treatments-xml/data/03/8C/A4/038CA47F72632077C6B57FDC9230F8D0.xml
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<document id="2E7BD122C87DF0CEEB72526F67601B07" ID-DOI="10.11646/zootaxa.5384.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10376440" ID-ZooBank="0211978A-E124-4D9F-9A92-B565AA7B7891" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1702549873658" checkinUser="plazi" docAuthor="Yu, Kun, Maddison, Wayne P. &amp; Zhang, Junxia" docDate="2023" docId="038CA47F72632077C6B57FDC9230F8D0" docLanguage="en" docName="zootaxa.5384.1.1.pdf" docOrigin="Zootaxa 5384 (1)" docSource="https://mapress.com/zt/article/download/zootaxa.5384.1.1/52479" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Orcevia bokoblin Yu, Maddison &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFB5DC0772662072C6227F449060FFD6" masterDocTitle="Taxonomic revision of Orcevia Thorell, 1890, with description of fifteen new species (Araneae, Salticidae, Euophryini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="6" updateTime="1702998765381" updateUser="guilherme">
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<mods:title id="231BF7EA56C0B9685F864F37CC4151E6">Taxonomic revision of Orcevia Thorell, 1890, with description of fifteen new species (Araneae, Salticidae, Euophryini)</mods:title>
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<mods:namePart id="E82DF8DB0EE67A0CF562F0AD249D8483">Yu, Kun</mods:namePart>
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<mods:affiliation id="47C558AF36752B8622C98B2A8239D1B5">Key Laboratory of Zoological Systematics and Application of Hebei Province, Hebei Basic Science Center for Biotic Interaction, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China. &amp; 13468690210 @ 163. com; https: // orcid. org / 0000 - 0002 - 5517 - 4484</mods:affiliation>
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<mods:namePart id="A4640FD74290A0C61BA5ABBA31B83DD3">Maddison, Wayne P.</mods:namePart>
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<mods:affiliation id="30F6068E95E768580F0DADBBBE7370A0">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada &amp; wayne. maddison @ ubc. ca; https: // orcid. org / 0000 - 0003 - 4953 - 4575</mods:affiliation>
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<mods:namePart id="3B3252829245A7A1EFEB013E30D5AD88">Zhang, Junxia</mods:namePart>
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<mods:affiliation id="50B50216958AFE9457938C2F2BF4FDD4">Key Laboratory of Zoological Systematics and Application of Hebei Province, Hebei Basic Science Center for Biotic Interaction, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China. &amp; jxzhang 1976 @ 163. com; https: // orcid. org / 0000 - 0003 - 2179 - 3954 Corresponding author</mods:affiliation>
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<taxonomicName id="4C256EEA72632077C6B57FDC92F7FF64" authority="Yu, Maddison &amp; Zhang" authorityName="Yu, Maddison &amp; Zhang" authorityYear="2023" box="[151,663,152,179]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bokoblin" status="sp. nov.">
<emphasis id="B951C97B72632077C6B57FDC9107FF65" bold="true" box="[151,359,152,179]" italics="true" pageId="5" pageNumber="6">Orcevia bokoblin</emphasis>
Yu, Maddison &amp; Zhang
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,
<taxonomicNameLabel id="A262740072632077C4877FDC9362FF64" box="[677,770,152,178]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(波克ĸ尔¤)
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<figureCitation id="131E09EC72632077C6B57FF890BCFF00" box="[151,220,188,214]" captionStart="FIGURES 112" captionStartId="19.[151,264,1823,1847]" captionTargetBox="[151,1436,181,1803]" captionTargetId="figure-17@19.[151,1436,181,1803]" captionTargetPageId="19" captionText="FIGURES 112. Comparison of male palpal bulbs of some euophryines, in ventral view. 18. Orcevia spp. [O. bokoblin Yu, Maddison &amp; Zhang sp. nov. (1), O. nietzschei Yu, Maddison &amp; Zhang sp. nov. (2), O. jinping Yu &amp; Zhang, sp. nov. (3), O. mercuryi Yu, Maddison &amp; Zhang, sp. nov. (4), O. shuyuani Yu &amp; Zhang, sp. nov. (5), O. zabkai Yu, Maddison &amp; Zhang, sp. nov. (6), O. yahaha Yu, Maddison &amp; Zhang, sp. nov. (7) and O. wuliang Yu &amp; Zhang, sp. nov. (8)]; 9. Laufeia concava Zhang &amp; Maddison, 2012; 10. Amoenema erhai Yu &amp; Zhang, 2023; 11. Lokina nyuewa Yu &amp; Zhang, 2023; 12. Chalcovietnamicus vietnamensis (Żabka, 1985). Black dotted lines indicate retrolateral edge of membrane between salticid radix and tegulum, white arrows point to the retrolateral extension, and yellow lines indicate the proximal margin of tegulum." figureDoi="http://doi.org/10.5281/zenodo.10376442" httpUri="https://zenodo.org/record/10376442/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
,
<figureCitation id="131E09EC72632077C6C87FF89150FF00" box="[234,304,188,214]" captionStart-0="FIGURES 2429" captionStart-1="FIGURES 3039" captionStart-2="FIGURES 4044" captionStartId-0="22.[151,264,1437,1461]" captionStartId-1="23.[151,264,1883,1907]" captionStartId-2="24.[151,264,1912,1936]" captionTargetBox-0="[332,1255,181,1414]" captionTargetBox-1="[225,1362,181,1860]" captionTargetBox-2="[196,1392,151,1889]" captionTargetId-0="figure-22@22.[332,1255,181,1414]" captionTargetId-1="figure-17@23.[225,1362,181,1860]" captionTargetId-2="figure-22@24.[196,1392,151,1889]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionText-0="FIGURES 2429. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., living males (2427) and females (2829), from Lambir Hills National Park (24, 27, 29) and Mulu National Park (2526, 28), Sarawak, show variation of coloration. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." captionText-1="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." captionText-2="FIGURES 4044. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., male palp (40), RTA (41), epigynum (42) and vulva (43 44), in ventral (40, 42), retrolateral (41), dorsal (43) and top (44) view.Abbreviations:AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; TL, tegular lobe." figureDoi-0="http://doi.org/10.5281/zenodo.10376450" figureDoi-1="http://doi.org/10.5281/zenodo.10376452" figureDoi-2="http://doi.org/10.5281/zenodo.10376454" httpUri-0="https://zenodo.org/record/10376450/files/figure.png" httpUri-1="https://zenodo.org/record/10376452/files/figure.png" httpUri-2="https://zenodo.org/record/10376454/files/figure.png" pageId="5" pageNumber="6">2444</figureCitation>
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:
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, leg.
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&amp;
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.
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:
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~
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), same data as holotype
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<location id="8EFA43B272632077C4567E349342FE5C" LSID="urn:lsid:plazi:treatment:038CA47F72632077C6B57FDC9230F8D0:8EFA43B272632077C4567E349342FE5C" box="[628,802,368,395]" country="Malaysia" county="Cave Trail" latitude="4.0582" longLatPrecision="7" longitude="114.8288" municipality="Clearwater Cave" name="Mulu Nat. Pk." pageId="5" pageNumber="6" stateProvince="Sarawak">Mulu Nat. Pk.</location>
,
<location id="8EFA43B272632077C5117E349459FE5C" LSID="urn:lsid:plazi:treatment:038CA47F72632077C6B57FDC9230F8D0:8EFA43B272632077C5117E349459FE5C" box="[819,1081,368,394]" country="Malaysia" county="Cave Trail" latitude="4.0582" longLatPrecision="7" longitude="114.8288" municipality="Clearwater Cave" name="Clearwater Cave Trail" pageId="5" pageNumber="6" stateProvince="Sarawak">Clearwater Cave Trail</location>
, 4.0582
<geoCoordinate id="EE1173AE72632077C2B87E349498FE5D" box="[1178,1272,368,395]" degrees="4.0584" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="-4.0584">
<geoCoordinate id="EE1173AE72632077C28B7E349575FE5D" box="[1193,1301,368,395]" degrees="4.0584" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="4.0584">4.0584°N</geoCoordinate>
</geoCoordinate>
, 114.8285
<geoCoordinate id="EE1173AE72632077C6B57ED0917FFE78" box="[151,287,404,430]" degrees="114.8288" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="114.8288">114.8288°E</geoCoordinate>
,
<quantity id="4CDDB88C72632077C70A7ED09100FE79" box="[296,352,404,431]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="60.0">
<elevation id="0008F25A72632077C70A7ED09100FE79" box="[296,352,404,431]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="60.0">60 m</elevation>
</quantity>
elev.,
<date id="FF9B33A972632077C7857ED09233FE78" box="[423,595,404,430]" pageId="5" pageNumber="6" value="2012-03-14">
<collectingDate id="EFDFCA4172632077C7857ED09233FE78" box="[423,595,404,430]" pageId="5" pageNumber="6" value="2012-03-14">14 March 2012</collectingDate>
</date>
, leg.
<collectorName id="26D070BF72632077C4A97ED19356FE78" box="[651,822,404,431]" pageId="5" pageNumber="6">W.P. Maddison</collectorName>
,
<collectorName id="26D070BF72632077C5627ED193AAFE78" box="[832,970,404,431]" pageId="5" pageNumber="6">E.K. Piascik</collectorName>
&amp;
<collectorName id="26D070BF72632077C5C87ED0945DFE78" box="[1002,1085,404,430]" pageId="5" pageNumber="6">A. Ang</collectorName>
,
<specimenCode id="DB83BD1272632077C2657ED0948BFE78" box="[1095,1259,404,430]" pageId="5" pageNumber="6">WPM#19-030</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B4D1F3472632077C2D77ED094EEFE20" collectingDate="2012-04-02" collectionCode="UBCZ" collectorName="W. P. Maddison &amp; E. K. Piascik" country="Malaysia" elevation="200" latitude="4.2022" location="Lambir Hills Nat. Pk." longLatPrecision="7" longitude="114.0283" pageId="5" pageNumber="6" specimenCode="KYU-SAL734, WPM#12-123" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="9D23DEE072632077C2D77ED0957EFE78" box="[1269,1310,404,430]" count="2" pageId="5" pageNumber="6" type="male">2♂</specimenCount>
<specimenCount id="9D23DEE072632077C33C7ED0955BFE78" box="[1310,1339,404,430]" count="1" pageId="5" pageNumber="6" type="female">1♀</specimenCount>
(
<collectionCode id="ED348DAC72632077C36A7ED095F8FE78" box="[1352,1432,404,430]" country="Canada" name="University of British Columbia, Spencer Museum" pageId="5" pageNumber="6">UBCZ</collectionCode>
;
<specimenCode id="DB83BD1272632077C6B57EFC9121FE04" box="[151,321,440,466]" pageId="5" pageNumber="6">KYU-SAL734</specimenCode>
),
<collectingCountry id="F33255F972632077C7777EFC918BFE04" box="[341,491,440,466]" name="Malaysia" pageId="5" pageNumber="6">MALAYSIA</collectingCountry>
:
<collectingRegion id="49E1DB8B72632077C7D57EFC923AFE04" box="[503,602,440,466]" pageId="5" pageNumber="6">Sarawak</collectingRegion>
,
<location id="8EFA43B272632077C4457EFC9301FE04" LSID="urn:lsid:plazi:treatment:038CA47F72632077C6B57FDC9230F8D0:8EFA43B272632077C4457EFC9301FE04" box="[615,865,440,467]" country="Malaysia" latitude="4.2022" longLatPrecision="7" longitude="114.0283" name="Lambir Hills Nat. Pk." pageId="5" pageNumber="6" stateProvince="Sarawak">Lambir Hills Nat. Pk.</location>
,
<location id="8EFA43B272632077C54C7EFC9422FE04" LSID="urn:lsid:plazi:treatment:038CA47F72632077C6B57FDC9230F8D0:8EFA43B272632077C54C7EFC9422FE04" box="[878,1090,440,466]" country="Malaysia" latitude="4.2022" longLatPrecision="7" longitude="114.0283" name="Lepoh-Ridan Trail" pageId="5" pageNumber="6" stateProvince="Sarawak">Lepoh-Ridan Trail</location>
, 4.2022
<geoCoordinate id="EE1173AE72632077C2BD7EFC949DFE04" box="[1183,1277,440,466]" degrees="4.2028" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="-4.2028">
<geoCoordinate id="EE1173AE72632077C28C7EFC9579FE04" box="[1198,1305,440,466]" degrees="4.2028" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="4.2028">4.2028°N</geoCoordinate>
</geoCoordinate>
, 114.0279
<geoCoordinate id="EE1173AE72632077C6B57E98917EFE20" box="[151,286,476,502]" degrees="114.0283" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="114.0283">
<geoCoordinate id="EE1173AE72632077C6B57E98916DFE20" box="[151,269,476,502]" pageId="5" pageNumber="6">114.0283°</geoCoordinate>
E
</geoCoordinate>
,
<quantity id="4CDDB88C72632077C70B7E989110FE21" box="[297,368,476,503]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="m" value="200.0">
<elevation id="0008F25A72632077C70B7E989110FE21" box="[297,368,476,503]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="m" value="200.0">200 m</elevation>
</quantity>
elev.,
<date id="FF9B33A972632077C79B7E98922AFE20" box="[441,586,476,502]" pageId="5" pageNumber="6" value="2012-04-02">
<collectingDate id="EFDFCA4172632077C79B7E98922AFE20" box="[441,586,476,502]" pageId="5" pageNumber="6" value="2012-04-02">2 April 2012</collectingDate>
</date>
, leg.
<collectorName id="26D070BF72632077C4A67E99934EFE20" box="[644,814,476,503]" pageId="5" pageNumber="6">W.P. Maddison</collectorName>
&amp;
<collectorName id="26D070BF72632077C5737E999380FE20" box="[849,992,476,503]" pageId="5" pageNumber="6">E.K. Piascik</collectorName>
,
<specimenCode id="DB83BD1272632077C5C87E9894EEFE20" box="[1002,1166,476,502]" pageId="5" pageNumber="6">WPM#12-123</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272632077C6E57D44910FFD50" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8B9A156972632077C6E57D44910FFD50" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
<emphasis id="B951C97B72632077C6E57D44912EFDCC" bold="true" box="[199,334,512,538]" pageId="5" pageNumber="6">Etymology.</emphasis>
The specific epithet is from “Bokoblin”, a small piggy monster in the game
<emphasis id="B951C97B72632077C2897D449105FDE8" italics="true" pageId="5" pageNumber="6">The Legend of Zelda: Breath of the Wild</emphasis>
, developed and published by Nintendo. In the Master Mode of this game, Bokoblin commonly has either silver or gold coloration, corresponding to two color-forms observed in the males of the new species; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272632077C6E57DD495EBFCEC" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B9A156972632077C6E57DD495EBFCEC" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
<emphasis id="B951C97B72632077C6E57DD49122FD7C" bold="true" box="[199,322,656,682]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
It resembles
<taxonomicName id="4C256EEA72632077C7F27DD5923DFD7C" authority="Yu &amp; Maddison &amp; Zhang, 2023" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" box="[464,605,656,683]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nietzschei" status="sp. nov.">
<emphasis id="B951C97B72632077C7F27DD5923DFD7C" box="[464,605,656,683]" italics="true" pageId="5" pageNumber="6">O. nietzschei</emphasis>
</taxonomicName>
<emphasis id="B951C97B72632077C4407DD492D9FD7C" bold="true" box="[610,697,656,682]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A262740072632077C4407DD492D9FD7C" box="[610,697,656,682]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the small and screw-like embolus, but males can be distinguished by the presence of a small ventro-basal femoral bump on palp (
<figureCitation id="131E09EC72632077C55C7DF093B5FD18" box="[894,981,692,718]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 36</figureCitation>
; vs. VFB is absent in
<taxonomicName id="4C256EEA72632077C2FF7DF19510FD18" authority="Yu &amp; Maddison &amp; Zhang, 2023" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" box="[1245,1392,692,719]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nietzschei" status="sp. nov.">
<emphasis id="B951C97B72632077C2FF7DF19510FD18" box="[1245,1392,692,719]" italics="true" pageId="5" pageNumber="6">O. nietzschei</emphasis>
</taxonomicName>
<emphasis id="B951C97B72632077C3597DF090A8FD24" bold="true" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A262740072632077C3597DF090A8FD24" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and the revolute tip of RTA (
<figureCitation id="131E09EC72632077C4067D9C92C6FD24" box="[548,678,728,754]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Figs 32, 34</figureCitation>
,
<figureCitation id="131E09EC72632077C4917D9C92B2FD24" box="[691,722,728,754]" captionStart="FIGURES 4044" captionStartId="24.[151,264,1912,1936]" captionTargetBox="[196,1392,151,1889]" captionTargetId="figure-22@24.[196,1392,151,1889]" captionTargetPageId="24" captionText="FIGURES 4044. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., male palp (40), RTA (41), epigynum (42) and vulva (43 44), in ventral (40, 42), retrolateral (41), dorsal (43) and top (44) view.Abbreviations:AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.10376454" httpUri="https://zenodo.org/record/10376454/files/figure.png" pageId="5" pageNumber="6">41</figureCitation>
; vs. tip of RTA is flat in
<taxonomicName id="4C256EEA72632077C5CB7D9D941AFD24" authority="Yu &amp; Maddison &amp; Zhang, 2023" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" box="[1001,1146,728,755]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nietzschei" status="sp. nov.">
<emphasis id="B951C97B72632077C5CB7D9D941AFD24" box="[1001,1146,728,755]" italics="true" pageId="5" pageNumber="6">O. nietzschei</emphasis>
</taxonomicName>
<emphasis id="B951C97B72632077C2A07D9C94BCFD24" bold="true" box="[1154,1244,728,754]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A262740072632077C2A07D9C94BCFD24" box="[1154,1244,728,754]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); females can be distinguished by the smoother spermathecae and the accessory glands that are visible in dorsal view of vulva (
<figureCitation id="131E09EC72632077C34D7DB890D6FCEC" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 38</figureCitation>
; vs. spermathecae are relatively rough and AGs are invisible in dorsal view of vulva in
<taxonomicName id="4C256EEA72632077C2AB7C659579FCEC" authority="Yu &amp; Maddison &amp; Zhang, 2023" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" box="[1161,1305,800,827]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nietzschei" status="sp. nov.">
<emphasis id="B951C97B72632077C2AB7C659579FCEC" box="[1161,1305,800,827]" italics="true" pageId="5" pageNumber="6">O. nietzschei</emphasis>
</taxonomicName>
<emphasis id="B951C97B72632077C3037C64951AFCEC" bold="true" box="[1313,1402,800,826]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A262740072632077C3037C64951AFCEC" box="[1313,1402,800,826]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272632077C6E57C0094DDF934" pageId="5" pageNumber="6" type="description">
<paragraph id="8B9A156972632077C6E57C00921DFBE1" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
<emphasis id="B951C97B72632077C6E57C009139FC88" bold="true" box="[199,345,836,862]" pageId="5" pageNumber="6">Description.</emphasis>
Male. Habitus as shown in
<figureCitation id="131E09EC72632077C4B67C009287FC88" box="[660,743,836,862]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 30</figureCitation>
. Measurements of
<typeStatus id="549EABCB72632077C59E7C01947CFC89" box="[956,1052,837,863]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
(one of KYU-SAL734): carapace length 1.62; abdomen length 1.41; measurements of eyes:AME 0.37, ALE 0.25, PME 0.08, PLE 0.21; measurements of legs: I 3.46 (1.02, 0.63, 0.88, 0.51, 0.42), II 2.44 (0.79, 0.40, 0.50, 0.43, 0.32), III 2.59 (0.88, 0.36, 0.48, 0.52, 0.35), IV 2.72 (0.86, 0.37, 0.56, 0.54, 0.39); leg formula 1432. Carapace dark, widest at front and gradually narrower towards back. Cephalon and dorsal abdomen covered with dense yellow (
<figureCitation id="131E09EC72632077C5E67C909474FC39" box="[964,1044,980,1007]" captionStart="FIGURES 2429" captionStartId="22.[151,264,1437,1461]" captionTargetBox="[332,1255,181,1414]" captionTargetId="figure-22@22.[332,1255,181,1414]" captionTargetPageId="22" captionText="FIGURES 2429. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., living males (2427) and females (2829), from Lambir Hills National Park (24, 27, 29) and Mulu National Park (2526, 28), Sarawak, show variation of coloration. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." figureDoi="http://doi.org/10.5281/zenodo.10376450" httpUri="https://zenodo.org/record/10376450/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
) or pale grey (
<figureCitation id="131E09EC72632077C2967C909564FC38" box="[1204,1284,980,1006]" captionStart="FIGURES 2429" captionStartId="22.[151,264,1437,1461]" captionTargetBox="[332,1255,181,1414]" captionTargetId="figure-22@22.[332,1255,181,1414]" captionTargetPageId="22" captionText="FIGURES 2429. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., living males (2427) and females (2829), from Lambir Hills National Park (24, 27, 29) and Mulu National Park (2526, 28), Sarawak, show variation of coloration. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." figureDoi="http://doi.org/10.5281/zenodo.10376450" httpUri="https://zenodo.org/record/10376450/files/figure.png" pageId="5" pageNumber="6">Fig. 24</figureCitation>
) setae; dorsal abdomen with dark patches, at least on median part of each side; setae near eye lens slightly red; legs I slightly darker than legs IIIV, setae on legs sparse.
</paragraph>
<paragraph id="8B9A156972632077C6E57B0492CDFB3C" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
Palp (
<figureCitation id="131E09EC72632077C72B7B0491EBFB8C" box="[265,395,1088,1114]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Figs 3234</figureCitation>
,
<figureCitation id="131E09EC72632077C7B57B049182FB8C" box="[407,482,1088,1114]" captionStart="FIGURES 4044" captionStartId="24.[151,264,1912,1936]" captionTargetBox="[196,1392,151,1889]" captionTargetId="figure-22@24.[196,1392,151,1889]" captionTargetPageId="24" captionText="FIGURES 4044. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., male palp (40), RTA (41), epigynum (42) and vulva (43 44), in ventral (40, 42), retrolateral (41), dorsal (43) and top (44) view.Abbreviations:AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.10376454" httpUri="https://zenodo.org/record/10376454/files/figure.png" pageId="5" pageNumber="6">4041</figureCitation>
): embolus short and screw-like, embolic disc low, around base of embolus (
<figureCitation id="131E09EC72632077C3187B0495EEFB8D" box="[1338,1422,1088,1115]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 35</figureCitation>
); lobe-like protrusion of tegulum relatively wide, width as ca. 1/2 of width of tegulum, with small mastoid tip well defined from other part; tegular lobe well defined, with prolateral and retrolateral constrictions close to tegulum (
<figureCitation id="131E09EC72632077C6BD7BE89098FB10" box="[159,248,1196,1222]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Figs 32</figureCitation>
,
<figureCitation id="131E09EC72632077C7247BE89146FB10" box="[262,294,1196,1222]" captionStart="FIGURES 4044" captionStartId="24.[151,264,1912,1936]" captionTargetBox="[196,1392,151,1889]" captionTargetId="figure-22@24.[196,1392,151,1889]" captionTargetPageId="24" captionText="FIGURES 4044. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., male palp (40), RTA (41), epigynum (42) and vulva (43 44), in ventral (40, 42), retrolateral (41), dorsal (43) and top (44) view.Abbreviations:AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.10376454" httpUri="https://zenodo.org/record/10376454/files/figure.png" pageId="5" pageNumber="6">40</figureCitation>
); RTA bent medially, distal part thin and lamina-like, forming a revolute structure (
<figureCitation id="131E09EC72632077C2C07BE89557FB10" box="[1250,1335,1196,1222]" captionStart="FIGURES 4044" captionStartId="24.[151,264,1912,1936]" captionTargetBox="[196,1392,151,1889]" captionTargetId="figure-22@24.[196,1392,151,1889]" captionTargetPageId="24" captionText="FIGURES 4044. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., male palp (40), RTA (41), epigynum (42) and vulva (43 44), in ventral (40, 42), retrolateral (41), dorsal (43) and top (44) view.Abbreviations:AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.10376454" httpUri="https://zenodo.org/record/10376454/files/figure.png" pageId="5" pageNumber="6">Fig. 41</figureCitation>
); ventral femoral bump small and tubercle-like (
<figureCitation id="131E09EC72632077C46E7B9492C0FB3C" box="[588,672,1232,1258]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="8B9A156972632077C6E57BB092F9FA49" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
Female. Habitus as shown in
<figureCitation id="131E09EC72632077C40F7BB092E5FAD8" box="[557,645,1268,1294]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 31</figureCitation>
. Measurements of
<typeStatus id="549EABCB72632077C54B7BB193A9FAD9" box="[873,969,1269,1295]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
(KYU-SAL734): carapace length 1.51; abdomen length 1.46; measurements of eyes: AME 0.35, ALE 0.22, PME 0.06, PLE 0.19; measurements of legs: I 2.30 (0.74, 0.42, 0.52, 0.36, 0.26), II 2.04 (0.70, 0.36, 0.42, 0.30, 0.26), III 2.27 (0.76, 0.30, 0.47, 0.43, 0.31), IV 2.59 (0.83, 0.34, 0.55, 0.54, 0.33); leg formula 4132. Body dark, setae on dorsal body sparser than male (
<figureCitation id="131E09EC72632077C3487A249082FA48" captionStart="FIGURES 2429" captionStartId="22.[151,264,1437,1461]" captionTargetBox="[332,1255,181,1414]" captionTargetId="figure-22@22.[332,1255,181,1414]" captionTargetPageId="22" captionText="FIGURES 2429. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., living males (2427) and females (2829), from Lambir Hills National Park (24, 27, 29) and Mulu National Park (2526, 28), Sarawak, show variation of coloration. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." figureDoi="http://doi.org/10.5281/zenodo.10376450" httpUri="https://zenodo.org/record/10376450/files/figure.png" pageId="5" pageNumber="6">Figs 2829</figureCitation>
). Legs I slightly darker than legs IIIV.
</paragraph>
<paragraph id="8B9A156972632077C6E57AEC949AF984" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
Epigynum (
<figureCitation id="131E09EC72632077C76E7AEC91C1FA14" box="[332,417,1448,1474]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Fig. 37</figureCitation>
): atria close to each other, with ratio of distance between atria to diameter of atria about 1: 3 ~ 1: 2; atria far from genital furrow, with ratio of distance between lower edge of atria and genital furrow to diameter of atria about 2: 1. Vulva (
<figureCitation id="131E09EC72632077C4187AB492A0F9DC" box="[570,704,1520,1546]" captionStart="FIGURES 3039" captionStartId="23.[151,264,1883,1907]" captionTargetBox="[225,1362,181,1860]" captionTargetId="figure-17@23.[225,1362,181,1860]" captionTargetPageId="23" captionText="FIGURES 3039. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., paratypes, male from Lambir Hills N. P. (30); male and female from Mulu N. P. (3139). 3031. Habitus; 3234. Palp; 35. Embolic complex; 36. Palpal femur; 37. Epigynum; 3839. Vulva; in dorsal (3031, 38), ventral (32, 37), prolateral (33), retrolateral (34, 36) and top (35, 39) view. Abbreviations: AG, accessory gland; ED, embolic disc; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376452" httpUri="https://zenodo.org/record/10376452/files/figure.png" pageId="5" pageNumber="6">Figs 3839</figureCitation>
): starting part of copulatory ducts wide, with section connecting spermathecae narrowed; accessory glands tubular and pointing sideways, attached on joint of copulatory ducts and spermathecae; spermathecae kidney-shaped, fertilization ducts attached on mound on spermathecae.
</paragraph>
<paragraph id="8B9A156972632077C6E5791894DDF934" blockId="5.[151,1437,260,1798]" pageId="5" pageNumber="6">
<emphasis id="B951C97B72632077C6E57918915FF9A0" bold="true" box="[199,319,1628,1654]" pageId="5" pageNumber="6">Variation.</emphasis>
Males from Lambir Hills Nat. Pk. have dense pale grey setae on dorsal body, whereas these setae are yellow in males from Mulu Nat. Pk.; setae on dorsal body of female are relatively sparse in the population of Lambir Hills Nat. Pk., but obviously dense in the population of Mulu Nat. Pk. (
<figureCitation id="131E09EC72632077C58B79E0944CF968" box="[937,1068,1700,1726]" captionStart="FIGURES 2429" captionStartId="22.[151,264,1437,1461]" captionTargetBox="[332,1255,181,1414]" captionTargetId="figure-22@22.[332,1255,181,1414]" captionTargetPageId="22" captionText="FIGURES 2429. Orcevia bokoblin Yu, Maddison &amp; Zhang, sp. nov., living males (2427) and females (2829), from Lambir Hills National Park (24, 27, 29) and Mulu National Park (2526, 28), Sarawak, show variation of coloration. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." figureDoi="http://doi.org/10.5281/zenodo.10376450" httpUri="https://zenodo.org/record/10376450/files/figure.png" pageId="5" pageNumber="6">Figs 2829</figureCitation>
). These variations are considered intraspecific since no difference in genitalia has been observed between these two populations.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272632077C6E579A89230F8D0" box="[199,592,1772,1798]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8B9A156972632077C6E579A89230F8D0" blockId="5.[151,1437,260,1798]" box="[199,592,1772,1798]" pageId="5" pageNumber="6">
<emphasis id="B951C97B72632077C6E579A89101F8D0" bold="true" box="[199,353,1772,1798]" pageId="5" pageNumber="6">Distribution.</emphasis>
<collectingCountry id="F33255F972632077C74579A891AEF8D0" box="[359,462,1772,1798]" name="Malaysia" pageId="5" pageNumber="6">Malaysia</collectingCountry>
(
<collectingCountry id="F33255F972632077C7FF79A89224F8D0" box="[477,580,1772,1798]" name="Malaysia" pageId="5" pageNumber="6">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>