640 lines
58 KiB
XML
640 lines
58 KiB
XML
<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3619.3.5" ID-GBIF-Dataset="1557e4fb-c16a-447b-a152-f4a08a8739b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="217371" ID-ZooBank="4FB7A97C-D25F-41C8-8C62-2664E6E6F148" checkinTime="1460578250636" checkinUser="plazi" docAuthor="De Frias Martins, António M., Brito, Carlos P. & Backeljau, Thierry" docDate="2013" docId="915F87CB1238FFEA519A121BFDE8C780" docLanguage="en" docName="zt03619p368.pdf" docOrigin="Zootaxa 3619 (3)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@4300e240" docStyleName="zootaxa.2013.journal_article" docTitle="Drouetia Gude 1911" docType="treatment" docVersion="7" lastPageNumber="364" masterDocId="6D66FFB3122AFFFF510D1036FF9AC01C" masterDocTitle="Oxychilus (Drouetia) viridescens (Gastropoda: Pulmonata: Oxychilidae), a new species from Santa Maria, Açores, and a review of the subgenus" masterLastPageNumber="368" masterPageNumber="343" pageNumber="361" updateTime="1635533516141" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Oxychilus (Drouetia) viridescens (Gastropoda: Pulmonata: Oxychilidae), a new species from Santa Maria, Açores, and a review of the subgenus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Frias Martins, António M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brito, Carlos P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Backeljau, Thierry</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>3619</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>343</mods:start>
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<mods:end>368</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3619.3.5</mods:identifier>
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<mods:identifier type="GBIF-Dataset">1557e4fb-c16a-447b-a152-f4a08a8739b8</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">217371</mods:identifier>
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<mods:identifier type="ZooBank">4FB7A97C-D25F-41C8-8C62-2664E6E6F148</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5619903" ID-GBIF-Taxon="119560740" ID-Zenodo-Dep="5619903" LSID="urn:lsid:plazi:treatment:915F87CB1238FFEA519A121BFDE8C780" httpUri="http://treatment.plazi.org/id/915F87CB1238FFEA519A121BFDE8C780" lastPageId="21" lastPageNumber="364" pageId="18" pageNumber="361">
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<subSubSection pageId="18" pageNumber="361" type="nomenclature">
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<paragraph blockId="18.[151,955,557,583]" box="[151,955,557,583]" pageId="18" pageNumber="361">
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<heading bold="true" box="[151,955,557,583]" fontSize="11" level="1" pageId="18" pageNumber="361" reason="1">
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<emphasis bold="true" box="[151,955,557,583]" pageId="18" pageNumber="361">
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Reviewing
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<taxonomicName authority="Gude, 1911" authorityName="Gude" authorityYear="1911" box="[288,543,557,583]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis bold="true" box="[288,396,557,583]" italics="true" pageId="18" pageNumber="361">Drouetia</emphasis>
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Gude,
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<number box="[486,543,557,583]" pageId="18" pageNumber="361" value="1911.0">1911</number>
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</taxonomicName>
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and
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<taxonomicName authority="Riedel, 1964" authorityName="Riedel" authorityYear="1964" box="[605,955,557,583]" class="Gastropoda" family="Oxychilidae" genus="Atlantoxychilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis bold="true" box="[605,795,557,583]" italics="true" pageId="18" pageNumber="361">Atlantoxychilus</emphasis>
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Riedel,
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<number box="[897,955,557,583]" pageId="18" pageNumber="361" value="1964.0">1964</number>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="18.[151,1436,621,2032]" pageId="18" pageNumber="361">
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The genus with the highest number of endemic taxa in the Azorean biota is the molluscan
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<taxonomicName box="[1169,1281,621,645]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis box="[1169,1281,621,645]" italics="true" pageId="18" pageNumber="361">Oxychilus</emphasis>
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</taxonomicName>
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, of which the subgenus
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<taxonomicName box="[263,363,657,680]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis box="[263,363,657,680]" italics="true" pageId="18" pageNumber="361">Drouetia</emphasis>
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</taxonomicName>
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accounts for
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<number box="[519,533,655,679]" pageId="18" pageNumber="361" value="8.0">8</number>
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of the
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<number box="[616,644,655,679]" pageId="18" pageNumber="361" value="15.0">15</number>
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||
hitherto described endemic species of that genus. However,
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<taxonomicName box="[1336,1436,657,680]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis box="[1336,1436,657,680]" italics="true" pageId="18" pageNumber="361">Drouetia</emphasis>
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</taxonomicName>
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was first monotypic (Gude
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<number box="[471,529,690,714]" pageId="18" pageNumber="361" value="1911.0">1911</number>
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), for its
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<typeStatus box="[633,681,691,715]" pageId="18" pageNumber="361">type</typeStatus>
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species,
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<taxonomicName box="[789,959,691,715]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlantica">
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<emphasis box="[789,959,691,715]" italics="true" pageId="18" pageNumber="361">Helix atlantica</emphasis>
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</taxonomicName>
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, was considered to live throughout most islands of the Azores (Morelet
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<number box="[500,560,725,749]" pageId="18" pageNumber="361" value="1860.0">1860</number>
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). It was only through Riedel’s (
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<number box="[913,971,725,749]" pageId="18" pageNumber="361" value="1964.0">1964</number>
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) anatomical research that the diversity of
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<taxonomicName box="[151,251,761,784]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis box="[151,251,761,784]" italics="true" pageId="18" pageNumber="361">Drouetia</emphasis>
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</taxonomicName>
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started to be noticed, and Martins (
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<number box="[682,741,759,783]" pageId="18" pageNumber="361" value="2005.0">2005</number>
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) has provided evidence that the number of known species remains understated. Besides the three species from Santa Maria sufficiently dealt with above, five more species are known: two from São Miguel,
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<emphasis box="[534,731,829,853]" italics="true" pageId="18" pageNumber="361">
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O. (D.)
