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<mods:title id="A11A9F5A447235958CA40425F0F04E38">A new species of Neocompsa (Neoibidionini), and a new species and new combinations in Tillomorphini (Cerambycidae, Cerambycinae)</mods:title>
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<emphasis id="B92ED975D34AFFC16BDFED4AFE03D892" bold="true" box="[151,387,364,391]" italics="true" pageId="5" pageNumber="339">Tetranodus reticeps</emphasis>
(
<bibRefCitation id="EFCB7896D34AFFC16ADBED4AFDA3D892" author="Bates, H. W." box="[403,547,364,390]" pageId="5" pageNumber="339" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates, 1880</bibRefCitation>
)
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(
<figureCitation id="136119E2D34AFFC16BD7EDB6FEA7D8BE" box="[159,295,399,426]" captionStart="FIGURES 15–21. 15–19" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[178,1413,277,1887]" captionTargetId="figure@7.[169,1418,273,1895]" captionTargetPageId="7" captionText="FIGURES 15–21. 15–19) Tetranodus reticeps, female 1: 15) Dorsal habitus; 16) Ventral habitus; 17) Lateral habitus; 18) Head, frontal view; 19) Pronotal sculpturing. 20) Tetranodus reticeps, female 2, dorsal habitus. 21) Tetranodus mexicanus, holotype female, pronotal sculpturing (currently, junior synonym of Tetranodus niveicollis Linell, 1897)." figureDoi="http://doi.org/10.5281/zenodo.3698869" httpUri="https://zenodo.org/record/3698869/files/figure.png" pageId="5" pageNumber="339">Figs. 15–20</figureCitation>
,
<figureCitation id="136119E2D34AFFC16A7BEDB6FEFAD8BE" box="[307,378,400,426]" captionStart="FIGURES 22–30. 22–26" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[191,1396,185,1919]" captionTargetId="figure@10.[185,1402,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–30. 22–26) Pentanodes nicaraguensis, holotype female: 22) Dorsal habitus; 23) Ventral habitus; 24) Lateral habitus; 25) Head, frontal view; 26) Labial palpomere III. 27–30) Euderces reticeps, holotype: 27) Dorsal habitus; 28) Lateral habitus; 29) Basal area of the elytra, lateral view; 30) Labels." figureDoi="http://doi.org/10.5281/zenodo.3698871" httpUri="https://zenodo.org/record/3698871/files/figure.png" pageId="5" pageNumber="339">27–30</figureCitation>
)
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<treatmentCitationGroup id="AB4A2249D34AFFC16BDFEDE9FA15D8F3" box="[151,1429,462,487]" pageId="5" pageNumber="339">
<treatmentCitation id="0AFB2376D34AFFC16BDFEDE9FE71D8F3" author="Bates, H. W." box="[151,497,462,487]" page="59" pageId="5" pageNumber="339" year="1880">
<taxonomicName id="4C5A7EE4D34AFFC16BDFEDE9FE71D8F3" ID-CoL="6GV2W" authority="Bates, 1880: 59" authorityName="Bates" authorityPageNumber="59" authorityYear="1880" box="[151,497,462,487]" class="Insecta" family="Cerambycidae" genus="Euderces" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="339" phylum="Arthropoda" rank="species" species="reticeps">
<emphasis id="B92ED975D34AFFC16BDFEDE9FECAD8F3" box="[151,330,463,487]" italics="true" pageId="5" pageNumber="339">Euderces reticeps</emphasis>
<bibRefCitation id="EFCB7896D34AFFC16A18EDE9FE71D8F3" author="Bates, H. W." box="[336,497,462,487]" pageId="5" pageNumber="339" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates, 1880: 59</bibRefCitation>
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;
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<bibRefCitation id="EFCB7896D34AFFC16AB4EDE9FD3AD8F3" author="Lameere, A. A." box="[508,698,463,487]" pageId="5" pageNumber="339" pagination="1 - 78" refId="ref10727" refString="Lameere, A. A. (1883) Liste des cerambycides, decrits posterieurement au catalogue de Munich. Annales de la Societe Entomologique de Belgique, 26, 1 - 78." type="journal article" year="1883">Lameere, 1883: 41</bibRefCitation>
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(cat.);
<treatmentCitation id="0AFB2376D34AFFC1684AEDE8FC62D8F3" author="Aurivillius, C." box="[770,994,462,487]" page="419" pageId="5" pageNumber="339" year="1912">
<bibRefCitation id="EFCB7896D34AFFC1684AEDE8FC62D8F3" author="Aurivillius, C." box="[770,994,462,487]" pageId="5" pageNumber="339" refId="ref10200" refString="Aurivillius, C. (1912) Coleopterorum Catalogus. Pars 39. Cerambycidae: Cerambycinae. W. Junk, Berlin, 574 pp." type="book" year="1912">Aurivillius, 1912: 419</bibRefCitation>
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(cat.);
<treatmentCitation id="0AFB2376D34AFFC16F63EDE8FA9CD8F3" author="Blackwelder, R. E." box="[1067,1308,462,487]" page="582" pageId="5" pageNumber="339" year="1946">
<bibRefCitation id="EFCB7896D34AFFC16F63EDE8FA9CD8F3" author="Blackwelder, R. E." box="[1067,1308,462,487]" pageId="5" pageNumber="339" pagination="551 - 763" refId="ref10377" refString="Blackwelder, R. E. (1946) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 4. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 582</bibRefCitation>
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(checklist).
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<paragraph id="8BE50567D34AFFC16BDFEDC8FEE6DB76" blockId="5.[151,1437,462,610]" pageId="5" pageNumber="339">
<treatmentCitationGroup id="AB4A2249D34AFFC16BDFEDC8FEE6DB76" pageId="5" pageNumber="339">
<taxonomicName id="4C5A7EE4D34AFFC16BDFEDC8FEE0DB11" ID-CoL="55Q7T" baseAuthorityName="Bates" baseAuthorityYear="1880" box="[151,352,494,518]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="339" phylum="Arthropoda" rank="species" species="reticeps">
<emphasis id="B92ED975D34AFFC16BDFEDC8FEE0DB11" box="[151,352,494,518]" italics="true" pageId="5" pageNumber="339">Tetranodus reticeps</emphasis>
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;
<treatmentCitation id="0AFB2376D34AFFC16A38EDCBFDC8DB12" author="Chemsak, J. A." box="[368,584,493,518]" page="306" pageId="5" pageNumber="339" year="1969">
<bibRefCitation id="EFCB7896D34AFFC16A38EDCBFDC8DB12" author="Chemsak, J. A." box="[368,584,493,518]" pageId="5" pageNumber="339" pagination="303 - 317" refId="ref10487" refString="Chemsak, J. A. (1969) Records and descriptions of Mexican and Central American Tillomorphini (Coleoptera: Cerambycidae). The Pan-Pacific Entomologist, 45 (4), 303 - 317." type="journal article" year="1969">Chemsak, 1969: 306</bibRefCitation>
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;
<treatmentCitation id="0AFB2376D34AFFC1691DEDC8FD40DB12" author="Chemsak, J. A." box="[597,704,494,518]" page="124" pageId="5" pageNumber="339" year="1977">
<bibRefCitation id="EFCB7896D34AFFC1691DEDC8FD40DB12" author="Chemsak, J. A." box="[597,704,494,518]" pageId="5" pageNumber="339" pagination="124 - 125" refId="ref10524" refString="Chemsak, J. A. (1977) New neotropical Tillomorphini in the genus Tetranodus Linell (Coleoptera, Cerambycidae). The Pan- Pacific Entomologist, 53 (2), 124 - 125." type="journal article" year="1977">1977: 124</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFB2376D34AFFC16984EDCBFC52DB12" author="Chemsak, J. A. &amp; Linsley, E. G. &amp; Noguera, F. A." box="[716,978,493,518]" page="73" pageId="5" pageNumber="339" year="1992">
<bibRefCitation id="EFCB7896D34AFFC16984EDCBFC52DB12" author="Chemsak, J. A. &amp; Linsley, E. G. &amp; Noguera, F. A." box="[716,978,493,518]" pageId="5" pageNumber="339" refId="ref10561" refString="Chemsak, J. A., Linsley, E. G. &amp; Noguera, F. A. (1992) Listados faunisticos de Mexico. II. Los Cerambycidae y Disteniidae de Norteamerica, Centroamerica y las Indias Occidentales (Coleoptera). Universidad Nacional Autonoma de Mexico, Mexico, Distrito Federal, 204 pp." type="book" year="1992">
Chemsak
<emphasis id="B92ED975D34AFFC1687BEDC8FCE9DB12" box="[819,873,494,518]" italics="true" pageId="5" pageNumber="339">et al.</emphasis>
, 1992: 73
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(checklist);
<treatmentCitation id="0AFB2376D34AFFC16F1DEDC8FA87DB12" author="Monne, M. A." box="[1109,1287,494,518]" page="72" pageId="5" pageNumber="339" year="1993">
<bibRefCitation id="EFCB7896D34AFFC16F1DEDC8FA87DB12" author="Monne, M. A." box="[1109,1287,494,518]" pageId="5" pageNumber="339" refId="ref11191" refString="Monne, M. A. (1993) Catalogue of the Cerambycidae (Coleoptera) of the Western Hemisphere. Part IX. Subfamily Cerambycinae: Tribes Clytini, Anaglyptini, Tillomorphini and Cleomenini. Sociedade Brasileira de Entomologia, Sao Paulo, 131 pp." type="book" year="1993">Monné, 1993: 72</bibRefCitation>
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(cat.);
<treatmentCitation id="0AFB2376D34AFFC16E1CEDC8FE30DB30" author="Monne, M. A. &amp; Giesbert, E. F." page="121" pageId="5" pageNumber="339" year="1994">
<bibRefCitation id="EFCB7896D34AFFC16E1CEDC8FE30DB30" author="Monne, M. A. &amp; Giesbert, E. F." pageId="5" pageNumber="339" refId="ref11387" refString="Monne, M. A. &amp; Giesbert, E. F. (1994) Checklist of the Cerambycidae and Disteniidae (Coleoptera) of the Western Hemisphere. Wolfsgarden Books, Burbank, 409 pp." type="book" year="1994">Monné &amp; Giesbert, 1994: 121</bibRefCitation>
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(checklist);
<treatmentCitation id="0AFB2376D34AFFC1697AEE2BFC03DB30" author="Noguera, F. A. &amp; Chemsak, J. A." box="[562,899,524,549]" page="401" pageId="5" pageNumber="339" year="1996">
<bibRefCitation id="EFCB7896D34AFFC1697AEE2BFC03DB30" author="Noguera, F. A. &amp; Chemsak, J. A." box="[562,899,524,549]" pageId="5" pageNumber="339" pagination="381 - 409" refId="ref11465" refString="Noguera, F. A. &amp; Chemsak, J. A. (1996) Cerambycidae (Coleoptera). In: Llorente Bousquets, J., Garcia Aldrete, A. N. &amp; Soriano, E. (Eds.), Biodiversidad, taxonomia, y biogeografia de artropodos de Mexico: Hacia una sintesis de su conocimiento. Universidad Nacional Autonoma de Mexico, Mexico, pp. 381 - 409." type="book chapter" year="1996">Noguera &amp; Chemsak, 1996: 401</bibRefCitation>
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(cat.);
<treatmentCitation id="0AFB2376D34AFFC16887EE2AFB0DDB31" author="Monne, M. A." box="[975,1165,524,549]" page="554" pageId="5" pageNumber="339" year="2005">
<bibRefCitation id="EFCB7896D34AFFC16887EE2AFB0DDB31" author="Monne, M. A." box="[975,1165,524,549]" pageId="5" pageNumber="339" pagination="1 - 765" refId="ref11238" refString="Monne, M. A. (2005) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical Region. Part I. Subfamily Cerambycinae. Zootaxa, 946 (1), 1 - 765. https: // doi. org / 10.11646 / zootaxa. 946.1.1" type="journal article" year="2005">Monné, 2005: 554</bibRefCitation>
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(cat.);
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<bibRefCitation id="EFCB7896D34AFFC16F92EE2BFA1CDB30" author="Hovore, F. T." box="[1242,1436,524,549]" pageId="5" pageNumber="339" pagination="363 - 378" refId="ref10677" refString="Hovore, F. T. (2006) The Cerambycidae (Coleoptera) of Guatemala. In: Cano, E. (Ed.), Biodiversidad de Guatemala. Vol. 1. Universidad del Valle de Guatemala, Guatemala, pp. 363 - 378." type="book chapter" year="2006">Hovore, 2006: 374</bibRefCitation>
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(distr.);
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<bibRefCitation id="EFCB7896D34AFFC16A51EE0DFDC4DB57" author="Monne, M. A. &amp; Hovore, F. T." box="[281,580,554,579]" pageId="5" pageNumber="339" refId="ref11425" refString="Monne, M. A. &amp; Hovore, F. T. (2006) Checklist of the Cerambycidae, or longhorned wood-boring beetles, of the Western Hemisphere. BioQuip Publications, Rancho Dominguez, 394 pp." type="book" year="2006">Monné &amp; Hovore, 2006: 133</bibRefCitation>
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(checklist);
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<bibRefCitation id="EFCB7896D34AFFC1698DEE0CFC1FDB57" author="Swift, I. &amp; Bezark, L. G. &amp; Nearns, E. H. &amp; Solis, A. &amp; Hovore, F. T." box="[709,927,554,579]" pageId="5" pageNumber="339" pagination="1 - 68" refId="ref11837" refString="Swift, I., Bezark, L. G., Nearns, E. H., Solis, A. &amp; Hovore, F. T. (2010) Checklist of the Cerambycidae (Coleoptera) of Costa Rica. Insecta Mundi, 0131, 1 - 68." type="book chapter" year="2010">
Swift
<emphasis id="B92ED975D34AFFC1684CEE0AFCB1DB57" box="[772,817,555,579]" italics="true" pageId="5" pageNumber="339">et al</emphasis>
., 2010: 30
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(distr.);
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<bibRefCitation id="EFCB7896D34AFFC168B1EE0DFB35DB57" author="Monne, M. A." box="[1017,1205,555,579]" pageId="5" pageNumber="339" refId="ref11344" refString="Monne, M. A. (2019) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical region. Part I. Subfamily Cerambycinae. Available from: http: // cerambyxcat. com / (accessed 9 November 2019)" type="url" year="2019">Monné, 2019: 827</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0AFB2376D34AFFC16E48EE0CFF6DDB76" author="Bezark, L. G." page="177" pageId="5" pageNumber="339" year="2019">
<bibRefCitation id="EFCB7896D34AFFC16E48EE0CFF6DDB76" author="Bezark, L. G." pageId="5" pageNumber="339" pagination="2019" refId="ref10319" refString="Bezark, L. G. (2019 b) Checklist of the Oxypeltidae, Vesperidae, Disteniidae and Cerambycidae (Coleoptera) of the Western Hemisphere. 2019 Edition. Updated through 31 December 2018. Available from: www. bezbycids. com / checklists / WestHemiCerambycidae 2019. pdf (accessed 9 November 2019)" type="journal article" year="2019">Bezark, 2019b: 177</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34056ECD34AFFC26BDFEEACFE71DBBF" lastPageId="6" lastPageNumber="340" pageId="5" pageNumber="339" type="description">
<paragraph id="8BE50567D34AFFC16BDFEEACFC8CDA69" blockId="5.[151,1437,650,2045]" pageId="5" pageNumber="339">
<emphasis id="B92ED975D34AFFC16BDFEEACFE29DBB1" bold="true" box="[151,425,650,677]" pageId="5" pageNumber="339">Redescription. Female.</emphasis>
Integument mostly dark brown, almost black; parts of frons, clypeus and labrum dark reddish brown; scape, pedicel, antennomeres III–IV, basal third of V, base of VI dark reddish brown; antennomere VI and basal 2/3 of VIII reddish brown; remaining surface of antennomeres V, VI, and VIII dark brown; antennomeres IX–XI dark brown; sides of prothorax mostly dark reddish brown; elytra with large, transverse pale yellow band slightly before middle, from epipleural margin to suture; trochanter and peduncle of femora, and apex of femora reddish brown (gradually darker toward apex of peduncle); tibiae slightly reddish brown on base and apex; tarsomeres reddish brown, except tarsomeres V mostly dark brown.
</paragraph>
<paragraph id="8BE50567D34AFFC16B8FEFA1FE78DF3D" blockId="5.[151,1437,650,2045]" pageId="5" pageNumber="339">
<emphasis id="B92ED975D34AFFC16B8FEFA1FE8EDAB5" bold="true" box="[199,270,903,929]" pageId="5" pageNumber="339">Head.</emphasis>
Frons, vertex, and area behind eyes coarsely, densely punctate (punctures partially confluent on vertex and behind eyes); with short, sparse, decumbent white setae; with a few long, erect white setae behind lower eye lobes (more light yellowish-brown depending on light intensity). Genae moderately finely, densely, confluently punctate, except smooth apex; with short, sparse, decumbent white setae, except glabrous apex. Antennal tubercles with sculpturing and setae as on frons, except smooth, glabrous apex. Postclypeus with sculpturing and setae as on frons on central area, smooth, glabrous laterally; with a few long, erect yellowish-brown setae on each side of wide central area. Labrum with short, sparse, bristly yellowish-brown setae, and one long, erect yellowish-brown seta on each side. Mandibles with a few long, erect yellowish-brown setae on outer surface. Gulamentum smooth, glabrous on posterior half, transversely striate, with a few short white setae, and a few long, erect white setae on anterior half (especially on anterior third). Upper eye lobes present, very distinct; distance between upper eye lobes 0.95 times length of scape (0.55 times distance between outer margins of eyes); in frontal view, distance between lower eye lobes 1.12 times length of scape (0.60 times distance between outer margins of eyes). Antennae 1.5 times elytral length (only
<typeStatus id="54E1BBC5D34AFFC16A3DE911FE57DC45" box="[373,471,1335,1361]" pageId="5" pageNumber="339" type="holotype">holotype</typeStatus>
measured), almost reaching elytral apex. Scape and antennomeres with short, decumbent white setae, slightly more abundant from antennomere V; ventral surface of scape, pedicel and antennomeres III–VI whit long, erect, sparse white setae (more light yellowish-brown depending on light intensity); dorsal surface of scape, and dorsal apex of antennomeres III–X with a few long, erect white setae (more light yellowish-brown depending on light intensity), gradually shorter toward X. Antennal formula (ratio) based on length of antennomere III (only
<typeStatus id="54E1BBC5D34AFFC16BB4E9CDFEDEDF11" box="[252,350,1515,1541]" pageId="5" pageNumber="339" type="holotype">holotype</typeStatus>
measured): scape = 0.68; pedicel = 0.17; IV = 0.51; V = 0.71; VI = 0.68; VII = 0.54; VIII = 0.46; IX = 0.34; X = 0.29; XI = 0.40.
</paragraph>
<paragraph id="8BE50567D34AFFC26B8FEA15FE89D8A4" blockId="5.[151,1437,650,2045]" lastBlockId="6.[151,1437,153,1547]" lastPageId="6" lastPageNumber="340" pageId="5" pageNumber="339">
<emphasis id="B92ED975D34AFFC16B8FEA15FEA7DF59" bold="true" box="[199,295,1587,1613]" pageId="5" pageNumber="339">Thorax.</emphasis>
Prothorax longer than wider; sides with two constrictions, anterior one narrow, slightly rounded between then, distinctly rounded from second anterior constriction to posterior seventh, then strongly narrowed toward apex (nearly straight on posterior seventh). Pronotum in lateral view, distinctly elevated from posterior margin to before middle; coarsely alveolate, with one minute whitish seta on center of each alveolus; with dense, transverse white pubescent band close to posterior margin; with long, erect, sparse yellowish-white setae on anterior third and sides of middle third.
<collectingRegion id="499ECB85D34AFFC16AF4EAC1FE79DE15" box="[444,505,1767,1793]" country="Haiti" name="Sud-Est" pageId="5" pageNumber="339">Sides</collectingRegion>
of prothorax micropunctate on wide area close to pronotum, with coarse punctures interspersed, smooth close to prosternum; with a few short white setae. Prosternum moderately coarsely, shallowly, sparsely punctate; with short, very sparse white setae. Prosternal process with its narrowest area 0.16 times width of procoxal cavity; with short white pubescence slightly more abundant than on prosternum. Mesoventrite with short, sparse white setae, slightly denser laterally. Mesanepisternum with yellowish-white pubescence, gradually denser toward elytron; mesepimeron with short, sparse white setae. Metanepisternum with short, decumbent, moderately abundant white setae. Metaventrite with white pubescence laterally close to metacoxal cavities, with long, erect white setae on remaining surface. Scutellum with sparse yellowish-white pubescence on anterior area, dense, slight longer on posterior area.
<emphasis id="B92ED975D349FFC26AB9ECBFFDC4D9A7" bold="true" box="[497,580,153,179]" pageId="6" pageNumber="340">Elytra.</emphasis>
Basal gibbosity nearly absent; surface coarsely, sparsely punctate (punctures slightly finer and more abundant toward apex); pale yellow area tumid; most punctures of posterior 2/3 with short yellowish-white setae, gradually longer toward apex; some punctures on entire surface with long, erect yellowishwhite setae, slightly more abundant on posterior quarter.
<emphasis id="B92ED975D349FFC2686EED23FCE5D80B" bold="true" box="[806,869,261,287]" pageId="6" pageNumber="340">Legs.</emphasis>
Femora strongly pedunculate-clavate; with short, sparse, decumbent white setae (sparser on profemora), with long, erect yellowish-white setae interspersed. Tibiae with short pubescence not obscuring integument, white, sparser basally, gradually denser, yellower toward apex, especially ventrally on posterior third, with long, erect white setae interspersed (more yellowish depending on light intensity).
</paragraph>
<paragraph id="8BE50567D349FFC26B8FED9EFC68DB08" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
<emphasis id="B92ED975D349FFC26B8FED9EFEC3D8C7" bold="true" box="[199,323,440,467]" pageId="6" pageNumber="340">Abdomen.</emphasis>
Ventrites finely, sparsely punctate centrally, slightly more abundant laterally; with short, sparse, decumbent white setae, distinctly more abundant laterally from ventrite II, with long, erect yellowish-white setae interspersed; apex of ventrite V rounded (slightly more projected centrally).
</paragraph>
<paragraph id="8BE50567D349FFC26B8FEE02FC36DB2B" blockId="6.[151,1437,153,1547]" box="[199,950,548,575]" pageId="6" pageNumber="340">
<emphasis id="B92ED975D349FFC26B8FEE02FEB8DB2B" bold="true" box="[199,312,548,575]" pageId="6" pageNumber="340">Variation</emphasis>
.
<collectingRegion id="499ECB85D349FFC26A0EEE03FE03DB2B" box="[326,387,549,575]" country="Haiti" name="Sud-Est" pageId="6" pageNumber="340">Sides</collectingRegion>
of prothorax from reddish-brown to almost black.
</paragraph>
<paragraph id="8BE50567D349FFC26B8FEE6EFE71DBBF" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
<emphasis id="B92ED975D349FFC26B8FEE6EFE19DB77" bold="true" box="[199,409,584,611]" pageId="6" pageNumber="340">Dimensions (mm)</emphasis>
, females. Total length, 4.85–6.35; prothoracic length, 1.25–1.65; anterior prothoracic width, 0.80–1.10; posterior prothoracic width, 0.65–0.90; maximum prothoracic width, 1.00–1.35; humeral width, 1.05– 1.40; elytral length, 2.95–3.70.
</paragraph>
</subSubSection>
<subSubSection id="C34056ECD349FFC26B8FEE92FE73DA03" pageId="6" pageNumber="340" type="materials_examined">
<paragraph id="8BE50567D349FFC26B8FEE92FE73DA03" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
<emphasis id="B92ED975D349FFC26B8FEE92FE34DBDB" bold="true" box="[199,436,692,719]" pageId="6" pageNumber="340">Material examined.</emphasis>
<materialsCitation id="3B320F3AD349FFC26AF7EE93FD3FDBE7" ID-GBIF-Occurrence="2574624374" collectionCode="VI" country="Honduras" elevation="780" latitude="14.0" location="Zamorano University area" longLatPrecision="7740" longitude="-87.0" pageId="6" pageNumber="340" specimenCount="5" specimenCount-female="5" stateProvince="Francisco Morazan">
<collectingCountry id="F34D45F7D349FFC26AF7EE93FDDBDBDB" box="[447,603,693,719]" name="Honduras" pageId="6" pageNumber="340">HONDURAS</collectingCountry>
(
<emphasis id="B92ED975D349FFC26926EE93FCE0DBDB" bold="true" box="[622,864,693,719]" pageId="6" pageNumber="340">new country record</emphasis>
),
<emphasis id="B92ED975D349FFC26833EE90FBDCDBDB" box="[891,1116,693,719]" italics="true" pageId="6" pageNumber="340">
<collectingRegion id="499ECB85D349FFC26833EE90FBDCDBDB" box="[891,1116,693,719]" country="Honduras" name="Francisco Morazan" pageId="6" pageNumber="340">Francisco Morazán</collectingRegion>
</emphasis>
:
<location id="8E8553BCD349FFC26F27EE93FA1DDBDB" LSID="urn:lsid:plazi:treatment:03F3B471D34AFFC26BDFED4AFD6EDF1F:8E8553BCD349FFC26F27EE93FA1DDBDB" box="[1135,1437,693,719]" country="Honduras" latitude="14.0" longLatPrecision="7740" longitude="-87.0" name="Zamorano University area" pageId="6" pageNumber="340" stateProvince="Francisco Morazan">Zamorano University area</location>
(
<geoCoordinate id="EE6E63A0D349FFC26BD7EEFFFF71DBE7" box="[159,241,729,755]" degrees="14.0" direction="north" orientation="latitude" pageId="6" pageNumber="340" precision="5555" value="14.0">14.0ºN</geoCoordinate>
,
<geoCoordinate id="EE6E63A0D349FFC26BB5EEFFFED4DBE7" box="[253,340,729,755]" degrees="87.0" direction="west" orientation="longitude" pageId="6" pageNumber="340" precision="5555" value="-87.0">87.0ºW</geoCoordinate>
;
<quantity id="4CA2A882D349FFC26A28EEFEFE62DBE7" box="[352,482,728,755]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="8.1" metricValueMin="7.5" pageId="6" pageNumber="340" unit="m" value="780.0" valueMax="810.0" valueMin="750.0">
<elevation id="0077E254D349FFC26A28EEFEFE62DBE7" box="[352,482,728,755]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="8.1" metricValueMin="7.5" pageId="6" pageNumber="340" unit="m" value="780.0" valueMax="810.0" valueMin="750.0">750–810 m</elevation>
</quantity>
),
<specimenCount id="9D5CCEEED349FFC26ABFEEFEFDE7DBE7" box="[503,615,728,755]" pageId="6" pageNumber="340" type="female">5 females</specimenCount>
,
<date id="FFE423A7D349FFC2693AEEFFFD3FDBE7" box="[626,703,729,755]" pageId="6" pageNumber="340" value="2018-06-01" valueMax="2018-06-06" valueMin="2018-06-01">
1–6.
<collectionCode id="ED4B9DA2D349FFC269D6EEFFFD3FDBE7" box="[670,703,729,755]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="340">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B320F3AD349FFC269F7EEFFFE70DA03" ID-GBIF-Occurrence="2574624370" collectionCode="DHCO, MZSP, JVCO" collectorName="E. van den Berghe &amp; J. Vlasak &amp; B. Raber &amp; D. Heffern" country="Honduras" pageId="6" pageNumber="340" specimenCount="1">
.2018,
<collectorName id="26AF60B1D349FFC26846EEFFFC61DBE7" box="[782,993,729,755]" pageId="6" pageNumber="340">E. van den Berghe</collectorName>
,
<collectorName id="26AF60B1D349FFC268A5EEFFFBD7DBE7" box="[1005,1111,729,755]" pageId="6" pageNumber="340">J. Vlasak</collectorName>
,
<collectorName id="26AF60B1D349FFC26F2BEEFFFB49DBE7" box="[1123,1225,729,755]" pageId="6" pageNumber="340">B. Raber</collectorName>
, &amp;
<collectorName id="26AF60B1D349FFC26FBAEEFFFAECDBE7" box="[1266,1388,729,755]" pageId="6" pageNumber="340">D. Heffern</collectorName>
col. (3,
<collectionCode id="ED4B9DA2D349FFC26BF5EEDBFE90DA03" box="[189,272,765,791]" pageId="6" pageNumber="340">DHCO</collectionCode>
; 1
<collectionCode id="ED4B9DA2D349FFC26A78EEDBFEFDDA03" box="[304,381,765,791]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="340" type="Museum">MZSP</collectionCode>
; 1
<collectionCode id="ED4B9DA2D349FFC26AD5EEDBFE68DA03" box="[413,488,765,791]" pageId="6" pageNumber="340">JVCO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34056ECD349FFC26B8FEF07FD6EDF1F" pageId="6" pageNumber="340" type="discussion">
<paragraph id="8BE50567D349FFC26B8FEF07FEC8DDF8" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
<emphasis id="B92ED975D349FFC26B8FEF07FEBBDA2F" bold="true" box="[199,315,801,827]" pageId="6" pageNumber="340">Remarks.</emphasis>
<bibRefCitation id="EFCB7896D349FFC26A0BEF07FE7EDA2F" author="Chemsak, J. A." box="[323,510,801,827]" pageId="6" pageNumber="340" pagination="124 - 125" refId="ref10524" refString="Chemsak, J. A. (1977) New neotropical Tillomorphini in the genus Tetranodus Linell (Coleoptera, Cerambycidae). The Pan- Pacific Entomologist, 53 (2), 124 - 125." type="journal article" year="1977">Chemsak (1977)</bibRefCitation>
provided a key to species of
<taxonomicName id="4C5A7EE4D349FFC26804EF07FBDDDA2F" authority="Linell, 1896" authorityName="Linell" authorityYear="1896" box="[844,1117,801,827]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B92ED975D349FFC26804EF07FC49DA2F" box="[844,969,801,827]" italics="true" pageId="6" pageNumber="340">Tetranodus</emphasis>
Linell, 1896
</taxonomicName>
, and reported in the alternative of couplet “1”: “Elytra gibbose at base or with basal punctures coarse, dense, without broad longitudinal spaces between rows of punctures”, leading to alternative of couplet “2”; and “Elytra with basal punctures fine, sparse, arranged in rows separated by broad longitudinal glabrous spaces, color shining black, antennae paler”, leading to
<taxonomicName id="4C5A7EE4D349FFC26EC0EFA8FE0FDADF" authority="(Bates, 1880)" baseAuthorityName="Bates" baseAuthorityYear="1880" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="reticeps">
<emphasis id="B92ED975D349FFC26EC0EFA8FF6FDADF" italics="true" pageId="6" pageNumber="340">T. reticeps</emphasis>
(
<bibRefCitation id="EFCB7896D349FFC26BB5EF97FE06DADF" author="Bates, H. W." box="[253,390,945,971]" pageId="6" pageNumber="340" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates, 1880</bibRefCitation>
)
</taxonomicName>
. However, according to
<bibRefCitation id="EFCB7896D349FFC269D5EF97FCAFDADF" author="Bates, H. W." box="[669,815,945,971]" pageId="6" pageNumber="340" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates (1880)</bibRefCitation>
, “pedibus elytrisque rufo-castaneis [legs and elytra reddish-brown].” Examination of photographs of a
<typeStatus id="54E1BBC5D349FFC269FEEFF3FC8DDAFB" box="[694,781,981,1007]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
(
<figureCitation id="136119E2D349FFC26855EFF3FC26DAFB" box="[797,934,981,1007]" captionStart="FIGURES 22–30. 22–26" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[191,1396,185,1919]" captionTargetId="figure@10.[185,1402,181,1919]" captionTargetPageId="10" captionText="FIGURES 22–30. 22–26) Pentanodes nicaraguensis, holotype female: 22) Dorsal habitus; 23) Ventral habitus; 24) Lateral habitus; 25) Head, frontal view; 26) Labial palpomere III. 27–30) Euderces reticeps, holotype: 27) Dorsal habitus; 28) Lateral habitus; 29) Basal area of the elytra, lateral view; 30) Labels." figureDoi="http://doi.org/10.5281/zenodo.3698871" httpUri="https://zenodo.org/record/3698871/files/figure.png" pageId="6" pageNumber="340">Figs. 27–30</figureCitation>
), shows that the description by
<bibRefCitation id="EFCB7896D349FFC26E44EFF3FA1CDAFB" author="Bates, H. W." box="[1292,1436,981,1007]" pageId="6" pageNumber="340" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates (1880)</bibRefCitation>
is accurate because the basal gibbosity of the elytra is absent and the color is evidently not black. As we did not find morphological differences between the
<typeStatus id="54E1BBC5D349FFC2691DE83BFD2CDD23" box="[597,684,1053,1079]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
and the specimens examined, except for the color, we believe that all of them belong to the same species. The color of the antennomeres in the
<typeStatus id="54E1BBC5D349FFC26F4FE867FBDEDD4F" box="[1031,1118,1089,1115]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
follows the same pattern of that in the darker specimens, although to a slight degree. The elytral punctures in the
<typeStatus id="54E1BBC5D349FFC26F13E843FB32DD6B" box="[1115,1202,1125,1151]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
appear to be coarser and sparser when seen in dorsal view. However, the concentration of punctures in the darker specimens is slightly variable (not different in the
<typeStatus id="54E1BBC5D349FFC26A91E88BFDB4DDD3" box="[473,564,1197,1223]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
), and the size is not different too (more evident when the
<typeStatus id="54E1BBC5D349FFC26FF0E88BFA8FDDD3" box="[1208,1295,1197,1223]" pageId="6" pageNumber="340" type="syntype">syntype</typeStatus>
is examined in lateral view).
</paragraph>
<paragraph id="8BE50567D349FFC26B8FE8D3FACCDC8B" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
Although it has been reported that
<taxonomicName id="4C5A7EE4D349FFC2691EE8D0FD4BDC1B" baseAuthorityName="Bates" baseAuthorityYear="1880" box="[598,715,1270,1295]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="reticeps">
<emphasis id="B92ED975D349FFC2691EE8D0FD4BDC1B" box="[598,715,1270,1295]" italics="true" pageId="6" pageNumber="340">T. reticeps</emphasis>
</taxonomicName>
was described based on a single specimen (
<emphasis id="B92ED975D349FFC26F8DE8D0FB6DDC1B" box="[1221,1261,1270,1295]" italics="true" pageId="6" pageNumber="340">e.g.</emphasis>
<bibRefCitation id="EFCB7896D349FFC26FBDE8D3FED8DC27" author="Tavakilian, G. L. &amp; Chevillotte, H." pageId="6" pageNumber="340" refId="ref11890" refString="Tavakilian, G. L. &amp; Chevillotte, H. (2019) Titan: base de donnees internationales sur les Cerambycidae ou Longicornes. Available from: http: // titan. gbif. fr / index. html (accessed 9 November 2019)" type="url" year="2019">Tavakilian and Chevillotte 2019</bibRefCitation>
),
<bibRefCitation id="EFCB7896D349FFC26A25E93FFE7EDC27" author="Bates, H. W." box="[365,510,1305,1331]" pageId="6" pageNumber="340" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates (1880)</bibRefCitation>
made very clear that he had more specimens: “… and by the form of the thorax, which is somewhat constricted near its anterior margin and very greatly narrowed, as well as depressed, at the base, the middle portion being dilated (more so in some examples than others) and convex.” Accordingly, the species was described based on
<typeStatus id="54E1BBC5D349FFC26A3BE9A3FE59DC8B" box="[371,473,1413,1439]" pageId="6" pageNumber="340" type="syntype">syntypes</typeStatus>
. Apparently, the error is due to the single dimension provided by
<bibRefCitation id="EFCB7896D349FFC26FFDE9A3FAC7DC8B" author="Bates, H. W." box="[1205,1351,1413,1439]" pageId="6" pageNumber="340" pagination="17 - 152" refId="ref10226" refString="Bates, H. W. (1880) Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Published for the editors by R. H. Porter, London, 136 pp. [pp. 17 - 152]" type="book chapter" year="1880">Bates (1880)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE50567D349FFC26BDFE98FFD6EDF1F" blockId="6.[151,1437,153,1547]" pageId="6" pageNumber="340">
<taxonomicName id="4C5A7EE4D349FFC26BDFE98FFEF4DCD7" baseAuthorityName="Bates" baseAuthorityYear="1880" box="[151,372,1449,1475]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="reticeps">
<emphasis id="B92ED975D349FFC26BDFE98FFEF4DCD7" box="[151,372,1449,1475]" italics="true" pageId="6" pageNumber="340">Tetranodus reticeps</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5A7EE4D349FFC26944E98CFCE7DCD7" authority="Chemsak, 1969" authorityName="Chemsak" authorityYear="1969" box="[524,871,1449,1475]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="angulicollis">
<emphasis id="B92ED975D349FFC26944E98CFD2CDCD7" box="[524,684,1449,1475]" italics="true" pageId="6" pageNumber="340">T. angulicollis</emphasis>
<bibRefCitation id="EFCB7896D349FFC269FDE98FFCE7DCD7" author="Chemsak, J. A." box="[693,871,1449,1475]" pageId="6" pageNumber="340" pagination="303 - 317" refId="ref10487" refString="Chemsak, J. A. (1969) Records and descriptions of Mexican and Central American Tillomorphini (Coleoptera: Cerambycidae). The Pan-Pacific Entomologist, 45 (4), 303 - 317." type="journal article" year="1969">Chemsak, 1969</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5A7EE4D349FFC2683CE98CFA80DCD7" authority="Chemsak &amp; Linsley, 1988" authorityName="Chemsak &amp; Linsley" authorityYear="1988" box="[884,1280,1449,1475]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="copei">
<emphasis id="B92ED975D349FFC2683CE98CFC4DDCD7" box="[884,973,1450,1475]" italics="true" pageId="6" pageNumber="340">T. copei</emphasis>
Chemsak &amp; Linsley, 1988
</taxonomicName>
,
<taxonomicName id="4C5A7EE4D349FFC26E44E98CFEA4DCF3" authority="Linell, 1897" authorityName="Linell" authorityYear="1897" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="niveicollis">
<emphasis id="B92ED975D349FFC26E44E98CFA1CDCD7" box="[1292,1436,1449,1475]" italics="true" pageId="6" pageNumber="340">T. niveicollis</emphasis>
Linell, 1897
</taxonomicName>
,
<taxonomicName id="4C5A7EE4D349FFC26A67E9E8FDFDDCF3" authority="Chemsak, 1969" authorityName="Chemsak" authorityYear="1969" box="[303,637,1485,1511]" class="Insecta" family="Cerambycidae" genus="Tetranodus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B92ED975D349FFC26A67E9E8FE43DCF3" box="[303,451,1486,1511]" italics="true" pageId="6" pageNumber="340">T. rugipennis</emphasis>
<bibRefCitation id="EFCB7896D349FFC26A83E9EBFDFDDCF3" author="Chemsak, J. A." box="[459,637,1485,1511]" pageId="6" pageNumber="340" pagination="303 - 317" refId="ref10487" refString="Chemsak, J. A. (1969) Records and descriptions of Mexican and Central American Tillomorphini (Coleoptera: Cerambycidae). The Pan-Pacific Entomologist, 45 (4), 303 - 317." type="journal article" year="1969">Chemsak, 1969</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5A7EE4D349FFC269F0E9E8FB8ADCF3" authority="Chemsak, 1977" authorityName="Chemsak" authorityYear="1977" box="[696,1034,1485,1511]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="340" phylum="Arthropoda" rank="species" species="tropipennis">
<emphasis id="B92ED975D349FFC269F0E9E8FCD4DCF3" box="[696,852,1486,1511]" italics="true" pageId="6" pageNumber="340">T. tropipennis</emphasis>
<bibRefCitation id="EFCB7896D349FFC26813E9EBFB8ADCF3" author="Chemsak, J. A." box="[859,1034,1485,1511]" pageId="6" pageNumber="340" pagination="124 - 125" refId="ref10524" refString="Chemsak, J. A. (1977) New neotropical Tillomorphini in the genus Tetranodus Linell (Coleoptera, Cerambycidae). The Pan- Pacific Entomologist, 53 (2), 124 - 125." type="journal article" year="1977">Chemsak, 1977</bibRefCitation>
</taxonomicName>
by the alveolate pronotum, while it is longitudinally striate in these species (
<emphasis id="B92ED975D349FFC26917E9D4FD07DF1F" box="[607,647,1522,1547]" italics="true" pageId="6" pageNumber="340">e.g.</emphasis>
<figureCitation id="136119E2D349FFC269C6E9D7FD61DF1F" box="[654,737,1521,1547]" captionStart="FIGURES 15–21. 15–19" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[178,1413,277,1887]" captionTargetId="figure@7.[169,1418,273,1895]" captionTargetPageId="7" captionText="FIGURES 15–21. 15–19) Tetranodus reticeps, female 1: 15) Dorsal habitus; 16) Ventral habitus; 17) Lateral habitus; 18) Head, frontal view; 19) Pronotal sculpturing. 20) Tetranodus reticeps, female 2, dorsal habitus. 21) Tetranodus mexicanus, holotype female, pronotal sculpturing (currently, junior synonym of Tetranodus niveicollis Linell, 1897)." figureDoi="http://doi.org/10.5281/zenodo.3698869" httpUri="https://zenodo.org/record/3698869/files/figure.png" pageId="6" pageNumber="340">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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