428 lines
52 KiB
XML
428 lines
52 KiB
XML
<document id="F91EBD63285FE82D155CCAB438557F3E" ID-CLB-Dataset="28504" ID-DOI="10.5281/zenodo.2579565" ID-GBIF-Dataset="2ffd1299-7a53-4ce9-868c-64e82f6fc053" ID-ISSN="1942-1354" ID-Zenodo-Dep="2579565" ID-ZooBank="urn:lsid:zoobank.org:pub:600D70CF-9674-44C6-87FF-DE1B0D2CAA4D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1551440540580" checkinUser="PlaziZenodoSync" docAuthor="Paulsen, M. J." docDate="2019" docId="BC3287C6FFC6FFC1E4AAFA02FD2EFCB3" docLanguage="en" docName="insectaMundi.684.pdf" docOrigin="Insecta Mundi 2019 (684)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ochodaeus meandrus Paulsen 2019, new species" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="400BFFBEFFCFFFCDE46AFF9BFFF4FFF0" masterDocTitle="Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="14" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1551337755633" originalUpdateUser="PlaziZenodoSync" pageNumber="8" updateTime="1698700379637" updateUser="ExternalLinkService">
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<mods:mods id="86F48B9CA02564E55B6091FA8BE93ED4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58C072ECC715B4AF1239535064228840">
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<mods:title id="9437FDF236632CF36FDA192D1EA471FF">Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="BEFD68C16F134BC4E130C3774EF922CB" type="personal">
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<mods:roleTerm id="232F7401F4CC89E47FCFFEC3E49FA06E">Author</mods:roleTerm>
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<mods:namePart id="513A7DD8D7268C828069D79D5E9C080B">Paulsen, M. J.</mods:namePart>
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<mods:affiliation id="00E5B840A52BF4DC26662ADD5B0A5974">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:typeOfResource id="E1F8628BABCD9387242D07A03DC8483E">text</mods:typeOfResource>
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<mods:title id="55F7603755E5E714F22B77CEBB21BE65">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="BB4472BAC5DA84028E0F2E89AE496458">
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<mods:date id="2D9439D389B23E4551FFE5D5E76F561C">2019</mods:date>
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<mods:number id="E19C11AAED5E073F8D49298101FB13AE">2019-01-18</mods:number>
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<mods:number id="47F608605C758D556AF3ABB76A04F542">2019</mods:number>
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<mods:detail id="DB1F79E3777900B2D3A92DF9D94CC574" type="issue">
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<mods:number id="C9EF216B6E04FC5AAF387EC74994005C">684</mods:number>
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<mods:identifier id="82A1E473BEA28C57A8171298E7BC175D" type="DOI">10.5281/zenodo.2579565</mods:identifier>
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<mods:identifier id="0805DBF9C81038D25BEC17B76CB6C05D" type="ISSN">1942-1354</mods:identifier>
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<treatment id="BC3287C6FFC6FFC1E4AAFA02FD2EFCB3" ID-DOI="http://doi.org/10.5281/zenodo.5315610" ID-GBIF-Taxon="154006305" ID-Zenodo-Dep="5315610" LSID="urn:lsid:plazi:treatment:BC3287C6FFC6FFC1E4AAFA02FD2EFCB3" httpUri="http://treatment.plazi.org/id/BC3287C6FFC6FFC1E4AAFA02FD2EFCB3" lastPageId="12" lastPageNumber="11" pageId="9" pageNumber="8">
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<subSubSection id="7C81655BFFC6FFC4E4AAFA02FCFBFA41" box="[192,783,1433,1458]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="342436D0FFC6FFC4E4AAFA02FCFBFA41" blockId="9.[192,783,1433,1458]" box="[192,783,1433,1458]" pageId="9" pageNumber="8">
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<heading id="6F6C81BCFFC6FFC4E4AAFA02FCFBFA41" bold="true" box="[192,783,1433,1458]" fontSize="10" level="4" pageId="9" pageNumber="8" reason="2">
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<emphasis id="06EFEAC2FFC6FFC4E4AAFA02FCFBFA41" bold="true" box="[192,783,1433,1458]" pageId="9" pageNumber="8">
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<taxonomicName id="F39B4D53FFC6FFC4E4AAFA02FDAAFA42" ID-CoL="48DDB" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[192,606,1433,1458]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="meandrus" status="sp. nov.">
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<emphasis id="06EFEAC2FFC6FFC4E4AAFA02FE12FA41" bold="true" box="[192,486,1433,1457]" italics="true" pageId="9" pageNumber="8">Ochodaeus meandrus</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="1DDC57B9FFC6FFC4E606FA01FCFBFA41" box="[620,783,1433,1458]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7C81655BFFC6FFC7E4AAFA51FCCDFCC4" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="materials_examined">
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<paragraph id="342436D0FFC6FFC4E4AAFA51FCBCF9B3" blockId="9.[192,1440,1482,1929]" pageId="9" pageNumber="8">
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<emphasis id="06EFEAC2FFC6FFC4E4AAFA51FE7EFA12" bold="true" box="[192,394,1482,1506]" pageId="9" pageNumber="8">Type material.</emphasis>
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<materialsCitation id="84F33C8DFFC6FFC4E5FBFA51FCBCF9B3" ID-GBIF-Occurrence="2005648082" collectingDate="2002-04-21" collectorName="M. Irwin" country="Madagascar" county="Province" elevation="1110" location="Ranomafana" longitude="47.369835" municipality="Parc National" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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<typeStatus id="EB208872FFC6FFC4E5FBFA51FE0BFA12" box="[401,511,1482,1506]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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male (CASC;
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<figureCitation id="ACA02A55FFC6FFC4E6C4FA51FD00FA12" box="[686,756,1482,1506]" captionStart="Figures 2–3" captionStartId="8.[192,287,1848,1870]" captionTargetBox="[334,1282,213,1825]" captionTargetId="figure@8.[320,1311,199,1833]" captionTargetPageId="8" captionText="Figures 2–3. Madagascar Ochodaeus species. 2) Ochodaeus polypollicatus Paulsen, new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view, arrow indicating second pollex. c) Labels. 3) Ochodaeus meandrus Paulsen new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view. c) Labels." httpUri="https://zenodo.org/record/2579569/files/figure.png" pageId="9" pageNumber="8">Fig. 3</figureCitation>
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), labeled: a) “
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<collectingCountry id="4C8C7640FFC6FFC4E7F6FA51FB93FA12" box="[924,1127,1482,1506]" name="Madagascar" pageId="9" pageNumber="8">MADAGASCAR</collectingCountry>
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:
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<collectingCounty id="DD454E5CFFC6FFC4E018FA51FB29FA12" box="[1138,1245,1482,1506]" pageId="9" pageNumber="8">Province</collectingCounty>
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/
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<collectingRegion id="F65FF832FFC6FFC4E099FA51FA6FFA12" box="[1267,1435,1482,1506]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="8">Fianarantsoa</collectingRegion>
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,
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<collectingMunicipality id="D440ACAAFFC6FFC4E4AAFA70FE9FF9F3" box="[192,363,1515,1539]" pageId="9" pageNumber="8">Parc National</collectingMunicipality>
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/
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<location id="3144600BFFC6FFC4E515FA70FDEAF9F3" LSID="urn:lsid:plazi:treatment:BC3287C6FFC6FFC1E4AAFA02FD2EFCB3:3144600BFFC6FFC4E515FA70FDEAF9F3" box="[383,542,1515,1539]" country="Madagascar" county="Province" longitude="47.369835" municipality="Parc National" name="Ranomafana" pageId="9" pageNumber="8" stateProvince="Fianarantsoa">Ranomafana</location>
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,
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<location id="3144600BFFC6FFC4E640FA70FD4DF9F3" LSID="urn:lsid:plazi:treatment:BC3287C6FFC6FFC1E4AAFA02FD2EFCB3:3144600BFFC6FFC4E640FA70FD4DF9F3" box="[554,697,1515,1539]" country="Madagascar" county="Province" longitude="47.369835" municipality="Parc National" name="Vohiparara" pageId="9" pageNumber="8" stateProvince="Fianarantsoa">Vohiparara</location>
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, / at broken bridge, el
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<elevation id="BFB6D1E3FFC6FFC4E7AFFA70FBEBF9F2" box="[965,1055,1515,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="m" value="1110.0">
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<quantity id="F3639B35FFC6FFC4E7AFFA70FBEBF9F2" box="[965,1055,1515,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="m" value="1110.0">1110 m</quantity>
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</elevation>
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/ 8–15
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<collectingDate id="5061E9F8FFC6FFC4E01FFA70FADDF9F3" box="[1141,1321,1515,1539]" pageId="9" pageNumber="8" value="2002-04-21">
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<date id="40251010FFC6FFC4E01FFA70FADDF9F3" box="[1141,1321,1515,1539]" pageId="9" pageNumber="8" value="2002-04-21">April 2002 / 21</date>
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</collectingDate>
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° 13.57′ S,
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<geoCoordinate id="51AF5017FFC6FFC4E4AAF990FEBAF9D3" box="[192,334,1546,1571]" direction="east" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="47.369835">47° 22.19′ E</geoCoordinate>
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” b) “coll:
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<collectorName id="996E5306FFC6FFC4E5D1F997FDD3F9D3" box="[443,551,1547,1571]" pageId="9" pageNumber="8">M. Irwin</collectorName>
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, R. Harin’Hala /
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<location id="3144600BFFC6FFC4E684F990FBC0F9D3" LSID="urn:lsid:plazi:treatment:BC3287C6FFC6FFC1E4AAFA02FD2EFCB3:3144600BFFC6FFC4E684F990FBC0F9D3" box="[750,1076,1547,1571]" country="Madagascar" county="Province" longitude="47.369835" municipality="Parc National" name="California Acad of Sciences" pageId="9" pageNumber="8" stateProvince="Fianarantsoa">California Acad of Sciences</location>
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/
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<collectingMethod id="EDDA4EC7FFC6FFC4E02DF990FB2BF9D3" box="[1095,1247,1547,1571]" pageId="9" pageNumber="8">malaise trap</collectingMethod>
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in high altitude / rainforest, MA-02-09A-24”; c) “CASENT / 8055264”.
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</materialsCitation>
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</paragraph>
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<paragraph id="342436D0FFC6FFC7E482F9D7FCCDFCC4" blockId="9.[192,1440,1482,1929]" lastBlockId="10.[192,1441,217,884]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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One male, two female
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<typeStatus id="EB208872FFC6FFC4E591F9D6FD81F995" box="[507,629,1613,1637]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
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labeled: a, b as
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<typeStatus id="EB208872FFC6FFC4E756F9D7FC53F994" box="[828,935,1612,1636]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
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; c) “CASENT / #” (# = 8055260, 8055262, 8055300). One female
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<typeStatus id="EB208872FFC6FFC4E5B9F9F6FDB4F975" box="[467,576,1645,1669]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
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labeled: a) “
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<collectingCountry id="4C8C7640FFC6FFC4E6B3F9F6FC50F975" box="[729,932,1645,1669]" name="Madagascar" pageId="9" pageNumber="8">MADAGASCAR</collectingCountry>
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: Province /
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<collectingRegion id="F65FF832FFC6FFC4E05BF9F6FB2DF975" box="[1073,1241,1645,1669]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="8">Fianarantsoa</collectingRegion>
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, Parc National / Ranomafana, Vohiparara, / at broken bridge, el
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<quantity id="F3639B35FFC6FFC4E690F916FCA7F955" box="[762,851,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="m" value="1110.0">1110 m</quantity>
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/ 6–15
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<date id="40251010FFC6FFC4E7CEF916FB7BF955" box="[932,1167,1677,1701]" pageId="9" pageNumber="8" value="2001-12-21">December 2001 / 21</date>
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° 13.5 7′ S,
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<geoCoordinate id="51AF5017FFC6FFC4E162F916FA61F955" box="[1288,1429,1676,1701]" direction="east" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="47.369835">47° 22.19′ E</geoCoordinate>
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” b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise trap in high altitude / rainforest, MA-02-09A-06”; c) “CASENT / 8055632”. One female
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<typeStatus id="EB208872FFC6FFC4E73CF954FC37F917" box="[854,963,1743,1767]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
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labeled: a) “
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<collectingCountry id="4C8C7640FFC6FFC4E034F955FADDF916" box="[1118,1321,1742,1766]" name="Madagascar" pageId="9" pageNumber="8">MADAGASCAR</collectingCountry>
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: Province /
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<collectingRegion id="F65FF832FFC6FFC4E4A5F975FE83F8F6" box="[207,375,1774,1798]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="8">Fianarantsoa</collectingRegion>
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, Parc National / Ranomafana, Vohiparara, / at broken bridge, el
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<quantity id="F3639B35FFC6FFC4E0E5F974FB1DF8F6" box="[1167,1257,1775,1799]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="m" value="1110.0">1110 m</quantity>
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/ 19–26
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<date id="40251010FFC6FFC4E13AF975FED9F8D7" pageId="9" pageNumber="8" value="2001-03-21">
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March 2001 /
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||
<geoCoordinate id="51AF5017FFC6FFC4E565F894FED9F8D7" box="[271,301,1807,1831]" direction="south" orientation="latitude" pageId="9" pageNumber="8" precision="9" value="-21.226166">21</geoCoordinate>
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||
</date>
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° 13.57′ S,
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<geoCoordinate id="51AF5017FFC6FFC4E5C0F894FDCEF8D7" box="[426,570,1806,1831]" direction="east" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="47.369835">47° 22.19′ E</geoCoordinate>
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” b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise trap in high altitude / rainforest, MA-02-09A-21”; c) “CASENT / 8055999”. Two male
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||
<typeStatus id="EB208872FFC6FFC4E0C2F8ABFAD4F8B8" box="[1192,1312,1840,1864]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
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labeled: a) “Two male
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<typeStatus id="EB208872FFC6FFC4E529F8CBFE4FF898" box="[323,443,1872,1896]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
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labeled: a) “
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<collectingCountry id="4C8C7640FFC6FFC4E620F8CBFCE7F898" box="[586,787,1872,1896]" name="Madagascar" pageId="9" pageNumber="8">MADAGASCAR</collectingCountry>
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: Province /
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<collectingRegion id="F65FF832FFC6FFC4E7FCF8CBFBCFF898" box="[918,1083,1872,1896]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="8">Fianarantsoa</collectingRegion>
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, Parc National / Ranomafana, Vohiparara, / at broken bridge, el
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<quantity id="F3639B35FFC6FFC4E627F8EAFD50F878" box="[589,676,1905,1929]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="m" value="1110.0">1110 m</quantity>
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/ 19–26
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<date id="40251010FFC6FFC4E76EF8EBFC36F879" box="[772,962,1904,1929]" pageId="9" pageNumber="8" value="2001-03-21">
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||
March 2001 /
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<geoCoordinate id="51AF5017FFC6FFC4E7CEF8EAFC36F879" box="[932,962,1905,1929]" direction="south" orientation="latitude" pageId="9" pageNumber="8" precision="9" value="-21.226166">21</geoCoordinate>
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</date>
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° 13.57′ S,
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<geoCoordinate id="51AF5017FFC6FFC4E050F8EAFB30F879" box="[1082,1220,1903,1929]" direction="east" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="47.369835">47° 22.19′ E</geoCoordinate>
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” b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise trap in high altitude / rainforest, MA-02-09A-22”; c) “CASENT / 8055999”. Two male, two female
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||
<typeStatus id="EB208872FFC5FFC7E696FF61FC82FEE2" box="[764,886,250,274]" pageId="10" pageNumber="9" type="paratype">paratypes</typeStatus>
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||
labeled: a) “
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<collectingCountry id="4C8C7640FFC5FFC7E060FF61FB21FEE2" box="[1034,1237,250,274]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
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: Province / Fian- arantsoa, Parc National /Ranomafana, radio tower / at forest edge, elev
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<quantity id="F3639B35FFC5FFC7E041FE81FB71FEC1" box="[1067,1157,282,306]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="10" pageNumber="9" unit="m" value="1130.0">1130 m</quantity>
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/ 19–26
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||
<date id="40251010FFC5FFC7E081FE81FF1AFEA2" pageId="10" pageNumber="9" value="2002-02-21">
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February 2002 /
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<geoCoordinate id="51AF5017FFC5FFC7E4BAFEA1FF1AFEA2" box="[208,238,314,338]" direction="south" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="-21.250834">21</geoCoordinate>
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||
</date>
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° 15.05′ S,
|
||
<geoCoordinate id="51AF5017FFC5FFC7E51EFEA1FDFEFEA2" box="[372,522,312,338]" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="47.407166">47° 24.43′ E</geoCoordinate>
|
||
”; b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise, mixed tropical / forest MA-02-09B-17”; c) “CASENT / #” (# = 8055520, 8055521, 8055522, 8055544). One male
|
||
<typeStatus id="EB208872FFC5FFC7E56BFEE0FE9AFE63" box="[257,366,379,403]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
labeled: a) “
|
||
<collectingCountry id="4C8C7640FFC5FFC7E594FEE1FD3CFE62" box="[510,712,378,402]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
: Province /
|
||
<collectingRegion id="F65FF832FFC5FFC7E727FEE1FC00FE62" box="[845,1012,378,402]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="9">Fianarantsoa</collectingRegion>
|
||
, Parc National / Ranomafana, radio tower / at forest edge, elev
|
||
<quantity id="F3639B35FFC5FFC7E660FE01FD90FE42" box="[522,612,410,434]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="10" pageNumber="9" unit="m" value="1130.0">1130 m</quantity>
|
||
/ 20 Sept–5
|
||
<date id="40251010FFC5FFC7E696FE01FC6AFE42" box="[764,926,410,434]" pageId="10" pageNumber="9" value="2006-10-21">
|
||
Oct 2006 /
|
||
<geoCoordinate id="51AF5017FFC5FFC7E7EAFE01FC6AFE42" box="[896,926,410,434]" direction="south" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="-21.250834">21</geoCoordinate>
|
||
</date>
|
||
° 15.05′ S,
|
||
<geoCoordinate id="51AF5017FFC5FFC7E075FE01FB46FE42" box="[1055,1202,409,434]" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="47.407166">47° 24.43′ E</geoCoordinate>
|
||
”; b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise, mixed tropical / forest MA-02-09B-160”. One female
|
||
<typeStatus id="EB208872FFC5FFC7E571FE40FE7CFE03" box="[283,392,475,499]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
labeled: a) “
|
||
<collectingCountry id="4C8C7640FFC5FFC7E64FFE40FD04FE03" box="[549,752,475,499]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
: Province /
|
||
<collectingRegion id="F65FF832FFC5FFC7E7EAFE40FBDCFE03" box="[896,1064,475,499]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="9">Fianarantsoa</collectingRegion>
|
||
, Parc National / Ranomafana, radio tower / at forest edge, elev
|
||
<quantity id="F3639B35FFC5FFC7E624FE60FD5CFDE2" box="[590,680,507,531]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="10" pageNumber="9" unit="m" value="1130.0">1130 m</quantity>
|
||
/ 1–11
|
||
<date id="40251010FFC5FFC7E76AFE60FC00FDE3" box="[768,1012,507,531]" pageId="10" pageNumber="9" value="2006-11-21">
|
||
November 2006 /
|
||
<geoCoordinate id="51AF5017FFC5FFC7E7BCFE60FC00FDE3" box="[982,1012,507,531]" direction="south" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="-21.250834">21</geoCoordinate>
|
||
</date>
|
||
° 15.05′ S,
|
||
<geoCoordinate id="51AF5017FFC5FFC7E01EFE60FAF2FDE3" box="[1140,1286,505,531]" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="47.407166">47° 24.43′ E</geoCoordinate>
|
||
”; b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise, mixed tropical / forest MA-02-09B-163”. One male
|
||
<typeStatus id="EB208872FFC5FFC7E553FDA0FE52FDA3" box="[313,422,571,595]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
labeled: a) “
|
||
<collectingCountry id="4C8C7640FFC5FFC7E65DFDA0FCF6FDA3" box="[567,770,571,595]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
: Province /
|
||
<collectingRegion id="F65FF832FFC5FFC7E7E2FDA0FBC4FDA3" box="[904,1072,571,595]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="9">Fianarantsoa</collectingRegion>
|
||
, Ranomafana / JIRAMA water works / 21–24
|
||
<date id="40251010FFC5FFC7E51AFDC0FD94FD83" box="[368,608,603,627]" pageId="10" pageNumber="9" value="2001-12-21">
|
||
December 2001 /
|
||
<geoCoordinate id="51AF5017FFC5FFC7E628FDC0FD94FD83" box="[578,608,603,627]" direction="south" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="-21.2485">21</geoCoordinate>
|
||
</date>
|
||
° 14.91′ S,
|
||
<geoCoordinate id="51AF5017FFC5FFC7E6B5FDC0FC84FD83" box="[735,880,602,627]" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="47.452168">47° 27.13′ E</geoCoordinate>
|
||
”; b) “coll: M. Irwin, R. Harin’Hala / California Acad of Sciences / malaise trap near river / elev
|
||
<quantity id="F3639B35FFC5FFC7E779FDE0FC95FD63" box="[787,865,635,659]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="9" unit="m" value="690.0">690 m</quantity>
|
||
, MA-02-09D-08”; c) “CASENT / 8056011”. One female
|
||
<typeStatus id="EB208872FFC5FFC7E572FD07FE71FD44" box="[280,389,668,692]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
labeled: a) “
|
||
<collectingCountry id="4C8C7640FFC5FFC7E673FD07FD10FD44" box="[537,740,668,692]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
: Province /
|
||
<collectingRegion id="F65FF832FFC5FFC7E701FD00FBE7FD43" box="[875,1043,667,691]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="9">Fianarantsoa</collectingRegion>
|
||
, SE of Fandriana, / Ranomena,
|
||
<date id="40251010FFC5FFC7E1FBFD00FEE6FD23" pageId="10" pageNumber="9">5 March</date>
|
||
, 2000 /
|
||
<geoCoordinate id="51AF5017FFC5FFC7E500FD27FDF2FD24" box="[362,518,698,724]" direction="south" orientation="latitude" pageId="10" pageNumber="9" precision="15" value="-20.393612">20° 23′ 37″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="51AF5017FFC5FFC7E67BFD27FD5BFD24" box="[529,687,698,724]" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="15" value="47.6525">47° 39′ 09″ E</geoCoordinate>
|
||
”; b) “M.E. Irwin & E. I. Schlinger / California Acad. of Sciences / el.
|
||
<quantity id="F3639B35FFC5FFC7E49FFD47FEA5FD03" box="[245,337,732,756]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="10" pageNumber="9" unit="m" value="1345.0">1345 m</quantity>
|
||
coll. code: SI-010”; c) “CASENT / 8003859”. One
|
||
<typeStatus id="EB208872FFC5FFC7E7C5FD47FBE8FD04" box="[943,1052,732,756]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
(NHM; on glueboard, unsexed) labeled: a) “
|
||
<collectingCountry id="4C8C7640FFC5FFC7E538FD67FDE8FCE4" box="[338,540,764,788]" name="Madagascar" pageId="10" pageNumber="9">MADAGASCAR</collectingCountry>
|
||
, 2007 / Ranomafana Nat. Park / near Ranomafana vill. / M. Trýzna leg.,
|
||
<date id="40251010FFC5FFC7E4AAFC87FEEFFCC4" box="[192,283,796,820]" pageId="10" pageNumber="9">26.-31.i</date>
|
||
(sic)”; b) “BMNH (E) /
|
||
<date id="40251010FFC5FFC7E65AFC87FD66FCC4" box="[560,658,796,820]" pageId="10" pageNumber="9" value="2016" valueMax="2044">2016-44</date>
|
||
/ M. Trýzna”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C81655BFFC5FFC7E482FCA7FE1AFC84" pageId="10" pageNumber="9" type="discussion">
|
||
<paragraph id="342436D0FFC5FFC7E482FCA7FE1AFC84" blockId="10.[192,1441,217,884]" pageId="10" pageNumber="9">
|
||
All paratypes (
|
||
<emphasis id="06EFEAC2FFC5FFC7E5F6FCA7FE58FCA3" box="[412,428,828,851]" italics="true" pageId="10" pageNumber="9">n</emphasis>
|
||
= 6 male, 9 female, 1 unsexed) with label: on yellow paper, “
|
||
<taxonomicName id="F39B4D53FFC5FFC7E0E1FCA7FAE4FCA4" authorityName="Dejean" authorityYear="1821" box="[1163,1296,828,852]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06EFEAC2FFC5FFC7E0E1FCA7FAE4FCA4" box="[1163,1296,828,852]" italics="true" pageId="10" pageNumber="9">Ochodaeus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="F39B4D53FFC5FFC7E14FFCA7FA54FCA4" authorityName="Paulsen" authorityYear="2019" box="[1317,1440,828,852]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="meandrus">
|
||
<emphasis id="06EFEAC2FFC5FFC7E14FFCA7FA54FCA4" box="[1317,1440,828,852]" italics="true" pageId="10" pageNumber="9">meandrus</emphasis>
|
||
</taxonomicName>
|
||
/ Paulsen / PARATYPE”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C81655BFFC5FFC7E4AAFC17FDE5F948" pageId="10" pageNumber="9" type="description">
|
||
<paragraph id="342436D0FFC5FFC7E4AAFC17FDE1F9D7" blockId="10.[192,1440,908,1575]" pageId="10" pageNumber="9">
|
||
<emphasis id="06EFEAC2FFC5FFC7E4AAFC17FE9FFC54" bold="true" box="[192,363,908,932]" pageId="10" pageNumber="9">Description.</emphasis>
|
||
Holotype male (
|
||
<figureCitation id="ACA02A55FFC5FFC7E62BFC17FD6FFC54" box="[577,667,908,932]" captionStart="Figures 2–3" captionStartId="8.[192,287,1848,1870]" captionTargetBox="[334,1282,213,1825]" captionTargetId="figure@8.[320,1311,199,1833]" captionTargetPageId="8" captionText="Figures 2–3. Madagascar Ochodaeus species. 2) Ochodaeus polypollicatus Paulsen, new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view, arrow indicating second pollex. c) Labels. 3) Ochodaeus meandrus Paulsen new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view. c) Labels." httpUri="https://zenodo.org/record/2579569/files/figure.png" pageId="10" pageNumber="9">Fig. 3a</figureCitation>
|
||
).
|
||
<taxonomicName id="F39B4D53FFC5FFC7E6DFFC17FCCBFC54" authorityName="" authorityYear="1758" box="[693,831,908,932]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
: Scarabaeoidea:
|
||
<taxonomicName id="F39B4D53FFC5FFC7E079FC17FB41FC54" authorityName="Mulsant & Rey" authorityYear="1871" box="[1043,1205,908,932]" class="Insecta" family="Ochodaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="family">Ochodaeidae</taxonomicName>
|
||
.
|
||
<emphasis id="06EFEAC2FFC5FFC7E0A9FC17FADAFC54" bold="true" box="[1219,1326,908,932]" pageId="10" pageNumber="9">Length:</emphasis>
|
||
7.4 mm.
|
||
<emphasis id="06EFEAC2FFC5FFC7E4AAFC37FEEEFC34" bold="true" box="[192,282,940,964]" pageId="10" pageNumber="9">Width:</emphasis>
|
||
3.9 mm.
|
||
<emphasis id="06EFEAC2FFC5FFC7E5E1FC37FE2BFC34" bold="true" box="[395,479,940,964]" pageId="10" pageNumber="9">Color:</emphasis>
|
||
Everywhere uniformly dark reddish brown; surface nitid.
|
||
<emphasis id="06EFEAC2FFC5FFC7E0C3FC37FB0DFC34" bold="true" box="[1193,1273,940,964]" pageId="10" pageNumber="9">Head:</emphasis>
|
||
Surface lacking tubercles, irregularly punctate; punctures mixed, moderately large and setose, separated by 1–2 diameters. Eyes large, globose; lacking ventral projection of canthus. Antennal club large, subequal to eye in size. Mandibles broadly scooped, shortened (shorter than head), apices falcate; left mandible with large triangular, subacute internal tooth; right mandible with complimenting tooth obsolete, second tooth larger, subacute. Frons weakly convex; surface punctate behind frontoclypeal suture. Clypeus flat with disc relatively long (1/4 as long medially as wide), subtrapezoidal; anterior margin beaded, bead roughly circular but wavy (
|
||
<figureCitation id="ACA02A55FFC5FFC7E63AFB16FD53FB56" box="[592,679,1165,1190]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="10" pageNumber="9">Fig. 18</figureCitation>
|
||
). Mentum flat emarginated by round fovea in apical half (
|
||
<figureCitation id="ACA02A55FFC5FFC7E11AFB16FF25FB35" captionStart="Figures 4–12" captionStartId="11.[192,285,1819,1841]" captionTargetBox="[192,1440,213,1793]" captionTargetId="figure@11.[192,1440,213,1793]" captionTargetPageId="11" captionText="Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="10" pageNumber="9">Fig. 6</figureCitation>
|
||
).
|
||
<emphasis id="06EFEAC2FFC5FFC7E48FFB35FE8CFB36" bold="true" box="[229,376,1198,1222]" pageId="10" pageNumber="9">Pronotum:</emphasis>
|
||
Form convex. Surface densely punctate, appearing smooth between punctures laterally, disc with slight indication of tiled granules; punctation mixed, predominantly large setose punctures with few smaller glabrous punctures interspersed.
|
||
<emphasis id="06EFEAC2FFC5FFC7E729FB75FC56FAF6" bold="true" box="[835,930,1262,1286]" pageId="10" pageNumber="9">Elytra:</emphasis>
|
||
Form convex, relatively elongate, elytra together longer than wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures large, separated by 2 diameters, lacking setae. Intervals irregularly punctate; punctures small, setose, with tubercle anteriorly.
|
||
<emphasis id="06EFEAC2FFC5FFC7E133FAD5FA54FA96" bold="true" box="[1369,1440,1358,1382]" pageId="10" pageNumber="9">Legs:</emphasis>
|
||
Protibia tridentate externally, internal side with short, ventrally directed pollex <1/3 length of apical spur (
|
||
<figureCitation id="ACA02A55FFC5FFC7E563FA15FE94FA56" box="[265,352,1422,1446]" captionStart="Figures 2–3" captionStartId="8.[192,287,1848,1870]" captionTargetBox="[334,1282,213,1825]" captionTargetId="figure@8.[320,1311,199,1833]" captionTargetPageId="8" captionText="Figures 2–3. Madagascar Ochodaeus species. 2) Ochodaeus polypollicatus Paulsen, new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view, arrow indicating second pollex. c) Labels. 3) Ochodaeus meandrus Paulsen new species, male holotype. a) Dorsal habitus. b) Right front leg, dorsal view. c) Labels." httpUri="https://zenodo.org/record/2579569/files/figure.png" pageId="10" pageNumber="9">Fig. 3b</figureCitation>
|
||
). Profemur with strong apical tooth. Mesofemur and metafemur with isolated subapical tooth not strongly connected to apex.
|
||
<emphasis id="06EFEAC2FFC5FFC7E61FFA35FCABFA36" bold="true" box="[629,863,1454,1478]" pageId="10" pageNumber="9">Venter/Abdomen:</emphasis>
|
||
Metasternum and abdominal segments sparsely punctate; punctures large with long, erect, golden setae. Pygidium roughly punctate. Stridulatory peg short, capitate (
|
||
<figureCitation id="ACA02A55FFC5FFC7E514FA74FE20F9F7" box="[382,468,1519,1543]" captionStart="Figures 4–12" captionStartId="11.[192,285,1819,1841]" captionTargetBox="[192,1440,213,1793]" captionTargetId="figure@11.[192,1440,213,1793]" captionTargetPageId="11" captionText="Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="10" pageNumber="9">Fig. 12</figureCitation>
|
||
).
|
||
<emphasis id="06EFEAC2FFC5FFC7E58DFA74FD46F9F7" bold="true" box="[487,690,1519,1543]" pageId="10" pageNumber="9">Male genitalia:</emphasis>
|
||
Internal sac with narrow arrowhead-shaped patch of sclerites between parameres (
|
||
<figureCitation id="ACA02A55FFC5FFC7E5AAF994FDFCF9D7" box="[448,520,1551,1575]" captionStart="Figures 4–12" captionStartId="11.[192,285,1819,1841]" captionTargetBox="[192,1440,213,1793]" captionTargetId="figure@11.[192,1440,213,1793]" captionTargetPageId="11" captionText="Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="10" pageNumber="9">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="342436D0FFC5FFC7E4AAF9A4FDE5F948" blockId="10.[192,1440,1599,1720]" pageId="10" pageNumber="9">
|
||
<emphasis id="06EFEAC2FFC5FFC7E4AAF9A4FE3FF9A7" bold="true" box="[192,459,1599,1623]" pageId="10" pageNumber="9">Paratype variation.</emphasis>
|
||
Sexual dimorphism is not readily apparent in this species. The acute, apical tooth on the profemur is not present in females or smaller males, and the subapical tooth on the mesofemur is also obsolete on most female specimens. The subapical tooth on the metafemur may be reduced but is present on all specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C81655BFFC5FFC1E4AAF94BFDF7FEC2" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="9" type="discussion">
|
||
<paragraph id="342436D0FFC5FFC1E4AAF94BFDF7FEC2" blockId="10.[192,1441,1743,1928]" lastBlockId="12.[192,1440,217,434]" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="9">
|
||
<emphasis id="06EFEAC2FFC5FFC7E4AAF94BFEB7F918" bold="true" box="[192,323,1744,1768]" pageId="10" pageNumber="9">Remarks.</emphasis>
|
||
This species may be confused with smaller males and females of
|
||
<emphasis id="06EFEAC2FFC5FFC7E0E4F94BFA92F917" box="[1166,1382,1743,1767]" italics="true" pageId="10" pageNumber="9">
|
||
<taxonomicName id="F39B4D53FFC5FFC7E0E4F94BFA96F917" authorityName="Paulsen" authorityYear="2019" box="[1166,1378,1743,1767]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="polypollicatus">O. polypollicatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The much shorter, ventrally-directed pollex, less furrowed mentum, and isolated subapical tooth on the metafemur are all consistent differences and diagnostic. Males of this species lack the setal patch on the last abdominal segment, and second pollex of most male
|
||
<emphasis id="06EFEAC2FFC5FFC7E7DAF8ABFB72F8B8" box="[944,1158,1840,1864]" italics="true" pageId="10" pageNumber="9">
|
||
<taxonomicName id="F39B4D53FFC5FFC7E7DAF8ABFB76F8B8" authorityName="Paulsen" authorityYear="2019" box="[944,1154,1840,1864]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="polypollicatus">O. polypollicatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The best character for specimens of both sexes of
|
||
<taxonomicName id="F39B4D53FFC5FFC7E662F8CBFD52F898" authorityName="Paulsen" authorityYear="2019" box="[520,678,1872,1896]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="meandrus">
|
||
<emphasis id="06EFEAC2FFC5FFC7E662F8CBFD52F898" box="[520,678,1872,1896]" italics="true" pageId="10" pageNumber="9">O. meandrus</emphasis>
|
||
</taxonomicName>
|
||
is the wavy appearance of the clypeal bead when viewed from directly above. Positioning the specimen in the correct plane so that the light reflects on the crest of the bead is necessary to best see the wavy form. The clypeal bead of
|
||
<taxonomicName id="F39B4D53FFC3FFC1E780FF41FB3FFF01" authorityName="Paulsen" authorityYear="2019" box="[1002,1227,217,241]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="modopunctatus">
|
||
<emphasis id="06EFEAC2FFC3FFC1E780FF41FB3FFF01" box="[1002,1227,217,241]" italics="true" pageId="12" pageNumber="11">O. modopunctatus</emphasis>
|
||
</taxonomicName>
|
||
may also appear wavy, but that species has no indication of granules on the pronotum and a mentum with a furrow that becomes acute posteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="60E46658FFC4FFC6E4AAF880FC6EF878" httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="11" pageNumber="10" startId="11.[192,285,1819,1841]" targetBox="[192,1440,213,1793]" targetPageId="11">
|
||
<paragraph id="342436D0FFC4FFC6E4AAF880FC6EF878" blockId="11.[192,1440,1819,1928]" pageId="11" pageNumber="10">
|
||
<emphasis id="06EFEAC2FFC4FFC6E4AAF880FEAAF8C0" bold="true" box="[192,350,1819,1841]" pageId="11" pageNumber="10">Figures 4–12.</emphasis>
|
||
Additional characters of Madagascar
|
||
<taxonomicName id="F39B4D53FFC4FFC6E69AF880FC92F8C0" authorityName="Dejean" authorityYear="1821" box="[752,870,1819,1840]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06EFEAC2FFC4FFC6E69AF880FC92F8C0" box="[752,870,1819,1840]" italics="true" pageId="11" pageNumber="10">Ochodaeus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="06EFEAC2FFC4FFC6E7ABF880FC06F8C1" bold="true" box="[961,1010,1819,1841]" pageId="11" pageNumber="10">4–6)</emphasis>
|
||
Mentum, shading indicates foveae.
|
||
<emphasis id="06EFEAC2FFC4FFC6E105F887FA54F8C1" bold="true" box="[1391,1440,1819,1841]" pageId="11" pageNumber="10">7–9)</emphasis>
|
||
Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres.
|
||
<emphasis id="06EFEAC2FFC4FFC6E139F8A3FA54F8BE" bold="true" box="[1363,1440,1848,1870]" pageId="11" pageNumber="10">10–12)</emphasis>
|
||
Right stridulatory peg, dorsal view.
|
||
<emphasis id="06EFEAC2FFC4FFC6E639F8CEFD44F89B" bold="true" box="[595,688,1877,1899]" pageId="11" pageNumber="10">4, 7, 10)</emphasis>
|
||
<taxonomicName id="F39B4D53FFC4FFC6E6D2F8CEFC10F89B" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[696,996,1877,1899]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="modopunctatus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC4FFC6E6D2F8CEFC75F89A" box="[696,897,1877,1898]" italics="true" pageId="11" pageNumber="10">
|
||
O.
|
||
<taxonomicName id="F39B4D53FFC4FFC6E6B2F8CEFC75F89A" box="[728,897,1877,1898]" pageId="11" pageNumber="10" rank="species" species="modopunctatus">modopunctatus</taxonomicName>
|
||
</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC4FFC6E79AF8CDFB82F89B" box="[1008,1142,1877,1899]" pageId="11" pageNumber="10" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
<emphasis id="06EFEAC2FFC4FFC6E0EBF8CEFB2AF89B" bold="true" box="[1153,1246,1877,1899]" pageId="11" pageNumber="10">5, 8, 11)</emphasis>
|
||
<taxonomicName id="F39B4D53FFC4FFC6E08CF8CEFEEFF878" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="polypollicatus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC4FFC6E08CF8CEFA54F89A" box="[1254,1440,1877,1898]" italics="true" pageId="11" pageNumber="10">
|
||
O.
|
||
<taxonomicName id="F39B4D53FFC4FFC6E16DF8CEFA54F89A" box="[1287,1440,1877,1898]" pageId="11" pageNumber="10" rank="species" species="polypollicatus">polypollicatus</taxonomicName>
|
||
</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC4FFC6E54CF8E9FE5FF878" box="[294,427,1906,1928]" pageId="11" pageNumber="10" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
<emphasis id="06EFEAC2FFC4FFC6E5DFF8E9FDFBF878" bold="true" box="[437,527,1906,1928]" pageId="11" pageNumber="10">6, 9, 12)</emphasis>
|
||
<taxonomicName id="F39B4D53FFC4FFC6E67CF8E9FCF2F878" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[534,774,1906,1928]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="meandrus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC4FFC6E67CF8E9FD50F877" box="[534,676,1906,1927]" italics="true" pageId="11" pageNumber="10">O. meandrus</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC4FFC6E77BF8E9FC62F878" box="[785,918,1906,1928]" pageId="11" pageNumber="10" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7C81655BFFC3FFC1E482FEA1FD72FE42" pageId="12" pageNumber="11" type="etymology">
|
||
<paragraph id="342436D0FFC3FFC1E482FEA1FD72FE42" blockId="12.[192,1440,217,434]" pageId="12" pageNumber="11">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FEA1FE77FEA2" bold="true" box="[232,387,314,338]" pageId="12" pageNumber="11">Etymology.</emphasis>
|
||
The name is formed from “
|
||
<taxonomicName id="F39B4D53FFC3FFC1E6A2FEA1FCB0FEA2" authorityName="Paulsen" authorityYear="2019" box="[712,836,314,338]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="meandrus">meandrus</taxonomicName>
|
||
”, a noun in apposition derived classically from the Meandros River of antiquity and used then figuratively for ‘a wavy border’ in embroidery and continuing into modern usage as ‘meandering’. It is a reference to the wavy margin on the clypeus that is the most recognizable character of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C81655BFFC3FFC1E4AAFE51FA82FE12" box="[192,1398,458,482]" pageId="12" pageNumber="11" type="distribution">
|
||
<paragraph id="342436D0FFC3FFC1E4AAFE51FA82FE12" blockId="12.[192,1398,458,482]" box="[192,1398,458,482]" pageId="12" pageNumber="11">
|
||
<emphasis id="06EFEAC2FFC3FFC1E4AAFE51FE87FE12" bold="true" box="[192,371,458,482]" pageId="12" pageNumber="11">Distribution.</emphasis>
|
||
The species is known only from the vicinity of Ranomafana National Park (
|
||
<figureCitation id="ACA02A55FFC3FFC1E17BFE51FA9DFE12" box="[1297,1385,458,482]" captionStart="Figures 19–20" captionStartId="14.[192,287,1222,1244]" captionTargetBox="[192,1440,213,1188]" captionTargetId="figure@14.[192,1440,213,1188]" captionTargetPageId="14" captionText="Figures 19–20. Distributions of new species of Madagascar Ochodaeus. 19) Blue squares = O. modopunctatus Paulsen, new species; orange circles = O. polypollicatus Paulsen, new species. 20) White circles = O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579575/files/figure.png" pageId="12" pageNumber="11">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C81655BFFC3FFC1E4AAFD81FD2EFCB3" pageId="12" pageNumber="11" type="synonymic_list">
|
||
<paragraph id="342436D0FFC3FFC1E4AAFD81FCF7FDC2" blockId="12.[192,771,538,562]" box="[192,771,538,562]" pageId="12" pageNumber="11">
|
||
<emphasis id="06EFEAC2FFC3FFC1E4AAFD81FCF7FDC2" bold="true" box="[192,771,538,562]" pageId="12" pageNumber="11">Generic Catalog of the Madagascar Fauna</emphasis>
|
||
</paragraph>
|
||
<paragraph id="342436D0FFC3FFC1E482FDD1FD69FD53" blockId="12.[232,824,586,835]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FDD1FD02FD93" authority="Paulian, 1959: 129" authorityName="Paulian" authorityPageNumber="129" authorityYear="1959" box="[232,758,586,611]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="isoanalensis">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FDD1FDFEFD92" box="[232,522,586,610]" italics="true" pageId="12" pageNumber="11">Ochodaeus isoanalensis</emphasis>
|
||
<bibRefCitation id="500A4B21FFC3FFC1E67BFDD0FD4BFD92" author="Paulian, R." box="[529,703,586,611]" journalOrPublisher="Revue Francaise d'Ento- mologie" pageId="12" pageNumber="11" pagination="128 - 133" part="26" refId="ref10312" refString="Paulian, R. 1959. A propos des Ochodaeitae malgaches [Col. Scarabaeidae]. Revue Francaise d'Ento- mologie 26 (3): 128 - 133." title="A propos des Ochodaeitae malgaches [Col. Scarabaeidae]" type="journal article" year="1959">Paulian, 1959</bibRefCitation>
|
||
: 129
|
||
</taxonomicName>
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FDF1FDAFFD73" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[232,603,618,643]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="meandrus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FDF1FE1BFD72" box="[232,495,618,642]" italics="true" pageId="12" pageNumber="11">Ochodaeus meandrus</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC3FFC1E602FDF0FD03FD73" box="[616,759,619,643]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FD11FD69FD53" authority="Klug, 1832: 164" authorityName="Klug" authorityPageNumber="164" authorityYear="1832" box="[232,669,650,675]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="miliaris">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FD11FE22FD52" box="[232,470,650,674]" italics="true" pageId="12" pageNumber="11">Ochodaeus miliaris</emphasis>
|
||
<bibRefCitation id="500A4B21FFC3FFC1E5B7FD10FD92FD53" author="Klug, J. C. F." box="[477,614,651,675]" journalOrPublisher="Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin" pageId="12" pageNumber="11" pagination="91 - 223" part="1832" refId="ref10158" refString="Klug, J. C. F. 1832. Bericht uber eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin 1832 (1): 91 - 223." title="Bericht uber eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung Coleoptera" type="journal article" year="1832">Klug, 1832</bibRefCitation>
|
||
: 164
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="342436D0FFC3FFC1E57AFD30FCC6FD33" blockId="12.[232,824,586,835]" box="[272,818,683,707]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="F39B4D53FFC3FFC1E57AFD30FCC6FD33" authority="Fairmaire, 1868: 785" authorityName="Fairmaire" authorityPageNumber="785" authorityYear="1868" box="[272,818,683,707]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="cannellinus">
|
||
<emphasis id="06EFEAC2FFC3FFC1E57AFD30FDDEFD33" box="[272,554,683,707]" italics="true" pageId="12" pageNumber="11">Ochodaeus cannellinus</emphasis>
|
||
<bibRefCitation id="500A4B21FFC3FFC1E65BFD30FD0FFD33" author="Fairmaire, L." box="[561,763,683,707]" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="12" pageNumber="11" pagination="753 - 820" part="4" refId="ref9979" refString="Fairmaire, L. 1868. Notes sur les coleopteres receuillis par Charles Coquerel a Madagascar et sur les cotes d'Afrique. Annales de la Societe Entomologique de France 4 (8): 753 - 820." title="Notes sur les coleopteres receuillis par Charles Coquerel a Madagascar et sur les cotes d'Afrique" type="journal article" year="1868">Fairmaire, 1868</bibRefCitation>
|
||
: 785
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="342436D0FFC3FFC1E57AFD50FD2EFCB3" blockId="12.[232,824,586,835]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="F39B4D53FFC3FFC1E57AFD50FCD6FD13" authority="Fairmaire, 1868: 785" authorityName="Fairmaire" authorityPageNumber="785" authorityYear="1868" box="[272,802,715,739]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="infuscatus">
|
||
<emphasis id="06EFEAC2FFC3FFC1E57AFD50FDEEFD13" box="[272,538,715,739]" italics="true" pageId="12" pageNumber="11">Ochodaeus infuscatus</emphasis>
|
||
<bibRefCitation id="500A4B21FFC3FFC1E648FD50FD18FD13" author="Fairmaire, L." box="[546,748,715,739]" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="12" pageNumber="11" pagination="753 - 820" part="4" refId="ref9979" refString="Fairmaire, L. 1868. Notes sur les coleopteres receuillis par Charles Coquerel a Madagascar et sur les cotes d'Afrique. Annales de la Societe Entomologique de France 4 (8): 753 - 820." title="Notes sur les coleopteres receuillis par Charles Coquerel a Madagascar et sur les cotes d'Afrique" type="journal article" year="1868">Fairmaire, 1868</bibRefCitation>
|
||
: 785
|
||
</taxonomicName>
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FD70FD68FCF3" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[232,668,747,771]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="modopunctatus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FD70FDC4FCF3" box="[232,560,747,771]" italics="true" pageId="12" pageNumber="11">Ochodaeus modopunctatus</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC3FFC1E6C3FD77FCCCFCF3" box="[681,824,747,771]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FC90FD7EFCD3" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[232,650,779,803]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="polypollicatus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FC90FDEAFCD3" box="[232,542,779,803]" italics="true" pageId="12" pageNumber="11">Ochodaeus polypollicatus</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC3FFC1E6FDFC97FCD2FCD3" box="[663,806,779,803]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
|
||
<taxonomicName id="F39B4D53FFC3FFC1E482FCB0FD2EFCB3" authority="Paulian, 1976: 151" authorityName="Paulian" authorityPageNumber="151" authorityYear="1976" box="[232,730,811,835]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="06EFEAC2FFC3FFC1E482FCB0FE19FCB3" box="[232,493,811,835]" italics="true" pageId="12" pageNumber="11">Ochodaeus pygmaeus</emphasis>
|
||
<bibRefCitation id="500A4B21FFC3FFC1E59FFCB0FD57FCB3" author="Paulian, R." box="[501,675,811,835]" journalOrPublisher="Nouvelle Revue d'Entomologie" pageId="12" pageNumber="11" pagination="139 - 152" part="6" refId="ref10342" refString="Paulian, R. 1976. Les Ochodaeidae (Col. Scarab.) de Madagascar. Nouvelle Revue d'Entomologie 6 (2): 139 - 152." title="Les Ochodaeidae (Col. Scarab.) de Madagascar" type="journal article" year="1976">Paulian, 1976</bibRefCitation>
|
||
: 151
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |