509 lines
63 KiB
XML
509 lines
63 KiB
XML
<document id="20C6311DDEB77F918C5E69A1E0616DCA" ID-CLB-Dataset="21869" ID-DOI="10.5852/ejt.2016.236" ID-GBIF-Dataset="2dfee902-a7fc-472b-bf1d-27029b3504f3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3847840" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590436812908" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2016" docId="C146B323FFBCFFEFFD4C9FD8FE3AF17B" docLanguage="en" docName="ejt-2016-236.pdf.imf" docOrigin="European Journal of Taxonomy 236" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella davisae Herbert 2016, sp. nov." docType="treatment" docUuid="DDCAA18B-CC50-4EC1-B63B-28ABAF6904C2" docUuidSource="ZooBank" docVersion="10" lastPageNumber="7" masterDocId="3D7FCB5BFFBEFFE9FFC0994BFF82F57F" masterDocTitle="New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="3" updateTime="1698841246942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3BFDF886C9E260FD60F0938F13C4D801">New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)</mods:title>
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<mods:name id="D86B0E67903712BC3F3E3BB65CF3EE83" type="personal">
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<mods:roleTerm id="7784F7359E753D8E6AC603AB3DB38AD3">Author</mods:roleTerm>
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<mods:namePart id="4921A8CD243741BD28EF1301F05943E3">Herbert, David G.</mods:namePart>
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<mods:typeOfResource id="59A7116F141F5F5FF5327D154BCF1484">text</mods:typeOfResource>
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<mods:title id="38B11D8D15F44BE2FC4A864D5C0CC50A">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="7B79F62312A9B8D031D65708E09579EA">
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<mods:date id="7A4679C0FFD3806F6948D5F0A4C7C327">2016</mods:date>
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<mods:number id="09573785F516F816C45A9695C116225E">2016-10-12</mods:number>
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</mods:detail>
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<mods:detail id="4DAD960112E3840C8188F655731E8341" type="volume">
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<mods:number id="5C1585FF2AFA4FE8DCACA0A17CAD7553">236</mods:number>
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<mods:classification id="C9080F85FD08FF56C6528A39CC93CC82">journal article</mods:classification>
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<mods:identifier id="8F5316977C9DD0DF9C90DA55B6BA3DFD" type="CLB-Dataset">21869</mods:identifier>
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<mods:identifier id="63E7D96A0CE9D1D9A8B5AE0D2D3F8E17" type="DOI">10.5852/ejt.2016.236</mods:identifier>
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<mods:identifier id="DDB58EA9764F6639970210AF725EEE23" type="GBIF-Dataset">2dfee902-a7fc-472b-bf1d-27029b3504f3</mods:identifier>
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<mods:identifier id="37B32B84E587C84FF0485A45E395DE88" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="E9BD2BF71E7569DAEF1F8A36730AD22D" type="Zenodo-Dep">3847840</mods:identifier>
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<mods:identifier id="D49DD4EBE1595C783A65200E23291BCD" type="ZooBank">B51BF718-79F5-47F5-8740-BA181CE88257</mods:identifier>
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</mods:mods>
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<treatment id="C146B323FFBCFFEFFD4C9FD8FE3AF17B" ID-DOI="http://doi.org/10.5281/zenodo.3854746" ID-GBIF-Taxon="164282650" ID-Zenodo-Dep="3854746" LSID="urn:lsid:zoobank.org:act:DDCAA18B-CC50-4EC1-B63B-28ABAF6904C2" httpUri="http://treatment.plazi.org/id/C146B323FFBCFFEFFD4C9FD8FE3AF17B" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
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<subSubSection id="01F551BEFFBCFFEBFD4C9FD8FB55F3AE" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="49500235FFBCFFEBFD4C9FD8FC2AF3D1" blockId="2.[349,1239,1683,1780]" box="[652,936,1683,1710]" pageId="2" pageNumber="3">
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<heading id="1218B559FFBCFFEBFD4C9FD8FC2AF3D1" box="[652,936,1683,1710]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="8EEF79B6FFBCFFEBFD4C9FD8FCC5F3D1" ID-CoL="3HLQ2" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[652,839,1683,1710]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
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||
<emphasis id="7B9BDE27FFBCFFEBFD4C9FD8FCC5F3D1" bold="true" box="[652,839,1683,1710]" italics="true" pageId="2" pageNumber="3">Gulella davisae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="60A8635CFFBCFFEBFC8E9FDFFC2AF3D1" box="[846,936,1684,1710]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="49500235FFBCFFEBFE9D9FFDFB55F3AE" blockId="2.[349,1239,1683,1780]" box="[349,1239,1718,1745]" pageId="2" pageNumber="3">
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<uri id="3D7E0E37FFBCFFEBFE9D9FFDFB55F3AE" box="[349,1239,1718,1745]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="3D4938E0FFBCFFEBFDBB9FFDFB55F3AE" box="[635,1239,1718,1745]" pageId="2" pageNumber="3">DDCAA18B-CC50-4EC1-B63B-28ABAF6904C2</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="01F551BEFFBCFFEBFD279F91FCCFF38B" box="[743,845,1754,1780]" pageId="2" pageNumber="3" type="description">
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<paragraph id="49500235FFBCFFEBFD279F91FCCFF38B" blockId="2.[349,1239,1683,1780]" box="[743,845,1754,1780]" pageId="2" pageNumber="3">
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<figureCitation id="D1D41EB0FFBCFFEBFD279F91FCCFF38B" box="[743,845,1754,1780]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1928,1954]" captionStartId-1="5.[189,232,1964,1990]" captionTargetBox-0="[192,1396,881,1877]" captionTargetBox-1="[196,1385,1293,1916]" captionTargetId-0="figure@3.[189,1399,877,1878]" captionTargetId-1="figure@5.[189,1399,1285,1923]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Fig. 1. Gulella davisae sp. nov. A–D. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086). E. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091)." captionText-1="Fig. 2. Gulella davisae sp. nov., genitalia, paratype (NMSA P0269/T4090). A. Distal portion of genital system. B. Internal structure of penis." figureDoi-0="http://doi.org/10.5281/zenodo.3847844" figureDoi-1="http://doi.org/10.5281/zenodo.3847848" httpUri-0="https://zenodo.org/record/3847844/files/figure.png" httpUri-1="https://zenodo.org/record/3847848/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="01F551BEFFBCFFEBFF7D9E50FCEFF2F7" pageId="2" pageNumber="3" type="etymology">
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||
<paragraph id="49500235FFBCFFEBFF7D9E50FEC6F24A" blockId="2.[189,324,1819,1845]" box="[189,324,1819,1845]" pageId="2" pageNumber="3">
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||
<heading id="1218B559FFBCFFEBFF7D9E50FEC6F24A" bold="true" box="[189,324,1819,1845]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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||
<emphasis id="7B9BDE27FFBCFFEBFF7D9E50FEC6F24A" bold="true" box="[189,324,1819,1845]" pageId="2" pageNumber="3">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="49500235FFBCFFEBFF7D9E01FCEFF2F7" blockId="2.[189,1398,1866,1928]" pageId="2" pageNumber="3">
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||
Named for Linda Davis, manager of the Mollusca collection at the
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<collectingRegion id="8B2BCCD7FFBCFFEBFC169E01FB0CF21B" box="[982,1166,1866,1892]" country="South Africa" name="KwaZulu-Natal" pageId="2" pageNumber="3">KwaZulu-Natal</collectingRegion>
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Museum and a key member of the institution’s malacological field work team.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="01F551BEFFBDFFEDFF7D9840FB51F660" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
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||
<paragraph id="49500235FFBDFFEAFF7D9840FE28F45A" blockId="3.[189,426,267,293]" box="[189,426,267,293]" pageId="3" pageNumber="4">
