName>
36, 
<collectionCode id="36A6AE24512BFF9F5A36DCB3FB37F8C4" box="[1131,1212,1774,1797]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="5">MNHN</collectionCode>
<specimenCode id="00119E9A512BFF9F5A99DCB3FA95F8C7" box="[1220,1310,1774,1798]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="5">RS8787</specimenCode>
;
</materialsCitation>
<materialsCitation id="E0DF3CBC512BFF9F5B77DCB3FB82F8A4" ID-GBIF-Occurrence="3758760304" collectedFrom="found dead on level limestone rock and sand" collectingDate="1980-11-10" collectionCode="NHMB" collectorName="M. D. Gallagher" latitude="19.933332" location="Jiddat Al Harasis" longLatPrecision="1270" longitude="57.1" municipality="Yalooni" pageId="6" pageNumber="5" specimenCode="MDG6020" specimenCount="1" specimenCount-female="1">
<specimenCount id="46B1FD68512BFF9F5B77DCB3FAD8F8C4" box="[1322,1363,1774,1798]" pageId="6" pageNumber="5" type="female">1 ♀</specimenCount>
fragment, 
<collectingMunicipality id="B06CAC9B512BFF9F5D6DDD53FC0DF8E7" box="[816,902,1806,1830]" pageId="6" pageNumber="5">Yalooni</collectingMunicipality>
, 
<location id="5568603A512BFF9F5DCDDD53FBCCF8E7" LSID="urn:lsid:plazi:treatment:D81E87F7512BFF8E5D94D991FD8BFEA0:5568603A512BFF9F5DCDDD53FBCCF8E7" box="[912,1095,1806,1830]" country="Oman" latitude="19.933332" longLatPrecision="1270" longitude="57.1" municipality="Yalooni" name="Jiddat Al Harasis" pageId="6" pageNumber="5" stateProvince="Zufar">Jiddat Al Harasis</location>
, 
<collectedFrom id="E3AB751C512BFF9F5A0DDD53FC1BF887" pageId="6" pageNumber="5">found dead on level limestone rock and sand</collectedFrom>
, 
<geoCoordinate id="35835026512BFF9F5DC6DD73FC79F887" box="[923,1010,1838,1862]" degrees="19" direction="north" minutes="56" orientation="latitude" pageId="6" pageNumber="5" precision="925" value="19.933332">19°56'N</geoCoordinate>
<geoCoordinate id="35835026512BFF9F5DA4DD73FBDAF887" box="[1017,1105,1838,1862]" degrees="57" direction="east" minutes="06" orientation="longitude" pageId="6" pageNumber="5" precision="925" value="57.1">57°06'E</geoCoordinate>
, 
<date id="24091021512BFF9F5A06DD73FB5CF887" box="[1115,1239,1838,1862]" pageId="6" pageNumber="5" value="1980-11-10">
<collectingDate id="344DE9C9512BFF9F5A06DD73FB5CF887" box="[1115,1239,1838,1862]" pageId="6" pageNumber="5" value="1980-11-10">10.XI.1980</collectingDate>
</date>
, leg. 
<collectorName id="FD425337512BFF9F5B52DD73FA35F887" box="[1295,1470,1838,1862]" pageId="6" pageNumber="5">M. D. Gallagher</collectorName>
, 
<specimenCode id="00119E9A512BFF9F5D6DDD13FC2DF8A7" box="[816,934,1870,1894]" pageId="6" pageNumber="5">MDG6020</specimenCode>
, 
<collectionCode id="36A6AE24512BFF9F5DECDD13FB8FF8A4" box="[945,1028,1870,1893]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="6" pageNumber="5">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="18AD656A512BFF9F5D6DDDD3FBB8F827" pageId="6" pageNumber="5" type="distribution">
<paragraph id="500836E1512BFF9F5D6DDDD3FBB8F827" blockId="6.[816,1474,1934,2022]" pageId="6" pageNumber="5">
DISTRIBUTION. 
<collectingCountry id="28A07671512BFF9F5D86DDD3FBBBF867" box="[987,1072,1934,1958]" name="Oman" pageId="6" pageNumber="5">OMAN</collectingCountry>
. Ranging from the inland plateau of Jiddat al-Harasis in central Oman to the Nejd Desert of 
<collectingRegion id="9273F803512BFF9F5D6DDD93FCF6F827" box="[816,893,1998,2022]" country="Oman" name="Zufar" pageId="6" pageNumber="5">Dhofar</collectingRegion>
, southern Oman.
</paragraph>
</subSubSection>
<caption id="04C86669512AFF9E5E29DC24FD25F96D" ID-DOI="http://doi.org/10.5281/zenodo.6511473" ID-Zenodo-Dep="6511473" httpUri="https://zenodo.org/record/6511473/files/figure.png" pageId="7" pageNumber="6" startId="7.[116,193,1657,1679]" targetBox="[116,1473,234,1632]" targetPageId="7">
<paragraph id="500836E1512AFF9E5E29DC24FD25F96D" blockId="7.[116,1475,1657,1709]" pageId="7" pageNumber="6">
<emphasis id="62C3EAF3512AFF9E5E29DC24FE89F94F" bold="true" box="[116,258,1657,1679]" pageId="7" pageNumber="6">Figures 11–12</emphasis>
: 
<taxonomicName id="97B74D62512AFF9E5F52DC27FE66F94E" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[271,493,1658,1679]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF3512AFF9E5F52DC27FE66F94E" box="[271,493,1658,1679]" italics="true" pageId="7" pageNumber="6">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788512AFF9E5FAEDC27FDF8F94E" box="[499,627,1658,1679]" pageId="7" pageNumber="6" rank="species">
<emphasis id="62C3EAF3512AFF9E5FAEDC27FD9CF94E" bold="true" box="[499,535,1658,1679]" pageId="7" pageNumber="6">gen</emphasis>
. et 
<emphasis id="62C3EAF3512AFF9E5C61DC27FDD8F94E" bold="true" box="[572,595,1658,1679]" pageId="7" pageNumber="6">sp</emphasis>
. 
<emphasis id="62C3EAF3512AFF9E5C02DC27FDE7F94E" bold="true" box="[607,620,1658,1679]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="18AD656A512AFF9E5E2CDC96FF54F8A2" pageId="7" pageNumber="6" type="etymology">
<paragraph id="500836E1512AFF9E5E2CDC96FF54F8A2" blockId="7.[113,771,1739,1891]" pageId="7" pageNumber="6">
ETYMOLOGY. The specific epithet is a patronym dedicated to naturalist J. Neil Barnes, Airworks Ltd, Salalah, 
<collectingCountry id="28A07671512AFF9E5CD8DCB6FD42F8C2" box="[645,713,1771,1795]" name="Oman" pageId="7" pageNumber="6">Oman</collectingCountry>
, who first collected this species in 1980 from around the Thumrait airbase, and contributed other important specimens of 
<collectingCountry id="28A07671512AFF9E5C9FDD76FC88F882" box="[706,771,1835,1859]" name="Oman" pageId="7" pageNumber="6">Oman</collectingCountry>
scorpions.
</paragraph>
</subSubSection>
<subSubSection id="18AD656A512AFF9E5E2CDDD6FBEDF8C2" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="500836E1512AFF9E5E2CDDD6FBEDF8C2" blockId="7.[113,771,1931,2019]" lastBlockId="7.[816,1474,1739,1795]" pageId="7" pageNumber="6">
DIAGNOSIS. A member of 
<taxonomicName id="97B74D62512AFF9E5FDDDDD6FD94F862" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[384,543,1931,1955]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="62C3EAF3512AFF9E5FDDDDD6FD94F862" box="[384,543,1931,1955]" italics="true" pageId="7" pageNumber="6">Trypanothacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788512AFF9E5C79DDD6FDE4F863" box="[548,623,1931,1955]" pageId="7" pageNumber="6" rank="genus">
<emphasis id="62C3EAF3512AFF9E5C79DDD6FDC7F862" bold="true" box="[548,588,1931,1955]" pageId="7" pageNumber="6">gen</emphasis>
. 
<emphasis id="62C3EAF3512AFF9E5C04DDD6FDE3F862" bold="true" box="[601,616,1931,1955]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
differentiated from its congener as follows: pedipalp femur L/W 
<specimenCount id="46B1FD68512AFF9E5CCCDDF6FD2EF803" box="[657,677,1963,1986]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
2.6–3.1, 
<specimenCount id="46B1FD68512AFF9E5E2CDD96FF0EF823" box="[113,133,1995,2018]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
2.5–2.7, patella L/W 
<specimenCount id="46B1FD68512AFF9E5F24DD96FE06F823" box="[377,397,1995,2018]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
2.5–2.7, 
<specimenCount id="46B1FD68512AFF9E5FABDD96FD81F823" box="[502,522,1995,2018]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
2.4–2.7, chela L/W 
<specimenCount id="46B1FD68512AFF9E5CB2DD96FC88F823" box="[751,771,1995,2018]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
3.5–4.0, 
<specimenCount id="46B1FD68512AFF9E5DD2DC96FC28F923" box="[911,931,1739,1762]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
4.4–4.6; metasoma IV L/D 
<specimenCount id="46B1FD68512AFF9E5A88DC96FB62F923" box="[1237,1257,1739,1762]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
2.0–2.2, 
<specimenCount id="46B1FD68512AFF9E5B12DC96FAE8F923" box="[1359,1379,1739,1762]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
1.9–2.0, V L/D 
<specimenCount id="46B1FD68512AFF9E5D26DCB6FC04F8C3" box="[891,911,1771,1794]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
2.7–3.0, 
<specimenCount id="46B1FD68512AFF9E5DAEDCB6FB8CF8C3" box="[1011,1031,1771,1794]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
2.5–2.9.
</paragraph>
</subSubSection>
<subSubSection id="18AD656A512AFF8D5D6DDD76FEB4FCA2" lastPageId="20" lastPageNumber="19" pageId="7" pageNumber="6" type="description">
<paragraph id="500836E1512AFF9E5D6DDD76FBF8F8A2" blockId="7.[816,1475,1835,2019]" pageId="7" pageNumber="6">
DESCRIPTION. Length of adults: 
<specimenCount id="46B1FD68512AFF9E5AD6DD76FB14F883" box="[1163,1183,1835,1858]" pageId="7" pageNumber="6" type="male">♂</specimenCount>
<quantity id="974F9B04512AFF9E5AFADD76FA95F883" box="[1191,1310,1835,1859]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.1" metricValueMin="4.0" pageId="7" pageNumber="6" unit="mm" value="45.5" valueMax="51.0" valueMin="40.0">40–51 mm</quantity>
, 
<specimenCount id="46B1FD68512AFF9E5B77DD76FAB5F883" box="[1322,1342,1835,1858]" pageId="7" pageNumber="6" type="female">♀</specimenCount>
<quantity id="974F9B04512AFF9E5B1BDD76FA35F883" box="[1350,1470,1835,1859]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.7" metricValueMin="4.3" pageId="7" pageNumber="6" unit="mm" value="50.0" valueMax="57.0" valueMin="43.0">43–57 mm</quantity>
; habitus as shown in 
<figureCitation id="C88C2A64512AFF9E5A54DD16FBE4F8A2" box="[1033,1135,1867,1891]" captionStart="Figures 1–4" captionStartId="3.[113,190,1967,1989]" captionTargetBox="[132,1458,266,1903]" captionTargetId="figure-7@3.[113,1474,253,1912]" captionTargetPageId="3" captionText="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511465" httpUri="https://zenodo.org/record/6511465/files/figure.png" pageId="7" pageNumber="6">Figs. 1–4</figureCitation>
.
</paragraph>
<paragraph id="500836E1512AFF915D6DDD36FE99FB07" blockId="7.[816,1475,1835,2019]" lastBlockId="8.[113,772,1070,2022]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<emphasis id="62C3EAF3512AFF9E5D6DDD36FC20F842" bold="true" box="[816,939,1899,1923]" pageId="7" pageNumber="6">Coloration</emphasis>
(
<figureCitation id="C88C2A64512AFF9E5DE1DD36FBBAF842" box="[956,1073,1899,1923]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–10" captionStartId-0="3.[113,190,1967,1989]" captionStartId-1="5.[113,190,1457,1479]" captionTargetBox-0="[132,1458,266,1903]" captionTargetBox-1="[113,1474,223,1433]" captionTargetId-0="figure-7@3.[113,1474,253,1912]" captionTargetId-1="figure-291@5.[113,1474,223,1449]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." figureDoi-0="http://doi.org/10.5281/zenodo.6511465" figureDoi-1="http://doi.org/10.5281/zenodo.6511467" httpUri-0="https://zenodo.org/record/6511465/files/figure.png" httpUri-1="https://zenodo.org/record/6511467/files/figure.png" pageId="7" pageNumber="6">Figs. 1–10</figureCitation>
, 
<figureCitation id="C88C2A64512AFF9E5A63DD36FB0FF842" box="[1086,1156,1899,1923]" captionStart="Figures 15–22" captionStartId="10.[113,190,1958,1980]" captionTargetBox="[120,1471,258,1929]" captionTargetId="figure-11@10.[113,1474,234,1934]" captionTargetPageId="10" captionText="Figures 15–22: Trypanothacus barnesi gen. et sp. n. Figures 15, 20–22. Male holotype, metasoma V and telson lateral view (15), metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Figures. 16–19. Female paratype from type locality, metasoma V and telson lateral view (16), metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Scale bars: 4 mm (17–19, 20–21)." figureDoi="http://doi.org/10.5281/zenodo.6511479" httpUri="https://zenodo.org/record/6511479/files/figure.png" pageId="7" pageNumber="6">15–22</figureCitation>
, 
<figureCitation id="C88C2A64512AFF9E5ACDDD36FB5EF842" box="[1168,1237,1899,1923]" captionStart="Figures 29–46" captionStartId="12.[115,192,1871,1893]" captionTargetBox="[131,1463,242,1849]" captionTargetId="figure-164@12.[113,1471,234,1854]" captionTargetPageId="12" captionText="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi="http://doi.org/10.5281/zenodo.6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="7" pageNumber="6">29–42</figureCitation>
, 
<figureCitation id="C88C2A64512AFF9E5ABFDD36FAACF842" box="[1250,1319,1899,1923]" captionStart="Figures 71–77" captionStartId="15.[113,190,1918,1940]" captionTargetBox="[113,1460,257,1882]" captionTargetId="figure-8@15.[110,1474,236,1899]" captionTargetPageId="15" captionText="Figures 71–77: Trypanothacus barnesi gen. et sp. n. Figures 71–73. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. Figures 74–77. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77)." figureDoi="http://doi.org/10.5281/zenodo.6511493" httpUri="https://zenodo.org/record/6511493/files/figure.png" pageId="7" pageNumber="6">71–72</figureCitation>
, 
<figureCitation id="C88C2A64512AFF9E5B69DD36FAFCF842" box="[1332,1399,1899,1923]" captionStart-0="Figures 84–85" captionStart-1="Figures 86–89" captionStartId-0="17.[116,193,1988,2010]" captionStartId-1="18.[113,190,1969,1991]" captionTargetBox-0="[113,1474,248,1949]" captionTargetBox-1="[114,1474,236,1939]" captionTargetId-0="figure-35@17.[113,1474,248,1949]" captionTargetId-1="figure-12@18.[114,1474,234,1939]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figures 84–85: Trypanothacus barnesi gen. et sp. n., in vivo habitus, male holotype (84) and female paratype from type locality (85)." captionText-1="Figures 86–89: Trypanothacus barnesi gen. et sp. n.. Figures 86–87. Male (86) and female (87) from Thumrait, photos by J. N. Barnes. Figures 88–89. Type locality, Oman, S of Schelim, 18.041070°N 55.612262°E." figureDoi-0="http://doi.org/10.5281/zenodo.6511498" figureDoi-1="http://doi.org/10.5281/zenodo.6511503" httpUri-0="https://zenodo.org/record/6511498/files/figure.png" httpUri-1="https://zenodo.org/record/6511503/files/figure.png" pageId="7" pageNumber="6">84–87</figureCitation>
). Base color yellow or yellowish brown to orange; chelicerae yellow with or without reticulation, dentition reddish to black; median eyes may be surrounded by moderate fuscosity, with weak fuscosity extending forward towards lateral eyes; articular condyles of pedipalp chela fingers and legs reddish brown; macrosetae of body and appendages dark reddish brown; denticles on pedipalp chela fingers and aculeus of telson dark reddish brown.
</paragraph>
<paragraph id="500836E15125FF915CF6DAB5FB60FC7D" pageId="8" pageNumber="7">
<table id="22B7C441512500665EFCDABAFAD2FC7D" box="[161,1369,231,956]" gridcols="4" gridrows="19" pageId="8" pageNumber="7">
<tr id="EE8734A3512500665EFCDABAFAD2FEC3" box="[161,1369,231,258]" gridrow="0" pageId="8" pageNumber="7" rowspan-0="1" rowspan-1="1">
<th id="AD565DDF512500665CF6DABAFC59FEC3" box="[683,978,231,258]" gridcol="2" gridrow="0" pageId="8" pageNumber="7">
<taxonomicName id="97B74D625125FF915CF6DAB5FCADFEC3" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[683,806,232,258]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35125FF915CF6DAB5FCADFEC3" bold="true" box="[683,806,232,258]" italics="true" pageId="8" pageNumber="7">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885125FF915D6ADABAFC59FEC0" box="[823,978,231,258]" pageId="8" pageNumber="7" rank="species">
<emphasis id="62C3EAF35125FF915D6ADABAFCE0FEC0" bold="true" box="[823,875,231,257]" pageId="8" pageNumber="7">gen.</emphasis>
et 
<emphasis id="62C3EAF35125FF915DD2DABAFC59FEC0" bold="true" box="[911,978,231,257]" pageId="8" pageNumber="7">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<th id="AD565DDF512500665A6FDABAFAD2FEC3" box="[1074,1369,231,258]" gridcol="3" gridrow="0" pageId="8" pageNumber="7">
<taxonomicName id="97B74D625125FF915A6FDAB5FB26FEC3" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[1074,1197,232,258]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35125FF915A6FDAB5FB26FEC3" bold="true" box="[1074,1197,232,258]" italics="true" pageId="8" pageNumber="7">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885125FF915AE2DABAFAD2FEC0" box="[1215,1369,231,258]" pageId="8" pageNumber="7" rank="species">
<emphasis id="62C3EAF35125FF915AE2DABAFB79FEC0" bold="true" box="[1215,1266,231,257]" pageId="8" pageNumber="7">gen.</emphasis>
et 
<emphasis id="62C3EAF35125FF915B4BDABAFAD2FEC0" bold="true" box="[1302,1369,231,257]" pageId="8" pageNumber="7">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
</tr>
<tr id="EE8734A3512500665EFCDB53FAD2FEE8" box="[161,1369,270,297]" gridrow="1" pageId="8" pageNumber="7" rowspan-1="1">
<th id="AD565DDF512500665EFCDB53FEFBFEE8" box="[161,368,270,297]" gridcol="0" gridrow="1" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915EFCDB4CFEFBFEE8" bold="true" box="[161,368,273,297]" pageId="8" pageNumber="7">Dimensions (MM)</emphasis>
</th>
<td id="AD565DDF512500665CF6DB53FC59FEE8" box="[683,978,270,297]" gridcol="2" gridrow="1" pageId="8" pageNumber="7">
♂ 
<emphasis id="62C3EAF35125FF915D4ADB53FC08FEE9" bold="true" box="[791,899,270,296]" pageId="8" pageNumber="7">holotype</emphasis>
</td>
<td id="AD565DDF512500665A6FDB53FAD2FEE8" box="[1074,1369,270,297]" gridcol="3" gridrow="1" pageId="8" pageNumber="7">
♀ 
<emphasis id="62C3EAF35125FF915AC1DB53FA86FEE9" bold="true" box="[1180,1293,270,296]" pageId="8" pageNumber="7">paratype</emphasis>
</td>
</tr>
<tr id="EE8734A3512500665EFCDB6AFAD2FE8E" box="[161,1369,311,335]" gridrow="2" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDB6AFEFBFE8E" box="[161,368,311,335]" gridcol="0" gridrow="2" pageId="8" pageNumber="7">Carapace</th>
<td id="AD565DDF512500665F81DB6AFDC9FE8E" box="[476,578,311,335]" gridcol="1" gridrow="2" pageId="8" pageNumber="7">L / W</td>
<td id="AD565DDF512500665CF6DB6AFC59FE8E" box="[683,978,311,335]" gridcol="2" gridrow="2" pageId="8" pageNumber="7">4.849 / 5.733</td>
<td id="AD565DDF512500665A6FDB6AFAD2FE8E" box="[1074,1369,311,335]" gridcol="3" gridrow="2" pageId="8" pageNumber="7">6.126 / 6.952</td>
</tr>
<tr id="EE8734A3512500665EFCDB00FAD2FEB7" box="[161,1369,349,374]" gridrow="3" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDB00FEFBFEB7" box="[161,368,349,374]" gridcol="0" gridrow="3" pageId="8" pageNumber="7">Mesosoma</th>
<td id="AD565DDF512500665F81DB00FDC9FEB7" box="[476,578,349,374]" gridcol="1" gridrow="3" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6DB00FC59FEB7" box="[683,978,349,374]" gridcol="2" gridrow="3" pageId="8" pageNumber="7">9.519</td>
<td id="AD565DDF512500665A6FDB00FAD2FEB7" box="[1074,1369,349,374]" gridcol="3" gridrow="3" pageId="8" pageNumber="7">21.43</td>
</tr>
<tr id="EE8734A3512500665EFCDBD9FAD2FE5C" box="[161,1369,388,413]" gridrow="4" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDBD9FEFBFE5C" box="[161,368,388,413]" gridcol="0" gridrow="4" pageId="8" pageNumber="7">Tergite VII</th>
<td id="AD565DDF512500665F81DBD9FDC9FE5C" box="[476,578,388,413]" gridcol="1" gridrow="4" pageId="8" pageNumber="7">L / W</td>
<td id="AD565DDF512500665CF6DBD9FC59FE5C" box="[683,978,388,413]" gridcol="2" gridrow="4" pageId="8" pageNumber="7">2.843 / 4.934</td>
<td id="AD565DDF512500665A6FDBD9FAD2FE5C" box="[1074,1369,388,413]" gridcol="3" gridrow="4" pageId="8" pageNumber="7">3.960 / 6.755</td>
</tr>
<tr id="EE8734A3512500665EFCDBF6FAD2FE05" box="[161,1369,427,452]" gridrow="5" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDBF6FEFBFE05" box="[161,368,427,452]" gridcol="0" gridrow="5" pageId="8" pageNumber="7">Metasoma + telson</th>
<td id="AD565DDF512500665F81DBF6FDC9FE05" box="[476,578,427,452]" gridcol="1" gridrow="5" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6DBF6FC59FE05" box="[683,978,427,452]" gridcol="2" gridrow="5" pageId="8" pageNumber="7">27.599</td>
<td id="AD565DDF512500665A6FDBF6FAD2FE05" box="[1074,1369,427,452]" gridcol="3" gridrow="5" pageId="8" pageNumber="7">29.225</td>
</tr>
<tr id="EE8734A3512500665EFCDB8CFAD2FE2B" box="[161,1369,465,490]" gridrow="6" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDB8CFEFBFE2B" box="[161,368,465,490]" gridcol="0" gridrow="6" pageId="8" pageNumber="7">Segment I</th>
<td id="AD565DDF512500665F81DB8CFDC9FE2B" box="[476,578,465,490]" gridcol="1" gridrow="6" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6DB8CFC59FE2B" box="[683,978,465,490]" gridcol="2" gridrow="6" pageId="8" pageNumber="7">3.701 / 3.252 / 2.793</td>
<td id="AD565DDF512500665A6FDB8CFAD2FE2B" box="[1074,1369,465,490]" gridcol="3" gridrow="6" pageId="8" pageNumber="7">3.758 / 3.702 / 3.197</td>
</tr>
<tr id="EE8734A3512500665EFCDBA5FAD2FDD0" box="[161,1369,504,529]" gridrow="7" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCDBA5FEFBFDD0" box="[161,368,504,529]" gridcol="0" gridrow="7" pageId="8" pageNumber="7">Segment II</th>
<td id="AD565DDF512500665F81DBA5FDC9FDD0" box="[476,578,504,529]" gridcol="1" gridrow="7" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6DBA5FC59FDD0" box="[683,978,504,529]" gridcol="2" gridrow="7" pageId="8" pageNumber="7">3.947 / 3.058 / 2.850</td>
<td id="AD565DDF512500665A6FDBA5FAD2FDD0" box="[1074,1369,504,529]" gridcol="3" gridrow="7" pageId="8" pageNumber="7">4.207 / 3.451 / 3.290</td>
</tr>
<tr id="EE8734A3512500665EFCD842FAD2FDF9" box="[161,1369,543,568]" gridrow="8" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD842FEFBFDF9" box="[161,368,543,568]" gridcol="0" gridrow="8" pageId="8" pageNumber="7">Segment III</th>
<td id="AD565DDF512500665F81D842FDC9FDF9" box="[476,578,543,568]" gridcol="1" gridrow="8" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6D842FC59FDF9" box="[683,978,543,568]" gridcol="2" gridrow="8" pageId="8" pageNumber="7">4.194 / 2.939 / 2.930</td>
<td id="AD565DDF512500665A6FD842FAD2FDF9" box="[1074,1369,543,568]" gridcol="3" gridrow="8" pageId="8" pageNumber="7">4.344 / 3.332 / 3.253</td>
</tr>
<tr id="EE8734A3512500665EFCD81BFAD2FD9E" box="[161,1369,582,607]" gridrow="9" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD81BFEFBFD9E" box="[161,368,582,607]" gridcol="0" gridrow="9" pageId="8" pageNumber="7">Segment IV</th>
<td id="AD565DDF512500665F81D81BFDC9FD9E" box="[476,578,582,607]" gridcol="1" gridrow="9" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6D81BFC59FD9E" box="[683,978,582,607]" gridcol="2" gridrow="9" pageId="8" pageNumber="7">5.120 / 2.791 / 2.530</td>
<td id="AD565DDF512500665A6FD81BFAD2FD9E" box="[1074,1369,582,607]" gridcol="3" gridrow="9" pageId="8" pageNumber="7">5.340 / 3.137 / 2.852</td>
</tr>
<tr id="EE8734A3512500665EFCD831FAD2FD44" box="[161,1369,620,645]" gridrow="10" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD831FEFBFD44" box="[161,368,620,645]" gridcol="0" gridrow="10" pageId="8" pageNumber="7">Segment V</th>
<td id="AD565DDF512500665F81D831FDC9FD44" box="[476,578,620,645]" gridcol="1" gridrow="10" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6D831FC59FD44" box="[683,978,620,645]" gridcol="2" gridrow="10" pageId="8" pageNumber="7">5.839 / 2.439 / 2.024</td>
<td id="AD565DDF512500665A6FD831FAD2FD44" box="[1074,1369,620,645]" gridcol="3" gridrow="10" pageId="8" pageNumber="7">6.098 / 2.915 / 2.394</td>
</tr>
<tr id="EE8734A3512500665EFCD8CEFAD2FD6D" box="[161,1369,659,684]" gridrow="11" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD8CEFEFBFD6D" box="[161,368,659,684]" gridcol="0" gridrow="11" pageId="8" pageNumber="7">Telson</th>
<td id="AD565DDF512500665F81D8CEFDC9FD6D" box="[476,578,659,684]" gridcol="1" gridrow="11" pageId="8" pageNumber="7">L / W / D</td>
<td id="AD565DDF512500665CF6D8CEFC59FD6D" box="[683,978,659,684]" gridcol="2" gridrow="11" pageId="8" pageNumber="7">4.798 / 1.983 / 2.069</td>
<td id="AD565DDF512500665A6FD8CEFAD2FD6D" box="[1074,1369,659,684]" gridcol="3" gridrow="11" pageId="8" pageNumber="7">5.478 / 2.612 / 2.383</td>
</tr>
<tr id="EE8734A3512500665EFCD8E7FAD2FD12" box="[161,1369,698,723]" gridrow="12" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD8E7FEFBFD12" box="[161,368,698,723]" gridcol="0" gridrow="12" pageId="8" pageNumber="7">Pedipalp</th>
<td id="AD565DDF512500665F81D8E7FDC9FD12" box="[476,578,698,723]" gridcol="1" gridrow="12" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6D8E7FC59FD12" box="[683,978,698,723]" gridcol="2" gridrow="12" pageId="8" pageNumber="7">16.036</td>
<td id="AD565DDF512500665A6FD8E7FAD2FD12" box="[1074,1369,698,723]" gridcol="3" gridrow="12" pageId="8" pageNumber="7">15.891</td>
</tr>
<tr id="EE8734A3512500665EFCD8BCFAD2FD38" box="[161,1369,737,761]" gridrow="13" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD8BCFEFBFD38" box="[161,368,737,761]" gridcol="0" gridrow="13" pageId="8" pageNumber="7">Femur</th>
<td id="AD565DDF512500665F81D8BCFDC9FD38" box="[476,578,737,761]" gridcol="1" gridrow="13" pageId="8" pageNumber="7">L / W</td>
<td id="AD565DDF512500665CF6D8BCFC59FD38" box="[683,978,737,761]" gridcol="2" gridrow="13" pageId="8" pageNumber="7">4.136 / 1.354</td>
<td id="AD565DDF512500665A6FD8BCFAD2FD38" box="[1074,1369,737,761]" gridcol="3" gridrow="13" pageId="8" pageNumber="7">4.001 / 1.600</td>
</tr>
<tr id="EE8734A3512500665EFCD95AFAD2FCE1" box="[161,1369,775,800]" gridrow="14" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD95AFEFBFCE1" box="[161,368,775,800]" gridcol="0" gridrow="14" pageId="8" pageNumber="7">Patella</th>
<td id="AD565DDF512500665F81D95AFDC9FCE1" box="[476,578,775,800]" gridcol="1" gridrow="14" pageId="8" pageNumber="7">L / W</td>
<td id="AD565DDF512500665CF6D95AFC59FCE1" box="[683,978,775,800]" gridcol="2" gridrow="14" pageId="8" pageNumber="7">4.733 / 1.807</td>
<td id="AD565DDF512500665A6FD95AFAD2FCE1" box="[1074,1369,775,800]" gridcol="3" gridrow="14" pageId="8" pageNumber="7">4.743 / 1.948</td>
</tr>
<tr id="EE8734A3512500665EFCD973FAD2FC86" box="[161,1369,814,839]" gridrow="15" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD973FEFBFC86" box="[161,368,814,839]" gridcol="0" gridrow="15" pageId="8" pageNumber="7">Chela</th>
<td id="AD565DDF512500665F81D973FDC9FC86" box="[476,578,814,839]" gridcol="1" gridrow="15" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6D973FC59FC86" box="[683,978,814,839]" gridcol="2" gridrow="15" pageId="8" pageNumber="7">7.167</td>
<td id="AD565DDF512500665A6FD973FAD2FC86" box="[1074,1369,814,839]" gridcol="3" gridrow="15" pageId="8" pageNumber="7">7.147</td>
</tr>
<tr id="EE8734A3512500665EFCD908FAD2FCAF" box="[161,1369,853,878]" gridrow="16" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD908FEFBFCAF" box="[161,368,853,878]" gridcol="0" gridrow="16" pageId="8" pageNumber="7">Manus</th>
<td id="AD565DDF512500665F81D908FDC9FCAF" box="[476,578,853,878]" gridcol="1" gridrow="16" pageId="8" pageNumber="7">W / D</td>
<td id="AD565DDF512500665CF6D908FC59FCAF" box="[683,978,853,878]" gridcol="2" gridrow="16" pageId="8" pageNumber="7">1.797 / 1.901</td>
<td id="AD565DDF512500665A6FD908FAD2FCAF" box="[1074,1369,853,878]" gridcol="3" gridrow="16" pageId="8" pageNumber="7">1.600 / 1.836</td>
</tr>
<tr id="EE8734A3512500665EFCD926FAD2FC55" box="[161,1369,891,916]" gridrow="17" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD926FEFBFC55" box="[161,368,891,916]" gridcol="0" gridrow="17" pageId="8" pageNumber="7">Movable finger</th>
<td id="AD565DDF512500665F81D926FDC9FC55" box="[476,578,891,916]" gridcol="1" gridrow="17" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6D926FC59FC55" box="[683,978,891,916]" gridcol="2" gridrow="17" pageId="8" pageNumber="7">3.929</td>
<td id="AD565DDF512500665A6FD926FAD2FC55" box="[1074,1369,891,916]" gridcol="3" gridrow="17" pageId="8" pageNumber="7">4.166</td>
</tr>
<tr id="EE8734A3512500665EFCD9F9FAD2FC7D" box="[161,1369,932,956]" gridrow="18" pageId="8" pageNumber="7">
<th id="AD565DDF512500665EFCD9F9FEFBFC7D" box="[161,368,932,956]" gridcol="0" gridrow="18" pageId="8" pageNumber="7">Total</th>
<td id="AD565DDF512500665F81D9F9FDC9FC7D" box="[476,578,932,956]" gridcol="1" gridrow="18" pageId="8" pageNumber="7">L</td>
<td id="AD565DDF512500665CF6D9F9FC59FC7D" box="[683,978,932,956]" gridcol="2" gridrow="18" pageId="8" pageNumber="7">41.967</td>
<td id="AD565DDF512500665A6FD9F9FAD2FC7D" box="[1074,1369,932,956]" gridcol="3" gridrow="18" pageId="8" pageNumber="7">56.781</td>
</tr>
</table>
</paragraph>
<caption id="04C866695125FF915E2DD986FD87FBCF" ID-Table-UUID="04C866695125FF915E2DD986FD87FBCF" httpUri="http://table.plazi.org/id/04C866695125FF915E2DD986FD87FBCF" pageId="8" pageNumber="7" startId="8.[112,168,987,1008]" targetBox="[161,1369,231,956]" targetIsTable="true" targetPageId="8">
<paragraph id="500836E15125FF915E2DD986FD87FBCF" blockId="8.[112,1432,987,1038]" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915E2DD986FF30FC31" bold="true" box="[112,187,987,1008]" pageId="8" pageNumber="7">Table 1</emphasis>
. Comparative measurements of adults of 
<taxonomicName id="97B74D625125FF915C11D986FCA3FC31" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[588,808,987,1008]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35125FF915C11D986FCA3FC31" box="[588,808,987,1008]" italics="true" pageId="8" pageNumber="7">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885125FF915D73D986FC27FC31" box="[814,940,987,1008]" pageId="8" pageNumber="7" rank="species">
<emphasis id="62C3EAF35125FF915D73D986FCD9FC31" bold="true" box="[814,850,987,1008]" pageId="8" pageNumber="7">gen</emphasis>
. et 
<emphasis id="62C3EAF35125FF915D2BD986FC06FC31" bold="true" box="[886,909,987,1008]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="62C3EAF35125FF915DC5D986FC2EFC31" bold="true" box="[920,933,987,1008]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<paragraph id="500836E15125FF915E2CDE93FE94F947" blockId="8.[113,772,1070,2022]" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915E2CDE93FF54FB27" bold="true" box="[113,223,1230,1254]" pageId="8" pageNumber="7">Carapace</emphasis>
(
<figureCitation id="C88C2A645125FF915EAFDE93FED7FB27" box="[242,348,1230,1254]" captionStart="Figures 5–10" captionStartId="5.[113,190,1457,1479]" captionTargetBox="[113,1474,223,1433]" captionTargetId="figure-291@5.[113,1474,223,1449]" captionTargetPageId="5" captionText="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." figureDoi="http://doi.org/10.5281/zenodo.6511467" httpUri="https://zenodo.org/record/6511467/files/figure.png" pageId="8" pageNumber="7">Figs. 5–6</figureCitation>
, 
<figureCitation id="C88C2A645125FF915F36DE93FE0DFB27" box="[363,390,1230,1254]" captionStart="Figures 11–12" captionStartId="7.[116,193,1657,1679]" captionTargetBox="[116,1473,234,1632]" captionTargetId="figure-143@7.[114,1475,234,1632]" captionTargetPageId="7" captionText="Figures 11–12: Trypanothacus barnesi gen. et sp. n. Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6511473" httpUri="https://zenodo.org/record/6511473/files/figure.png" pageId="8" pageNumber="7">11</figureCitation>
, 
<figureCitation id="C88C2A645125FF915FCBDE93FE38FB27" box="[406,435,1230,1254]" captionStart="Figures 13–14" captionStartId="9.[113,190,1589,1611]" captionTargetBox="[113,1474,223,1572]" captionTargetId="figure-7@9.[113,1474,223,1574]" captionTargetPageId="9" captionText="Figures 13–14: Trypanothacus barnesi gen. et sp. n. Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6511477" httpUri="https://zenodo.org/record/6511477/files/figure.png" pageId="8" pageNumber="7">13</figureCitation>
). Trapezoidal, anterior margin straight or almost straight, with weak fine denticulation, 6–12 marginal macrosetae, bordered with row of coarse granules; surface with sparse to moderately dense coarse and fine granulation; anterior median carinae indicated by coarse granules, other carinae of carapace obsolete; lateral areas of interocular triangle with coarse granules, area between anterior median carinae smooth; median ocular tubercle smooth, except for a few posterior granules; median eyes large, well separated; posterolateral areas of interocular triangle, median postocular area and posterior marginal furrow smooth; posterior margin bordered by row of coarse granules; 5 lateral eyes (3 larger, 2 smaller; in ‘ 
<typeStatus id="8F0C88435125FF915FF9DC13FE5CF9A7" box="[420,471,1614,1638]" pageId="8" pageNumber="7">type</typeStatus>
5’ configuration of 
<bibRefCitation id="34264B105125FF915CF0DC13FE98F947" author="Loria &amp; Prendini" firstAuthor="Loria" pageId="8" pageNumber="7" pagination="112913" refId="ref14951" refString="LORIA, S. F. &amp; L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9 (12): e 112913. doi: 10.1371 / journal. pone. 0112913." type="journal article" year="2014">Loria &amp; Prendini, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="500836E15125FF915E2CDCD3FE24F807" blockId="8.[113,772,1070,2022]" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915E2CDCD3FF54F967" bold="true" box="[113,223,1678,1702]" pageId="8" pageNumber="7">Chelicera</emphasis>
(
<figureCitation id="C88C2A645125FF915EB0DCD3FEE4F967" box="[237,367,1678,1702]" captionStart="Figures 71–77" captionStartId="15.[113,190,1918,1940]" captionTargetBox="[113,1460,257,1882]" captionTargetId="figure-8@15.[110,1474,236,1899]" captionTargetPageId="15" captionText="Figures 71–77: Trypanothacus barnesi gen. et sp. n. Figures 71–73. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. Figures 74–77. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77)." figureDoi="http://doi.org/10.5281/zenodo.6511493" httpUri="https://zenodo.org/record/6511493/files/figure.png" pageId="8" pageNumber="7">Figs. 71–73</figureCitation>
). Fingers with typical buthid dentition (
<bibRefCitation id="34264B105125FF915E24DCF3FE9EF907" author="Vachon" box="[121,277,1710,1734]" firstAuthor="Vachon" pageId="8" pageNumber="7" pagination="161 - 166" refId="ref15851" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
); fixed finger with large distal denticle, one subdistal denticle and two basal denticles fused into bicusp, two denticles on ventral surface, one at level of bicusp, other slightly proximal to subdistal denticle; dorsal margin of movable finger with 5 denticles: one large distal denticle, medium-sized subdistal denticle, large medial denticle, and two small, partially fused basal denticles; ventral margin with 3 denticles: one large distal denticle, and two smaller denticles in medial and basal positions.
</paragraph>
<paragraph id="500836E15125FF915E2CDD93FC1EF867" blockId="8.[113,772,1070,2022]" lastBlockId="8.[816,1475,1070,2022]" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915E2CDD93FF62F827" bold="true" box="[113,233,1998,2022]" pageId="8" pageNumber="7">Mesosoma</emphasis>
(
<figureCitation id="C88C2A645125FF915EA7DD93FEE9F827" box="[250,354,1998,2022]" captionStart="Figures 1–4" captionStartId="3.[113,190,1967,1989]" captionTargetBox="[132,1458,266,1903]" captionTargetId="figure-7@3.[113,1474,253,1912]" captionTargetPageId="3" captionText="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511465" httpUri="https://zenodo.org/record/6511465/files/figure.png" pageId="8" pageNumber="7">Figs. 1–4</figureCitation>
, 
<figureCitation id="C88C2A645125FF915F33DD93FE12F827" box="[366,409,1998,2022]" captionStart="Figures 5–10" captionStartId="5.[113,190,1457,1479]" captionTargetBox="[113,1474,223,1433]" captionTargetId="figure-291@5.[113,1474,223,1449]" captionTargetPageId="5" captionText="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." figureDoi="http://doi.org/10.5281/zenodo.6511467" httpUri="https://zenodo.org/record/6511467/files/figure.png" pageId="8" pageNumber="7">6, 8</figureCitation>
, 
<figureCitation id="C88C2A645125FF915FFADD93FE61F827" box="[423,490,1998,2022]" captionStart-0="Figures 11–12" captionStart-1="Figures 13–14" captionStartId-0="7.[116,193,1657,1679]" captionStartId-1="9.[113,190,1589,1611]" captionTargetBox-0="[116,1473,234,1632]" captionTargetBox-1="[113,1474,223,1572]" captionTargetId-0="figure-143@7.[114,1475,234,1632]" captionTargetId-1="figure-7@9.[113,1474,223,1574]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="Figures 11–12: Trypanothacus barnesi gen. et sp. n. Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm." captionText-1="Figures 13–14: Trypanothacus barnesi gen. et sp. n. Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6511473" figureDoi-1="http://doi.org/10.5281/zenodo.6511477" httpUri-0="https://zenodo.org/record/6511473/files/figure.png" httpUri-1="https://zenodo.org/record/6511477/files/figure.png" pageId="8" pageNumber="7">11–14</figureCitation>
). 
<emphasis id="62C3EAF35125FF915C5DDD92FDDFF827" box="[512,596,1999,2022]" italics="true" pageId="8" pageNumber="7">Tergites</emphasis>
I–VI tricarinate with median carina and anteriorly diverging pair of lateral carinae; carinae coarsely granular, the lateral pair on the tergites I–II possibly indistinct; all carinae short with only ca. 2–5 granules, confined to posterior half or third of tergite; lateral flanks of tergites with sparse to moderately dense, coarse granulation on posterior half, posterior margins bordered with row of granules; other tergite surfaces smooth to faintly shagreened; tergite VII pentacarinate, median carina a granulated hump, lateral carinae well developed, coarsely granular; posterior margin of tergite VII without rim of granules, intercarinal surfaces smooth or with sparse, coarse granulation; coxae mostly smooth, with few sparse isolated patches of granules, coxal margins smooth or with weak granulation; sternum subtriangular, smooth, with deep posteromedian pit; 
<emphasis id="62C3EAF35125FF915A1FDFB2FB17F9C7" box="[1090,1180,1519,1542]" italics="true" pageId="8" pageNumber="7">sternites</emphasis>
with smooth surfaces and posterior margins, III–VI without carinae, VII with 2 pairs of weak to moderate, smooth to granulated carinae; 
<emphasis id="62C3EAF35125FF915B25DC73FC28F9A7" italics="true" pageId="8" pageNumber="7">sternal chaetotaxy</emphasis>
: sternite III-VI posterior margins conspicuously setose, bearing approximately 20 macrosetae, III bearing about a dozen macrosetae on medial surface, IV-VII with 4 medial and 2 lateral macrosetae, VII with single anterior macroseta on each of the 4 carinae, plus a lateral pair; 
<emphasis id="62C3EAF35125FF915A81DC92FAB8F927" box="[1244,1331,1743,1766]" italics="true" pageId="8" pageNumber="7">pectines</emphasis>
with anterior margins extending to midpoint (
<specimenCount id="46B1FD685125FF915AF3DCB3FB48F8C7" box="[1198,1219,1774,1798]" pageId="8" pageNumber="7" type="male">♂</specimenCount>
) or to approximately proximal 1/6 (
<specimenCount id="46B1FD685125FF915D91DD53FC6BF8E4" box="[972,992,1806,1829]" pageId="8" pageNumber="7" type="female">♀</specimenCount>
) of trochanter IV, with 3 marginal lamellae, 4–9 middle lamellae; lamellae and fulcra bear numerous short, fine, dark macrosetae; pectine basal piece and genital opercula smooth with fine macrosetae; pectinal tooth counts, 
<specimenCount id="46B1FD685125FF915B03DD33FAF9F844" box="[1374,1394,1902,1925]" pageId="8" pageNumber="7" type="male">♂</specimenCount>
22–26, 
<specimenCount id="46B1FD685125FF915D6DDDD3FCCFF864" box="[816,836,1934,1957]" pageId="8" pageNumber="7" type="female">♀</specimenCount>
17–21.
</paragraph>
<paragraph id="500836E15125FF905D6DDDF3FD29F83E" blockId="8.[816,1475,1070,2022]" lastBlockId="9.[113,772,1671,2047]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="62C3EAF35125FF915D6DDDF3FB92F807" bold="true" box="[816,1049,1966,1990]" pageId="8" pageNumber="7">Hemispermatophore</emphasis>
(
<figureCitation id="C88C2A645125FF915A7ADDF3FB21F807" box="[1063,1194,1966,1990]" captionStart="Figures 71–77" captionStartId="15.[113,190,1918,1940]" captionTargetBox="[113,1460,257,1882]" captionTargetId="figure-8@15.[110,1474,236,1899]" captionTargetPageId="15" captionText="Figures 71–77: Trypanothacus barnesi gen. et sp. n. Figures 71–73. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. Figures 74–77. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77)." figureDoi="http://doi.org/10.5281/zenodo.6511493" httpUri="https://zenodo.org/record/6511493/files/figure.png" pageId="8" pageNumber="7">Figs. 74–77</figureCitation>
). Flagelliform, moderately elongated, trunk ca. 6 times length of capsule region as measured from proximal flagellum base; flagellum well separated from capsule lobes, pars recta short, ca. 2.7 times length of capsule and lobes, 50% length of trunk, with weak fin along anterior margin; pars reflecta long, narrow, gradually tapered, hyaline, 0.88 times length of trunk; capsule region with 4 lobes arranged in ‘3 +1’ configuration, posterior lobe the largest, a lanceolate lamina with pointed apex; median lobe smallest, acuminate, attached to base of posterior lobe along short, suture, but a sclerotized carina not clearly visible; anterior lobe of intermediate length, laminate, apically acuminate; basal lobe a small, rounded knob-like process situated well proximal from point of splitting of posterior and median lobes.
</paragraph>
<caption id="04C866695124FF905E2CDC68FD29F9A9" ID-DOI="http://doi.org/10.5281/zenodo.6511477" ID-Zenodo-Dep="6511477" httpUri="https://zenodo.org/record/6511477/files/figure.png" pageId="9" pageNumber="8" startId="9.[113,190,1589,1611]" targetBox="[113,1474,223,1572]" targetPageId="9">
<paragraph id="500836E15124FF905E2CDC68FD29F9A9" blockId="9.[113,1474,1589,1641]" pageId="9" pageNumber="8">
<emphasis id="62C3EAF35124FF905E2CDC68FE8AF98B" bold="true" box="[113,257,1589,1611]" pageId="9" pageNumber="8">Figures 13–14</emphasis>
: 
<taxonomicName id="97B74D625124FF905F53DC6BFE61F98A" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[270,490,1590,1611]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35124FF905F53DC6BFE61F98A" box="[270,490,1590,1611]" italics="true" pageId="9" pageNumber="8">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885124FF905FACDC6BFDE4F98A" box="[497,623,1590,1611]" pageId="9" pageNumber="8" rank="species">
<emphasis id="62C3EAF35124FF905FACDC6BFD9EF98A" bold="true" box="[497,533,1590,1611]" pageId="9" pageNumber="8">gen</emphasis>
. et 
<emphasis id="62C3EAF35124FF905C64DC6BFDDBF98A" bold="true" box="[569,592,1590,1611]" pageId="9" pageNumber="8">sp</emphasis>
. 
<emphasis id="62C3EAF35124FF905C01DC6BFDE2F98A" bold="true" box="[604,617,1590,1611]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="500836E15124FF8A5D6DDCDAFD6DF830" blockId="9.[816,1475,1671,2047]" lastBlockId="19.[113,772,1305,2033]" lastPageId="19" lastPageNumber="18" pageId="9" pageNumber="8">
<emphasis id="62C3EAF35124FF905D6DDCDAFBA1F95E" bold="true" box="[816,1066,1671,1695]" pageId="9" pageNumber="8">Metasoma and telson</emphasis>
(
<figureCitation id="C88C2A645124FF905A62DCDAFB4EF95E" box="[1087,1221,1671,1695]" captionStart-0="Figures 15–22" captionStart-1="Figures 23–28" captionStartId-0="10.[113,190,1958,1980]" captionStartId-1="11.[116,193,1958,1980]" captionTargetBox-0="[120,1471,258,1929]" captionTargetBox-1="[113,1474,234,1927]" captionTargetId-0="figure-11@10.[113,1474,234,1934]" captionTargetId-1="figure-69@11.[111,1474,234,1930]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figures 15–22: Trypanothacus barnesi gen. et sp. n. Figures 15, 20–22. Male holotype, metasoma V and telson lateral view (15), metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Figures. 16–19. Female paratype from type locality, metasoma V and telson lateral view (16), metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Scale bars: 4 mm (17–19, 20–21)." captionText-1="Figures 23–28: Trypanothacus barnesi gen. et sp. n. Figures 23–25. Male paratype from Thumrait (VII.1997), metasoma and telson dorsal (23), lateral (24) and ventral (25) views. Figures 26–28. Female paratype from Yalooni (XII.1988), metasoma and telson dorsal (26), lateral (27) and ventral (28) views. UV fluorescence. Scale bar: 4 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6511479" figureDoi-1="http://doi.org/10.5281/zenodo.6511481" httpUri-0="https://zenodo.org/record/6511479/files/figure.png" httpUri-1="https://zenodo.org/record/6511481/files/figure.png" pageId="9" pageNumber="8">Figs. 15–28</figureCitation>
). 
<emphasis id="62C3EAF35124FF905A80DCD5FAC1F95E" box="[1245,1354,1672,1695]" italics="true" pageId="9" pageNumber="8">Metasoma</emphasis>
I with 10 granulated or crenulated carinae, median lateral carinae complete, ventromedian carinae may be weak or obsolete in males; II–III with 10 granulated or crenulated carinae, median lateral carinae incomplete, indicated by ca. 5–14 granules on posterior 1/3 to 2/3 of segment; ventromedian and ventrolateral carinae on II–III stronger in females, with conspicuously enlarged, dentate or lobate granules, increasing in size posteriorly; coarsely granulose anterior ventral margins present on III–IV; IV with 8 granulated or crenulated carinae; V with 5 carinae, dorsolateral carinae weak, granulated, ventrolateral carinae strong, irregularly crenulated with several enlarged, lobate granules that become larger posteriorly; ventromedian carina of V composed of series of large granules interspersed with finer granulation; intercarinal surfaces of I–IV smooth or almost smooth dorsally, almost smooth or with sparse granules dorsolaterally and laterally; ventrolateral and ventral surfaces of I smooth, of II–III almost smooth with sparse fine granulation, of IV with weak, fine granulation or shagreened; segment V smooth dorsally with scattered small granules, with weak fine granulation laterally, slightly denser fine granulation ventrally; lateral anal arch divided into 2–3 lobes; ventral anal arch armed with regular series of ca. 5–7 coarse granules; 
<emphasis id="62C3EAF3513EFF8A5C99DC04FC89F9B0" box="[708,770,1625,1649]" italics="true" pageId="19" pageNumber="18">telson</emphasis>
with distinctly bulbous vesicle bearing coarse granules on ventral surface; granulation more dense in males, more sparse in females, with smooth areas; lateral surface of vesicle with sparser granulation, dorsal surface smooth; aculeus equal to or shorter than vesicle in length, moderately curved; subaculear tubercle absent; 
<emphasis id="62C3EAF3513EFF8A5F7BDD44FE1CF8F0" box="[294,407,1817,1841]" italics="true" pageId="19" pageNumber="18">chaetotaxy</emphasis>
: metasomal segments and telson sparsely setose; long macrosetae located along carinae on metasoma I–IV, typically 2–3 per carina; V with several setae along dorsolateral carina, ca. 6 setae in longitudinal series on lower lateral surface, several setae on ventral surface associated with enlarged granules; telson vesicle with a dozen or more long macrosetae scattered over ventral and ventrolateral surfaces.
</paragraph>
<caption id="04C866695127FF935E2CDDFBFB35F837" ID-DOI="http://doi.org/10.5281/zenodo.6511479" ID-Zenodo-Dep="6511479" httpUri="https://zenodo.org/record/6511479/files/figure.png" pageId="10" pageNumber="9" startId="10.[113,190,1958,1980]" targetBox="[120,1471,258,1929]" targetPageId="10">
<paragraph id="500836E15127FF935E2CDDFBFB35F837" blockId="10.[113,1474,1958,2038]" pageId="10" pageNumber="9">
<emphasis id="62C3EAF35127FF935E2CDDFBFE8AF87A" bold="true" box="[113,257,1958,1980]" pageId="10" pageNumber="9">Figures 15–22</emphasis>
: 
<taxonomicName id="97B74D625127FF935F53DDFBFE60F87A" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[270,491,1958,1979]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35127FF935F53DDFBFE60F87A" box="[270,491,1958,1979]" italics="true" pageId="10" pageNumber="9">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885127FF935FACDDFBFDE4F87D" box="[497,623,1958,1980]" pageId="10" pageNumber="9" rank="species">
<emphasis id="62C3EAF35127FF935FACDDFBFD9EF87A" bold="true" box="[497,533,1958,1979]" pageId="10" pageNumber="9">gen</emphasis>
. et 
<emphasis id="62C3EAF35127FF935C64DDFBFDDBF87A" bold="true" box="[569,592,1958,1979]" pageId="10" pageNumber="9">sp</emphasis>
. 
<emphasis id="62C3EAF35127FF935C01DDFBFDE2F87A" bold="true" box="[604,617,1958,1979]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="62C3EAF35127FF935C2BDDFBFD6AF87A" bold="true" box="[630,737,1958,1980]" pageId="10" pageNumber="9">Figures 15</emphasis>
, 
<emphasis id="62C3EAF35127FF935CB0DDFBFCA2F87A" bold="true" box="[749,809,1958,1980]" pageId="10" pageNumber="9">20–22</emphasis>
. Male holotype, metasoma V and telson lateral view (15), metasoma and telson lateral (20), ventral (21), and dorsal (22) views. 
<emphasis id="62C3EAF35127FF935CFBDD9EFD78F818" bold="true" box="[678,755,1987,2009]" pageId="10" pageNumber="9">Figures</emphasis>
. 
<emphasis id="62C3EAF35127FF935CA3DD9EFCB1F819" bold="true" box="[766,826,1987,2008]" pageId="10" pageNumber="9">16–19</emphasis>
. Female paratype from type locality, metasoma V and telson lateral view (16), metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Scale bars: 4 mm (17–19, 20–21).
</paragraph>
</caption>
<caption id="04C866695126FF925E29DDFBFD5EF834" ID-DOI="http://doi.org/10.5281/zenodo.6511481" ID-Zenodo-Dep="6511481" httpUri="https://zenodo.org/record/6511481/files/figure.png" pageId="11" pageNumber="10" startId="11.[116,193,1958,1980]" targetBox="[113,1474,234,1927]" targetPageId="11">
<paragraph id="500836E15126FF925E29DDFBFD5EF834" blockId="11.[116,1474,1958,2038]" pageId="11" pageNumber="10">
<emphasis id="62C3EAF35126FF925E29DDFBFE8FF87A" bold="true" box="[116,260,1958,1980]" pageId="11" pageNumber="10">Figures 23–28</emphasis>
: 
<taxonomicName id="97B74D625126FF925F4CDDFBFE64F87A" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[273,495,1958,1979]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35126FF925F4CDDFBFE64F87A" box="[273,495,1958,1979]" italics="true" pageId="11" pageNumber="10">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885126FF925FABDDFBFDFDF87D" box="[502,630,1958,1980]" pageId="11" pageNumber="10" rank="species">
<emphasis id="62C3EAF35126FF925FABDDFBFD91F87A" bold="true" box="[502,538,1958,1979]" pageId="11" pageNumber="10">gen</emphasis>
. et 
<emphasis id="62C3EAF35126FF925C62DDFBFDDDF87A" bold="true" box="[575,598,1958,1979]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="62C3EAF35126FF925C3EDDFBFDFBF87A" bold="true" box="[611,624,1958,1979]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="62C3EAF35126FF925C20DDFBFC86F87A" bold="true" box="[637,781,1958,1980]" pageId="11" pageNumber="10">Figures 23–25</emphasis>
. Male paratype from Thumrait (VII.1997), metasoma and telson dorsal (23), lateral (24) and ventral (25) views. 
<emphasis id="62C3EAF35126FF925C5CDD9EFD1BF819" bold="true" box="[513,656,1987,2009]" pageId="11" pageNumber="10">Figures 26–28</emphasis>
. Female paratype from Yalooni (XII.1988), metasoma and telson dorsal (26), lateral (27) and ventral (28) views. UV fluorescence. Scale bar: 4 mm.
</paragraph>
</caption>
<caption id="04C866695121FF955E2EDD12FA33F834" ID-DOI="http://doi.org/10.5281/zenodo.6511485" ID-Zenodo-Dep="6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="12" pageNumber="11" startId="12.[115,192,1871,1893]" targetBox="[131,1463,242,1849]" targetPageId="12">
<paragraph id="500836E15121FF955E2EDD12FA33F834" blockId="12.[115,1473,1871,2037]" pageId="12" pageNumber="11">
<emphasis id="62C3EAF35121FF955E2EDD12FE8AF8A5" bold="true" box="[115,257,1871,1893]" pageId="12" pageNumber="11">Figures 29–46</emphasis>
: 
<emphasis id="62C3EAF35121FF955F50DD12FE10F8A5" bold="true" box="[269,411,1871,1893]" pageId="12" pageNumber="11">Figures 29–42</emphasis>
: 
<taxonomicName id="97B74D625121FF955FFADD12FD08F8A5" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[423,643,1871,1892]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35121FF955FFADD12FD08F8A5" box="[423,643,1871,1892]" italics="true" pageId="12" pageNumber="11">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885121FF955CD5DD12FC88F8A4" box="[648,771,1871,1893]" pageId="12" pageNumber="11" rank="species">
<emphasis id="62C3EAF35121FF955CD5DD12FD27F8A5" bold="true" box="[648,684,1871,1892]" pageId="12" pageNumber="11">gen</emphasis>
. et 
<emphasis id="62C3EAF35121FF955C93DD12FD6EF8A5" bold="true" box="[718,741,1871,1892]" pageId="12" pageNumber="11">sp</emphasis>
. 
<emphasis id="62C3EAF35121FF955CB2DD12FD77F8A5" bold="true" box="[751,764,1871,1892]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="62C3EAF35121FF955D55DD12FC1DF8A5" bold="true" box="[776,918,1871,1893]" pageId="12" pageNumber="11">Figures 29–40</emphasis>
. Pedipalp segments. 
<emphasis id="62C3EAF35121FF955A02DD12FB66F8A5" bold="true" box="[1119,1261,1871,1893]" pageId="12" pageNumber="11">Figures 29–38</emphasis>
. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. 
<emphasis id="62C3EAF35121FF955B16DDD4FF00F87B" bold="true" pageId="12" pageNumber="11">Figures 39– 40</emphasis>
. Female paratype from the type locality, chela dorsal (39) and external (40) views. 
<emphasis id="62C3EAF35121FF955DECDDF8FBCBF87B" bold="true" box="[945,1088,1957,1979]" pageId="12" pageNumber="11">Figures 41–42</emphasis>
. Telson, male holotype (41) and female paratype from type locality (42), lateral views. 
<emphasis id="62C3EAF35121FF955C63DD9FFD45F816" bold="true" box="[574,718,1986,2008]" pageId="12" pageNumber="11">Figures 43–44</emphasis>
: 
<taxonomicName id="97B74D625121FF955C86DD9EFCDBF816" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" baseAuthorityYear="2019" box="[731,848,1986,2008]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF35121FF955C86DD9EFCDBF816" box="[731,848,1986,2008]" italics="true" pageId="12" pageNumber="11">T. buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885121FF955D0ADD9EFC25F819" box="[855,942,1987,2008]" pageId="12" pageNumber="11" rank="species">
<emphasis id="62C3EAF35121FF955D0ADD9EFC04F819" bold="true" box="[855,911,1987,2008]" pageId="12" pageNumber="11">comb</emphasis>
. 
<emphasis id="62C3EAF35121FF955DC6DD9EFC23F819" bold="true" box="[923,936,1987,2008]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, pedipalp chela dorsal, male holotype (43) and female paratype (44). 
<emphasis id="62C3EAF35121FF955EA2DD82FE05F835" bold="true" box="[255,398,2015,2037]" pageId="12" pageNumber="11">Figures 45–46</emphasis>
: 
<taxonomicName id="97B74D625121FF955FC6DD82FCA4F834" authority="Kovarik et al., 2016" authorityName="Kovarik, Lowe et Stahlavsky" authorityYear="2016" box="[411,815,2015,2037]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="stockmanni">
<emphasis id="62C3EAF35121FF955FC6DD82FDECF835" box="[411,615,2015,2036]" italics="true" pageId="12" pageNumber="11">Buthacus stockmanni</emphasis>
<bibRefCitation id="34264B105121FF955C30DD82FCA4F834" author="KOVARIK, F. &amp; G. LOWE &amp; K. B. RANAWANA &amp; D. HOFEREK &amp; V. A. SANJEEWA JAYARATHNE &amp; J. PLISKOVA &amp; F. STAHLAVSKY" box="[621,815,2015,2037]" journalOrPublisher="Euscorpius" pageId="12" pageNumber="11" pagination="1 - 133" part="220" refId="ref14587" refString="KOVARIK, F., G. LOWE, K. B. RANAWANA, D. HOFEREK, V. A. SANJEEWA JAYARATHNE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016 a. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n .. Euscorpius, 220: 1 - 133." title="Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n .." type="journal article" year="2016">Kovařík et al., 2016</bibRefCitation>
</taxonomicName>
, telson, male holotype (45) and female paratype (46), lateral views.
</paragraph>
</caption>
<caption id="04C866695120FF945E2CDDFAFC56F837" ID-DOI="http://doi.org/10.5281/zenodo.6511487" ID-Zenodo-Dep="6511487" httpUri="https://zenodo.org/record/6511487/files/figure.png" pageId="13" pageNumber="12" startId="13.[113,190,1959,1981]" targetBox="[113,1474,234,1930]" targetPageId="13">
<paragraph id="500836E15120FF945E2CDDFAFC56F837" blockId="13.[113,1474,1959,2038]" pageId="13" pageNumber="12">
<emphasis id="62C3EAF35120FF945E2CDDFAFF74F87D" bold="true" box="[113,255,1959,1981]" pageId="13" pageNumber="12">Figures 47–58</emphasis>
: 
<taxonomicName id="97B74D625120FF945F56DDFAFE6CF87D" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[267,487,1959,1980]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35120FF945F56DDFAFE6CF87D" box="[267,487,1959,1980]" italics="true" pageId="13" pageNumber="12">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885120FF945FB1DDFAFDEDF87D" box="[492,614,1959,1980]" pageId="13" pageNumber="12" rank="species">
<emphasis id="62C3EAF35120FF945FB1DDFAFD9BF87D" bold="true" box="[492,528,1959,1980]" pageId="13" pageNumber="12">gen</emphasis>
. et 
<emphasis id="62C3EAF35120FF945C6CDDFAFDC3F87D" bold="true" box="[561,584,1959,1980]" pageId="13" pageNumber="12">sp</emphasis>
. 
<emphasis id="62C3EAF35120FF945C0EDDFAFDEBF87D" bold="true" box="[595,608,1959,1980]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
Pedipalp segments of male paratype from Thumrait (VII.1997), chela dorsal (47), external (48), ventral (49) and internal (50) views. Patella dorsal (51), external (52), ventral (53) and internal (54) views. Femur and trochanter dorsal (55), external (56), ventral (57) and internal (58) views. UV fluorescence. Scale bar: 2 mm.
</paragraph>
</caption>
<caption id="04C866695123FF975E29DD2EFB12F83D" ID-DOI="http://doi.org/10.5281/zenodo.6511489" ID-Zenodo-Dep="6511489" httpUri="https://zenodo.org/record/6511489/files/figure.png" pageId="14" pageNumber="13" startId="14.[116,193,1907,1929]" targetBox="[124,1469,237,1889]" targetPageId="14">
<paragraph id="500836E15123FF975E29DD2EFB12F83D" blockId="14.[116,1477,1907,2044]" pageId="14" pageNumber="13">
<emphasis id="62C3EAF35123FF975E29DD2EFE88F849" bold="true" box="[116,259,1907,1929]" pageId="14" pageNumber="13">Figures 59–70</emphasis>
: 
<taxonomicName id="97B74D625123FF975F52DD2EFE67F849" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[271,492,1907,1928]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35123FF975F52DD2EFE67F849" box="[271,492,1907,1928]" italics="true" pageId="14" pageNumber="13">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885123FF975FAFDD2EFDE4F848" box="[498,623,1907,1929]" pageId="14" pageNumber="13" rank="species">
<emphasis id="62C3EAF35123FF975FAFDD2EFD9DF849" bold="true" box="[498,534,1907,1928]" pageId="14" pageNumber="13">gen</emphasis>
. et 
<emphasis id="62C3EAF35123FF975C64DD2EFDDBF849" bold="true" box="[569,592,1907,1928]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="62C3EAF35123FF975C01DD2EFDE2F849" bold="true" box="[604,617,1907,1928]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
Pedipalp segments of female paratypes, chela dorsal (59), external (60), ventral (61) and internal (62) views. Patella dorsal (63), external (64), ventral (65) and internal (66) views. Femur and trochanter dorsal (67), external (68), ventral (69) and internal (70) views. Dorsal views (59, 63, 67): paratype female from Thumrait (XI.1989) (with normal dorsointernal carina on femur). External (60, 64, 68), ventral (61, 65, 69) and internal (62, 66, 70) views: paratype female from Yalooni (XII.1988) (with malformed dorsointernal carina on femur). UV fluorescence. Scale bars: 2 mm (59, 63, 67), 2 mm (60–62, 64–66, 69–70).
</paragraph>
</caption>
<caption id="04C866695122FF965E2CDD23FEBEF82A" ID-DOI="http://doi.org/10.5281/zenodo.6511493" ID-Zenodo-Dep="6511493" httpUri="https://zenodo.org/record/6511493/files/figure.png" pageId="15" pageNumber="14" startId="15.[113,190,1918,1940]" targetBox="[113,1460,257,1882]" targetPageId="15">
<paragraph id="500836E15122FF965E2CDD23FEBEF82A" blockId="15.[113,1472,1918,2027]" pageId="15" pageNumber="14">
<emphasis id="62C3EAF35122FF965E2CDD23FE8AF855" bold="true" box="[113,257,1918,1940]" pageId="15" pageNumber="14">Figures 71–77</emphasis>
: 
<taxonomicName id="97B74D625122FF965F53DD23FE60F852" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[270,491,1918,1939]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF35122FF965F53DD23FE60F852" box="[270,491,1918,1939]" italics="true" pageId="15" pageNumber="14">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885122FF965FACDD22FDFBF855" box="[497,624,1919,1940]" pageId="15" pageNumber="14" rank="species">
<emphasis id="62C3EAF35122FF965FACDD22FD9EF855" bold="true" box="[497,533,1919,1940]" pageId="15" pageNumber="14">gen</emphasis>
. et 
<emphasis id="62C3EAF35122FF965C64DD22FDDBF855" bold="true" box="[569,592,1919,1940]" pageId="15" pageNumber="14">sp</emphasis>
. 
<emphasis id="62C3EAF35122FF965C00DD22FDE1F855" bold="true" box="[605,618,1919,1940]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="62C3EAF35122FF965C2BDD23FC8DF852" bold="true" box="[630,774,1918,1940]" pageId="15" pageNumber="14">Figures 71–73</emphasis>
. Paratype male, right chelicera, dorsal (71), ventral (72) and ventral UV fluorescence (73) views. Scale bar: 1 mm. 
<emphasis id="62C3EAF35122FF965C59DDC6FD19F871" bold="true" box="[516,658,1947,1969]" pageId="15" pageNumber="14">Figures 74–77</emphasis>
. Holotype male, right hemispermatophore, capsule region with lobes in anterior (74), convex compressed (75) and posterior (76) views, and whole hemispermatophore in convex view (77). Scale bars: 1 mm (71–73), 500 µm (74–76), 1 mm (77).
</paragraph>
</caption>
<caption id="04C86669513DFF895E2CDD35FA21F830" ID-DOI="http://doi.org/10.5281/zenodo.6511496" ID-Zenodo-Dep="6511496" httpUri="https://zenodo.org/record/6511496/files/figure.png" pageId="16" pageNumber="15" startId="16.[113,188,1896,1918]" targetBox="[181,1394,252,1873]" targetPageId="16">
<paragraph id="500836E1513DFF895E2CDD35FA21F830" blockId="16.[112,1475,1896,2033]" pageId="16" pageNumber="15">
<emphasis id="62C3EAF3513DFF895E2CDD35FF77F8BC" bold="true" box="[113,252,1896,1918]" pageId="16" pageNumber="15">Figures 78–83</emphasis>
: Hemispermatophore capsule regions and lobes of buthid genera presumed to be closely related to 
<taxonomicName id="97B74D62513DFF895AC7DD35FAAFF8BC" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[1178,1316,1896,1917]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="62C3EAF3513DFF895AC7DD35FAAFF8BC" box="[1178,1316,1896,1917]" italics="true" pageId="16" pageNumber="15">Trypanothacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513DFF895B77DD35FAEDF8BC" box="[1322,1382,1896,1917]" pageId="16" pageNumber="15" rank="genus">
<emphasis id="62C3EAF3513DFF895B77DD35FAC6F8BC" bold="true" box="[1322,1357,1896,1917]" pageId="16" pageNumber="15">gen</emphasis>
. 
<emphasis id="62C3EAF3513DFF895B04DD35FAEDF8BC" bold="true" box="[1369,1382,1896,1917]" pageId="16" pageNumber="15">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="62C3EAF3513DFF895B2ADD35FF20F858" bold="true" pageId="16" pageNumber="15">Figures 78–79</emphasis>
: 
<taxonomicName id="97B74D62513DFF895EEBDDD8FD4EF85B" authority="Kovarik, Lowe et Stahlavsky, 2016" authorityName="Kovarik, Lowe et Stahlavsky" authorityYear="2016" box="[182,709,1924,1948]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="stockmanni">
<emphasis id="62C3EAF3513DFF895EEBDDD8FEF0F85B" box="[182,379,1925,1946]" italics="true" pageId="16" pageNumber="15">Buthacus stockmanni</emphasis>
<bibRefCitation id="34264B10513DFF895F22DDD8FD4EF85B" author="KOVARIK, F. &amp; G. LOWE &amp; F. STAHLAVSKY" box="[383,709,1924,1948]" journalOrPublisher="Euscorpius" pageId="16" pageNumber="15" pagination="1 - 18" part="236" refId="ref14669" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2016 b. Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae). Euscorpius, 236: 1 - 18." title="Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae)" type="journal article" year="2016">Kovařík, Lowe et Šťáhlavský, 2016</bibRefCitation>
</taxonomicName>
(topoparatype male, N. of Zag, Morocco). 
<emphasis id="62C3EAF3513DFF895A17DDD9FB59F858" bold="true" box="[1098,1234,1924,1946]" pageId="16" pageNumber="15">Figures 80–81</emphasis>
: 
<taxonomicName id="97B74D62513DFF895A80DDD8FE0DF877" authority="Levy, Amitai et Shulov, 1973" authorityName="Levy, Amitai et Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="nigroaculeatus">
<emphasis id="62C3EAF3513DFF895A80DDD8FA48F85B" box="[1245,1475,1925,1946]" italics="true" pageId="16" pageNumber="15">Buthacus nigroaculeatus</emphasis>
<bibRefCitation id="34264B10513DFF895E2DDDFFFE0DF877" author="LEVY, G. &amp; P. AMITAI &amp; A. SHULOV" box="[112,390,1953,1975]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="16" pageNumber="15" pagination="113 - 140" part="52" refId="ref14867" refString="LEVY, G., P. AMITAI &amp; A. SHULOV. 1973. New scorpions from Israel, Jordan and Arabia. Zoological Journal of the Linnean Society, 52 (2): 113 - 140." title="New scorpions from Israel, Jordan and Arabia" type="journal article" year="1973">Levy, Amitai et Shulov, 1973</bibRefCitation>
</taxonomicName>
(Wadi Muqshin, Montesar, Oman, 30.III.1989, leg. A.S. Gardner). 
<emphasis id="62C3EAF3513DFF895A5ADDFCFB18F877" bold="true" box="[1031,1171,1953,1975]" pageId="16" pageNumber="15">Figures 82–83</emphasis>
: 
<taxonomicName id="97B74D62513DFF895AFCDDFCFEC6F812" authority="Levy, Amitai et Shulov, 1973" authorityName="Levy, Amitai et Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Vachoniolus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="globimanus">
<emphasis id="62C3EAF3513DFF895AFCDDFCFA0CF877" box="[1185,1415,1953,1974]" italics="true" pageId="16" pageNumber="15">Vachoniolus globimanus</emphasis>
<bibRefCitation id="34264B10513DFF895BD3DDFFFEC6F812" author="LEVY, G. &amp; P. AMITAI &amp; A. SHULOV" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="16" pageNumber="15" pagination="113 - 140" part="52" refId="ref14867" refString="LEVY, G., P. AMITAI &amp; A. SHULOV. 1973. New scorpions from Israel, Jordan and Arabia. Zoological Journal of the Linnean Society, 52 (2): 113 - 140." title="New scorpions from Israel, Jordan and Arabia" type="journal article" year="1973">Levy, Amitai et Shulov, 1973</bibRefCitation>
</taxonomicName>
(Ramlat As Sahmah, Oman, 7.X.1994, leg. G. Lowe &amp; M.D. Gallagher). All figures right capsule except Figures 78–79 (mirrored left). Anterior (78, 81–82) and convex compressed (79–80, 83) views. Scale bars: 400 µm (78–79), 400 µm (80–81), 400 µm (82–83).
</paragraph>
</caption>
<caption id="04C86669513CFF885E29DD99FAF7F81B" ID-DOI="http://doi.org/10.5281/zenodo.6511498" ID-Zenodo-Dep="6511498" box="[116,1404,1988,2010]" httpUri="https://zenodo.org/record/6511498/files/figure.png" pageId="17" pageNumber="16" startId="17.[116,193,1988,2010]" targetBox="[113,1474,248,1949]" targetPageId="17">
<paragraph id="500836E1513CFF885E29DD99FAF7F81B" blockId="17.[116,1404,1988,2010]" box="[116,1404,1988,2010]" pageId="17" pageNumber="16">
<emphasis id="62C3EAF3513CFF885E29DD99FE88F818" bold="true" box="[116,259,1988,2010]" pageId="17" pageNumber="16">Figures 84–85</emphasis>
: 
<taxonomicName id="97B74D62513CFF885F52DD99FE67F818" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[271,492,1988,2009]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF3513CFF885F52DD99FE67F818" box="[271,492,1988,2009]" italics="true" pageId="17" pageNumber="16">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513CFF885FAFDD99FDE5F81B" box="[498,622,1988,2010]" pageId="17" pageNumber="16" rank="species">
<emphasis id="62C3EAF3513CFF885FAFDD99FD9DF818" bold="true" box="[498,534,1988,2009]" pageId="17" pageNumber="16">gen</emphasis>
. et 
<emphasis id="62C3EAF3513CFF885C64DD99FDDBF818" bold="true" box="[569,592,1988,2009]" pageId="17" pageNumber="16">sp</emphasis>
. 
<emphasis id="62C3EAF3513CFF885C01DD99FDE2F818" bold="true" box="[604,617,1988,2009]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
, in vivo habitus, male holotype (84) and female paratype from type locality (85).
</paragraph>
</caption>
<caption id="04C86669513FFF8B5E2CDDECFCE4F825" ID-DOI="http://doi.org/10.5281/zenodo.6511503" ID-Zenodo-Dep="6511503" httpUri="https://zenodo.org/record/6511503/files/figure.png" pageId="18" pageNumber="17" startId="18.[113,190,1969,1991]" targetBox="[114,1474,236,1939]" targetPageId="18">
<paragraph id="500836E1513FFF8B5E2CDDECFCE4F825" blockId="18.[113,1475,1969,2020]" pageId="18" pageNumber="17">
<emphasis id="62C3EAF3513FFF8B5E2CDDECFE88F807" bold="true" box="[113,259,1969,1991]" pageId="18" pageNumber="17">Figures 86–89</emphasis>
: 
<taxonomicName id="97B74D62513FFF8B5F4FDDECFE7AF807" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[274,497,1969,1990]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n">
<emphasis id="62C3EAF3513FFF8B5F4FDDECFE7AF807" box="[274,497,1969,1990]" italics="true" pageId="18" pageNumber="17">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513FFF8B5FA4DDEFFDFCF806" box="[505,631,1970,1991]" pageId="18" pageNumber="17" rank="species">
<emphasis id="62C3EAF3513FFF8B5FA4DDEFFD96F806" bold="true" box="[505,541,1970,1991]" pageId="18" pageNumber="17">gen</emphasis>
. et 
<emphasis id="62C3EAF3513FFF8B5C18DDEFFDD7F806" bold="true" box="[581,604,1970,1991]" pageId="18" pageNumber="17">sp</emphasis>
. 
<emphasis id="62C3EAF3513FFF8B5C37DDEFFDFCF806" bold="true" box="[618,631,1970,1991]" pageId="18" pageNumber="17">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="62C3EAF3513FFF8B5CD1DDECFC96F807" bold="true" box="[652,797,1969,1991]" pageId="18" pageNumber="17">Figures 86–87</emphasis>
. Male (86) and female (87) from Thumrait, photos by J. N. Barnes. 
<emphasis id="62C3EAF3513FFF8B5E2CDD93FE8BF822" bold="true" box="[113,256,1998,2020]" pageId="18" pageNumber="17">Figures 88–89</emphasis>
. Type locality, Oman, S of Schelim, 18.041070°N 55.612262°E.
</paragraph>
</caption>
<paragraph id="500836E1513EFF8A5E26DAA4FA2DFB6D" pageId="19" pageNumber="18">
<table id="22B7C441513E00665E26DAAAFA2CFB6D" box="[123,1447,247,1196]" gridcols="5" gridrows="25" pageId="19" pageNumber="18">
<tr id="EE8734A3513E00665E26DAAAFA2CFED3" box="[123,1447,247,274]" gridrow="0" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DAAAFDB0FED3" box="[123,571,247,274]" gridcol="0" gridrow="0" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5E26DAA4FF5CFED3" bold="true" box="[123,215,249,274]" pageId="19" pageNumber="18">RATIO</emphasis>
</th>
<th id="AD565DDF513E00665C2ADAAAFC8FFED3" box="[631,772,247,274]" gridcol="1" gridrow="0" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5CC6DAAAFD6BFED1" bold="true" box="[667,736,247,272]" pageId="19" pageNumber="18">range</emphasis>
</th>
<th id="AD565DDF513E00665D6BDAAAFC62FED3" box="[822,1001,247,274]" gridcol="2" gridrow="0" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5D61DAAAFC68FED1" bold="true" box="[828,995,247,272]" pageId="19" pageNumber="18">mean ± sd (N)</emphasis>
</th>
<th id="AD565DDF513E00665A74DAAAFB3DFED3" box="[1065,1206,247,274]" gridcol="3" gridrow="0" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5A10DAAAFB19FED1" bold="true" box="[1101,1170,247,272]" pageId="19" pageNumber="18">range</emphasis>
</th>
<th id="AD565DDF513E00665AA9DAAAFA2CFED3" box="[1268,1447,247,274]" gridcol="4" gridrow="0" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5AA4DAAAFA2BFED1" bold="true" box="[1273,1440,247,272]" pageId="19" pageNumber="18">mean ± sd (N)</emphasis>
</th>
</tr>
<tr id="EE8734A3513E00665E26DB7DFA2CFEF7" box="[123,1447,288,310]" gridrow="1" pageId="19" pageNumber="18" rowspan-0="1">
<td id="AD565DDF513E00665C2ADB7DFC8FFEF7" box="[631,772,288,310]" gridcol="1" gridrow="1" pageId="19" pageNumber="18">♂</td>
<td id="AD565DDF513E00665D6BDB7DFC62FEF7" box="[822,1001,288,310]" gridcol="2" gridrow="1" pageId="19" pageNumber="18">♂</td>
<td id="AD565DDF513E00665A74DB7DFB3DFEF7" box="[1065,1206,288,310]" gridcol="3" gridrow="1" pageId="19" pageNumber="18">♀</td>
<td id="AD565DDF513E00665AA9DB7DFA2CFEF7" box="[1268,1447,288,310]" gridcol="4" gridrow="1" pageId="19" pageNumber="18">♀</td>
</tr>
<tr id="EE8734A3513E00665E26DB1AFA2CFE9C" box="[123,1447,327,349]" gridrow="2" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DB1AFDB0FE9C" box="[123,571,327,349]" gridcol="0" gridrow="2" pageId="19" pageNumber="18">carapace W/L</th>
<td id="AD565DDF513E00665C2ADB1AFC8FFE9C" box="[631,772,327,349]" gridcol="1" gridrow="2" pageId="19" pageNumber="18">1.098 – 1.182</td>
<td id="AD565DDF513E00665D6BDB1AFC62FE9C" box="[822,1001,327,349]" gridcol="2" gridrow="2" pageId="19" pageNumber="18">1.137 ± 0.037 (4)</td>
<td id="AD565DDF513E00665A74DB1AFB3DFE9C" box="[1065,1206,327,349]" gridcol="3" gridrow="2" pageId="19" pageNumber="18">1.135 – 1.252</td>
<td id="AD565DDF513E00665AA9DB1AFA2CFE9C" box="[1268,1447,327,349]" gridcol="4" gridrow="2" pageId="19" pageNumber="18">1.211 ± 0.052 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DB36FA2CFE40" box="[123,1447,363,385]" gridrow="3" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DB36FDB0FE40" box="[123,571,363,385]" gridcol="0" gridrow="3" pageId="19" pageNumber="18">pedipalp femur L/ carapace L</th>
<td id="AD565DDF513E00665C2ADB36FC8FFE40" box="[631,772,363,385]" gridcol="1" gridrow="3" pageId="19" pageNumber="18">0.811 – 0.876</td>
<td id="AD565DDF513E00665D6BDB36FC62FE40" box="[822,1001,363,385]" gridcol="2" gridrow="3" pageId="19" pageNumber="18">0.841 ± 0.029 (4)</td>
<td id="AD565DDF513E00665A74DB36FB3DFE40" box="[1065,1206,363,385]" gridcol="3" gridrow="3" pageId="19" pageNumber="18">0.653 – 0.739</td>
<td id="AD565DDF513E00665AA9DB36FA2CFE40" box="[1268,1447,363,385]" gridcol="4" gridrow="3" pageId="19" pageNumber="18">0.706 ± 0.039 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DBCCFA2CFE66" box="[123,1447,401,423]" gridrow="4" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DBCCFDB0FE66" box="[123,571,401,423]" gridcol="0" gridrow="4" pageId="19" pageNumber="18">pedipalp femur L/W</th>
<td id="AD565DDF513E00665C2ADBCCFC8FFE66" box="[631,772,401,423]" gridcol="1" gridrow="4" pageId="19" pageNumber="18">2.609 – 3.055</td>
<td id="AD565DDF513E00665D6BDBCCFC62FE66" box="[822,1001,401,423]" gridcol="2" gridrow="4" pageId="19" pageNumber="18">2.805 ± 0.185 (4)</td>
<td id="AD565DDF513E00665A74DBCCFB3DFE66" box="[1065,1206,401,423]" gridcol="3" gridrow="4" pageId="19" pageNumber="18">2.501 – 2.647</td>
<td id="AD565DDF513E00665AA9DBCCFA2CFE66" box="[1268,1447,401,423]" gridcol="4" gridrow="4" pageId="19" pageNumber="18">2.574 ± 0.080 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DBEBFA2CFE0D" box="[123,1447,438,460]" gridrow="5" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DBEBFDB0FE0D" box="[123,571,438,460]" gridcol="0" gridrow="5" pageId="19" pageNumber="18">pedipalp patella L/W</th>
<td id="AD565DDF513E00665C2ADBEBFC8FFE0D" box="[631,772,438,460]" gridcol="1" gridrow="5" pageId="19" pageNumber="18">2.543 – 2.653</td>
<td id="AD565DDF513E00665D6BDBEBFC62FE0D" box="[822,1001,438,460]" gridcol="2" gridrow="5" pageId="19" pageNumber="18">2.603 ± 0.046 (4)</td>
<td id="AD565DDF513E00665A74DBEBFB3DFE0D" box="[1065,1206,438,460]" gridcol="3" gridrow="5" pageId="19" pageNumber="18">2.435 – 2.723</td>
<td id="AD565DDF513E00665AA9DBEBFA2CFE0D" box="[1268,1447,438,460]" gridcol="4" gridrow="5" pageId="19" pageNumber="18">2.539 ± 0.130 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DB86FA2CFE30" box="[123,1447,475,497]" gridrow="6" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DB86FDB0FE30" box="[123,571,475,497]" gridcol="0" gridrow="6" pageId="19" pageNumber="18">pedipalp chela L/W</th>
<td id="AD565DDF513E00665C2ADB86FC8FFE30" box="[631,772,475,497]" gridcol="1" gridrow="6" pageId="19" pageNumber="18">3.557 – 3.988</td>
<td id="AD565DDF513E00665D6BDB86FC62FE30" box="[822,1001,475,497]" gridcol="2" gridrow="6" pageId="19" pageNumber="18">3.739 ± 0.212 (4)</td>
<td id="AD565DDF513E00665A74DB86FB3DFE30" box="[1065,1206,475,497]" gridcol="3" gridrow="6" pageId="19" pageNumber="18">4.467 – 4.589</td>
<td id="AD565DDF513E00665AA9DB86FA2CFE30" box="[1268,1447,475,497]" gridcol="4" gridrow="6" pageId="19" pageNumber="18">4.525 ± 0.052 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D85DFA2CFDD7" box="[123,1447,512,534]" gridrow="7" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D85DFDB0FDD7" box="[123,571,512,534]" gridcol="0" gridrow="7" pageId="19" pageNumber="18">pedipalp movable finger L/ manus ventral L</th>
<td id="AD565DDF513E00665C2AD85DFC8FFDD7" box="[631,772,512,534]" gridcol="1" gridrow="7" pageId="19" pageNumber="18">0.958 – 1.249</td>
<td id="AD565DDF513E00665D6BD85DFC62FDD7" box="[822,1001,512,534]" gridcol="2" gridrow="7" pageId="19" pageNumber="18">1.109 ± 0.122 (4)</td>
<td id="AD565DDF513E00665A74D85DFB3DFDD7" box="[1065,1206,512,534]" gridcol="3" gridrow="7" pageId="19" pageNumber="18">1.502 – 2.025</td>
<td id="AD565DDF513E00665AA9D85DFA2CFDD7" box="[1268,1447,512,534]" gridcol="4" gridrow="7" pageId="19" pageNumber="18">1.645 ± 0.254 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D878FA2CFDFA" box="[123,1447,549,571]" gridrow="8" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D878FDB0FDFA" box="[123,571,549,571]" gridcol="0" gridrow="8" pageId="19" pageNumber="18">pedipalp fixed finger L/ manus ventral L</th>
<td id="AD565DDF513E00665C2AD878FC8FFDFA" box="[631,772,549,571]" gridcol="1" gridrow="8" pageId="19" pageNumber="18">0.853 – 1.023</td>
<td id="AD565DDF513E00665D6BD878FC62FDFA" box="[822,1001,549,571]" gridcol="2" gridrow="8" pageId="19" pageNumber="18">0.908 ± 0.077 (4)</td>
<td id="AD565DDF513E00665A74D878FB3DFDFA" box="[1065,1206,549,571]" gridcol="3" gridrow="8" pageId="19" pageNumber="18">1.200 – 1.251</td>
<td id="AD565DDF513E00665AA9D878FA2CFDFA" box="[1268,1447,549,571]" gridcol="4" gridrow="8" pageId="19" pageNumber="18">1.228 ± 0.023 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D817FA2CFDA1" box="[123,1447,586,608]" gridrow="9" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D817FDB0FDA1" box="[123,571,586,608]" gridcol="0" gridrow="9" pageId="19" pageNumber="18">pedipalp manus W/D</th>
<td id="AD565DDF513E00665C2AD817FC8FFDA1" box="[631,772,586,608]" gridcol="1" gridrow="9" pageId="19" pageNumber="18">0.888 – 0.947</td>
<td id="AD565DDF513E00665D6BD817FC62FDA1" box="[822,1001,586,608]" gridcol="2" gridrow="9" pageId="19" pageNumber="18">0.919 ± 0.027 (4)</td>
<td id="AD565DDF513E00665A74D817FB3DFDA1" box="[1065,1206,586,608]" gridcol="3" gridrow="9" pageId="19" pageNumber="18">0.842 – 0.900</td>
<td id="AD565DDF513E00665AA9D817FA2CFDA1" box="[1268,1447,586,608]" gridcol="4" gridrow="9" pageId="19" pageNumber="18">0.866 ± 0.026 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D832FA2CFD44" box="[123,1447,623,645]" gridrow="10" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D832FDB0FD44" box="[123,571,623,645]" gridcol="0" gridrow="10" pageId="19" pageNumber="18">pedipalp manus ventral L/ manus W</th>
<td id="AD565DDF513E00665C2AD832FC8FFD44" box="[631,772,623,645]" gridcol="1" gridrow="10" pageId="19" pageNumber="18">1.714 – 1.879</td>
<td id="AD565DDF513E00665D6BD832FC62FD44" box="[822,1001,623,645]" gridcol="2" gridrow="10" pageId="19" pageNumber="18">1.779 ± 0.071 (4)</td>
<td id="AD565DDF513E00665A74D832FB3DFD44" box="[1065,1206,623,645]" gridcol="3" gridrow="10" pageId="19" pageNumber="18">1.287 – 1.889</td>
<td id="AD565DDF513E00665AA9D832FA2CFD44" box="[1268,1447,623,645]" gridcol="4" gridrow="10" pageId="19" pageNumber="18">1.696 ± 0.276 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D8C9FA2CFD6B" box="[123,1447,660,682]" gridrow="11" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D8C9FDB0FD6B" box="[123,571,660,682]" gridcol="0" gridrow="11" pageId="19" pageNumber="18">metasoma I L/W</th>
<td id="AD565DDF513E00665C2AD8C9FC8FFD6B" box="[631,772,660,682]" gridcol="1" gridrow="11" pageId="19" pageNumber="18">1.111 – 1.138</td>
<td id="AD565DDF513E00665D6BD8C9FC62FD6B" box="[822,1001,660,682]" gridcol="2" gridrow="11" pageId="19" pageNumber="18">1.120 ± 0.013 (4)</td>
<td id="AD565DDF513E00665A74D8C9FB3DFD6B" box="[1065,1206,660,682]" gridcol="3" gridrow="11" pageId="19" pageNumber="18">0.985 – 1.086</td>
<td id="AD565DDF513E00665AA9D8C9FA2CFD6B" box="[1268,1447,660,682]" gridcol="4" gridrow="11" pageId="19" pageNumber="18">1.032 ± 0.043 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D8E4FA2CFD0E" box="[123,1447,697,719]" gridrow="12" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D8E4FDB0FD0E" box="[123,571,697,719]" gridcol="0" gridrow="12" pageId="19" pageNumber="18">metasoma II L/W</th>
<td id="AD565DDF513E00665C2AD8E4FC8FFD0E" box="[631,772,697,719]" gridcol="1" gridrow="12" pageId="19" pageNumber="18">1.291 – 1.418</td>
<td id="AD565DDF513E00665D6BD8E4FC62FD0E" box="[822,1001,697,719]" gridcol="2" gridrow="12" pageId="19" pageNumber="18">1.356 ± 0.052 (4)</td>
<td id="AD565DDF513E00665A74D8E4FB3DFD0E" box="[1065,1206,697,719]" gridcol="3" gridrow="12" pageId="19" pageNumber="18">1.203 – 1.299</td>
<td id="AD565DDF513E00665AA9D8E4FA2CFD0E" box="[1268,1447,697,719]" gridcol="4" gridrow="12" pageId="19" pageNumber="18">1.255 ± 0.051 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D883FA2CFD35" box="[123,1447,734,756]" gridrow="13" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D883FDB0FD35" box="[123,571,734,756]" gridcol="0" gridrow="13" pageId="19" pageNumber="18">metasoma III L/W</th>
<td id="AD565DDF513E00665C2AD883FC8FFD35" box="[631,772,734,756]" gridcol="1" gridrow="13" pageId="19" pageNumber="18">1.405 – 1.526</td>
<td id="AD565DDF513E00665D6BD883FC62FD35" box="[822,1001,734,756]" gridcol="2" gridrow="13" pageId="19" pageNumber="18">1.455 ± 0.053 (4)</td>
<td id="AD565DDF513E00665A74D883FB3DFD35" box="[1065,1206,734,756]" gridcol="3" gridrow="13" pageId="19" pageNumber="18">1.304 – 1.387</td>
<td id="AD565DDF513E00665AA9D883FA2CFD35" box="[1268,1447,734,756]" gridcol="4" gridrow="13" pageId="19" pageNumber="18">1.357 ± 0.039 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D95EFA2CFCD8" box="[123,1447,771,793]" gridrow="14" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D95EFDB0FCD8" box="[123,571,771,793]" gridcol="0" gridrow="14" pageId="19" pageNumber="18">metasoma IV L/W</th>
<td id="AD565DDF513E00665C2AD95EFC8FFCD8" box="[631,772,771,793]" gridcol="1" gridrow="14" pageId="19" pageNumber="18">1.753 – 1.887</td>
<td id="AD565DDF513E00665D6BD95EFC62FCD8" box="[822,1001,771,793]" gridcol="2" gridrow="14" pageId="19" pageNumber="18">1.818 ± 0.057 (4)</td>
<td id="AD565DDF513E00665A74D95EFB3DFCD8" box="[1065,1206,771,793]" gridcol="3" gridrow="14" pageId="19" pageNumber="18">1.702 – 1.785</td>
<td id="AD565DDF513E00665AA9D95EFA2CFCD8" box="[1268,1447,771,793]" gridcol="4" gridrow="14" pageId="19" pageNumber="18">1.751 ± 0.036 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D97AFA2CFCFC" box="[123,1447,807,829]" gridrow="15" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D97AFDB0FCFC" box="[123,571,807,829]" gridcol="0" gridrow="15" pageId="19" pageNumber="18">metasoma V L/W</th>
<td id="AD565DDF513E00665C2AD97AFC8FFCFC" box="[631,772,807,829]" gridcol="1" gridrow="15" pageId="19" pageNumber="18">2.256 – 2.500</td>
<td id="AD565DDF513E00665D6BD97AFC62FCFC" box="[822,1001,807,829]" gridcol="2" gridrow="15" pageId="19" pageNumber="18">2.389 ± 0.101 (4)</td>
<td id="AD565DDF513E00665A74D97AFB3DFCFC" box="[1065,1206,807,829]" gridcol="3" gridrow="15" pageId="19" pageNumber="18">2.092 – 2.286</td>
<td id="AD565DDF513E00665AA9D97AFA2CFCFC" box="[1268,1447,807,829]" gridcol="4" gridrow="15" pageId="19" pageNumber="18">2.178 ± 0.099 (3)</td>
</tr>
<tr id="EE8734A3513E00665E26D911FA2CFCA3" box="[123,1447,844,866]" gridrow="16" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D911FDB0FCA3" box="[123,571,844,866]" gridcol="0" gridrow="16" pageId="19" pageNumber="18">metasoma I L/D</th>
<td id="AD565DDF513E00665C2AD911FC8FFCA3" box="[631,772,844,866]" gridcol="1" gridrow="16" pageId="19" pageNumber="18">1.284 – 1.352</td>
<td id="AD565DDF513E00665D6BD911FC62FCA3" box="[822,1001,844,866]" gridcol="2" gridrow="16" pageId="19" pageNumber="18">1.316 ± 0.029 (4)</td>
<td id="AD565DDF513E00665A74D911FB3DFCA3" box="[1065,1206,844,866]" gridcol="3" gridrow="16" pageId="19" pageNumber="18">1.147 – 1.315</td>
<td id="AD565DDF513E00665AA9D911FA2CFCA3" box="[1268,1447,844,866]" gridcol="4" gridrow="16" pageId="19" pageNumber="18">1.210 ± 0.074 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D92CFA2CFC46" box="[123,1447,881,903]" gridrow="17" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D92CFDB0FC46" box="[123,571,881,903]" gridcol="0" gridrow="17" pageId="19" pageNumber="18">metasoma II L/D</th>
<td id="AD565DDF513E00665C2AD92CFC8FFC46" box="[631,772,881,903]" gridcol="1" gridrow="17" pageId="19" pageNumber="18">1.385 – 1.569</td>
<td id="AD565DDF513E00665D6BD92CFC62FC46" box="[822,1001,881,903]" gridcol="2" gridrow="17" pageId="19" pageNumber="18">1.512 ± 0.086 (4)</td>
<td id="AD565DDF513E00665A74D92CFB3DFC46" box="[1065,1206,881,903]" gridcol="3" gridrow="17" pageId="19" pageNumber="18">1.279 – 1.414</td>
<td id="AD565DDF513E00665AA9D92CFA2CFC46" box="[1268,1447,881,903]" gridcol="4" gridrow="17" pageId="19" pageNumber="18">1.359 ± 0.067 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D9CBFA2CFC6D" box="[123,1447,918,940]" gridrow="18" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D9CBFDB0FC6D" box="[123,571,918,940]" gridcol="0" gridrow="18" pageId="19" pageNumber="18">metasoma III L/D</th>
<td id="AD565DDF513E00665C2AD9CBFC8FFC6D" box="[631,772,918,940]" gridcol="1" gridrow="18" pageId="19" pageNumber="18">1.431 – 1.610</td>
<td id="AD565DDF513E00665D6BD9CBFC62FC6D" box="[822,1001,918,940]" gridcol="2" gridrow="18" pageId="19" pageNumber="18">1.556 ± 0.083 (4)</td>
<td id="AD565DDF513E00665A74D9CBFB3DFC6D" box="[1065,1206,918,940]" gridcol="3" gridrow="18" pageId="19" pageNumber="18">1.335 – 1.528</td>
<td id="AD565DDF513E00665AA9D9CBFA2CFC6D" box="[1268,1447,918,940]" gridcol="4" gridrow="18" pageId="19" pageNumber="18">1.473 ± 0.092 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D9E6FA2CFC10" box="[123,1447,955,977]" gridrow="19" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D9E6FDB0FC10" box="[123,571,955,977]" gridcol="0" gridrow="19" pageId="19" pageNumber="18">metasoma IV L/D</th>
<td id="AD565DDF513E00665C2AD9E6FC8FFC10" box="[631,772,955,977]" gridcol="1" gridrow="19" pageId="19" pageNumber="18">1.997 – 2.191</td>
<td id="AD565DDF513E00665D6BD9E6FC62FC10" box="[822,1001,955,977]" gridcol="2" gridrow="19" pageId="19" pageNumber="18">2.100 ± 0.104 (4)</td>
<td id="AD565DDF513E00665A74D9E6FB3DFC10" box="[1065,1206,955,977]" gridcol="3" gridrow="19" pageId="19" pageNumber="18">1.872 – 2.031</td>
<td id="AD565DDF513E00665AA9D9E6FA2CFC10" box="[1268,1447,955,977]" gridcol="4" gridrow="19" pageId="19" pageNumber="18">1.955 ± 0.073 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26D9BDFA2CFC37" box="[123,1447,992,1014]" gridrow="20" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26D9BDFDB0FC37" box="[123,571,992,1014]" gridcol="0" gridrow="20" pageId="19" pageNumber="18">metasoma V L/D</th>
<td id="AD565DDF513E00665C2AD9BDFC8FFC37" box="[631,772,992,1014]" gridcol="1" gridrow="20" pageId="19" pageNumber="18">2.763 – 2.955</td>
<td id="AD565DDF513E00665D6BD9BDFC62FC37" box="[822,1001,992,1014]" gridcol="2" gridrow="20" pageId="19" pageNumber="18">2.846 ± 0.091 (4)</td>
<td id="AD565DDF513E00665A74D9BDFB3DFC37" box="[1065,1206,992,1014]" gridcol="3" gridrow="20" pageId="19" pageNumber="18">2.547 – 2.855</td>
<td id="AD565DDF513E00665AA9D9BDFA2CFC37" box="[1268,1447,992,1014]" gridcol="4" gridrow="20" pageId="19" pageNumber="18">2.737 ± 0.166 (3)</td>
</tr>
<tr id="EE8734A3513E00665E26DE58FA2CFBDA" box="[123,1447,1029,1051]" gridrow="21" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DE58FDB0FBDA" box="[123,571,1029,1051]" gridcol="0" gridrow="21" pageId="19" pageNumber="18">metasoma II W/ metasoma I W</th>
<td id="AD565DDF513E00665C2ADE58FC8FFBDA" box="[631,772,1029,1051]" gridcol="1" gridrow="21" pageId="19" pageNumber="18">0.940 – 0.977</td>
<td id="AD565DDF513E00665D6BDE58FC62FBDA" box="[822,1001,1029,1051]" gridcol="2" gridrow="21" pageId="19" pageNumber="18">0.954 ± 0.017 (4)</td>
<td id="AD565DDF513E00665A74DE58FB3DFBDA" box="[1065,1206,1029,1051]" gridcol="3" gridrow="21" pageId="19" pageNumber="18">0.932 – 0.973</td>
<td id="AD565DDF513E00665AA9DE58FA2CFBDA" box="[1268,1447,1029,1051]" gridcol="4" gridrow="21" pageId="19" pageNumber="18">0.944 ± 0.019 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DE77FA2CFB81" box="[123,1447,1066,1088]" gridrow="22" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DE77FDB0FB81" box="[123,571,1066,1088]" gridcol="0" gridrow="22" pageId="19" pageNumber="18">metasoma IV W/ metasoma I W</th>
<td id="AD565DDF513E00665C2ADE77FC8FFB81" box="[631,772,1066,1088]" gridcol="1" gridrow="22" pageId="19" pageNumber="18">0.858 – 0.912</td>
<td id="AD565DDF513E00665D6BDE77FC62FB81" box="[822,1001,1066,1088]" gridcol="2" gridrow="22" pageId="19" pageNumber="18">0.874 ± 0.026 (4)</td>
<td id="AD565DDF513E00665A74DE77FB3DFB81" box="[1065,1206,1066,1088]" gridcol="3" gridrow="22" pageId="19" pageNumber="18">0.847 – 0.884</td>
<td id="AD565DDF513E00665AA9DE77FA2CFB81" box="[1268,1447,1066,1088]" gridcol="4" gridrow="22" pageId="19" pageNumber="18">0.865 ± 0.015 (4)</td>
</tr>
<tr id="EE8734A3513E00665E26DE12FA2CFBA4" box="[123,1447,1103,1125]" gridrow="23" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DE12FDB0FBA4" box="[123,571,1103,1125]" gridcol="0" gridrow="23" pageId="19" pageNumber="18">metasoma V W/ metasoma I W</th>
<td id="AD565DDF513E00665C2ADE12FC8FFBA4" box="[631,772,1103,1125]" gridcol="1" gridrow="23" pageId="19" pageNumber="18">0.711 – 0.788</td>
<td id="AD565DDF513E00665D6BDE12FC62FBA4" box="[822,1001,1103,1125]" gridcol="2" gridrow="23" pageId="19" pageNumber="18">0.741 ± 0.036 (4)</td>
<td id="AD565DDF513E00665A74DE12FB3DFBA4" box="[1065,1206,1103,1125]" gridcol="3" gridrow="23" pageId="19" pageNumber="18">0.760 – 0.809</td>
<td id="AD565DDF513E00665AA9DE12FA2CFBA4" box="[1268,1447,1103,1125]" gridcol="4" gridrow="23" pageId="19" pageNumber="18">0.785 ± 0.024 (3)</td>
</tr>
<tr id="EE8734A3513E00665E26DE29FA2CFB6D" box="[123,1447,1140,1196]" gridrow="24" pageId="19" pageNumber="18">
<th id="AD565DDF513E00665E26DE29FDB0FB6D" box="[123,571,1140,1196]" gridcol="0" gridrow="24" pageId="19" pageNumber="18">leg III patella L/W pectine anterior margin L/ carapace L</th>
<td id="AD565DDF513E00665C2ADE29FC8FFB6D" box="[631,772,1140,1196]" gridcol="1" gridrow="24" pageId="19" pageNumber="18">3.017 – 3.336 0.940 – 1.242</td>
<td id="AD565DDF513E00665D6BDE29FC62FB6D" box="[822,1001,1140,1196]" gridcol="2" gridrow="24" pageId="19" pageNumber="18">3.133 ± 0.176 (3) 1.101 ± 0.128 (4)</td>
<td id="AD565DDF513E00665A74DE29FB3DFB6D" box="[1065,1206,1140,1196]" gridcol="3" gridrow="24" pageId="19" pageNumber="18">2.721 – 2.903 0.633 – 0.994</td>
<td id="AD565DDF513E00665AA9DE29FA2CFB6D" box="[1268,1447,1140,1196]" gridcol="4" gridrow="24" pageId="19" pageNumber="18">2.788 ± 0.100 (3) 0.816 ± 0.181 (3)</td>
</tr>
</table>
</paragraph>
<caption id="04C86669513EFF8A5E2CDE94FD7EFB3D" ID-Table-UUID="04C86669513EFF8A5E2CDE94FD7EFB3D" httpUri="http://table.plazi.org/id/04C86669513EFF8A5E2CDE94FD7EFB3D" pageId="19" pageNumber="18" startId="19.[113,169,1225,1246]" targetBox="[123,1447,247,1196]" targetIsTable="true" targetPageId="19">
<paragraph id="500836E1513EFF8A5E2CDE94FD7EFB3D" blockId="19.[113,1474,1225,1276]" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5E2CDE94FF36FB1F" bold="true" box="[113,189,1225,1246]" pageId="19" pageNumber="18">Table 2</emphasis>
. Variation in morphometric ratios of adult 
<taxonomicName id="97B74D62513EFF8A5C3EDE94FCCAFB1F" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[611,833,1225,1246]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="barnesi" status="gen. et sp. n.">
<emphasis id="62C3EAF3513EFF8A5C3EDE94FCCAFB1F" box="[611,833,1225,1246]" italics="true" pageId="19" pageNumber="18">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513EFF8A5D17DE94FC41FB1F" box="[842,970,1225,1246]" pageId="19" pageNumber="18" rank="species">
<emphasis id="62C3EAF3513EFF8A5D17DE94FCE6FB1F" bold="true" box="[842,877,1225,1246]" pageId="19" pageNumber="18">gen</emphasis>
. et 
<emphasis id="62C3EAF3513EFF8A5DCEDE94FC21FB1F" bold="true" box="[915,938,1225,1246]" pageId="19" pageNumber="18">sp</emphasis>
. 
<emphasis id="62C3EAF3513EFF8A5DEADE94FC4FFB1F" bold="true" box="[951,964,1225,1246]" pageId="19" pageNumber="18">n</emphasis>
.
</taxonomicNameLabel>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D), standard deviation (sd).
</paragraph>
</caption>
<paragraph id="500836E1513EFF8A5D6DDF44FBB3F830" blockId="19.[816,1475,1305,2033]" pageId="19" pageNumber="18">
<emphasis id="62C3EAF3513EFF8A5D6DDF44FC14FAF0" bold="true" box="[816,927,1305,1329]" pageId="19" pageNumber="18">Pedipalps</emphasis>
(
<figureCitation id="C88C2A64513EFF8A5DEADF44FBCAFAF3" box="[951,1089,1305,1330]" captionStart="Figures 29–46" captionStartId="12.[115,192,1871,1893]" captionTargetBox="[131,1463,242,1849]" captionTargetId="figure-164@12.[113,1471,234,1854]" captionTargetPageId="12" captionText="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi="http://doi.org/10.5281/zenodo.6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="19" pageNumber="18">Figs. 29–40</figureCitation>
, 
<figureCitation id="C88C2A64513EFF8A5A09DF47FB10FAF3" box="[1108,1179,1306,1330]" captionStart-0="Figures 47–58" captionStart-1="Figures 59–70" captionStartId-0="13.[113,190,1959,1981]" captionStartId-1="14.[116,193,1907,1929]" captionTargetBox-0="[113,1474,234,1930]" captionTargetBox-1="[124,1469,237,1889]" captionTargetId-0="figure-7@13.[113,1474,234,1932]" captionTargetId-1="figure-11@14.[113,1469,232,1890]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figures 47–58: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of male paratype from Thumrait (VII.1997), chela dorsal (47), external (48), ventral (49) and internal (50) views. Patella dorsal (51), external (52), ventral (53) and internal (54) views. Femur and trochanter dorsal (55), external (56), ventral (57) and internal (58) views. UV fluorescence. Scale bar: 2 mm." captionText-1="Figures 59–70: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of female paratypes, chela dorsal (59), external (60), ventral (61) and internal (62) views. Patella dorsal (63), external (64), ventral (65) and internal (66) views. Femur and trochanter dorsal (67), external (68), ventral (69) and internal (70) views. Dorsal views (59, 63, 67): paratype female from Thumrait (XI.1989) (with normal dorsointernal carina on femur). External (60, 64, 68), ventral (61, 65, 69) and internal (62, 66, 70) views: paratype female from Yalooni (XII.1988) (with malformed dorsointernal carina on femur). UV fluorescence. Scale bars: 2 mm (59, 63, 67), 2 mm (60–62, 64–66, 69–70)." figureDoi-0="http://doi.org/10.5281/zenodo.6511487" figureDoi-1="http://doi.org/10.5281/zenodo.6511489" httpUri-0="https://zenodo.org/record/6511487/files/figure.png" httpUri-1="https://zenodo.org/record/6511489/files/figure.png" pageId="19" pageNumber="18">47–70</figureCitation>
). Segments short, robust; 
<emphasis id="62C3EAF3513EFF8A5D6DDF67FCE5FA93" box="[816,878,1338,1362]" italics="true" pageId="19" pageNumber="18">femur</emphasis>
with 3 strong, granulated carinae: dorsoexternal, dorsointernal, and ventrointernal, other carinae obsolete; dorsal, lateral and ventral surfaces smooth except for few small solitary granules, internal surface smooth except for several solitary coarse granules; distal external surface with group of ca. dozen macrosetae; 
<emphasis id="62C3EAF3513EFF8A5A71DF84FBFDFA30" box="[1068,1142,1497,1521]" italics="true" pageId="19" pageNumber="18">patella</emphasis>
with 7 carinae: dorsointernal, internal and ventrointernal carinae coarsely granulose, internal carina with sparse, non-contiguous granules; dorsomedian and dorsoexternal carinae weak, smooth or almost smooth with traces of coarse granules; external and ventroexternal carinae weak, smooth; ventromedian carina obsolete, indicated by series of few granules; all intercarinal surfaces smooth; setation very sparse, with few large solitary macrosetae on carinae; 
<emphasis id="62C3EAF3513EFF8A5DD2DC84FC42F930" box="[911,969,1753,1777]" italics="true" pageId="19" pageNumber="18">chela</emphasis>
smooth, carinae obsolete with residual traces of carinae in females; few large macrosetae present, ventral surface of movable finger with numerous short macrosetae, increasing in density distally; dentate margin of movable and fixed fingers with 7–8 rows of granules, each with a single external and internal accessory granule, and 3–6 subterminal granules (typically 3–4 of them enlarged); trichobothrial pattern orthobothriotaxic 
<typeStatus id="8F0C8843513EFF8A5A18DDE7FBF8F813" box="[1093,1139,1978,2002]" pageId="19" pageNumber="18">type</typeStatus>
Aβ, chela fixed finger with 
<emphasis id="62C3EAF3513EFF8A5BFADDE4FA49F810" box="[1447,1474,1977,2001]" italics="true" pageId="19" pageNumber="18">db</emphasis>
near base of fixed finger.
</paragraph>
<paragraph id="500836E15139FF8D5E2CDAB6FEB4FCA2" blockId="20.[113,772,235,1635]" pageId="20" pageNumber="19">
<emphasis id="62C3EAF35139FF8D5E2CDAB6FF2DFEC2" bold="true" box="[113,166,235,259]" pageId="20" pageNumber="19">Legs</emphasis>
(
<figureCitation id="C88C2A645139FF8D5EE7DAB6FEAEFEC2" box="[186,293,235,259]" captionStart="Figures 1–4" captionStartId="3.[113,190,1967,1989]" captionTargetBox="[132,1458,266,1903]" captionTargetId="figure-7@3.[113,1474,253,1912]" captionTargetPageId="3" captionText="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511465" httpUri="https://zenodo.org/record/6511465/files/figure.png" pageId="20" pageNumber="19">Figs. 1–4</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5F69DAB6FEE5FEC2" box="[308,366,235,259]" captionStart="Figures 5–10" captionStartId="5.[113,190,1457,1479]" captionTargetBox="[113,1474,223,1433]" captionTargetId="figure-291@5.[113,1474,223,1449]" captionTargetPageId="5" captionText="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." figureDoi="http://doi.org/10.5281/zenodo.6511467" httpUri="https://zenodo.org/record/6511467/files/figure.png" pageId="20" pageNumber="19">9–10</figureCitation>
). Legs with long femora and robust patellae, tibiae and tarsi; femora with few solitary macrosetae; tibiae I–III with dorsal (retrosuperior) linear series of long macrosetae (= tibial ‘bristle comb’, 8–12 setae in sample of N=13 leg III segments), and shorter macrosetae on other surfaces; basitarsi I–III compressed, with two linear series of shorter ventral (proinferior and retroinferior) macrosetae, single linear series of longer dorsal (retrosuperior) macrosetae (= basitarsal ‘bristle comb’, 17–23 setae in sample of N=12 leg III segments); macrosetae long and thin in both sexes; leg IV without basitarsal compression, longer than legs I–III, long macrosetae more sparse, not arranged in linear series; leg I–IV femora and patella with indications of 4–6 carinae, which are usually obsolete; paired ventral carinae on femora granulate or denticulate, prolateral surfaces of femora with weak granulation localized on inferior proximal areas of I–II; tibial spur on leg IV strong, longer than spur on leg III; prolateral pedal spurs basally bifurcate, setose (leg III spur with 6–8 short macrosetae in sample of N=10 segments); retrolateral pedal spurs simple.
</paragraph>
</subSubSection>
<subSubSection id="18AD656A5139FF8D5E2CD937FD73FB22" pageId="20" pageNumber="19" type="diagnosis">
<paragraph id="500836E15139FF8D5E2CD937FD73FB22" blockId="20.[113,772,235,1635]" pageId="20" pageNumber="19">
<emphasis id="62C3EAF35139FF8D5E2CD937FEC6FC43" bold="true" box="[113,333,874,898]" pageId="20" pageNumber="19">Sexual dimorphism</emphasis>
. Compared to females, males have longer pectines (
<figureCitation id="C88C2A645139FF8D5E8AD9D6FEB7FC62" box="[215,316,907,931]" captionStart="Figures 1–4" captionStartId="3.[113,190,1967,1989]" captionTargetBox="[132,1458,266,1903]" captionTargetId="figure-7@3.[113,1474,253,1912]" captionTargetPageId="3" captionText="Figures 1–4: Trypanothacus barnesi gen. et sp. n. Figures 1–2. Male paratype from the type locality in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype from the type locality in dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511465" httpUri="https://zenodo.org/record/6511465/files/figure.png" pageId="20" pageNumber="19">Figs. 2, 4</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5F1AD9D6FEFAFC62" box="[327,369,907,931]" captionStart="Figures 5–10" captionStartId="5.[113,190,1457,1479]" captionTargetBox="[113,1474,223,1433]" captionTargetId="figure-291@5.[113,1474,223,1449]" captionTargetPageId="5" captionText="Figures 5–10: Trypanothacus barnesi gen. et sp. n. Figures 5, 7, 9. Male holotype, carapace and tergites I–V (5), coxosternal area and sternites (7), right legs I–IV, retrolateral aspect (9.I–IV). Figures 6, 8, 10. Female paratype from the type locality, carapace and tergites I–III (6), coxosternal area and sternites III–IV (8), right legs III–IV, retrolateral aspect (10.III–IV)." figureDoi="http://doi.org/10.5281/zenodo.6511467" httpUri="https://zenodo.org/record/6511467/files/figure.png" pageId="20" pageNumber="19">7–8</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5F21D9D6FE12FC62" box="[380,409,907,931]" captionStart="Figures 11–12" captionStartId="7.[116,193,1657,1679]" captionTargetBox="[116,1473,234,1632]" captionTargetId="figure-143@7.[114,1475,234,1632]" captionTargetPageId="7" captionText="Figures 11–12: Trypanothacus barnesi gen. et sp. n. Male paratype from Thumrait (VII.1997), carapace and tergites I–VII (11), coxosternal area and sternites (12). UV fluorescence. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6511473" httpUri="https://zenodo.org/record/6511473/files/figure.png" pageId="20" pageNumber="19">12</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5FF9D9D6FE4AFC62" box="[420,449,907,931]" captionStart="Figures 13–14" captionStartId="9.[113,190,1589,1611]" captionTargetBox="[113,1474,223,1572]" captionTargetId="figure-7@9.[113,1474,223,1574]" captionTargetPageId="9" captionText="Figures 13–14: Trypanothacus barnesi gen. et sp. n. Female paratype from Yalooni (XII.1988), carapace and tergites I–VII (13), coxosternal area and sternites (14). UV fluorescence. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6511477" httpUri="https://zenodo.org/record/6511477/files/figure.png" pageId="20" pageNumber="19">14</figureCitation>
), higher pectinal tooth counts, and broader pedipalp chelae; chela fingers are straight in both sexes, and males lack recess or scalloping of the proximal dentate margins (
<figureCitation id="C88C2A645139FF8D5F74D9B6FE71FBC2" box="[297,506,1003,1027]" captionStart="Figures 29–46" captionStartId="12.[115,192,1871,1893]" captionTargetBox="[131,1463,242,1849]" captionTargetId="figure-164@12.[113,1471,234,1854]" captionTargetPageId="12" captionText="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi="http://doi.org/10.5281/zenodo.6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="20" pageNumber="19">Figs. 29–31, 39–40</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5C58D9B7FDC0FBC3" box="[517,587,1002,1026]" captionStart="Figures 47–58" captionStartId="13.[113,190,1959,1981]" captionTargetBox="[113,1474,234,1930]" captionTargetId="figure-7@13.[113,1474,234,1932]" captionTargetPageId="13" captionText="Figures 47–58: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of male paratype from Thumrait (VII.1997), chela dorsal (47), external (48), ventral (49) and internal (50) views. Patella dorsal (51), external (52), ventral (53) and internal (54) views. Femur and trochanter dorsal (55), external (56), ventral (57) and internal (58) views. UV fluorescence. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6511487" httpUri="https://zenodo.org/record/6511487/files/figure.png" pageId="20" pageNumber="19">47–50</figureCitation>
, 
<figureCitation id="C88C2A645139FF8D5C08D9B7FD11FBC3" box="[597,666,1002,1026]" captionStart="Figures 59–70" captionStartId="14.[116,193,1907,1929]" captionTargetBox="[124,1469,237,1889]" captionTargetId="figure-11@14.[113,1469,232,1890]" captionTargetPageId="14" captionText="Figures 59–70: Trypanothacus barnesi gen. et sp. n. Pedipalp segments of female paratypes, chela dorsal (59), external (60), ventral (61) and internal (62) views. Patella dorsal (63), external (64), ventral (65) and internal (66) views. Femur and trochanter dorsal (67), external (68), ventral (69) and internal (70) views. Dorsal views (59, 63, 67): paratype female from Thumrait (XI.1989) (with normal dorsointernal carina on femur). External (60, 64, 68), ventral (61, 65, 69) and internal (62, 66, 70) views: paratype female from Yalooni (XII.1988) (with malformed dorsointernal carina on femur). UV fluorescence. Scale bars: 2 mm (59, 63, 67), 2 mm (60–62, 64–66, 69–70)." figureDoi="http://doi.org/10.5281/zenodo.6511489" httpUri="https://zenodo.org/record/6511489/files/figure.png" pageId="20" pageNumber="19">59–62</figureCitation>
, 
<tableCitation id="1D35035A5139FF8D5CF9D9B6FD7AFBC2" box="[676,753,1003,1027]" captionStart="Table 1" captionStartId="8.[112,168,987,1008]" captionTargetId="graphics-540@8.[155,1419,222,960]" captionTargetPageId="8" captionText="Table 1. Comparative measurements of adults of Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/04C866695125FF915E2DD986FD87FBCF" pageId="20" pageNumber="19" tableUuid="04C866695125FF915E2DD986FD87FBCF">Table 1</tableCitation>
); the male telson vesicle is slightly less bulbous, granulation on the carapace and tergites is more dense; sternite VII carinae of the male are weaker and smoother, and metasomal carinae are not as robust as in females, especially the ventromedian carinae on metasoma II–III; intercarinal granulation is more dense on the male metasoma. For data on morphometric sexual dimorphism, see under species diagnosis and 
<tableCitation id="1D35035A5139FF8D5CFFDE96FD7FFB22" box="[674,756,1227,1251]" captionStart="Table 2" captionStartId="19.[113,169,1225,1246]" captionTargetId="graphics-432@19.[117,1474,234,1210]" captionTargetPageId="19" captionText="Table 2. Variation in morphometric ratios of adult Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it" httpUri="http://table.plazi.org/id/04C86669513EFF8A5E2CDE94FD7EFB3D" pageId="20" pageNumber="19" tableUuid="04C86669513EFF8A5E2CDE94FD7EFB3D">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="18AD656A5139FF8D5E2CDEB6FF69F9A2" pageId="20" pageNumber="19" type="description">
<paragraph id="500836E15139FF8D5E2CDEB6FE23FAC2" blockId="20.[113,772,235,1635]" box="[113,424,1259,1283]" pageId="20" pageNumber="19">
<emphasis id="62C3EAF35139FF8D5E2CDEB6FE93FAC2" bold="true" box="[113,280,1259,1283]" pageId="20" pageNumber="19">Measurements</emphasis>
. See 
<tableCitation id="1D35035A5139FF8D5F0FDEB6FE29FAC2" box="[338,418,1259,1283]" captionStart="Table 1" captionStartId="8.[112,168,987,1008]" captionTargetId="graphics-540@8.[155,1419,222,960]" captionTargetPageId="8" captionText="Table 1. Comparative measurements of adults of Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/04C866695125FF915E2DD986FD87FBCF" pageId="20" pageNumber="19" tableUuid="04C866695125FF915E2DD986FD87FBCF">Table 1</tableCitation>
.
</paragraph>
<paragraph id="500836E15139FF8D5E2CDF57FD9CFAE2" blockId="20.[113,772,235,1635]" box="[113,535,1290,1315]" pageId="20" pageNumber="19">
<emphasis id="62C3EAF35139FF8D5E2CDF57FE0CFAE3" bold="true" box="[113,391,1290,1314]" pageId="20" pageNumber="19">Morphometric variation</emphasis>
. See 
<tableCitation id="1D35035A5139FF8D5F9CDF56FD98FAE2" box="[449,531,1291,1315]" captionStart="Table 2" captionStartId="19.[113,169,1225,1246]" captionTargetId="graphics-432@19.[117,1474,234,1210]" captionTargetPageId="19" captionText="Table 2. Variation in morphometric ratios of adult Trypanothacus barnesi gen. et sp. n. Abbreviations: length (L), width (W, in carapace it" httpUri="http://table.plazi.org/id/04C86669513EFF8A5E2CDE94FD7EFB3D" pageId="20" pageNumber="19" tableUuid="04C86669513EFF8A5E2CDE94FD7EFB3D">Table 2</tableCitation>
.
</paragraph>
<paragraph id="500836E15139FF8D5E2CDF76FF69F9A2" blockId="20.[113,772,235,1635]" pageId="20" pageNumber="19">
<emphasis id="62C3EAF35139FF8D5E2CDF76FF7EFA82" bold="true" box="[113,245,1323,1347]" pageId="20" pageNumber="19">Karyotypes</emphasis>
(
<figureCitation id="C88C2A645139FF8D5F5CDF76FE1FFA82" box="[257,404,1323,1347]" captionStart="Figures 111–113" captionStartId="26.[113,190,671,693]" captionTargetBox="[113,1474,235,649]" captionTargetId="figure-786@26.[113,1474,234,659]" captionTargetPageId="26" captionText="Figures 111–113: Trypanothacus barnesi gen. et sp. n., mitotic metaphase (111), postpachytene (112) and ideogram (113) (y axis - % of the chromosome length of the haploid set) of male holotype. Scale bar = 5 μm." figureDoi="http://doi.org/10.5281/zenodo.6511517" httpUri="https://zenodo.org/record/6511517/files/figure.png" pageId="20" pageNumber="19">Figs. 111–113</figureCitation>
). We analyzed 
<specimenCount id="46B1FD685139FF8D5C6DDF76FD12FA82" box="[560,665,1323,1347]" pageId="20" pageNumber="19" type="male">two males</specimenCount>
(
<typeStatus id="8F0C88435139FF8D5CF9DF76FC88FA82" box="[676,771,1323,1347]" pageId="20" pageNumber="19" type="holotype">holotype</typeStatus>
and 
<typeStatus id="8F0C88435139FF8D5EFFDF16FF75FAA2" box="[162,254,1355,1379]" pageId="20" pageNumber="19" type="paratype">paratype</typeStatus>
from type locality) using standard cytogenetic methods (e. g. 
<bibRefCitation id="34264B105139FF8D5F4ADF36FE7CFA42" author="KOVARIK, F. &amp; F. STAHLAVSKY &amp; T. KORINKOVA &amp; J. KRAL &amp; T. VAN DER ENDE" box="[279,503,1387,1411]" journalOrPublisher="Euscorpius" pageId="20" pageNumber="19" pagination="1 - 12" part="77" refId="ref14768" refString="KOVARIK, F., F. STAHLAVSKY. T. KORINKOVA, J. KRAL &amp; T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1 - 12." title="Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA" type="journal article" year="2009">Kovařík et al., 2009</bibRefCitation>
). The diploid number of both specimens is 26 chromosomes (
<figureCitation id="C88C2A645139FF8D5C58DFD6FD16FA62" box="[517,669,1419,1443]" captionStart="Figures 111–113" captionStartId="26.[113,190,671,693]" captionTargetBox="[113,1474,235,649]" captionTargetId="figure-786@26.[113,1474,234,659]" captionTargetPageId="26" captionText="Figures 111–113: Trypanothacus barnesi gen. et sp. n., mitotic metaphase (111), postpachytene (112) and ideogram (113) (y axis - % of the chromosome length of the haploid set) of male holotype. Scale bar = 5 μm." figureDoi="http://doi.org/10.5281/zenodo.6511517" httpUri="https://zenodo.org/record/6511517/files/figure.png" pageId="20" pageNumber="19">Figs. 111–112</figureCitation>
). We did not observe chiasmata during male meiosis and centromeres on chromosomes. These characteristics are typical for the scorpions from the family 
<taxonomicName id="97B74D625139FF8D5FD3DFB6FE65F9C2" authorityName="C. L. Koch" authorityYear="1837" box="[398,494,1515,1539]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
(e.g. 
<bibRefCitation id="34264B105139FF8D5C76DFB6FD73F9C2" author="Mattos" box="[555,760,1515,1539]" etAl="et al." firstAuthor="Mattos" pageId="20" pageNumber="19" pagination="121 - 136" refId="ref15306" refString="MATTOS, V. F., D. M. CELLA, L. S. CARVALHO, D. M. CANDIDO &amp; M. C. SCHNEIDER. 2013. Highchromosome variability and the presence of multivalent associations in buthid scorpions. Chromosome Research, 21: 121 - 136." type="journal article" year="2013">Mattos et al., 2013</bibRefCitation>
). The chromosomes of 
<taxonomicName id="97B74D625139FF8D5F01DC56FDD9F9E2" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[348,594,1547,1571]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5F01DC56FDD9F9E2" box="[348,594,1547,1571]" italics="true" pageId="20" pageNumber="19">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5C07DC56FD1CF9E3" box="[602,663,1547,1571]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5C07DC56FDF8F9E2" bold="true" box="[602,627,1547,1571]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5CDCDC56FD1BF9E2" bold="true" box="[641,656,1547,1571]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
gradually decrease in length from 8.69 % to 5.92 % of the halpoid set (
<figureCitation id="C88C2A645139FF8D5E24DC16FF5DF9A2" box="[121,214,1611,1635]" captionStart="Figures 111–113" captionStartId="26.[113,190,671,693]" captionTargetBox="[113,1474,235,649]" captionTargetId="figure-786@26.[113,1474,234,659]" captionTargetPageId="26" captionText="Figures 111–113: Trypanothacus barnesi gen. et sp. n., mitotic metaphase (111), postpachytene (112) and ideogram (113) (y axis - % of the chromosome length of the haploid set) of male holotype. Scale bar = 5 μm." figureDoi="http://doi.org/10.5281/zenodo.6511517" httpUri="https://zenodo.org/record/6511517/files/figure.png" pageId="20" pageNumber="19">Fig. 113</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="18AD656A5139FF8E5E2CDCD6FD8BFEA0" lastPageId="23" lastPageNumber="22" pageId="20" pageNumber="19" type="discussion">
<paragraph id="500836E15139FF8D5E2CDCD6FCFDFDA2" blockId="20.[113,772,1675,2051]" lastBlockId="20.[816,1474,235,611]" pageId="20" pageNumber="19">
AFFINITIES. In the diagnosis, we separated 
<taxonomicName id="97B74D625139FF8D5C11DCD6FD32F962" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[588,697,1675,1699]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5C11DCD6FD32F962" box="[588,697,1675,1699]" italics="true" pageId="20" pageNumber="19">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5C9EDCD6FC88F963" box="[707,771,1675,1699]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5C9EDCD6FD57F962" bold="true" box="[707,732,1675,1699]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5CB3DCD6FD76F962" bold="true" box="[750,765,1675,1699]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
from its similar allopatric congener, 
<taxonomicName id="97B74D625139FF8D5C79DCF6FD25F902" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" baseAuthorityYear="2019" box="[548,686,1707,1731]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF35139FF8D5C79DCF6FD25F902" box="[548,686,1707,1731]" italics="true" pageId="20" pageNumber="19">T. buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5CE3DCF6FF0CF923" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5CE3DCF6FD77F902" bold="true" box="[702,764,1707,1731]" pageId="20" pageNumber="19">comb</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5E2CDC96FF0BF922" bold="true" box="[113,128,1739,1763]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
by measurable differences in the shapes of the pedipalp and metasoma, indicated by non-overlapping ranges of morphometric ratios. 
<taxonomicName id="97B74D625139FF8D5F01DD56FE42F8E2" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[348,457,1803,1827]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5F01DD56FE42F8E2" box="[348,457,1803,1827]" italics="true" pageId="20" pageNumber="19">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5F8FDD56FD9AF8E3" box="[466,529,1803,1827]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5F8FDD56FE60F8E2" bold="true" box="[466,491,1803,1827]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5FA6DD56FD81F8E2" bold="true" box="[507,522,1803,1827]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
is a less robust, more slender scorpion with narrower pedipalps and posterior metasomal segments. Only limited numbers of specimens of 
<taxonomicName id="97B74D625139FF8D5ECCDD36FE9FF842" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" baseAuthorityYear="2019" box="[145,276,1899,1923]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF35139FF8D5ECCDD36FE9FF842" box="[145,276,1899,1923]" italics="true" pageId="20" pageNumber="19">T. buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5F43DD36FE09F843" box="[286,386,1899,1923]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5F43DD36FED7F842" bold="true" box="[286,348,1899,1923]" pageId="20" pageNumber="19">comb</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5F31DD36FEF0F842" bold="true" box="[364,379,1899,1923]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
were available to us for study and confirmation of species identity, and additional material may extend morphometric variation in this species. However, samples from around the 
<typeStatus id="8F0C88435139FF8D5FD3DD96FE37F822" box="[398,444,1995,2019]" pageId="20" pageNumber="19">type</typeStatus>
locality and a well separated western site (Wadi Turabah) both have robust segments, so the morphometric diagnosis is likely to remain valid. We hypothesize that 
<taxonomicName id="97B74D625139FF8D5DB4DB56FBDEFEE2" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[1001,1109,267,291]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5DB4DB56FBDEFEE2" box="[1001,1109,267,291]" italics="true" pageId="20" pageNumber="19">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5A03DB56FB17FEE3" box="[1118,1180,267,291]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5A03DB56FBFCFEE2" bold="true" box="[1118,1143,267,291]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5ADADB56FB1DFEE2" bold="true" box="[1159,1174,267,291]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
and 
<taxonomicName id="97B74D625139FF8D5A89DB56FADDFEE2" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" baseAuthorityYear="2019" box="[1236,1366,267,291]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF35139FF8D5A89DB56FADDFEE2" box="[1236,1366,267,291]" italics="true" pageId="20" pageNumber="19">T. buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5B02DB56FA49FEE3" box="[1375,1474,267,291]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5B02DB56FA16FEE2" bold="true" box="[1375,1437,267,291]" pageId="20" pageNumber="19">comb</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5BF0DB56FA37FEE2" bold="true" box="[1453,1468,267,291]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
are sister species that diverged after southern and northern populations were isolated from each other by formation of the Rub‘ al Khali (Empty Quarter) dune system during the Quaternary. Soft unstabilized sands may be an effective physical barrier blocking dispersal of ancestral 
<taxonomicName id="97B74D625139FF8D5B7EDBF6FA49FE02" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[1315,1474,427,451]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="62C3EAF35139FF8D5B7EDBF6FA49FE02" box="[1315,1474,427,451]" italics="true" pageId="20" pageNumber="19">Trypanothacus</emphasis>
</taxonomicName>
adapted to firmer substrates. This model of vicariant speciation by dune barriers has also been suggested for 
<taxonomicName id="97B74D625139FF8D5B2CDBB6FC3CFDE3" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="macroctenus">
<emphasis id="62C3EAF35139FF8D5B2CDBB6FC3CFDE3" italics="true" pageId="20" pageNumber="19">Leiurus macroctenus</emphasis>
</taxonomicName>
, a sympatric scorpion that may have substrate preferences similar to those of 
<taxonomicName id="97B74D625139FF8D5ADDD876FB67FD82" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[1152,1260,555,579]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5ADDD876FB67FD82" box="[1152,1260,555,579]" italics="true" pageId="20" pageNumber="19">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5AA9D876FABAFD83" box="[1268,1329,555,579]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5AA9D876FA86FD82" bold="true" box="[1268,1293,555,579]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5B46D876FAA1FD82" bold="true" box="[1307,1322,555,579]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
(
<bibRefCitation id="34264B105139FF8D5B1DD876FCE1FDA2" author="Lowe" etAl="et al." firstAuthor="Lowe" pageId="20" pageNumber="19" pagination="1 - 37" refId="ref15256" refString="LOWE, G., E. A. YAGMUR &amp; F. KOVARIK. 2014. A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula. Euscorpius, 100: 1 - 37." type="journal article" year="2014">Lowe et al., 2014</bibRefCitation>
).
</paragraph>
<paragraph id="500836E15139FF8D5D6DD8D6FAC2F962" blockId="20.[816,1475,651,2051]" pageId="20" pageNumber="19">
COMMENTS ON LOCALITIES AND LIFE STRATEGY.The 
<typeStatus id="8F0C88435139FF8D5B1DD8D6FAE5FD62" box="[1344,1390,651,675]" pageId="20" pageNumber="19">type</typeStatus>
locality is a flat desert plateau along route 42, the road between Shalim and Ash Shuwaymiyyah near the coast. The substrate consists of reddish soil alternating with bare rock, clay and sand. Shrubs and trees are scattered and provide only slight patches of humidity in an otherwise arid area. There are few if any loose rocks, except in disturbed areas where stones may be piled near the road by construction activity. The weather on both collection dates was mild at night (ca. 22 °C) with high humidity, barely any wind and no clouds to block the bright moon. At night, 
<taxonomicName id="97B74D625139FF8D5DBAD996FB54FC22" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[999,1247,971,995]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5DBAD996FB54FC22" box="[999,1247,971,995]" italics="true" pageId="20" pageNumber="19">Trypanothacus barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5AB4D996FAA3FC23" box="[1257,1320,971,995]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5AB4D996FA89FC22" bold="true" box="[1257,1282,971,995]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5B4FD996FAAAFC22" bold="true" box="[1298,1313,971,995]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
was the most active scorpion in this area. Males were highly active on open ground, whilst females were resting in ambush position at the entrances of their burrows. The burrows were located in sandy areas at the bases of shrubs or rocks, and only rarely on open ground. Only small juveniles to medium-sized immatures were found under rocks without a burrow. Far more males than females were observed on both nights. Remarkably, many were observed actively foraging and feeding on termites, close to the insect nests and trails. Several termites were captured and consumed simultaneously by one scorpion. Other potential prey, including various insects (
<taxonomicName id="97B74D625139FF8D5B1DDF76FBBAFAA2" authority=", Heteroptera, Coleoptera" authorityName="Heteroptera, Coleoptera" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="family">Formicidae, Heteroptera, Coleoptera</taxonomicName>
, 
<taxonomicName id="97B74D625139FF8D5A64DF16FB29FAA2" authorityName="Brunner von Wattenwyl" authorityYear="1882" box="[1081,1186,1355,1379]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="19" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
, lepismatids) and arachnids (ticks, spiders and pseudoscorpions) were abundant in the area. Mobile white mites were observed on many individuals of 
<taxonomicName id="97B74D625139FF8D5D0DDFF6FC36FA02" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[848,957,1451,1475]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF35139FF8D5D0DDFF6FC36FA02" box="[848,957,1451,1475]" italics="true" pageId="20" pageNumber="19">T. barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885139FF8D5D9ADFF6FB8DFA03" box="[967,1030,1451,1475]" pageId="20" pageNumber="19" rank="species">
<emphasis id="62C3EAF35139FF8D5D9ADFF6FC6BFA02" bold="true" box="[967,992,1451,1475]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="62C3EAF35139FF8D5DADDFF6FC74FA02" bold="true" box="[1008,1023,1451,1475]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
(cf. 
<figureCitation id="C88C2A645139FF8D5A60DFF6FB05FA03" box="[1085,1166,1450,1475]" captionStart="Figures 84–85" captionStartId="17.[116,193,1988,2010]" captionTargetBox="[113,1474,248,1949]" captionTargetId="figure-35@17.[113,1474,248,1949]" captionTargetPageId="17" captionText="Figures 84–85: Trypanothacus barnesi gen. et sp. n., in vivo habitus, male holotype (84) and female paratype from type locality (85)." figureDoi="http://doi.org/10.5281/zenodo.6511498" httpUri="https://zenodo.org/record/6511498/files/figure.png" pageId="20" pageNumber="19">Fig. 85</figureCitation>
, an adult female with mites on tergite II and carapace), but not on other arthropods in this habitat. Some individual scorpions were densely covered with mites, whereas others bore only a few. Other species of scorpions found in this area were: 
<taxonomicName id="97B74D625139FF8D5AEDDC76FC56F9A2" authority="(Olivier, 1807)" baseAuthorityName="Olivier" baseAuthorityYear="1807" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="62C3EAF35139FF8D5AEDDC76FA49F982" box="[1200,1474,1579,1603]" italics="true" pageId="20" pageNumber="19">Androctonus crassicauda</emphasis>
(Olivier, 1807)
</taxonomicName>
, 
<taxonomicName id="97B74D625139FF8D5DA9DC16FC44F942" authority="Lowe, Yagmur &amp; Kovarik, 2014" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="macroctenus">
<emphasis id="62C3EAF35139FF8D5DA9DC16FB54F9A3" box="[1012,1247,1611,1634]" italics="true" pageId="20" pageNumber="19">Leiurus macroctenus</emphasis>
<bibRefCitation id="34264B105139FF8D5AACDC16FC44F942" author="LOWE, G. &amp; E. A. YAGMUR &amp; F. KOVARIK" journalOrPublisher="Euscorpius" pageId="20" pageNumber="19" pagination="1 - 37" part="100" refId="ref15256" refString="LOWE, G., E. A. YAGMUR &amp; F. KOVARIK. 2014. A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula. Euscorpius, 100: 1 - 37." title="A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula" type="journal article" year="2014">Lowe, Yağmur &amp; Kovařík, 2014</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="97B74D625139FF8D5D81DC36FAC4F942" authority="Lowe, 2010" authorityName="Lowe" authorityYear="2010" box="[988,1359,1643,1667]" class="Arachnida" family="Buthidae" genus="Femtobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="shutuae">
<emphasis id="62C3EAF35139FF8D5D81DC36FB48F942" box="[988,1219,1643,1667]" italics="true" pageId="20" pageNumber="19">Femtobuthus shutuae</emphasis>
<bibRefCitation id="34264B105139FF8D5A96DC36FAC4F942" author="LOWE, G." box="[1227,1359,1643,1667]" journalOrPublisher="Euscorpius" pageId="20" pageNumber="19" pagination="1 - 22" part="96" refId="ref15167" refString="LOWE, G. 2010 a. A new species of Odontobuthus (Scorpiones: Buthidae) from northern Oman. Euscorpius, 96: 1 - 22." title="A new species of Odontobuthus (Scorpiones: Buthidae) from northern Oman" type="journal article" year="2010">Lowe, 2010</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="97B74D625139FF8D5B01DC36FA36F943" box="[1372,1469,1643,1667]" class="Arachnida" family="Diplocentridae" genus="Nebo" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="62C3EAF35139FF8D5B01DC36FA1FF942" box="[1372,1428,1643,1667]" italics="true" pageId="20" pageNumber="19">Nebo</emphasis>
sp.
</taxonomicName>
, and 
<taxonomicName id="97B74D625139FF8D5D03DCD6FACEF962" authority="(Simon, 1882)" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[862,1349,1675,1699]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="62C3EAF35139FF8D5D03DCD6FB29F963" box="[862,1186,1675,1699]" italics="true" pageId="20" pageNumber="19">Compsobuthus acutecarinatus</emphasis>
(Simon, 1882)
</taxonomicName>
.
</paragraph>
<paragraph id="500836E15139FF8F5D03DCF6FE77F98B" blockId="20.[816,1475,651,2051]" lastBlockId="22.[113,771,1331,1611]" lastPageId="22" lastPageNumber="21" pageId="20" pageNumber="19">
Both the type locality and the 
<typeStatus id="8F0C88435139FF8D5AF4DCF6FA8EF902" box="[1193,1285,1707,1731]" pageId="20" pageNumber="19" type="paratype">paratype</typeStatus>
collection site of Yalooni (Arabian Oryx sanctuary) are located on the Jiddat Al Harasis, an extensive stony limestone plateau in central 
<collectingCountry id="28A076715139FF8D5B27DCB6FA34F8C2" box="[1402,1471,1771,1795]" name="Oman" pageId="20" pageNumber="19">Oman</collectingCountry>
. The region is ecologically classified as coastal fog desert. Moisture is deposited by condensation of humid air swept inland from the sea during the summer monsoon (khareef). This sustains local vegetation and animal populations, including the scorpion fauna. The other collection sites around Thumrait are located on the Nejd Desert plain in the rain shadow of the Jabal Qara mountains. Without monsoon mist, the terrain is more arid, consisting mostly of sand and gravel flats with sporadic rocky outcrops. Drainage in some shallow wadi beds supports scattered trees and shrubs, with xerophytic herbs and grass as low ground cover. In this area, 
<taxonomicName id="97B74D62513BFF8F5E2CDFCEFF6BFA6A" authority="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský, 2019" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" box="[113,224,1427,1451]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
<emphasis id="62C3EAF3513BFF8F5E2CDFCEFF0BFA6B" box="[113,128,1427,1450]" italics="true" pageId="22" pageNumber="21">T</emphasis>
. 
<emphasis id="62C3EAF3513BFF8F5ECDDFCEFF6BFA6A" box="[144,224,1427,1451]" italics="true" pageId="22" pageNumber="21">barnesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513BFF8F5EB7DFCEFEA2FA6B" box="[234,297,1427,1451]" pageId="22" pageNumber="21" rank="species">
<emphasis id="62C3EAF3513BFF8F5EB7DFCEFE88FA6A" bold="true" box="[234,259,1427,1451]" pageId="22" pageNumber="21">sp</emphasis>
. 
<emphasis id="62C3EAF3513BFF8F5F4EDFCEFEA9FA6A" bold="true" box="[275,290,1427,1451]" pageId="22" pageNumber="21">n</emphasis>
.
</taxonomicNameLabel>
was reported to be present but uncommon (J.N. Barnes, personal communication, 
<date id="24091021513BFF8F5C4EDFEEFD1CFA0A" box="[531,663,1459,1483]" pageId="22" pageNumber="21" value="1992-07-05">05.VII.1992</date>
, 
<figureCitation id="C88C2A64513BFF8F5CFCDFEEFF04FA2A" captionStart="Figures 86–89" captionStartId="18.[113,190,1969,1991]" captionTargetBox="[114,1474,236,1939]" captionTargetId="figure-12@18.[114,1474,234,1939]" captionTargetPageId="18" captionText="Figures 86–89: Trypanothacus barnesi gen. et sp. n.. Figures 86–87. Male (86) and female (87) from Thumrait, photos by J. N. Barnes. Figures 88–89. Type locality, Oman, S of Schelim, 18.041070°N 55.612262°E." figureDoi="http://doi.org/10.5281/zenodo.6511503" httpUri="https://zenodo.org/record/6511503/files/figure.png" pageId="22" pageNumber="21">Figs. 86– 87</figureCitation>
). Other scorpions recorded from there were: 
<taxonomicName id="97B74D62513BFF8F5C21DF8EFF79F9CA" baseAuthorityName="Olivier" baseAuthorityYear="1807" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="62C3EAF3513BFF8F5C21DF8EFF79F9CA" italics="true" pageId="22" pageNumber="21">Androctonus crassicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="97B74D62513BFF8F5EA2DFAEFE56F9CB" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" box="[255,477,1523,1546]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="macroctenus">
<emphasis id="62C3EAF3513BFF8F5EA2DFAEFE56F9CB" box="[255,477,1523,1546]" italics="true" pageId="22" pageNumber="21">Leiurus macroctenus</emphasis>
</taxonomicName>
, 
<taxonomicName id="97B74D62513BFF8F5FB4DFAEFF2DF9EA" authority="Simon, 1910" authorityName="Simon" authorityYear="1910" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="innesi">
<emphasis id="62C3EAF3513BFF8F5FB4DFAEFD3AF9CB" box="[489,689,1523,1547]" italics="true" pageId="22" pageNumber="21">Orthochirus innesi</emphasis>
Simon, 1910
</taxonomicName>
and 
<taxonomicName id="97B74D62513BFF8F5EBDDC4EFD11F9EA" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[224,666,1555,1579]" class="Arachnida" family="Buthidae" genus="Xenobuthus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="anthracinus">
<emphasis id="62C3EAF3513BFF8F5EBDDC4EFE6CF9EA" box="[224,487,1555,1579]" italics="true" pageId="22" pageNumber="21">Xenobuthus anthracinus</emphasis>
(Pocock, 1895)
</taxonomicName>
. The first two of these were the most common.
</paragraph>
<caption id="04C866695138FF8C5E2CDDE4FCB4F82A" ID-DOI="http://doi.org/10.5281/zenodo.6511505" ID-Zenodo-Dep="6511505" httpUri="https://zenodo.org/record/6511505/files/figure.png" pageId="21" pageNumber="20" startId="21.[113,190,1977,1999]" targetBox="[131,1442,254,1923]" targetPageId="21">
<paragraph id="500836E15138FF8C5E2CDDE4FCB4F82A" blockId="21.[113,1474,1977,2028]" pageId="21" pageNumber="20">
<emphasis id="62C3EAF35138FF8C5E2CDDE4FE88F80F" bold="true" box="[113,259,1977,1999]" pageId="21" pageNumber="20">Figures 90–93</emphasis>
: 
<taxonomicName id="97B74D625138FF8C5F4EDDE4FD8DF80F" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Hendrixson" baseAuthorityYear="2006" box="[275,518,1977,1998]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF35138FF8C5F4EDDE4FD8DF80F" box="[275,518,1977,1998]" italics="true" pageId="21" pageNumber="20">Trypanothacus buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F057885138FF8C5C52DDE4FDE1F80F" box="[527,618,1977,1998]" pageId="21" pageNumber="20" rank="species">
<emphasis id="62C3EAF35138FF8C5C52DDE4FDCCF80F" bold="true" box="[527,583,1977,1998]" pageId="21" pageNumber="20">comb</emphasis>
. 
<emphasis id="62C3EAF35138FF8C5C0BDDE4FDE8F80F" bold="true" box="[598,611,1977,1998]" pageId="21" pageNumber="20">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="62C3EAF35138FF8C5C2EDDE4FC8FF80F" bold="true" box="[627,772,1977,1999]" pageId="21" pageNumber="20">Figures 90–91</emphasis>
. Male holotype in dorsal (90) and ventral (91) views. 
<emphasis id="62C3EAF35138FF8C5B77DDE4FA30F80F" bold="true" box="[1322,1467,1977,1999]" pageId="21" pageNumber="20">Figures 92–93</emphasis>
: Female paratype in dorsal (92) and ventral (93) views. Scale bars: 10 mm.
</paragraph>
</caption>
<caption id="04C86669513BFF8F5E2CDEB6FE12FADC" ID-DOI="http://doi.org/10.5281/zenodo.6511509" ID-Zenodo-Dep="6511509" httpUri="https://zenodo.org/record/6511509/files/figure.png" pageId="22" pageNumber="21" startId="22.[113,190,1259,1281]" targetBox="[113,1474,234,1247]" targetPageId="22">
<paragraph id="500836E1513BFF8F5E2CDEB6FE12FADC" blockId="22.[113,1474,1259,1310]" pageId="22" pageNumber="21">
<emphasis id="62C3EAF3513BFF8F5E2CDEB6FE8BFAC1" bold="true" box="[113,256,1259,1281]" pageId="22" pageNumber="21">Figures 94–95</emphasis>
: 
<taxonomicName id="97B74D62513BFF8F5F51DEB6FE77FAC1" authorityName="Lowe &amp; Kovařík &amp; Stockmann &amp; Šťáhlavský" authorityYear="2019" baseAuthorityName="Hendrixson" baseAuthorityYear="2006" box="[268,508,1259,1280]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
<emphasis id="62C3EAF3513BFF8F5F51DEB6FE77FAC1" box="[268,508,1259,1280]" italics="true" pageId="22" pageNumber="21">Trypanothacus buettikeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F05788513BFF8F5C5CDEB6FDDCFAC1" box="[513,599,1259,1280]" pageId="22" pageNumber="21" rank="species">
<emphasis id="62C3EAF3513BFF8F5C5CDEB6FDB2FAC1" bold="true" box="[513,569,1259,1280]" pageId="22" pageNumber="21">comb</emphasis>
. 
<emphasis id="62C3EAF3513BFF8F5C19DEB6FDDAFAC1" bold="true" box="[580,593,1259,1280]" pageId="22" pageNumber="21">n</emphasis>
.
</taxonomicNameLabel>
Male holotype, carapace and tergites I–IV (94), coxosternal area and sternites I–II (95). UV fluorescence. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="500836E1513BFF8F5E2CDC2EFBC2FA4A" blockId="22.[113,772,1651,2027]" lastBlockId="22.[816,1475,1331,2027]" pageId="22" pageNumber="21">
CAPTIVE REARING AND BREEDING.Two medium-sized immatures (
<specimenCount id="46B1FD68513BFF8F5E26DCCEFF16F96A" box="[123,157,1683,1707]" pageId="22" pageNumber="21" type="male">1♂</specimenCount>
, 
<specimenCount id="46B1FD68513BFF8F5EF2DCCEFF45F96A" box="[175,206,1683,1707]" pageId="22" pageNumber="21" type="female">1♀</specimenCount>
) were reared to maturity in captivity. They were housed in 
<quantity id="974F9B04513BFF8F5EB6DCEEFEB9F90B" box="[235,306,1715,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="21" unit="cm" value="11.0">11 cm</quantity>
× 
<quantity id="974F9B04513BFF8F5F04DCEEFE18F90A" box="[345,403,1715,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="21" unit="cm" value="11.0">11cm</quantity>
× 
<quantity id="974F9B04513BFF8F5FE1DCEEFE7DF90B" box="[444,502,1715,1739]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="21" unit="cm" value="6.0">6 cm</quantity>
plastic boxes with two perforated sides, filled with a thin layer of loose sand, and were provided with a small shelter. A burrowing substrate was not provided in order to more easily observe behavior. Average temperatures in summer were 32°C (day) and 24°C (night). In winter, a mild diapause was simulated with temperatures of about 25°C (day) and 20°C (night). One corner of the box was slightly moistened by spraying water once a week to provide drinking water. The water completely evaporated in the following day and conditions were kept dry for the rest of the week to prevent mycosis. Insects (
<taxonomicName id="97B74D62513BFF8F5A8FDF6EFA30FA8A" baseAuthorityName="Packard" baseAuthorityYear="1837" box="[1234,1467,1331,1355]" class="Insecta" family="Lepismatidae" genus="Thermobia" kingdom="Animalia" order="Zygentoma" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="domestica">
<emphasis id="62C3EAF3513BFF8F5A8FDF6EFA30FA8A" box="[1234,1467,1331,1355]" italics="true" pageId="22" pageNumber="21">Thermobia domestica</emphasis>
</taxonomicName>
, 
<taxonomicName id="97B74D62513BFF8F5D6DDF0EFB93FAAA" box="[816,1048,1363,1387]" class="Insecta" family="Blattidae" genus="Shelfordella" kingdom="Animalia" order="Blattodea" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="62C3EAF3513BFF8F5D6DDF0EFB93FAAA" box="[816,1048,1363,1387]" italics="true" pageId="22" pageNumber="21">Shelfordella lateralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="97B74D62513BFF8F5A07DF0EFAA2FAAA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1114,1321,1363,1387]" class="Insecta" family="Gryllidae" genus="Acheta" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="domesticus">
<emphasis id="62C3EAF3513BFF8F5A07DF0EFAA2FAAA" box="[1114,1321,1363,1387]" italics="true" pageId="22" pageNumber="21">Acheta domesticus</emphasis>
</taxonomicName>
) of adequate sizes were offered as prey.
</paragraph>
<paragraph id="500836E1513BFF8E5D03DFCEFD8BFEA0" blockId="22.[816,1475,1331,2027]" lastBlockId="23.[113,771,233,353]" lastPageId="23" lastPageNumber="22" pageId="22" pageNumber="21">
Three weeks after attaining maturity, the female was mated (
<date id="24091021513BFF8F5D22DFEEFC6AFA0A" box="[895,993,1459,1483]" pageId="22" pageNumber="21" value="2017-11">XI. 2017</date>
). Upon being introduced to the female, a male immediately initiated courtship without preliminary behavior like juddering. The promenade à deux began with the male grasping the female pedipalps, and continued with a series of back and forth pulling motions by both sexes (
<figureCitation id="C88C2A64513BFF8F5B40DC6EFAF2F98A" box="[1309,1401,1587,1611]" captionStart="Figures 114–116" captionStartId="27.[113,190,1990,2012]" captionTargetBox="[113,1474,234,1972]" captionTargetId="figure-7@27.[113,1474,234,1972]" captionTargetPageId="27" captionText="Figures 114–116: Trypanothacus barnesi gen. et sp. n., promenade à deux (courtship dance) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi="http://doi.org/10.5281/zenodo.6511519" httpUri="https://zenodo.org/record/6511519/files/figure.png" pageId="22" pageNumber="21">Fig. 114</figureCitation>
). Male pectinal movements occurred almost continually, without lengthy interruptions. Remarkably, both sexes engaged in sand scraping with legs I–III (
<figureCitation id="C88C2A64513BFF8F5A1BDCCEFB2DF96A" box="[1094,1190,1683,1707]" captionStart="Figures 114–116" captionStartId="27.[113,190,1990,2012]" captionTargetBox="[113,1474,234,1972]" captionTargetId="figure-7@27.[113,1474,234,1972]" captionTargetPageId="27" captionText="Figures 114–116: Trypanothacus barnesi gen. et sp. n., promenade à deux (courtship dance) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi="http://doi.org/10.5281/zenodo.6511519" httpUri="https://zenodo.org/record/6511519/files/figure.png" pageId="22" pageNumber="21">Fig. 115</figureCitation>
), sometimes settling their bodies into depressions they excavated in the loose sand to rest for a few seconds (
<figureCitation id="C88C2A64513BFF8F5DA4DC8EFBD8F92A" box="[1017,1107,1747,1771]" captionStart="Figures 114–116" captionStartId="27.[113,190,1990,2012]" captionTargetBox="[113,1474,234,1972]" captionTargetId="figure-7@27.[113,1474,234,1972]" captionTargetPageId="27" captionText="Figures 114–116: Trypanothacus barnesi gen. et sp. n., promenade à deux (courtship dance) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi="http://doi.org/10.5281/zenodo.6511519" httpUri="https://zenodo.org/record/6511519/files/figure.png" pageId="22" pageNumber="21">Fig. 116</figureCitation>
) before continuing the promenade à deux. Several episodes of cheliceral massage lasting several seconds were observed (
<figureCitation id="C88C2A64513BFF8F5A1CDD4EFB2AF8EA" box="[1089,1185,1811,1835]" captionStart="Figures 117–118" captionStartId="28.[113,190,1984,2006]" captionTargetBox="[113,1474,236,1962]" captionTargetId="figure-11@28.[113,1474,236,1964]" captionTargetPageId="28" captionText="Figures 117–118: Trypanothacus barnesi gen. et sp. n., cheliceral massage (117) and spermatophore deposition (118) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi="http://doi.org/10.5281/zenodo.6511521" httpUri="https://zenodo.org/record/6511521/files/figure.png" pageId="22" pageNumber="21">Fig. 117</figureCitation>
), but there was no sexual sting. After 17 min 48 s, a spermatophore was deposited by the male on a flat rock (
<figureCitation id="C88C2A64513BFF8F5A6EDD0EFB06F8AA" box="[1075,1165,1875,1899]" captionStart="Figures 117–118" captionStartId="28.[113,190,1984,2006]" captionTargetBox="[113,1474,236,1962]" captionTargetId="figure-11@28.[113,1474,236,1964]" captionTargetPageId="28" captionText="Figures 117–118: Trypanothacus barnesi gen. et sp. n., cheliceral massage (117) and spermatophore deposition (118) in a mating pair of individuals collected from the type locality. Photos taken in laboratory. Male is on the right." figureDoi="http://doi.org/10.5281/zenodo.6511521" httpUri="https://zenodo.org/record/6511521/files/figure.png" pageId="22" pageNumber="21">Fig. 118</figureCitation>
) and taken up by the female 3 s later. Both individuals then immediately separated, and the female began to display aggressive behavior towards the male, but did not attempt mate cannibalism. The female was subsequently reared under the same conditions as described above, and gave birth to 
<specimenCount id="46B1FD68513AFF8E5F23DAB4FD8BFEC0" box="[382,512,233,257]" pageId="23" pageNumber="22" type="juvenile">23 juveniles</specimenCount>
in the following season (
<date id="24091021513AFF8E5E24DB54FF63FEE0" box="[121,232,265,289]" pageId="23" pageNumber="22" value="2018-07">VII. 2018</date>
). These were separated from the parent one week after their first ecdysis, and successfully reared under the same conditions as the parental generation.
</paragraph>
</subSubSection>
</treatment>
</document>