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<taxonomicName box="[620,731,829,853]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
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</emphasis>
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and
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<emphasis box="[785,995,829,853]" italics="true" pageId="18" pageNumber="361">
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O. (D.)
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<taxonomicName box="[871,995,829,853]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
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</emphasis>
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de Winter,
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<number box="[1124,1183,829,853]" pageId="18" pageNumber="361" value="1989.0">1989</number>
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; two from Terceira,
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<emphasis italics="true" pageId="18" pageNumber="361">
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O. (D.)
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<taxonomicName authority="Martins, 1989" authorityName="Martins" authorityYear="1989" box="[207,281,865,888]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Miceui">miceui</taxonomicName>
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</emphasis>
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Martins,
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<number box="[397,453,863,887]" pageId="18" pageNumber="361" value="1989.0">1989</number>
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and
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<emphasis box="[515,699,864,888]" italics="true" pageId="18" pageNumber="361">
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O. (D.)
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<taxonomicName box="[609,699,864,888]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Furtadoi">furtadoi</taxonomicName>
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</emphasis>
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; one from Faial,
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<emphasis box="[907,1057,864,888]" italics="true" pageId="18" pageNumber="361">
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O.(D.)
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<taxonomicName authority="Riedel, 1964" authorityName="Riedel" authorityYear="1964" box="[990,1057,865,888]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Minor">minor</taxonomicName>
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</emphasis>
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Riedel,
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<number box="[1159,1219,863,887]" pageId="18" pageNumber="361" value="1964.0">1964</number>
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.
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<taxonomicName box="[1232,1406,864,888]" class="Gastropoda" family="Oxychilidae" genus="Atlantoxychilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
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<emphasis box="[1232,1406,864,888]" italics="true" pageId="18" pageNumber="361">Atlantoxychilus</emphasis>
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</taxonomicName>
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is represented by one, already mentioned species:
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<emphasis box="[686,898,899,923]" italics="true" pageId="18" pageNumber="361">
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O. (A.)
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<taxonomicName box="[771,894,899,923]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Spectabilis">spectabilis</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph blockId="18.[151,1436,621,2032]" pageId="18" pageNumber="361">
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<taxonomicName box="[199,565,933,958]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus" subGenus="Drouetia">
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||
<emphasis bold="true" box="[199,565,933,958]" italics="true" pageId="18" pageNumber="361">Oxychilus (Drouetia) atlanticus</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[581,661,933,957]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="18" pageNumber="361">
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Fig.
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<number box="[633,661,933,957]" pageId="18" pageNumber="361" value="18.0">18</number>
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</figureCitation>
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C,
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<number box="[694,767,933,957]" pageId="18" pageNumber="361" value="19.2">19, 20</number>
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A). The history of this taxon has been dealt with above (see
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<emphasis box="[151,249,968,992]" italics="true" pageId="18" pageNumber="361">Remarks</emphasis>
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). Martins (
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<number box="[374,432,967,991]" pageId="18" pageNumber="361" value="1991.0">1991</number>
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) first reported the anatomical variability of
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<emphasis box="[920,1117,968,992]" italics="true" pageId="18" pageNumber="361">
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O. (D.)
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<taxonomicName box="[1006,1117,968,992]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
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</emphasis>
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throughout São Miguel and, although calling attention to variability as an artifact of preservation, recognized the possibility of additional species, namely Riedel’s (
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<number box="[447,505,1037,1061]" pageId="18" pageNumber="361" value="1964.0">1964</number>
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) “small form” and the population of Sete Cidades. Nevertheless, Martins’ (
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<number box="[1369,1428,1037,1061]" pageId="18" pageNumber="361" value="2005.0">2005</number>
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) extended analysis interpreted those differences as allotopic, demic variability. However, further anatomical and morphometric research has revealed syntopic variability, with different shell and genitalia patterns existing sympatrically, thus suggesting the existence of various taxonomic units in São Miguel (Martins
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<emphasis box="[1252,1303,1141,1165]" italics="true" pageId="18" pageNumber="361">et al</emphasis>
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.
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<number box="[1320,1380,1141,1165]" pageId="18" pageNumber="361" value="2010.0">2010</number>
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), the description of which is presently under way.
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<emphasis box="[712,1011,1176,1200]" italics="true" pageId="18" pageNumber="361">
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<taxonomicName box="[712,824,1176,1200]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
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(D.)