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<heading id="1218B559FFBDFFEAFF7D9840FE28F45A" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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||
<emphasis id="7B9BDE27FFBDFFEAFF7D9840FE28F45A" bold="true" box="[189,426,267,293]" pageId="3" pageNumber="4">Material examined</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="49500235FFBDFFEAFF2A9871FEDEF42B" blockId="3.[189,1399,314,446]" box="[234,348,314,340]" pageId="3" pageNumber="4">
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||
<emphasis id="7B9BDE27FFBDFFEAFF2A9871FEDEF42B" bold="true" box="[234,348,314,340]" pageId="3" pageNumber="4">
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||
<typeStatus id="9654BC97FFBDFFEAFF2A9871FEDEF42B" box="[234,348,314,340]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
</paragraph>
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||
<paragraph id="49500235FFBDFFEAFF7D9816FBD4F4C1" blockId="3.[189,1399,314,446]" pageId="3" pageNumber="4">
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||
<materialsCitation id="F9870868FFBDFFEAFF7D9816FBD4F4C1" ID-GBIF-Occurrence="2625465303" collectingDate="2014-12-04" collectionCode="NMSA" collectorName="D. Herbert & L. Davis & M. Cole" country="South Africa" elevation="1700" latitude="-24.56128" location="Mariepskop Forest Reserve" longLatPrecision="1" longitude="30.86367" pageId="3" pageNumber="4" specimenCode="P0418" specimenCount="1" stateProvince="Mpumalanga" typeStatus="holotype">
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||
<collectingCountry id="31F842A5FFBDFFEAFF7D9816FE18F408" box="[189,410,349,375]" name="South Africa" pageId="3" pageNumber="4">SOUTH AFRICA</collectingCountry>
|
||
:
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||
<collectingRegion id="8B2BCCD7FFBDFFEAFE6A9816FDCAF408" box="[426,584,349,375]" country="South Africa" name="Mpumalanga" pageId="3" pageNumber="4">Mpumalanga</collectingRegion>
|
||
,
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||
<location id="4C3054EEFFBDFFEAFD999816FC2BF408" LSID="urn:lsid:plazi:treatment:C146B323FFBCFFEFFD4C9FD8FE3AF17B:4C3054EEFFBDFFEAFD999816FC2BF408" box="[601,937,349,375]" country="South Africa" latitude="-24.56128" longLatPrecision="1" longitude="30.86367" name="Mariepskop Forest Reserve" pageId="3" pageNumber="4" stateProvince="Mpumalanga">Mariepskop Forest Reserve</location>
|
||
,
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||
<geoCoordinate id="2CDB64F2FFBDFFEAFC7A9817FBD0F408" box="[954,1106,348,375]" degrees="24.56128" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-24.56128">
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||
24.56128°
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||
<collectionCode id="2FFE9AF0FFBDFFEAFB809816FBD0F408" box="[1088,1106,349,375]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="3" pageNumber="4" type="Herbarium">S</collectionCode>
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||
</geoCoordinate>
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||
,
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||
<geoCoordinate id="2CDB64F2FFBDFFEAFBA59817FB7DF408" box="[1125,1279,348,375]" degrees="30.86367" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="30.86367">
|
||
30.86367°
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||
<collectionCode id="2FFE9AF0FFBDFFEAFB2B9816FB7DF408" box="[1259,1279,349,375]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
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||
</geoCoordinate>
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||
,
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||
<quantity id="8E17AFD0FFBDFFEAFAD19816FAF3F408" box="[1297,1393,349,375]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="4" unit="m" value="1700.0">
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||
<elevation id="C2C2E506FFBDFFEAFAD19816FAF3F408" box="[1297,1393,349,375]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="4" unit="m" value="1700.0">1700 m</elevation>
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||
</quantity>
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||
, scrubby vegetation between road and forest, in leaf-litter,
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||
<collectorName id="E41A67E3FFBDFFEAFCA498CBFC67F4E5" box="[868,997,384,410]" pageId="3" pageNumber="4">D. Herbert</collectorName>
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||
,
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||
<collectorName id="E41A67E3FFBDFFEAFC3198CBFBD7F4E5" box="[1009,1109,384,410]" pageId="3" pageNumber="4">
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||
<collectionCode id="2FFE9AF0FFBDFFEAFC3198CBFB84F4E5" box="[1009,1030,384,410]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="3" pageNumber="4" type="Herbarium">L</collectionCode>
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||
. Davis
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||
</collectorName>
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||
&
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||
<collectorName id="E41A67E3FFBDFFEAFBBC98CBFB5DF4E5" box="[1148,1247,384,410]" pageId="3" pageNumber="4">
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||
<collectionCode id="2FFE9AF0FFBDFFEAFBBC98CBFB18F4E5" box="[1148,1178,384,410]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="3" pageNumber="4" type="Herbarium">M</collectionCode>
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||
. Cole
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||
</collectorName>
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||
, stn 14-26,
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||
<date id="3D5124F5FFBDFFEAFAA898CBFEBBF4C2" pageId="3" pageNumber="4" value="2014-12-04">
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||
<collectingDate id="2D15DD1DFFBDFFEAFAA898CBFEBBF4C2" pageId="3" pageNumber="4" value="2014-12-04">4 Dec. 2014</collectingDate>
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||
</date>
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||
, height
|
||
<quantity id="8E17AFD0FFBDFFEAFE5598E8FE70F4C1" box="[405,498,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="4" unit="mm" value="6.1">6.1 mm</quantity>
|
||
, diameter
|
||
<quantity id="8E17AFD0FFBDFFEAFDAB98E8FD47F4C1" box="[619,709,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="4" unit="mm" value="2.9">2.9 mm</quantity>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFD1498E8FCABF4C2" box="[724,809,419,445]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="1949AA4EFFBDFFEAFCF098E8FCFEF4C2" box="[816,892,419,445]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFCF098E8FCC2F4C2" box="[816,832,419,445]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="4" type="Herbarium">P</collectionCode>
|
||
0418
|
||
</specimenCode>
|
||
/
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFC4598E8FC15F4C2" box="[901,919,419,445]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
|
||
4086, dry shell).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBDFFEAFF2A98AEFD2FF480" blockId="3.[189,1399,484,828]" box="[234,685,484,511]" pageId="3" pageNumber="4">
|
||
<emphasis id="7B9BDE27FFBDFFEAFF2A98AEFEE5F480" bold="true" box="[234,359,485,511]" pageId="3" pageNumber="4">
|
||
<typeStatus id="9654BC97FFBDFFEAFF2A98AEFEE5F480" box="[234,359,485,511]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(listed from north to south)
|
||
</paragraph>
|
||
<paragraph id="49500235FFBDFFEDFF7D9B43FD6AF486" blockId="3.[189,1399,484,828]" lastBlockId="4.[189,1399,267,505]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<collectingCountry id="31F842A5FFBDFFEAFF7D9B43FE19F75D" box="[189,411,520,546]" name="South Africa" pageId="3" pageNumber="4">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="8B2BCCD7FFBDFFEAFE6D9B43FDC9F75D" box="[429,587,520,546]" country="South Africa" name="Mpumalanga" pageId="3" pageNumber="4">Mpumalanga</collectingRegion>
|
||
, same data as
|
||
<typeStatus id="9654BC97FFBDFFEAFCCD9B43FCF6F75D" box="[781,884,520,546]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFC4A9B43FC5DF75D" box="[906,991,520,546]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFC2D9B4CFC7FF75D" box="[1005,1021,519,546]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="4" type="Herbarium">P</collectionCode>
|
||
0245/
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFB829B4CFBD6F75D" box="[1090,1108,519,546]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
|
||
4088, 5 dry specimens;
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFF7D9B60FEAEF73A" box="[189,300,555,581]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="4" type="Museum">NHMUK</collectionCode>
|
||
20160037, one dry specimen;
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFD449B60FD62F739" box="[644,736,555,582]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">RMNH</collectionCode>
|
||
.5004141, one dry specimen); Mariepskop Forest Reserve,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFF7D9B05FED3F717" box="[189,337,590,617]" degrees="24.56374" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-24.56374">
|
||
24.56374°
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFEFF9B05FED3F717" box="[319,337,590,616]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="3" pageNumber="4" type="Herbarium">S</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFE9F9B05FE74F716" box="[351,502,590,617]" degrees="30.86293" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="30.86293">30.86293° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBDFFEAFDC49B05FDE2F716" box="[516,608,590,617]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="3" pageNumber="4" unit="m" value="1640.0">1640 m</quantity>
|
||
, northern mist-belt forest, in leaf-litter, D. Herbert, L. Davis & M. Cole, stn 14-22,
|
||
<date id="3D5124F5FFBDFFEAFE429B3AFD90F7F4" box="[386,530,625,652]" pageId="3" pageNumber="4" value="2014-12-03">3 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFDE19B3AFDF4F7F4" box="[545,630,625,651]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
|
||
P0269/T4090, three dry specimens, one in ethanol); Mariepskop Forest Reserve,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFE429BDFFD9BF7D0" box="[386,537,660,687]" degrees="24.56692" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-24.56692">24.56692° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFDEA9BDFFD3BF7D1" box="[554,697,660,686]" degrees="30.86482" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="30.86482">30.86482°E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBDFFEAFD089BDFFCA5F7D0" box="[712,807,660,687]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="4" unit="m" value="1520.0">1520 m</quantity>
|
||
, indigenous Afromontane forest, in leaf-litter on forest floor, J.L. Horn,
|
||
<date id="3D5124F5FFBDFFEAFE129BF3FDE6F7AD" box="[466,612,695,722]" pageId="3" pageNumber="4" value="2005-03-01">1 Mar. 2005</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFDB59BF3FD48F7AD" box="[629,714,696,722]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
|
||
W3501/T4087, three dry specimens); Mariepskop Forest Reserve,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFEE99B90FE2EF78A" box="[297,428,731,758]" degrees="24.5679" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="-24.5679">24.5679° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFE789B90FDBFF78A" box="[440,573,731,758]" degrees="30.8599" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="30.8599">30.8599° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBDFFEAFD889B90FD20F78A" box="[584,674,731,758]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="4" unit="m" value="1550.0">1550 m</quantity>
|
||
, northern mist-belt forest, in leaf-litter, D. Herbert, L. Davis & M. Cole, stn 14-26a,
|
||
<date id="3D5124F5FFBDFFEAFE6E9BB5FDBBF667" box="[430,569,766,793]" pageId="3" pageNumber="4" value="2014-12-04">4 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBDFFEAFD869BB5FD19F667" box="[582,667,766,792]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
|
||
P0240/T4092, one dry specimen); Mariepskop Forest Reserve, Bushpig Trail,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFEB09A6AFD86F644" box="[368,516,801,828]" degrees="24.56795" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-24.56795">24.56795° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBDFFEAFDD39A6AFD28F643" box="[531,682,801,828]" degrees="30.86138" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="30.86138">30.86138° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBDFFEAFD789A6AFC97F643" box="[696,789,801,828]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="4" unit="m" value="1520.0">1520 m</quantity>
|
||
, northern mist-belt forest, in leaf-litter, D. Herbert, L. Davis & M. Cole, stn 14-22a,
|
||
<date id="3D5124F5FFBAFFEDFD8C9840FD5CF45A" box="[588,734,267,294]" pageId="4" pageNumber="5" value="2014-12-03">3 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBAFFEDFD2F9840FCC6F45A" box="[751,836,267,293]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="4" pageNumber="5" type="Museum">NMSA</collectionCode>
|
||
P0262/T4089, one dry specimen); Mariepskop Forest Reserve, Picnic Trail,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFDE39865FD38F436" box="[547,698,302,329]" degrees="24.56847" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="-24.56847">24.56847° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFD0B9865FCE7F436" box="[715,869,302,329]" degrees="30.85920" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="30.8592">30.85920° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBAFFEDFCB69865FC57F436" box="[886,981,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.545" pageId="4" pageNumber="5" unit="m" value="1545.0">1545 m</quantity>
|
||
, northern mist-belt forest, in leaflitter, D. Herbert, L. Davis & M. Cole, stn 14-27,
|
||
<date id="3D5124F5FFBAFFEDFCCB9819FC19F413" box="[779,923,338,364]" pageId="4" pageNumber="5" value="2014-12-04">4 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBAFFEDFC6A9819FC7DF413" box="[938,1023,338,364]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="4" pageNumber="5" type="Museum">NMSA</collectionCode>
|
||
P0235/T4091, one dry juvenile specimen); Mariepskop Forest Reserve, east facing slope,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFCB1983EFB87F4F0" box="[881,1029,373,400]" degrees="24.57108" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="-24.57108">24.57108° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFBD3983EFB28F4F0" box="[1043,1194,373,399]" degrees="30.86014" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="30.86014">30.86014° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBAFFEDFB77983EFA96F4F0" box="[1207,1300,373,400]" metricMagnitude="3" metricUnit="m" metricValue="1.519" pageId="4" pageNumber="5" unit="m" value="1519.0">1519 m</quantity>
|
||
, leg. M. Cole,
|
||
<date id="3D5124F5FFBAFFEDFEC798D3FE2EF4CD" box="[263,428,408,435]" pageId="4" pageNumber="5" value="2013-12-10">10 Dec. 2013</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBAFFEDFE7F98D3FE7DF4CD" box="[447,511,408,434]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="4" pageNumber="5">ELM</collectionCode>
|
||
D17729/T035, one dry specimen); Mariepskop Forest Reserve, in forest beside road at
|
||
<quantity id="8E17AFD0FFBAFFEDFEAB98F0FD93F4A9" box="[363,529,443,470]" metricMagnitude="3" metricUnit="m" metricValue="1.61" metricValueMax="1.7" metricValueMin="1.52" pageId="4" pageNumber="5" unit="m" value="1610.0" valueMax="1700.0" valueMin="1520.0">1520–1700 m</quantity>
|
||
, leg. M. Cole,
|
||
<date id="3D5124F5FFBAFFEDFD0498F0FCF1F4AA" box="[708,883,443,470]" pageId="4" pageNumber="5" value="2014-12-03" valueMax="2014-12-04" valueMin="2014-12-03">3–4 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFBAFFEDFC4498F0FC46F4AA" box="[900,964,443,469]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="4" pageNumber="5">ELM</collectionCode>
|
||
D18022/T036, three dry specimens;
|
||
<collectionCode id="2FFE9AF0FFBAFFEDFF7D9894FF79F486" box="[189,251,479,505]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="4" pageNumber="5">ELM</collectionCode>
|
||
W3882/T037, two specimens in ethanol).
|
||
</paragraph>
|
||
<caption id="1D9052BDFFBDFFEAFF7D9EC3FE99F296" ID-DOI="http://doi.org/10.5281/zenodo.3847844" ID-Zenodo-Dep="3847844" httpUri="https://zenodo.org/record/3847844/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1928,1954]" targetBox="[192,1396,881,1877]" targetPageId="3">
|
||
<paragraph id="49500235FFBDFFEAFF7D9EC3FE99F296" blockId="3.[189,1398,1928,2025]" pageId="3" pageNumber="4">
|
||
<emphasis id="7B9BDE27FFBDFFEAFF7D9EC3FE8DF2DD" bold="true" box="[189,271,1928,1954]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
||
<taxonomicName id="8EEF79B6FFBDFFEAFEDA9EC3FE5AF2DD" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[282,472,1928,1954]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFBDFFEAFEDA9EC3FE5AF2DD" box="[282,472,1928,1954]" italics="true" pageId="3" pageNumber="4">Gulella davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFBDFFEAFE239EC2FDC0F2DC" box="[483,578,1929,1955]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="7B9BDE27FFBDFFEAFD8D9EC3FD04F2DD" bold="true" box="[589,646,1928,1954]" pageId="3" pageNumber="4">A–D</emphasis>
|
||
. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086).
|
||
<emphasis id="7B9BDE27FFBDFFEAFEE39EE7FEB5F2B9" bold="true" box="[291,311,1964,1990]" pageId="3" pageNumber="4">E</emphasis>
|
||
. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="49500235FFBAFFEDFF7D9B6BFEDFF745" blockId="4.[189,349,544,570]" box="[189,349,544,570]" pageId="4" pageNumber="5">
|
||
<heading id="1218B559FFBAFFEDFF7D9B6BFEDFF745" bold="true" box="[189,349,544,570]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="7B9BDE27FFBAFFEDFF7D9B6BFEDFF745" bold="true" box="[189,349,544,570]" pageId="4" pageNumber="5">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9B04FABEF716" blockId="4.[189,1340,590,617]" box="[189,1340,590,617]" pageId="4" pageNumber="5">
|
||
<collectingCountry id="31F842A5FFBAFFEDFF7D9B04FE16F716" box="[189,404,591,617]" name="South Africa" pageId="4" pageNumber="5">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="8B2BCCD7FFBAFFEDFE609B05FDBCF717" box="[416,574,590,616]" country="South Africa" name="Mpumalanga" pageId="4" pageNumber="5">Mpumalanga</collectingRegion>
|
||
, Mariepskop Forest Reserve,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFC5A9B05FBAFF716" box="[922,1069,590,617]" degrees="24.56128" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="-24.56128">24.56128° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFBAFFEDFBFA9B05FB4DF716" box="[1082,1231,590,617]" degrees="30.86367" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="30.86367">30.86367° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFBAFFEDFB1C9B04FABAF716" box="[1244,1336,591,617]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="5" unit="m" value="1700.0">1700 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9BDBFEE7F7D5" blockId="4.[189,357,656,682]" box="[189,357,656,682]" pageId="4" pageNumber="5">
|
||
<heading id="1218B559FFBAFFEDFF7D9BDBFEE7F7D5" bold="true" box="[189,357,656,682]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="7B9BDE27FFBAFFEDFF7D9BDBFEE7F7D5" bold="true" box="[189,357,656,682]" pageId="4" pageNumber="5">Identification</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9BF4FB51F660" blockId="4.[189,1398,702,799]" pageId="4" pageNumber="5">
|
||
A moderately sized species of
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFDEC9BF5FD06F7A7" authorityName="Pfeiffer" authorityYear="1856" box="[556,644,702,728]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFBAFFEDFDEC9BF5FD06F7A7" box="[556,644,702,728]" italics="true" pageId="4" pageNumber="5">Gulella</emphasis>
|
||
</taxonomicName>
|
||
with a strongly ribbed, cylindrical shell and five-fold apertural dentition including,
|
||
<emphasis id="7B9BDE27FFBAFFEDFE6F9BA8FD9CF783" box="[431,542,738,764]" italics="true" pageId="4" pageNumber="5">inter alia</emphasis>
|
||
, a tricuspid mid-labral complex and a tricuspid
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFBA29BA9FB54F783" box="[1122,1238,738,764]" pageId="4" pageNumber="5" rank="species" species="columella">columella</taxonomicName>
|
||
lamella; also distinctive is the almost complete peristome, interrupted only behind the parietal lamella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01F551BEFFBAFFECFF7D9A0DFC1FF763" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="49500235FFBAFFEDFF7D9A0DFECCF61F" blockId="4.[189,334,838,864]" box="[189,334,838,864]" pageId="4" pageNumber="5">
|
||
<heading id="1218B559FFBAFFEDFF7D9A0DFECCF61F" bold="true" box="[189,334,838,864]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="7B9BDE27FFBAFFEDFF7D9A0DFECCF61F" bold="true" box="[189,334,838,864]" pageId="4" pageNumber="5">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9A3EFDCFF353" blockId="4.[189,1399,885,1580]" pageId="4" pageNumber="5">
|
||
SHELL (
|
||
<figureCitation id="D1D41EB0FFBAFFEDFED49A3EFEDBF6F0" box="[276,345,885,911]" captionStart="Fig" captionStartId="3.[189,232,1928,1954]" captionTargetBox="[192,1396,881,1877]" captionTargetId="figure@3.[189,1399,877,1878]" captionTargetPageId="3" captionText="Fig. 1. Gulella davisae sp. nov. A–D. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086). E. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091)." figureDoi="http://doi.org/10.5281/zenodo.3847844" httpUri="https://zenodo.org/record/3847844/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
). Medium-sized, cylindrical; adult height
|
||
<quantity id="8E17AFD0FFBAFFEDFC8E9A3EFC62F6F0" box="[846,992,885,912]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="4" pageNumber="5" unit="mm" value="5.949999999999999" valueMax="6.3" valueMin="5.6">5.6–6.3 mm</quantity>
|
||
, diameter 2.8–3.0 mm; H:D 1.95– 2.21, with up to 8.0 whorls. Protoconch diameter
|
||
<quantity id="8E17AFD0FFBAFFEDFD2C9AD3FCFBF6CC" box="[748,889,920,947]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="4" pageNumber="5" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
|
||
, comprising 1.5–2.0 whorls beyond nucleus, sculptured with fine, close-set axial wrinkles, with an indistinct spiral element appearing in the last halfwhorl; junction with teleoconch distinct. Teleoconch comprising 5.5–6.0 whorls; whorls weakly convex, but usually with a distinct, albeit narrow, shoulder such that suture is indented; sculpture of closeset, prosocline, axial ribs. Aperture roundly quadrate, peristome flaring, almost complete, interrupted only behind parietal lamella; apertural dentition 5-fold (
|
||
<figureCitation id="D1D41EB0FFBAFFEDFC939D03FC2DF11D" box="[851,943,1096,1122]" captionStart="Fig" captionStartId="3.[189,232,1928,1954]" captionTargetBox="[192,1396,881,1877]" captionTargetId="figure@3.[189,1399,877,1878]" captionTargetPageId="3" captionText="Fig. 1. Gulella davisae sp. nov. A–D. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086). E. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091)." figureDoi="http://doi.org/10.5281/zenodo.3847844" httpUri="https://zenodo.org/record/3847844/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
|
||
): 1) a strong, vertical, slightly oblique parietal lamella; 2) a large mid-labral complex, itself with three smaller teeth, a simple peg-like upper one and below this a larger, in-running ridge with a large, roundly trigonal tooth at its outer end and a smaller one at its inner end, large outer tooth sometimes with an additional weak cusp on its lower margin; 3) a roundly trigonal basal tooth to left of centre; 4) a more deep-set, transverse, ridge-like, tooth (sometimes with two cusps) between basal tooth and mid-labral complex; 5) a large quadrate
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFAC39DB3FAF5F06D" box="[1283,1399,1272,1298]" pageId="4" pageNumber="5" rank="species" species="columella">columella</taxonomicName>
|
||
lamella with three denticles, the upper one usually horizontal and somewhat ridge-like, the middle one protruding furthest into aperture. In addition to the above, there may be a small sinular denticle delimiting a shallow notch behind parietal lamella. Mid-labral complex marked externally by a deep pit behind flared outer lip; a smaller pit underlies basal tooth;
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFCD69CCEFC08F0E0" box="[790,906,1413,1439]" pageId="4" pageNumber="5" rank="species" species="columella">columella</taxonomicName>
|
||
lip smooth. A juvenile of 2.25 teleoconch whorls exhibited a very small tooth on the upper
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFCC39CE3FCF5F0BD" box="[771,887,1448,1474]" pageId="4" pageNumber="5" rank="species" species="columella">columella</taxonomicName>
|
||
lip and a larger, transverse mid-basal tooth (
|
||
<figureCitation id="D1D41EB0FFBAFFEDFF069C80FEA3F09A" box="[198,289,1483,1509]" captionStart="Fig" captionStartId="3.[189,232,1928,1954]" captionTargetBox="[192,1396,881,1877]" captionTargetId="figure@3.[189,1399,877,1878]" captionTargetPageId="3" captionText="Fig. 1. Gulella davisae sp. nov. A–D. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086). E. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091)." figureDoi="http://doi.org/10.5281/zenodo.3847844" httpUri="https://zenodo.org/record/3847844/files/figure.png" pageId="4" pageNumber="5">Fig. 1E</figureCitation>
|
||
). Umbilicus open, drop-shaped and of moderate size, opening laterally some distance behind flared
|
||
<taxonomicName id="8EEF79B6FFBAFFEDFEC89CA5FEFEF377" box="[264,380,1518,1544]" pageId="4" pageNumber="5" rank="species" species="columella">columella</taxonomicName>
|
||
lip (
|
||
<figureCitation id="D1D41EB0FFBAFFEDFE739CA5FD93F376" box="[435,529,1518,1545]" captionStart="Fig" captionStartId="3.[189,232,1928,1954]" captionTargetBox="[192,1396,881,1877]" captionTargetId="figure@3.[189,1399,877,1878]" captionTargetPageId="3" captionText="Fig. 1. Gulella davisae sp. nov. A–D. Holotype, height 6.1 mm, diameter 2.9 mm (NMSA P0418/ T4086). E. Juvenile specimen showing apertural dentition, diameter 2.7 mm (paratype, NMSA P0235/ T4091)." figureDoi="http://doi.org/10.5281/zenodo.3847844" httpUri="https://zenodo.org/record/3847844/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
|
||
). Shell milky-white when fresh, usually with some superficial soil debris, particularly in sutural indentation.
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9F1FFB86F311" blockId="4.[189,1028,1619,1646]" box="[189,1028,1619,1646]" pageId="4" pageNumber="5">LIVING ANIMAL. Head-foot yellow, optic retractor muscles bright orange.</paragraph>
|
||
<paragraph id="49500235FFBAFFEDFF7D9FDEFAB5F267" blockId="4.[189,1399,1684,1817]" pageId="4" pageNumber="5">
|
||
RADULA. A single radula was available; corresponds to ‘Group A’ of
|
||
<bibRefCitation id="2D7E7FC4FFBAFFEDFC349FDFFB15F3D0" author="Aiken D. W." box="[1012,1175,1684,1711]" pageId="4" pageNumber="5" pagination="317 - 323" refId="ref13448" refString="Aiken D. W. 1981. Differentiation of the radula of South African species of the genus Gulella into three types (Gastropoda Pulmonata: Streptaxidae). Journal of Conchology 30: 317 - 323." type="journal article" year="1981">Aiken (1981)</bibRefCitation>
|
||
; formula 12 + 1 + 12; length ~
|
||
<quantity id="8E17AFD0FFBAFFEDFE8A9FF3FE2AF3AD" box="[330,424,1720,1746]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="5" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, with 87 V-shaped rows of teeth; total number of teeth ~2175; rachidian extremely small; inner eight laterals of more or less equal size (3–7 slightly larger) with stout, blade-like cusp; base-plate with an oblique anchoring peg; outer laterals progressively smaller, the twelfth minute.
|
||
</paragraph>
|
||
<paragraph id="49500235FFBAFFECFF7D9E0BFC1FF763" blockId="4.[189,1399,1855,2023]" lastBlockId="5.[189,1399,267,540]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
DISTAL GENITALIA (
|
||
<figureCitation id="D1D41EB0FFBAFFEDFE509E74FE55F225" box="[400,471,1855,1882]" captionStart="Fig" captionStartId="5.[189,232,1964,1990]" captionTargetBox="[196,1385,1293,1916]" captionTargetId="figure@5.[189,1399,1285,1923]" captionTargetPageId="5" captionText="Fig. 2. Gulella davisae sp. nov., genitalia, paratype (NMSA P0269/T4090). A. Distal portion of genital system. B. Internal structure of penis." figureDoi="http://doi.org/10.5281/zenodo.3847848" httpUri="https://zenodo.org/record/3847848/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
). Penis of moderate length (~
|
||
<quantity id="8E17AFD0FFBAFFEDFCF19E74FC13F225" box="[817,913,1855,1882]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="5" unit="mm" value="3.5">3.5 mm</quantity>
|
||
), subcylindrical, slightly broader distally, but somewhat constricted at junction with genital atrium; penial retractor undivided and attached to small penial caecum at penis apex. Lumen of penis with a single longitudinal pilaster running from area adjacent to penial caecum to base of penis; also with a raised subcircular pilaster near penis base; remainder of lumen with some weaker longitudinal folds and faint superficial striae; penial armature restricted to a small number of minute, trigonal hooks on apical portion of longitudinal pilaster. Vas deferens inserts laterally about halfway along penis, running toward penis base, then recurving along vagina and fusing with base of spermoviduct; vas deferens loosely adherent to penis base, but not fused to it; no evidence of a penis sheath or epiphallus. Atrium simple; vagina short and thin-walled; duct of gametolytic sac (bursa copulatrix) long, following course of spermoviduct to region of albumen gland; gametolytic sac itself narrow and elongate, scarcely broader than its duct; prostatic and oviductal portions of spermoviduct distinct; proximal portion of reproductive tract damaged and largely missing, but large fertilisation pouch–spermathecal complex remaining.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01F551BEFFBBFFECFF7D9B08FC5CF660" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="49500235FFBBFFECFF7D9B08FED4F722" blockId="5.[189,342,579,605]" box="[189,342,579,605]" pageId="5" pageNumber="6">
|
||
<heading id="1218B559FFBBFFECFF7D9B08FED4F722" bold="true" box="[189,342,579,605]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="7B9BDE27FFBBFFECFF7D9B08FED4F722" bold="true" box="[189,342,579,605]" pageId="5" pageNumber="6">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBBFFECFF7D9B39FAC8F7D0" blockId="5.[189,1399,626,688]" pageId="5" pageNumber="6">
|
||
A narrow-range endemic, currently recorded only from Mariepskop Forest Reserve on the edge of the Drakensberg Escarpment in northern
|
||
<collectingRegion id="8B2BCCD7FFBBFFECFDB49BDEFC90F7D0" box="[628,786,661,687]" country="South Africa" name="Mpumalanga" pageId="5" pageNumber="6">Mpumalanga</collectingRegion>
|
||
,
|
||
<collectingCountry id="31F842A5FFBBFFECFCDE9BDEFC35F7D0" box="[798,951,661,687]" name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
|
||
; at altitudes from
|
||
<quantity id="8E17AFD0FFBBFFECFB4A9BDEFAC4F7D0" box="[1162,1350,661,688]" metricMagnitude="3" metricUnit="m" metricValue="1.61" metricValueMax="1.7" metricValueMin="1.52" pageId="5" pageNumber="6" unit="m" value="1610.0" valueMax="1700.0" valueMin="1520.0">
|
||
<date id="3D5124F5FFBBFFECFB4A9BDEFAA7F7D0" box="[1162,1317,661,688]" pageId="5" pageNumber="6" value="1520" valueMax="1700">1520 to 1700</date>
|
||
m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="49500235FFBBFFECFF7D9B9DFE9FF78F" blockId="5.[189,285,726,752]" box="[189,285,726,752]" pageId="5" pageNumber="6">
|
||
<heading id="1218B559FFBBFFECFF7D9B9DFE9FF78F" bold="true" box="[189,285,726,752]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="7B9BDE27FFBBFFECFF7D9B9DFE9FF78F" bold="true" box="[189,285,726,752]" pageId="5" pageNumber="6">Habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBBFFECFF7D9A4EFC5CF660" blockId="5.[189,990,773,800]" box="[189,990,773,800]" pageId="5" pageNumber="6">
|
||
Northern mist-belt forest (
|
||
<bibRefCitation id="2D7E7FC4FFBBFFECFE309A4EFCBAF65F" author="Mucina L. & Rutherford M. C." box="[496,824,773,800]" pageId="5" pageNumber="6" refId="ref14287" refString="Mucina L. & Rutherford M. C. (eds) 2006. The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19, South African National Biodiversity Institute, Pretoria." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
), in leaf-litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01F551BEFFBBFFEFFF7D9A0CFE3AF17B" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="49500235FFBBFFECFF7D9A0CFEADF61E" blockId="5.[189,303,839,865]" box="[189,303,839,865]" pageId="5" pageNumber="6">
|
||
<heading id="1218B559FFBBFFECFF7D9A0CFEADF61E" bold="true" box="[189,303,839,865]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="7B9BDE27FFBBFFECFF7D9A0CFEADF61E" bold="true" box="[189,303,839,865]" pageId="5" pageNumber="6">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFBBFFEFFF7D9A3EFB58F437" blockId="5.[189,1399,885,1193]" lastBlockId="6.[189,1398,267,329]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
The size and overall facies of the shell is similar to that of
|
||
<taxonomicName id="8EEF79B6FFBBFFECFC6C9A3EFA83F6F0" authority="Bruggen, 2006" authorityName="Bruggen" authorityYear="2006" box="[940,1281,885,911]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="johannae">
|
||
<emphasis id="7B9BDE27FFBBFFECFC6C9A3EFBC3F6F0" box="[940,1089,885,911]" italics="true" pageId="5" pageNumber="6">G. johannae</emphasis>
|
||
Bruggen, 2006
|
||
</taxonomicName>
|
||
from the neighbouring Wolkberg massif. That species, like
|
||
<taxonomicName id="8EEF79B6FFBBFFECFCC09AD2FCFEF6CD" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[768,892,920,947]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFBBFFECFCC09AD2FCFEF6CD" box="[768,892,920,947]" italics="true" pageId="5" pageNumber="6">G. davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFBBFFECFC429AD2FC5FF6CC" box="[898,989,921,947]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, has an almost complete peristome, interrupted only behind the parietal lamella and similar apertural dentition in the juvenile. However, the adult apertural dentition of the present species differs considerably from that of
|
||
<taxonomicName id="8EEF79B6FFBBFFECFB959A94FB66F687" authorityName="Bruggen" authorityYear="2006" box="[1109,1252,990,1017]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="johannae">
|
||
<emphasis id="7B9BDE27FFBBFFECFB959A94FB66F687" box="[1109,1252,990,1017]" italics="true" pageId="5" pageNumber="6">G. johannae</emphasis>
|
||
</taxonomicName>
|
||
. In the latter this dentition is less complex, the mid-labral tooth is essentially a single entity rather than a complex of three denticles, there is no deep-set, ridge-like tooth to the right of the basal tooth and the columella lamella is a simple horizontal tooth. In addition, the shell of
|
||
<taxonomicName id="8EEF79B6FFBBFFECFC549D02FBA4F11D" authorityName="Bruggen" authorityYear="2006" box="[916,1062,1096,1123]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="johannae">
|
||
<emphasis id="7B9BDE27FFBBFFECFC549D02FBA4F11D" box="[916,1062,1096,1123]" italics="true" pageId="5" pageNumber="6">G. johannae</emphasis>
|
||
</taxonomicName>
|
||
attains a slightly larger size (height up to
|
||
<quantity id="8E17AFD0FFBBFFECFEA59D27FE44F1F9" box="[357,454,1132,1158]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="5" pageNumber="6" unit="mm" value="7.7">7.7 mm</quantity>
|
||
), is somewhat more slender (H:D 2.16–2.88), and the umbilicus is wider and basally orientated.
|
||
<taxonomicName id="8EEF79B6FFBBFFECFE5F9DC4FD7FF1D6" authority="Bruggen, 1980" authorityName="Bruggen" authorityYear="1980" box="[415,765,1166,1193]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="subSpecies" species="vicina" subSpecies="luci">
|
||
<emphasis id="7B9BDE27FFBBFFECFE5F9DC4FDC3F1D7" box="[415,577,1166,1193]" italics="true" pageId="5" pageNumber="6">G. vicina luci</emphasis>
|
||
Bruggen, 1980
|
||
</taxonomicName>
|
||
, from the eastern highlands of
|
||
<collectingCountry id="31F842A5FFBBFFECFBB39DC5FB76F1D7" box="[1139,1268,1166,1192]" name="Zimbabwe" pageId="5" pageNumber="6">Zimbabwe</collectingCountry>
|
||
, also has a ribbed shell and tricuspid labral and columella teeth, but it is larger (height
|
||
<quantity id="8E17AFD0FFB8FFEFFBF09840FB17F459" box="[1072,1173,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="7" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
|
||
), has distinct spiral sculpture on the protoconch and its peristome is broadly interrupted in the parietal region.
|
||
</paragraph>
|
||
<caption id="1D9052BDFFBBFFECFF7D9EE7FDF7F296" ID-DOI="http://doi.org/10.5281/zenodo.3847848" ID-Zenodo-Dep="3847848" httpUri="https://zenodo.org/record/3847848/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1964,1990]" targetBox="[196,1385,1293,1916]" targetPageId="5">
|
||
<paragraph id="49500235FFBBFFECFF7D9EE7FDF7F296" blockId="5.[189,1398,1964,2026]" pageId="5" pageNumber="6">
|
||
<emphasis id="7B9BDE27FFBBFFECFF7D9EE7FE88F2B9" bold="true" box="[189,266,1964,1990]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
|
||
<taxonomicName id="8EEF79B6FFBBFFECFED29EE7FE49F2B9" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[274,459,1964,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFBBFFECFED29EE7FE49F2B9" box="[274,459,1964,1990]" italics="true" pageId="5" pageNumber="6">Gulella davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFBBFFECFE129EE6FDB2F2B8" box="[466,560,1965,1991]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, genitalia, paratype (NMSA P0269/T4090).
|
||
<emphasis id="7B9BDE27FFBBFFECFBF99EE7FBCCF2B9" bold="true" box="[1081,1102,1964,1990]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Distal portion of genital system.
|
||
<emphasis id="7B9BDE27FFBBFFECFEDD9E9BFEB3F295" bold="true" box="[285,305,2000,2026]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Internal structure of penis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="49500235FFB8FFEFFF7D983BFE3AF17B" blockId="6.[189,1399,368,1028]" pageId="6" pageNumber="7">
|
||
The almost complete peristome shared by
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFD7C983BFCBEF4F5" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[700,828,368,394]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFB8FFEFFD7C983BFCBEF4F5" box="[700,828,368,394]" italics="true" pageId="6" pageNumber="7">G. davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFC86983BFC21F4F5" box="[838,931,368,394]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFC20983BFBF1F4F5" authorityName="Bruggen" authorityYear="2006" box="[992,1139,368,394]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="johannae">
|
||
<emphasis id="7B9BDE27FFB8FFEFFC20983BFBF1F4F5" box="[992,1139,368,394]" italics="true" pageId="6" pageNumber="7">G. johannae</emphasis>
|
||
</taxonomicName>
|
||
is an unusual feature suggesting that the two are closely related and perhaps not members of
|
||
<emphasis id="7B9BDE27FFB8FFEFFC3498D8FBFAF4D2" box="[1012,1144,403,429]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFC3498D8FBCEF4D2" authorityName="Pfeiffer" authorityYear="1856" box="[1012,1100,403,429]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus" sensu="stricto">Gulella</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFB9298DFFBFAF4D2" box="[1106,1144,404,429]" pageId="6" pageNumber="7" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Given that they occur in similar habitats on neighbouring mountain blocks, it seems likely that they are sister taxa derived from an ancestral stock that occurred on both blocks when forest cover was more extensive.
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFB349891FEECF768" authority="Bruggen, 2004" authorityName="Bruggen" authorityYear="2004" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="herberti">
|
||
<emphasis id="7B9BDE27FFB8FFEFFB349891FAF5F48B" box="[1268,1399,474,500]" italics="true" pageId="6" pageNumber="7">G. herberti</emphasis>
|
||
Bruggen, 2004
|
||
</taxonomicName>
|
||
, from southern Mpumalanga and
|
||
<collectingCountry id="31F842A5FFB8FFEFFD3798B6FCF6F768" box="[759,884,509,535]" name="Swaziland" pageId="6" pageNumber="7">Swaziland</collectingCountry>
|
||
, is another species with a similar peristome, but its shell is smaller (height ca 4.0 mm) and more globose, has a closed umbilicus and less complex apertural dentition. In
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFE7E9B0FFDBDF722" authorityName="Bruggen" authorityYear="2004" box="[446,575,579,606]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="herberti">
|
||
<emphasis id="7B9BDE27FFB8FFEFFE7E9B0FFDBDF722" box="[446,575,579,606]" italics="true" pageId="6" pageNumber="7">G. herberti</emphasis>
|
||
</taxonomicName>
|
||
the peristome is almost completely detached from the preceding whorl, a condition taken to the extreme in
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFDAD9B2CFC1AF7FF" authority="Herbert, 2002" authorityName="Herbert" authorityYear="2002" box="[621,920,614,641]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="salpinx">
|
||
<emphasis id="7B9BDE27FFB8FFEFFDAD9B2CFD6BF7FF" box="[621,745,614,641]" italics="true" pageId="6" pageNumber="7">G. salpinx</emphasis>
|
||
<bibRefCitation id="2D7E7FC4FFB8FFEFFD329B2DFC1AF7FF" author="Herbert D. G." box="[754,920,614,640]" pageId="6" pageNumber="7" pagination="125 - 138" refId="ref14121" refString="Herbert D. G. 2002. Gulella salpinx sp. n. a new, critically endangered, holoendemic species from the limestone deposits of the Marble Delta, KwaZulu-Natal, South Africa (Mollusca: Gastropoda: Streptaxidae). African Invertebrates 43: 125 - 138." type="journal article" year="2002">Herbert, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
from southern KwaZulu-Natal.
|
||
<bibRefCitation id="2D7E7FC4FFB8FFEFFADD9B2DFE89F7DB" author="Herbert D. G." pageId="6" pageNumber="7" pagination="125 - 138" refId="ref14121" refString="Herbert D. G. 2002. Gulella salpinx sp. n. a new, critically endangered, holoendemic species from the limestone deposits of the Marble Delta, KwaZulu-Natal, South Africa (Mollusca: Gastropoda: Streptaxidae). African Invertebrates 43: 125 - 138." type="journal article" year="2002">Herbert (2002)</bibRefCitation>
|
||
noted the similarities between
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFDBD9BC1FD75F7DB" authorityName="Herbert" authorityYear="2002" box="[637,759,650,676]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="salpinx">
|
||
<emphasis id="7B9BDE27FFB8FFEFFDBD9BC1FD75F7DB" box="[637,759,650,676]" italics="true" pageId="6" pageNumber="7">G. salpinx</emphasis>
|
||
</taxonomicName>
|
||
and the Mascarene genus
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFBF19BC1FF78F7B8" authority="Mollendorff, 1887" authorityName="Mollendorff" authorityYear="1887" class="Gastropoda" family="Streptaxidae" genus="Microstrophia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB8FFEFFBF19BC1FB5AF7DB" box="[1073,1240,650,676]" italics="true" pageId="6" pageNumber="7">Microstrophia</emphasis>
|
||
Möllendorff, 1887
|
||
</taxonomicName>
|
||
, and
|
||
<bibRefCitation id="2D7E7FC4FFB8FFEFFEF99BE6FE72F7B8" author="Rowson B." box="[313,496,685,711]" pageId="6" pageNumber="7" refId="ref14405" refString="Rowson B. 2010. Systematics and Diversity of the Streptaxidae (Gastropoda: Stylommatophora). PhD Thesis, University of Wales, Cardiff." type="book" year="2010">Rowson (2010)</bibRefCitation>
|
||
observed that both possess needle-like spines in the apical part of the penis. Subsequent molecular work (
|
||
<bibRefCitation id="2D7E7FC4FFB8FFEFFDD19B9BFCB1F794" author="Rowson B. & Herbert D. G." box="[529,819,720,747]" pageId="6" pageNumber="7" pagination="69 - 84" refId="ref14431" refString="Rowson B. & Herbert D. G. 2016. The type species and circumscription of the species-rich Afrotropical snail genus Gulella L. Pfeiffer, 1856, based on anatomical and mtDNA data (Mollusca: Eupulmonata: Streptaxidae). Archiv fur Molluskenkunde 145 (1): 69 - 84. http: // dx. doi. org / 10.1127 / arch. moll / 1869 - 0963 / 145 / 069 - 084" type="journal article" year="2016">Rowson & Herbert 2016</bibRefCitation>
|
||
) has shown that
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFC369B9BFBECF795" authorityName="Herbert" authorityYear="2002" box="[1014,1134,720,746]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="salpinx">
|
||
<emphasis id="7B9BDE27FFB8FFEFFC369B9BFBECF795" box="[1014,1134,720,746]" italics="true" pageId="6" pageNumber="7">G. salpinx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFB659B9BFACEF795" authorityName="Mollendorff" authorityYear="1887" box="[1189,1356,720,746]" class="Gastropoda" family="Streptaxidae" genus="Microstrophia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB8FFEFFB659B9BFACEF795" box="[1189,1356,720,746]" italics="true" pageId="6" pageNumber="7">Microstrophia</emphasis>
|
||
</taxonomicName>
|
||
are indeed related and that they cluster together with species of
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFCB69BB8FB88F672" authorityName="Pilsbry" authorityYear="1919" box="[886,1034,755,781]" class="Gastropoda" family="Streptaxidae" genus="Primigulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB8FFEFFCB69BB8FB88F672" box="[886,1034,755,781]" italics="true" pageId="6" pageNumber="7">Primigulella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFB829BB8FB57F672" authorityName="Rowson & Tatttersfield" authorityYear="2013" box="[1090,1237,755,781]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB8FFEFFB829BB8FB57F672" box="[1090,1237,755,781]" italics="true" pageId="6" pageNumber="7">Dadagulella</emphasis>
|
||
</taxonomicName>
|
||
rather than in
|
||
<emphasis id="7B9BDE27FFB8FFEFFF7D9A5DFEC2F64F" box="[189,320,790,816]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFF7D9A5DFE97F64F" authorityName="Pfeiffer" authorityYear="1856" box="[189,277,790,816]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus" sensu="lato">Gulella</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFEDE9A5DFEC2F64F" box="[286,320,790,816]" pageId="6" pageNumber="7" sensu="lato">s.l.</taxonomicNameLabel>
|
||
</emphasis>
|
||
However, such needle-like penial spines are not evident in
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFBC69A5CFB04F64F" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[1030,1158,790,817]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFB8FFEFFBC69A5CFB04F64F" box="[1030,1158,790,817]" italics="true" pageId="6" pageNumber="7">G. davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFB4E9A5CFB6EF64E" box="[1166,1260,791,817]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, suggesting that the apertural similarities might be superficial or convergent. The structure of the distal genitalia of
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFF7D9A16FEBEF608" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[189,316,861,887]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="davisae" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFB8FFEFFF7D9A16FEBEF608" box="[189,316,861,887]" italics="true" pageId="6" pageNumber="7">G. davisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFE859A16FE23F608" box="[325,417,861,887]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is unremarkable and is broadly comparable to that of many species of
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFB339A16FAC9F608" authorityName="Pfeiffer" authorityYear="1856" box="[1267,1355,861,887]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFB339A16FAC9F608" box="[1267,1355,861,887]" italics="true" pageId="6" pageNumber="7">Gulella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFA949A16FAF4F608" box="[1364,1398,861,887]" pageId="6" pageNumber="7" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFA949A16FAEDF608" box="[1364,1391,861,887]" italics="true" pageId="6" pageNumber="7">s.l</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Similarly, its ‘Group A’ radula is the most common
|
||
<typeStatus id="9654BC97FFB8FFEFFCEC9ACBFCDDF6E5" box="[812,863,896,922]" pageId="6" pageNumber="7">type</typeStatus>
|
||
of radula in
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFC3D9ACBFBD7F6E5" authorityName="Pfeiffer" authorityYear="1856" box="[1021,1109,896,922]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFC3D9ACBFBD7F6E5" box="[1021,1109,896,922]" italics="true" pageId="6" pageNumber="7">Gulella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFB9E9ACBFB02F6E5" box="[1118,1152,896,922]" pageId="6" pageNumber="7" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFB9E9ACBFBFBF6E5" box="[1118,1145,896,922]" italics="true" pageId="6" pageNumber="7">s.l</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and thus neither the radula nor the distal genitalia provide significant data regarding the relationships of the species. Further clarity in this regard must await analysis of molecular data and for the meantime I simply refer the species to
|
||
<taxonomicName id="8EEF79B6FFB8FFEFFEF79AA1FE0DF17B" authorityName="Pfeiffer" authorityYear="1856" box="[311,399,1002,1028]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFEF79AA1FE0DF17B" box="[311,399,1002,1028]" italics="true" pageId="6" pageNumber="7">Gulella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB8FFEFFE579AA1FE3AF17B" box="[407,440,1002,1028]" pageId="6" pageNumber="7" sensu="lato">
|
||
<emphasis id="7B9BDE27FFB8FFEFFE579AA1FE33F17B" box="[407,433,1002,1028]" italics="true" pageId="6" pageNumber="7">s.l</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |