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<mods:title id="14B92EF30222C6CFB89A9579437421D4">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n.</mods:title>
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Vachon, 1949
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<figureCitation id="ADF483FCFF9FFFC8FBE88083828DFA2D" box="[1077,1192,1487,1510]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–4" captionStart-10="Figure 60" captionStart-11="Figure 61–62" captionStart-12="Figures 63–73" captionStart-13="Figure 74–76" captionStart-14="Figure 77" captionStart-2="Figures 5–8" captionStart-3="Figures 9–18" captionStart-4="Figures 19–38" captionStart-5="Figures 39–46" captionStart-6="Figs" captionStart-7="Figures 52-54" captionStart-8="Figures 55–56" captionStart-9="Figures 57–59" captionStartId-0="3.[192,278,1806,1829]" captionStartId-1="5.[192,278,1745,1768]" captionStartId-10="15.[192,268,924,947]" captionStartId-11="16.[192,268,1757,1780]" captionStartId-12="17.[192,278,1624,1647]" captionStartId-13="18.[192,268,1806,1829]" captionStartId-14="19.[192,268,1203,1226]" captionStartId-2="6.[192,278,1766,1789]" captionStartId-3="7.[192,278,1590,1613]" captionStartId-4="8.[192,278,1710,1733]" captionStartId-5="10.[192,278,1590,1613]" captionStartId-6="11.[192,240,1526,1549]" captionStartId-7="12.[192,278,658,681]" captionStartId-8="13.[192,278,1806,1829]" captionStartId-9="14.[192,278,1710,1733]" captionTargetBox-0="[191,1440,203,1777]" captionTargetBox-1="[191,1440,167,1723]" captionTargetBox-10="[191,1437,191,894]" captionTargetBox-11="[191,1440,215,1722]" captionTargetBox-12="[191,1440,210,1576]" captionTargetBox-13="[191,1440,211,1777]" captionTargetBox-14="[201,1431,152,1168]" captionTargetBox-2="[212,1439,197,1711]" captionTargetBox-3="[191,1437,215,1561]" captionTargetBox-4="[200,1430,225,1668]" captionTargetBox-5="[201,1428,212,1546]" captionTargetBox-6="[200,1422,182,1490]" captionTargetBox-7="[191,1440,191,628]" captionTargetBox-8="[191,1440,214,1777]" captionTargetBox-9="[191,1440,192,1681]" captionTargetId-0="figure-6@3.[192,1439,204,1776]" captionTargetId-1="figure-6@5.[192,1439,168,1723]" captionTargetId-10="figure-6@15.[192,1437,192,893]" captionTargetId-11="figure-12@16.[192,1439,216,1721]" captionTargetId-12="figure-6@17.[192,1439,210,1574]" captionTargetId-13="figure-12@18.[192,1439,212,1776]" captionTargetId-14="figure-6@19.[192,1437,144,1173]" captionTargetId-2="figure-12@6.[192,1439,192,1728]" captionTargetId-3="figure-6@7.[192,1437,216,1560]" captionTargetId-4="figure-12@8.[192,1439,213,1680]" captionTargetId-5="figure-12@10.[192,1439,190,1560]" captionTargetId-6="figure-6@11.[192,1437,168,1497]" captionTargetId-7="figure-12@12.[192,1440,192,628]" captionTargetId-8="figure-6@13.[192,1439,214,1776]" captionTargetId-9="figure-12@14.[192,1439,193,1680]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="15" captionTargetPageId-11="16" captionTargetPageId-12="17" captionTargetPageId-13="18" captionTargetPageId-14="19" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="Figures 1–2: Compsobuthus abyssinicus, in vivo habitus. Male (1) in Ethiopia, locality No. 12EQ and female (2) in Ethiopia, locality No. 12EO." captionText-1="Figures 3–4: Compsobuthus abyssinicus. Female with juveniles after first ecdysis at locality 14EV (3) and the locality 14EV, Ethiopia, Oromia State, volcanic crater Fantale near Metahara, 09°00'56.2&quot;N 39°51'21&quot;E, 1050 m a.s.l. (4)." captionText-10="Figure 60: Compsobuthus eritreaensis sp. n., Locality 15EL, Eritrea, route Massawa to Gahtiela, 15°36'03.7&quot;N 39°16'38.4&quot;E, 115 m a.s.l." captionText-11="Figure 61–62: Compsobuthus somalilandus, male paratype in vivo habitus at type locality (61) and the type locality (62)." captionText-12="Figures 63–73: Compsobuthus werneri, male from Eritrea, locality 15EG, coxosternal area and sternites (63), carapace and tergites I–III (64), pedipalp chela dorsal (65) and external (66), pedipalp movable finger dentition (67), lateral view of telson (68), right legs III (69) and IV (70), retrolateral aspect, metasoma and telson, lateral (71), ventral (72), and dorsal (73) views." captionText-13="Figure 74–76: Compsobuthus werneri, two males in vivo habitus at locality 15EG (74–75) and the locality 15EG, Eritrea, Keren, 15°48'33&quot;N 38°28'14.6&quot;E, 1328 m a.s.l. (76)." captionText-14="Figure 77: Map showing the distribution of Compsobuthus in Eritrea, Ethiopia and Somaliland. Marks indicate sites checked during 2011–2016 expeditions (except for C. abyssinicus from Djibouti which is verified record)." captionText-2="Figures 5–8: Compsobuthus eritreaensis sp. n. Figures 5–6. Male holotype in dorsal (5) and ventral (6) views. Figures 7–8: Female paratype in dorsal (7) and ventral (8) aspects. Scale bar: 10 mm." captionText-3="Figures 9–18: Figures 9–14. Compsobuthus eritreaensis sp. n. Figures 9, 11, 13. Male holotype, carapace and tergites I–III (9), coxosternal area and sternites (11), and lateral view of telson (13). Figures 10, 12, 14. Female paratype, carapace and tergites I–III (10), coxosternal area and sternites (12), and lateral view of telson (14). Figures 15–16. C. abyssinicus, lateral view of telson of male (15) and female (16) from locality 11EW. Figures 17–18. C. somalilandus, lateral view of telson of paratype male (17) and female (18) from type locality." captionText-4="Figures 19–38: Figures 19–22. Compsobuthus abyssinicus, from locality 11EA. Figures 19–20. Female, pedipalp chela dorsal (19) and external (20). Figures 21–22. Male, pedipalp chela dorsal (21) and external (22). Figures 23–26. C. somalilandus, from type locality. Figures 23–24. Female paratype, pedipalp chela dorsal (23) and external (24). Figures 25–26. Male holotype, pedipalp chela dorsal (25) and external (26). Figures 27–38. Compsobuthus eritreaensis sp. n. Figures 27–28. Female paratype from type locality, pedipalp chela dorsal (27) and external (28). Figures 29–36. Male holotype from type locality, pedipalp segments. Chela dorsal (29), external (30), and ventral (31). Patella dorsal (32), external (33) and ventral (34). Femur dorsal (35). Movable finger dentition (36). The trichobothrial pattern is indicated in Figures 30–33 and 35. Figures 37–38. Male holotype, right legs III (37) and IV (38), retrolateral aspect." captionText-5="Figures 39–46: Figures 39–44. Compsobuthus eritreaensis sp. n., metasoma and telson. Figures 39–41. Male holotype, lateral (39), ventral (40), and dorsal (41). Figures 42–44. Female paratype from type locality, lateral (42), ventral (43), and dorsal (44). Scale bar: 10 mm. Figures 45–46. Compsobuthus abyssinicus, from locality 11EA, metasoma and lateral view of telson in male (45) and female (46)." captionText-6="Figs. 47–51: Compsobuthus eritreaensis sp. n., male paratype from type locality, left hemispermatophore. Figure 47. Dorsal view of capsule region, compressed to show form of lobes at base of flagellum. Scale bar: 500 µm. Figures 48–51. Whole hemispermatophore oriented to show capsule region in ventral (48), internal (49), dorsal (50) and external (51) views. Scale bar: 1 mm. Abbreviations: bl, basal lobe; el, external lobe; il, internal lobe; ml, median lobe." captionText-7="Figures 52-54: Compsobuthus eritreaensis sp. n., male paratype from type locality, chromosomes (2n = 22). Arrowheads indicate constrictions on one pair of chromosome. Figure 52. Male mitotic metaphase. Figure 53. Postpachytene. Figure 54. One sister metaphase II. Scale bar: 5 µm." captionText-8="Figures 55–56: Compsobuthus eritreaensis sp. n., female paratype with newborns from type locality (55) and the type locality 15EI, Eritrea, near Massawa, 15°36'58.7&quot;N 39°22'32.8&quot;E, 74 m a.s.l. (56)." captionText-9="Figures 57–59: Figures 57–58. Compsobuthus eritreaensis sp. n., paratypes in vivo habitus at locality 15EJ. Female (57) and male (58). Figure 59. Locality 15EJ, Eritrea, Dese Island, 15°26'39.2&quot;N 39°45'32.7&quot;E." pageId="2" pageNumber="1">Figs. 1–77</figureCitation>
)
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<taxonomicName id="F2CFE4FAFF9FFFC8FCE383458362F9EB" authority="Vachon, 1949: 93" authorityName="Vachon" authorityPageNumber="93" authorityYear="1949" box="[830,1351,1545,1568]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="07BB436BFF9FFFC8FCE3834585F8F9EB" box="[830,989,1545,1568]" italics="true" pageId="2" pageNumber="1">Compsobuthus</emphasis>
Vachon, 1949: 93 (1952: 213)
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; Fet &amp; Lowe, 2000: 124 (complete reference list until 1998); Kovařík, 2003a: 88 (in part); 
<bibRefCitation id="515EE288FF9FFFC8FB20830A85A8F9B7" author="KOVARIK" pageId="2" pageNumber="1" pagination="1 - 12" refId="ref8353" refString="KOVARIK, F., F. STAHLAVSKY, T. KORINKOVA, J. KRAL &amp; T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1 - 12." type="journal article" year="2009">Kovařík, 2009: 31</bibRefCitation>
; Kovařík, 2012: 1; Kovařík &amp; Ojanguren, 2013: 145–158, figs. 777–941.
</paragraph>
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. 
<taxonomicName id="F2CFE4FAFF9FFFC8FC0083F2837EF91E" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" box="[989,1371,1726,1749]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="07BB436BFF9FFFC8FC0083F282E8F91E" box="[989,1229,1726,1749]" italics="true" pageId="2" pageNumber="1">Buthus acutecarinatus</emphasis>
Simon, 1882
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.
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DIAGNOSIS. Total length 
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. Dorsal trichobothria of femur arranged in 
<emphasis id="07BB436BFF9FFFC8FB46825682ECF8FA" box="[1179,1225,1818,1841]" italics="true" pageId="2" pageNumber="1">beta</emphasis>
-configuration. Trichobothrium 
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d 
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of patella internal to dorsomedian carina. Trichobothrium 
<emphasis id="07BB436BFF9FFFC8FBC0821B821DF8A5" box="[1053,1080,1879,1902]" italics="true" pageId="2" pageNumber="1">db</emphasis>
on chela of pedipalp basal to 
<emphasis id="07BB436BFF9FFFC8FAAA821B83B0F8A5" box="[1399,1429,1879,1902]" italics="true" pageId="2" pageNumber="1">est</emphasis>
. Trichobothrium 
<emphasis id="07BB436BFF99FFCEFEA5858B87B4FF15" box="[376,401,199,222]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
located on fixed finger of chela. Pectines with fulcra. Pectinal teeth number 9–34. Tibial spurs present on third and fourth legs. Cheliceral fixed finger with two ventral denticles. Carapace with distinct carinae. Central lateral and posterior lateral carinae of carapace connected to form continuous linear series of granules extending to posterior margin. Carapace in lateral view with entire dorsal surface horizontal or nearly so. Dentate margin of pedipalp chela movable finger with distinct granules divided into 8–14 rows and 4 terminal granules and one basal terminal granule. Tergites I–VI of mesosoma bear three carinae projecting beyond posterior margin as distinct spiniform processes.
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