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<taxonomicName box="[901,1011,1176,1200]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
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</emphasis>
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needed, thus, to be unequivocally characterized. To this end, a
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||
<typeStatus box="[485,574,1211,1235]" pageId="18" pageNumber="361">neotype</typeStatus>
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was proposed so to restrict
|
||
<emphasis box="[903,1107,1211,1235]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
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<taxonomicName box="[996,1107,1211,1235]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
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</emphasis>
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||
to the large shell phenotype living in Sete Cidades, São Miguel (ICZN Case
|
||
<number box="[709,768,1245,1269]" pageId="18" pageNumber="361" value="3553.0">3553</number>
|
||
; Martins
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||
<emphasis box="[878,937,1245,1269]" italics="true" pageId="18" pageNumber="361">et al.</emphasis>
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<number box="[945,1005,1245,1269]" pageId="18" pageNumber="361" value="2011.0">2011</number>
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||
). However, this proposal is no longer needed, since the recent discovery of Drouët´s collection of Azorean material, with specimens from São Miguel, provided a suitable
|
||
<typeStatus box="[383,486,1314,1338]" pageId="18" pageNumber="361">lectotype</typeStatus>
|
||
for
|
||
<taxonomicName box="[540,711,1315,1339]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<emphasis box="[540,711,1315,1339]" italics="true" pageId="18" pageNumber="361">Helix atlantica</emphasis>
|
||
</taxonomicName>
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||
(MJSD.
|
||
<number box="[810,870,1314,1338]" pageId="18" pageNumber="361" value="2012.0">2012</number>
|
||
.CO.
|
||
<number box="[919,947,1314,1338]" pageId="18" pageNumber="361" value="68.0">68</number>
|
||
a;
|
||
<figureCitation box="[989,1075,1314,1338]" captionStart="FIGURE 19" captionStartId="21.[151,250,669,692]" captionTargetBox="[151,1436,205,630]" captionTargetId="figure@21.[151,1436,205,630]" captionTargetPageId="21" captionText="FIGURE 19. Lectotype of Helix atlantica Morelet & Drouët, Muséum Jardin des Sciences de Dijon, France (MJSD. 2012. CO. 68 a), São Miguel. Scale bar = 1 mm." httpUri="https://zenodo.org/record/217390/files/figure.png" pageId="18" pageNumber="361">
|
||
Fig.
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||
<number box="[1044,1075,1314,1338]" pageId="18" pageNumber="361" value="19.0">19</number>
|
||
</figureCitation>
|
||
). The specimen selected is the largest of a lot of
|
||
<number box="[346,374,1349,1373]" pageId="18" pageNumber="361" value="48.0">48</number>
|
||
(diameter 7.8 mm; height, 4.8 mm; number of whorls,
|
||
<number box="[989,1003,1349,1373]" pageId="18" pageNumber="361" value="6.0">6</number>
|
||
) and conforms with the measurements given in the original description (maximum diameter,
|
||
<quantity box="[769,838,1383,1408]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="361" unit="mm" value="8.0">8 mm</quantity>
|
||
; height,
|
||
<quantity box="[937,1007,1383,1408]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="361" unit="mm" value="5.0">5 mm</quantity>
|
||
; Morelet & Drouët
|
||
<number box="[1234,1294,1383,1407]" pageId="18" pageNumber="361" value="1857.0">1857</number>
|
||
). Due to the taxonomic and morphological complexity of
|
||
<taxonomicName box="[663,763,1420,1443]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[663,763,1420,1443]" italics="true" pageId="18" pageNumber="361">Drouetia</emphasis>
|
||
</taxonomicName>
|
||
in São Miguel and the importance of anatomy and locality for species identification (Martins
|
||
<number box="[549,682,1453,1477]" pageId="18" pageNumber="361" value="1991.2005">1991, 2005</number>
|
||
), the
|
||
<typeStatus box="[747,795,1454,1478]" pageId="18" pageNumber="361">type</typeStatus>
|
||
locality for
|
||
<taxonomicName authority="Morelet & Drouet" authorityName="Morelet & Drouet" box="[941,1328,1453,1477]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<emphasis box="[941,1111,1453,1477]" italics="true" pageId="18" pageNumber="361">Helix atlantica</emphasis>
|
||
Morelet & Drouët
|
||
</taxonomicName>
|
||
is herein restricted to Caldeira das Sete Cidades. Indistinguishable from the following species on the basis of shell morphology alone,
|
||
<emphasis box="[368,565,1523,1547]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[455,565,1523,1547]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
|
||
</emphasis>
|
||
has a darker border of the mantle and the foot sole is yellowish; however, it is readily identified anatomically, namely by the long epiphallus, the long and narrow bursa duct and large bursa, and, inside the penis, the reticulate ornamentation on the penial caecum and around the epiphallic pore.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="21" lastPageNumber="364" pageId="18" pageNumber="361" type="description">
|
||
<paragraph blockId="18.[151,1436,621,2032]" pageId="18" pageNumber="361">
|
||
<emphasis bold="true" box="[199,720,1626,1651]" pageId="18" pageNumber="361">
|
||
<emphasis bold="true" box="[199,513,1626,1651]" italics="true" pageId="18" pageNumber="361">
|
||
<taxonomicName box="[199,316,1626,1651]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
(D.)
|
||
<taxonomicName box="[384,513,1626,1651]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
de Winter,
|
||
<number box="[664,720,1626,1651]" pageId="18" pageNumber="361" value="1989.0">1989</number>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[740,824,1626,1650]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="18" pageNumber="361">
|
||
Fig.
|
||
<number box="[796,824,1626,1650]" pageId="18" pageNumber="361" value="18.0">18</number>
|
||
</figureCitation>
|
||
D,
|
||
<number box="[863,895,1626,1650]" pageId="18" pageNumber="361" value="20.0">20</number>
|
||
B). Conchologically indistinguishable from the previous species,
|
||
<emphasis box="[357,576,1661,1685]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[452,576,1661,1685]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
was proposed on the basis of the morphology of the penial complex (de Winter
|
||
<number box="[235,295,1695,1719]" pageId="18" pageNumber="361" value="1989.0">1989</number>
|
||
). Martins (
|
||
<number box="[418,476,1695,1719]" pageId="18" pageNumber="361" value="1991.0">1991</number>
|
||
) showed that the proportions of various structures of the penial complex, namely the penial caecum, can change as an artefact of preservation methods and, therefore, are by themselves unreliable diagnostic characters. For that reason
|
||
<emphasis box="[585,801,1765,1789]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[677,801,1765,1789]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
was considered a junior synonym of
|
||
<emphasis box="[1234,1436,1765,1789]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName authority="Martins 2005" authorityName="Martins" authorityYear="2005" box="[1325,1436,1765,1789]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
|
||
</emphasis>
|
||
(Martins
|
||
<number box="[256,317,1799,1823]" pageId="18" pageNumber="361" value="2005.0">2005</number>
|
||
). Recent research based mostly on the internal morphology of the penis (Martins
|
||
<emphasis box="[1257,1315,1800,1824]" italics="true" pageId="18" pageNumber="361">et al.</emphasis>
|
||
<number box="[1324,1382,1799,1823]" pageId="18" pageNumber="361" value="2010.0">2010</number>
|
||
) has provided evidence for the existence of various taxonomic units in São Miguel; restriction of
|
||
<emphasis box="[1205,1406,1835,1859]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[1295,1406,1835,1859]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Atlanticus">atlanticus</taxonomicName>
|
||
</emphasis>
|
||
to the anatomically different Sete Cidades population restores the validity of
|
||
<emphasis box="[1029,1250,1869,1893]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[1126,1250,1869,1893]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
. Although very variable, when compared with the previous species
|
||
<emphasis box="[741,956,1904,1928]" italics="true" pageId="18" pageNumber="361">
|
||
O. (D.)
|
||
<taxonomicName box="[832,956,1904,1928]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
shows a lighter blue border of the mantle and the foot is pinkish to orange, sometimes greenish; it is diagnosed by the short vagina, the membranous proximal penis with weak crests radiating from the epiphallic pore, the strongly muscular distal penis with strong pilasters inside.
|
||
<emphasis box="[332,628,2008,2032]" italics="true" pageId="18" pageNumber="361">
|
||
<taxonomicName box="[332,444,2008,2032]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
(D.)
|
||
<taxonomicName box="[504,628,2008,2032]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="361" phylum="Mollusca" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</emphasis>
|
||
is distributed throughout the entire island of São Miguel.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/217388/files/figure.png" pageId="19" pageNumber="362" targetBox="[258,1329,182,1920]" targetPageId="19">
|
||
<paragraph blockId="19.[151,1436,1945,2028]" pageId="19" pageNumber="362">
|
||
<emphasis bold="true" box="[151,292,1945,1968]" pageId="19" pageNumber="362">FIGURE 17.</emphasis>
|
||
Shell morphometry in
|
||
<taxonomicName box="[547,892,1945,1967]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="362" phylum="Mollusca" rank="species" species="Viridescens" status="n.sp." subGenus="Drouetia">
|
||
<emphasis box="[547,892,1945,1967]" italics="true" pageId="19" pageNumber="362">
|
||
Oxychilus
|
||
<taxonomicName box="[659,892,1945,1967]" pageId="19" pageNumber="362" rank="species" species="Viridescens" status="n.sp." subGenus="Drouetia">
|
||
(Drouetia)
|
||
<taxonomicName box="[778,892,1945,1967]" pageId="19" pageNumber="362" rank="species" species="Viridescens" status="n.sp.">viridescens</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[903,951,1946,1967]" pageId="19" pageNumber="362" rank="species">n.sp.</taxonomicNameLabel>
|
||
(*) and
|
||
<taxonomicName box="[1050,1353,1945,1967]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="362" phylum="Mollusca" rank="species" species="Brincki" subGenus="Drouetia">
|
||
<emphasis box="[1050,1353,1945,1967]" italics="true" pageId="19" pageNumber="362">
|
||
Oxychilus
|
||
<taxonomicName box="[1163,1353,1945,1967]" pageId="19" pageNumber="362" rank="species" species="Brincki" subGenus="Drouetia">
|
||
(Drouetia)
|
||
<taxonomicName box="[1281,1353,1945,1967]" pageId="19" pageNumber="362" rank="species" species="Brincki">brincki</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(o).
|
||
<emphasis bold="true" box="[1411,1429,1945,1966]" pageId="19" pageNumber="362">A</emphasis>
|
||
. Relationship di/#w.
|
||
<emphasis bold="true" box="[355,378,1976,1997]" pageId="19" pageNumber="362">B.</emphasis>
|
||
Relationship he/sp. Abbreviations: di, shell diameter; he, height of the shell; sp, height of the spire; #w, number of whorls. Holotype within square.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="20" pageNumber="363" targetBox="[175,1421,193,1865]" targetPageId="20">
|
||
<paragraph blockId="20.[151,1436,1881,2027]" pageId="20" pageNumber="363">
|
||
<emphasis bold="true" box="[151,289,1881,1904]" pageId="20" pageNumber="363">FIGURE 18.</emphasis>
|
||
Animals of species of
|
||
<taxonomicName box="[532,622,1883,1904]" class="Gastropoda" family="Oxychilidae" genus="Drouetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[532,622,1883,1904]" italics="true" pageId="20" pageNumber="363">Drouetia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[676,833,1882,1904]" class="Gastropoda" family="Oxychilidae" genus="Atlantoxychilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[676,833,1882,1904]" italics="true" pageId="20" pageNumber="363">Atlantoxychilus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" box="[848,866,1882,1903]" pageId="20" pageNumber="363">A</emphasis>
|
||
,
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[881,1267,1882,1904]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Viridescens" subGenus="Drouetia">
|
||
<emphasis box="[881,1221,1882,1904]" italics="true" pageId="20" pageNumber="363">
|
||
Oxychilus (
|
||
<taxonomicName box="[998,1090,1883,1904]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
)
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[1107,1221,1882,1904]" pageId="20" pageNumber="363" rank="species" species="Viridescens">viridescens</taxonomicName>
|
||
</emphasis>
|
||
, Sta
|
||
</taxonomicName>
|
||
7.
|
||
<emphasis bold="true" box="[1303,1320,1882,1903]" pageId="20" pageNumber="363">B</emphasis>
|
||
,
|
||
<taxonomicName authority="Sta" authorityName="Sta" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Agostinhoi" subGenus="Drouetia">
|
||
<taxonomicName class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Agostinhoi" subGenus="Drouetia">
|
||
<emphasis italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[1335,1436,1882,1904]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[151,371,1912,1934]" pageId="20" pageNumber="363" rank="species" species="Agostinhoi" subGenus="Drouetia">
|
||
(
|
||
<taxonomicName box="[158,251,1913,1934]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
)
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[264,371,1912,1934]" pageId="20" pageNumber="363" rank="species" species="Agostinhoi">agostinhoi</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, Sta
|
||
</taxonomicName>
|
||
7.
|
||
<emphasis bold="true" box="[445,463,1912,1935]" pageId="20" pageNumber="363">C</emphasis>
|
||
,
|
||
<taxonomicName box="[476,795,1912,1934]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Atlanticus" subGenus="Drouetia">
|
||
<emphasis box="[476,795,1912,1934]" italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[476,577,1912,1934]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName box="[582,795,1912,1934]" pageId="20" pageNumber="363" rank="species" species="Atlanticus" subGenus="Drouetia">
|
||
(
|
||
<taxonomicName box="[590,682,1913,1934]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
)
|
||
<taxonomicName box="[695,795,1912,1934]" pageId="20" pageNumber="363" rank="species" species="Atlanticus">atlanticus</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK 20100653), Sete Cidades, São Miguel.
|
||
<emphasis bold="true" box="[1304,1322,1913,1934]" pageId="20" pageNumber="363">D</emphasis>
|
||
,
|
||
<taxonomicName authority="Batalha, Sao Miguel" authorityName="Batalha, Sao Miguel" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Batalhanus" subGenus="Drouetia">
|
||
<taxonomicName authority="Batalha, Sao" authorityName="Batalha, Sao" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Batalhanus" subGenus="Drouetia">
|
||
<taxonomicName authority="Batalha" authorityName="Batalha" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Batalhanus" subGenus="Drouetia">
|
||
<taxonomicName class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Batalhanus" subGenus="Drouetia">
|
||
<emphasis italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[1335,1436,1912,1934]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName authority="Batalha, Sao Miguel" authorityName="Batalha, Sao Miguel" box="[151,381,1943,1965]" pageId="20" pageNumber="363" rank="species" species="Batalhanus" subGenus="Drouetia">
|
||
(Drouetia)
|
||
<taxonomicName authority="Batalha, Sao Miguel" authorityName="Batalha, Sao Miguel" box="[269,381,1943,1965]" pageId="20" pageNumber="363" rank="species" species="Batalhanus">batalhanus</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, Batalha
|
||
</taxonomicName>
|
||
, São
|
||
</taxonomicName>
|
||
Miguel
|
||
</taxonomicName>
|
||
(type locality).
|
||
<emphasis bold="true" box="[793,810,1944,1965]" pageId="20" pageNumber="363">E</emphasis>
|
||
,
|
||
<taxonomicName box="[827,1125,1943,1965]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Miceui" subGenus="Drouetia">
|
||
<emphasis box="[827,1125,1943,1965]" italics="true" pageId="20" pageNumber="363">
|
||
Oxychilus
|
||
<taxonomicName box="[939,1125,1944,1965]" pageId="20" pageNumber="363" rank="species" species="Miceui" subGenus="Drouetia">
|
||
(
|
||
<taxonomicName box="[947,1039,1944,1965]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
) miceui
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Caldeira de Santa Bárbara, Terceira.
|
||
<emphasis bold="true" box="[341,354,1974,1995]" pageId="20" pageNumber="363">F</emphasis>
|
||
,
|
||
<taxonomicName box="[369,674,1974,1996]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Furtadoi" subGenus="Drouetia">
|
||
<emphasis box="[369,674,1974,1996]" italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[369,470,1974,1996]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName box="[478,674,1974,1996]" pageId="20" pageNumber="363" rank="species" species="Furtadoi" subGenus="Drouetia">
|
||
(
|
||
<taxonomicName box="[485,577,1975,1996]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
)
|
||
<taxonomicName box="[593,674,1974,1996]" pageId="20" pageNumber="363" rank="species" species="Furtadoi">furtadoi</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
: paratype, Fonte da Telha, Posto Santo, Terceira.
|
||
<emphasis bold="true" box="[1189,1206,1973,1996]" pageId="20" pageNumber="363">G</emphasis>
|
||
,
|
||
<taxonomicName authority="Ribeirinha, Faial. H" authorityName="Ribeirinha, Faial. H" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Minor" subGenus="Drouetia">
|
||
<taxonomicName authority="Ribeirinha, Faial." authorityName="Ribeirinha, Faial." class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Minor" subGenus="Drouetia">
|
||
<taxonomicName authority="Ribeirinha, Faial" authorityName="Ribeirinha, Faial" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Minor" subGenus="Drouetia">
|
||
<taxonomicName authority="Ribeirinha" authorityName="Ribeirinha" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Minor" subGenus="Drouetia">
|
||
<taxonomicName class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Minor" subGenus="Drouetia">
|
||
<emphasis italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[1220,1321,1974,1996]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName authority="Ribeirinha, Faial" authorityName="Ribeirinha, Faial" pageId="20" pageNumber="363" rank="species" species="Minor" subGenus="Drouetia">
|
||
(
|
||
<taxonomicName box="[1336,1428,1975,1996]" genus="Drouetia" pageId="20" pageNumber="363" rank="genus">Drouetia</taxonomicName>
|
||
) minor
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, Ribeirinha
|
||
</taxonomicName>
|
||
, Faial
|
||
</taxonomicName>
|
||
.
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[406,426,2005,2026]" pageId="20" pageNumber="363">H</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[438,879,2005,2027]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Spectabilis" subGenus="Atlantoxychilus">
|
||
<taxonomicName box="[438,834,2005,2027]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="species" species="Spectabilis" subGenus="Atlantoxychilus">
|
||
<emphasis box="[438,834,2005,2027]" italics="true" pageId="20" pageNumber="363">
|
||
<taxonomicName box="[438,539,2005,2027]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="363" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[546,834,2005,2027]" pageId="20" pageNumber="363" rank="species" species="Spectabilis" subGenus="Atlantoxychilus">
|
||
(Atlantoxychilus)
|
||
<taxonomicName authority="Sta" authorityName="Sta" box="[726,834,2005,2027]" pageId="20" pageNumber="363" rank="species" species="Spectabilis">spectabilis</taxonomicName>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, Sta
|
||
</taxonomicName>
|
||
7.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://zenodo.org/record/217390/files/figure.png" pageId="21" pageNumber="364" targetBox="[151,1436,205,630]" targetPageId="21">
|
||
<paragraph blockId="21.[151,1436,669,722]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[151,300,669,692]" pageId="21" pageNumber="364">FIGURE 19.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName authority="Morelet & Drouet" authorityName="Morelet & Drouet" box="[478,871,669,691]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<taxonomicName authority="Morelet" authorityName="Morelet" box="[478,741,669,691]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<taxonomicName box="[478,642,669,691]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<emphasis box="[478,642,669,691]" italics="true" pageId="21" pageNumber="364">
|
||
<taxonomicName box="[478,533,669,691]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="genus">Helix</taxonomicName>
|
||
<taxonomicName authority="Morelet & Drouet" authorityName="Morelet & Drouet" box="[552,642,669,691]" pageId="21" pageNumber="364" rank="species" species="Atlantica">atlantica</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
Morelet
|
||
</taxonomicName>
|
||
& Drouët
|
||
</taxonomicName>
|
||
, Muséum Jardin des Sciences de Dijon, France (MJSD.2012.CO.68 a), São Miguel. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="21.[151,1436,766,1948]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,636,766,791]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,458,766,791]" italics="true" pageId="21" pageNumber="364">
|
||
<taxonomicName box="[199,318,766,791]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
(D.)
|
||
<taxonomicName authority="Martins, 1989" authorityName="Martins" authorityYear="1989" box="[379,458,766,791]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Miceui">miceui</taxonomicName>
|
||
</emphasis>
|
||
Martins,
|
||
<number box="[579,636,766,791]" pageId="21" pageNumber="364" value="1989.0">1989</number>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[651,744,767,791]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="21" pageNumber="364">
|
||
Figs.
|
||
<number box="[716,744,767,791]" pageId="21" pageNumber="364" value="18.0">18</number>
|
||
</figureCitation>
|
||
E,
|
||
<number box="[776,806,767,791]" pageId="21" pageNumber="364" value="21.0">21</number>
|
||
A). This species lives on the mountains of Terceira and is characterized by the yellow to whitish mantle spotted with small brown to blackish patches, contrasting with the dark-blue mantle border and neck, fading to light-blue toward the foot. The shell has a depressed spire, the last whorl a quadrangular profile, the aperture is slightly wider than that of the following species. Anatomically this species is characterized by a long, convoluted oviduct and moderately long vagina, a membranous proximal penis and penial caecum cumulatively as long as the muscular distal penis, both units usually separated by a constriction.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1436,766,1948]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,665,1028,1053]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,481,1028,1053]" italics="true" pageId="21" pageNumber="364">
|
||
<taxonomicName box="[199,318,1028,1053]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
(D.)
|
||
<taxonomicName authority="Martins, 1989" authorityName="Martins" authorityYear="1989" box="[386,481,1028,1053]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Furtadoi">furtadoi</taxonomicName>
|
||
</emphasis>
|
||
Martins,
|
||
<number box="[608,665,1028,1053]" pageId="21" pageNumber="364" value="1989.0">1989</number>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[684,779,1028,1052]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="21" pageNumber="364">
|
||
Figs.
|
||
<number box="[752,779,1028,1052]" pageId="21" pageNumber="364" value="18.0">18</number>
|
||
</figureCitation>
|
||
F,
|
||
<number box="[812,843,1028,1052]" pageId="21" pageNumber="364" value="21.0">21</number>
|
||
B). This species lives at lower altitude around the town of Angra do Heroísmo, Terceira. Conchologically similar to
|
||
<emphasis box="[943,1116,1065,1089]" italics="true" pageId="21" pageNumber="364">
|
||
O. (D.)
|
||
<taxonomicName box="[1040,1116,1066,1089]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Miceui">miceui</taxonomicName>
|
||
</emphasis>
|
||
, it has a slightly narrower aperture. The mantle has a greenish-brown background spotted with sparse, small whitish patches, the mantle border and neck light-blue, turning pinkish toward the foot. The reproductive anatomy is characterized by a long, narrow vagina, a short, membranous proximal penis and penial caecum, a long, narrow, muscular distal penis and an equally long penial sheath; the epiphallus is long, the distal third connected to the distal penis by muscular strands.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1436,766,1948]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,612,1289,1314]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,452,1289,1314]" italics="true" pageId="21" pageNumber="364">
|
||
<taxonomicName box="[199,316,1289,1314]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="genus">Oxychilus</taxonomicName>
|
||
(D.)
|
||
<taxonomicName authority="Riedel, 1964" authorityName="Riedel" authorityYear="1964" box="[382,452,1289,1314]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Minor">minor</taxonomicName>
|
||
</emphasis>
|
||
Riedel,
|
||
<number box="[556,612,1289,1314]" pageId="21" pageNumber="364" value="1964.0">1964</number>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[630,722,1289,1313]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="21" pageNumber="364">
|
||
Figs.
|
||
<number box="[696,722,1289,1313]" pageId="21" pageNumber="364" value="18.0">18</number>
|
||
</figureCitation>
|
||
G,
|
||
<number box="[758,788,1289,1313]" pageId="21" pageNumber="364" value="21.0">21</number>
|
||
C). This species was first referred by Morelet (
|
||
<number box="[1336,1395,1289,1313]" pageId="21" pageNumber="364" value="1860.0">1860</number>
|
||
) as variety “β
|
||
<taxonomicName box="[272,344,1327,1351]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Minor">minor</taxonomicName>
|
||
”, and it was Riedel (
|
||
<number box="[581,639,1327,1351]" pageId="21" pageNumber="364" value="1964.0">1964</number>
|
||
) who considered it a discrete taxon, attributing authorship and date to Morelet (
|
||
<number box="[256,317,1364,1388]" pageId="21" pageNumber="364" value="1860.0">1860</number>
|
||
). Bank
|
||
<emphasis box="[404,462,1364,1388]" italics="true" pageId="21" pageNumber="364">et al.</emphasis>
|
||
(
|
||
<number box="[477,538,1364,1388]" pageId="21" pageNumber="364" value="2002.0">2002</number>
|
||
), however, interpreted Morelet’s word “
|
||
<taxonomicName box="[990,1061,1364,1388]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Minor">minor</taxonomicName>
|
||
” not as a name but as part of the description, thus attributing authorship and date to Riedel (
|
||
<number box="[812,873,1401,1425]" pageId="21" pageNumber="364" value="1964.0">1964</number>
|
||
). The animal of
|
||
<emphasis box="[1057,1212,1402,1426]" italics="true" pageId="21" pageNumber="364">
|
||
O. (D.)
|
||
<taxonomicName box="[1145,1212,1403,1426]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Minor">minor</taxonomicName>
|
||
</emphasis>
|
||
is characteristically pink, the mantle mottled with white patches and rare dark spots, posterior tentacles dark-blue becoming lighter toward the base. The shell has a depressed spire and the aperture is somewhat wide. The most peculiar anatomic feature is the stout, strongly muscular penis without noticeable constriction, with strong pilasters running back to the penial caecum.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1436,766,1948]" pageId="21" pageNumber="364">
|
||
<taxonomicName authority="Milne-Edwards, 1885" authorityName="Milne-Edwards" authorityYear="1885" box="[199,959,1588,1613]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Spectabilis" subGenus="Atlantoxychilus">
|
||
<emphasis bold="true" box="[199,959,1588,1613]" pageId="21" pageNumber="364">
|
||
<emphasis bold="true" box="[199,665,1588,1613]" italics="true" pageId="21" pageNumber="364">Oxychilus (Atlantoxychilus) spectabilis</emphasis>
|
||
(Milne-Edwards,
|
||
<number box="[892,951,1588,1613]" pageId="21" pageNumber="364" value="1885.0">1885</number>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[977,1062,1588,1612]" captionStart="FIGURE 18" captionStartId="20.[151,250,1881,1904]" captionTargetBox="[175,1421,193,1865]" captionTargetId="figure@20.[175,1422,193,1867]" captionTargetPageId="20" captionText="FIGURE 18. Animals of species of Drouetia and Atlantoxychilus. A, Oxychilus (Drouetia) viridescens, Sta 7. B, Oxychilus (Drouetia) agostinhoi, Sta 7. C, Oxychilus (Drouetia) atlanticus (NHMUK 20100653), Sete Cidades, São Miguel. D, Oxychilus (Drouetia) batalhanus, Batalha, São Miguel (type locality). E, Oxychilus (Drouetia) miceui, paratype, Caldeira de Santa Bárbara, Terceira. F, Oxychilus (Drouetia) furtadoi: paratype, Fonte da Telha, Posto Santo, Terceira. G, Oxychilus (Drouetia) minor, Ribeirinha, Faial. H, Oxychilus (Atlantoxychilus) spectabilis, Sta 7." httpUri="https://zenodo.org/record/217389/files/figure.png" pageId="21" pageNumber="364">
|
||
Fig.
|
||
<number box="[1033,1062,1588,1612]" pageId="21" pageNumber="364" value="18.0">18</number>
|
||
</figureCitation>
|
||
H). Hausdorf (
|
||
<number box="[1234,1294,1588,1612]" pageId="21" pageNumber="364" value="1993.0">1993</number>
|
||
) considered
|
||
<taxonomicName box="[151,322,1626,1650]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Atlantica">
|
||
<emphasis box="[151,322,1626,1650]" italics="true" pageId="21" pageNumber="364">Helix atlantica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName authority="Morelet, 1860" authorityName="Morelet" authorityYear="1860" box="[331,674,1625,1650]" pageId="21" pageNumber="364" rank="variety" variety="spectabilis">
|
||
var.
|
||
<emphasis box="[380,503,1626,1650]" italics="true" pageId="21" pageNumber="364">spectabilis</emphasis>
|
||
Morelet,
|
||
<number box="[617,674,1625,1649]" pageId="21" pageNumber="364" value="1860.0">1860</number>
|
||
</taxonomicName>
|
||
to be a primary homonym of
|
||
<taxonomicName authority="Pfeiffer, 1844" authorityName="Pfeiffer" authorityYear="1844" box="[1021,1383,1625,1650]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Spectabilis">
|
||
<emphasis box="[1021,1213,1626,1650]" italics="true" pageId="21" pageNumber="364">Helix spectabilis</emphasis>
|
||
Pfeiffer,
|
||
<number box="[1323,1383,1625,1649]" pageId="21" pageNumber="364" value="1844.0">1844</number>
|
||
</taxonomicName>
|
||
, and introduced accordingly the new name
|
||
<emphasis box="[586,974,1663,1687]" italics="true" pageId="21" pageNumber="364">
|
||
Oxchilus (
|
||
<taxonomicName box="[703,884,1663,1687]" class="Gastropoda" family="Oxychilidae" genus="Atlantoxychilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="genus">Atlantoxychilus</taxonomicName>
|
||
) riedeli
|
||
</emphasis>
|
||
. However, Bank
|
||
<emphasis box="[1172,1221,1663,1687]" italics="true" pageId="21" pageNumber="364">et al</emphasis>
|
||
(
|
||
<number box="[1237,1297,1663,1687]" pageId="21" pageNumber="364" value="2002.0">2002</number>
|
||
) considered Hausdorf’s (
|
||
<number box="[294,353,1700,1724]" pageId="21" pageNumber="364" value="1993.0">1993</number>
|
||
) nomenclatorial action invalid, for Morelet’s (
|
||
<number box="[885,945,1700,1724]" pageId="21" pageNumber="364" value="1860.0">1860</number>
|
||
) indication of a “γ
|
||
<taxonomicName box="[1165,1338,1700,1724]" pageId="21" pageNumber="364" rank="variety" variety="spectabilis">var. spectabilis</taxonomicName>
|
||
” did not constitute attribution of a name to the variety, the word “
|
||
<taxonomicName box="[829,955,1737,1761]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="364" phylum="Mollusca" rank="species" species="Spectabilis">spectabilis</taxonomicName>
|
||
” instead being part of the description of variety γ. The name was validly introduced by Milne-Edwards (
|
||
<number box="[920,979,1775,1799]" pageId="21" pageNumber="364" value="1885.0">1885</number>
|
||
) as
|
||
<taxonomicName box="[1035,1269,1775,1799]" class="Orbiliomycetes" family="Orbiliaceae" genus="Hyalinia" kingdom="Fungi" order="Orbiliales" pageId="21" pageNumber="364" phylum="Ascomycota" rank="species" species="Spectabilis">
|
||
<emphasis box="[1035,1269,1775,1799]" italics="true" pageId="21" pageNumber="364">Hyalinia spectabilis</emphasis>
|
||
</taxonomicName>
|
||
, which is not preoccupied. Some aspects of this species were already dealt with, and only a brief morphological characterization will be added. The mantle is brown, crossed by elongated, irregular golden-brown blotches; border of mantle light-brown; neck dark-brown, sharply separated from the golden-brown foot which is surrounded by an intermittent brown rim.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |