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<document id="89FD09D5327020D4098BD14DD2D7789F" ID-DOI="10.5252/zoosystema2018v40a10" ID-GBIF-Dataset="ea3961a2-6410-4963-9e03-af59316cac6d" ID-ISSN="1638-9387" ID-Zenodo-Dep="3738229" ID-ZooBank="urn:lsid:zoobank.org:pub:F2681FDC-D9BE-48F8-918C-63CF67FF6F52" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585839718085" checkinUser="felipe" docAuthor="Nguyen, Minh &amp; Soto-Adames, Felipe N." docDate="2018" docId="720BFC4CFFEDFF82FC34FF501B95AE78" docLanguage="en" docName="zoosystema.40.10.179-196.pdf.imf" docOrigin="Zoosystema 40 (10)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Trogolaphysa nigromaculata subsp. pallida" docType="treatment" docVersion="7" lastPageNumber="190" masterDocId="8E328434FFE0FF8FFF9EFF881E0EAA48" masterDocTitle="Annotated checklist of Afrotropical Trogolaphysa Mills, 1938 (Hexapoda: Collembola: Paronellidae) and description of a new species from Madagascar" masterLastPageNumber="196" masterPageNumber="179" pageNumber="190" updateTime="1698735047691" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="252D3063D924C16865761A71ECCAB852">Annotated checklist of Afrotropical Trogolaphysa Mills, 1938 (Hexapoda: Collembola: Paronellidae) and description of a new species from Madagascar</mods:title>
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<mods:namePart id="4D2D4B5C6BF55BDBE903035FF9919FE8">Nguyen, Minh</mods:namePart>
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<mods:title id="D415C7A7C98BD8A45062CE75B5BC7CE3">Zoosystema</mods:title>
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<treatment id="720BFC4CFFEDFF82FC34FF501B95AE78" ID-DOI="http://doi.org/10.5281/zenodo.3811387" ID-GBIF-Taxon="163421415" ID-Zenodo-Dep="3811387" LSID="urn:lsid:plazi:treatment:720BFC4CFFEDFF82FC34FF501B95AE78" httpUri="http://treatment.plazi.org/id/720BFC4CFFEDFF82FC34FF501B95AE78" lastPageNumber="190" pageId="13" pageNumber="190">
<subSubSection id="B2B81ED1FFEDFF82FC34FF501AFFAB7B" pageId="13" pageNumber="190" type="nomenclature">
<paragraph id="FA1D4D5AFFEDFF82FC34FF501AFFAB7B" blockId="13.[846,1421,216,308]" pageId="13" pageNumber="190">
<heading id="A155FA36FFEDFF82FC34FF501AFFAB7B" centered="true" fontSize="11" level="2" pageId="13" pageNumber="190" reason="3">
<taxonomicName id="3DA236D9FFEDFF82FC34FF501A9FAB7C" authority="(Delamare Debouteville in Delamare Debouteville &amp; Pauliani, 1952)" baseAuthorityName="Delamare Debouteville in Delamare Debouteville &amp; Pauliani" baseAuthorityYear="1952" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FC34FF501B3CAABB" bold="true" box="[938,1330,216,243]" italics="true" pageId="13" pageNumber="190">Trogolaphysa nigromaculata pallida</emphasis>
(Delamare Debouteville
<emphasis id="C8D69148FFEDFF82FBC6FF721A60AB5B" box="[1112,1134,250,275]" italics="true" pageId="13" pageNumber="190">in</emphasis>
Delamare Debouteville &amp; Pauliani, 1952)
</taxonomicName>
n. comb
</heading>
</paragraph>
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<subSubSection id="B2B81ED1FFEDFF82FCB3FEDA1B65ABB6" pageId="13" pageNumber="190" type="reference_group">
<paragraph id="FA1D4D5AFFEDFF82FCB3FEDA1B79ABCC" blockId="13.[813,1455,338,389]" pageId="13" pageNumber="190">
<treatmentCitationGroup id="DAB26A74FFEDFF82FCB3FEDA1B79ABCC" pageId="13" pageNumber="190">
<taxonomicName id="3DA236D9FFEDFF82FCB3FEDA1A71ABCC" ID-CoL="5K8CS" authority="Delamare Debouteville in Delamare Debouteville &amp; Pauliani, 1952: 77" authorityName="Delamare Debouteville in Delamare Debouteville &amp; Pauliani" authorityPageNumber="77" authorityYear="1952" class="Collembola" family="Paronellidae" genus="Paronella" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FCB3FEDA1A5CAB22" box="[813,1106,338,362]" italics="true" pageId="13" pageNumber="190">Paronella nigromaculata pallida</emphasis>
Delamare Debouteville
<emphasis id="C8D69148FFEDFF82FAA7FEDA1B42AB21" box="[1337,1356,338,361]" italics="true" pageId="13" pageNumber="190">in</emphasis>
<treatmentCitation id="7B036B4BFFEDFF82FACCFEDA1A71ABCC" pageId="13" pageNumber="190">
<bibRefCitation id="9E3330ABFFEDFF82FACCFEDA1A71ABCC" author="DELAMARE DEBOUTTEVILLE C. &amp; PAULIAN R." pageId="13" pageNumber="190" pagination="62 - 77" refId="ref14006" refString="DELAMARE DEBOUTTEVILLE C. &amp; PAULIAN R. 1952. - Recherches sur la faune des nids et des terriers en Basse Cote d'Ivoire. Chapitre VIII, Etude systematique. I. - Collembola. Encyclopedie biogeographique et ecologique 8: 62 - 77" type="journal article" year="1952">Delamare Debouteville &amp; Pauliani, 1952: 77</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(Le Banco,
<collectingCountry id="82B50DCAFFEDFF82FB68FEE51B62ABCC" box="[1270,1388,364,389]" name="Ivory Coast" pageId="13" pageNumber="190">Ivory Coast</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA1D4D5AFFEDFF82FCB3FE141DB3AB86" blockId="13.[813,1455,412,462]" pageId="13" pageNumber="190">
<treatmentCitationGroup id="DAB26A74FFEDFF82FCB3FE141DB3AB86" pageId="13" pageNumber="190">
<taxonomicName id="3DA236D9FFEDFF82FCB3FE141BA5ABFC" ID-CoL="5K8CS" authority="Delamare Debouteville, 1951: 209" authorityName="Delamare Debouteville" authorityPageNumber="209" authorityYear="1951" box="[813,1451,412,436]" class="Collembola" family="Paronellidae" genus="Paronella" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FCB3FE141A5BABFC" box="[813,1109,412,436]" italics="true" pageId="13" pageNumber="190">Paronella nigromaculata pallida</emphasis>
<treatmentCitation id="7B036B4BFFEDFF82FBC4FE141BA5ABFC" box="[1114,1451,412,436]" pageId="13" pageNumber="190">
<bibRefCitation id="9E3330ABFFEDFF82FBC4FE141BA5ABFC" author="DELAMARE DEBOUTTEVILLE C." box="[1114,1451,412,436]" pageId="13" pageNumber="190" pagination="1072 - 1075" refId="ref13893" refString="DELAMARE DEBOUTTEVILLE C. 1951 a. - Nouveaux Paronelliens de la Cote d'Ivoire. Bulletin de l'Institut francaise d'Afrique noire 13: 1072 - 1075." type="journal article" year="1951">Delamare Debouteville, 1951: 209</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<emphasis id="C8D69148FFEDFF82FCB3FE3F1DB3AB86" box="[813,957,438,462]" italics="true" pageId="13" pageNumber="190">nomen nudum.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA1D4D5AFFEDFF82FCB3FE6F1B65ABB6" blockId="13.[813,1387,486,510]" box="[813,1387,486,510]" pageId="13" pageNumber="190">
<treatmentCitationGroup id="DAB26A74FFEDFF82FCB3FE6F1B65ABB6" box="[813,1387,486,510]" pageId="13" pageNumber="190">
<emphasis id="C8D69148FFEDFF82FCB3FE6F1A93ABB6" box="[813,1181,486,510]" italics="true" pageId="13" pageNumber="190">
<taxonomicName id="3DA236D9FFEDFF82FCB3FE6F1A58ABB6" authorityName="Nguyen &amp; Soto-Adames" authorityYear="2018" baseAuthorityName="Nguyen &amp; Soto-Adames" baseAuthorityYear="2018" box="[813,1110,486,510]" class="Entognatha" family="Paronellidae" genus="Dicranocentruga" higherTaxonomySource="CoL" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="190" phylum="Arthropoda" rank="species" species="nigromaculata">Dicranocentruga nigromaculata</taxonomicName>
pallida
</emphasis>
–
<treatmentCitation id="7B036B4BFFEDFF82FB29FE6E1B69ABB6" author="MITRA S. K." box="[1207,1383,486,510]" page="114" pageId="13" pageNumber="190" year="2002">
<bibRefCitation id="9E3330ABFFEDFF82FB29FE6E1B69ABB6" author="Mitra" box="[1207,1383,486,510]" firstAuthor="Mitra" pageId="13" pageNumber="190" pagination="105 - 116" refId="ref14625" refString="MITRA S. K. 2002 c. - Status of Dicranocentruga Wray, 1953 with the description of a new species (Collembola: Entomobryidae). Records of the Zoological Survey of India 100: 105 - 116." type="journal article" year="2002" yearSuffix="c">Mitra 2002c: 114</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2B81ED1FFEDFF82FCB3FD9D1A51A815" pageId="13" pageNumber="190" type="distribution">
<paragraph id="FA1D4D5AFFEDFF82FCB3FD9D1AC2A866" blockId="13.[813,1228,533,558]" box="[813,1228,533,558]" pageId="13" pageNumber="190">
<typeStatus id="2519F3F8FFEDFF82FCB3FD9D1D53A862" box="[813,861,533,557]" pageId="13" pageNumber="190">TYPE</typeStatus>
LOCALITY. —
<collectorName id="5757288CFFEDFF82FC75FD9E1A44A865" box="[1003,1098,534,557]" pageId="13" pageNumber="190">Le Banco</collectorName>
,
<collectingCountry id="82B50DCAFFEDFF82FBCDFD9E1AC6A866" box="[1107,1224,534,558]" name="Ivory Coast" pageId="13" pageNumber="190">Ivory Coast</collectingCountry>
.
</paragraph>
<paragraph id="FA1D4D5AFFEDFF82FCB3FDCD1A51A815" blockId="13.[813,1119,581,606]" box="[813,1119,581,606]" pageId="13" pageNumber="190">
DISTRIBUTION. —
<collectingCountry id="82B50DCAFFEDFF82FC78FDCE1A54A815" box="[998,1114,581,606]" name="Ivory Coast" pageId="13" pageNumber="190">Ivory Coast</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B81ED1FFEDFF82FCB3FD1F1B95AE78" pageId="13" pageNumber="190" type="discussion">
<paragraph id="FA1D4D5AFFEDFF82FCB3FD1F1D9FA8F8" blockId="13.[813,1456,663,1073]" box="[813,913,663,689]" pageId="13" pageNumber="190">REMARKS</paragraph>
<paragraph id="FA1D4D5AFFEDFF82FCB3FD3F1A23A999" blockId="13.[813,1456,663,1073]" pageId="13" pageNumber="190">
Delamare Debouteville (1951b) first mentions this taxon as a subspecies of
<taxonomicName id="3DA236D9FFEDFF82FC6FFD501A91A8B9" authorityName="Nguyen &amp; Soto-Adames" authorityYear="2018" baseAuthorityName="Nguyen &amp; Soto-Adames" baseAuthorityYear="2018" box="[1009,1183,727,753]" class="Entognatha" family="Paronellidae" genus="Dicranocentruga" higherTaxonomySource="CoL" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="190" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="C8D69148FFEDFF82FC6FFD501A91A8B9" box="[1009,1183,727,753]" italics="true" pageId="13" pageNumber="190">P. nigromaculata</emphasis>
</taxonomicName>
, but only as a name in a table (1951b: 209). Since no description is attached to this name,
<taxonomicName id="3DA236D9FFEDFF82FCEDFC901BA1A97A" authority="Delamare Debouteville, 1951 b" authorityName="Delamare Debouteville" authorityYear="1951" box="[883,1455,791,818]" class="Entognatha" family="Paronellidae" genus="Dicranocentruga" higherTaxonomySource="CoL" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FCEDFC901A69A979" box="[883,1127,791,817]" italics="true" pageId="13" pageNumber="190">P. nigromaculata pallida</emphasis>
Delamare Debouteville, 1951b
</taxonomicName>
is a
<emphasis id="C8D69148FFEDFF82FCC5FCB01DF9A919" box="[859,1015,823,849]" italics="true" pageId="13" pageNumber="190">nomen nudum</emphasis>
. Delamare Debouteville again used the combination
<taxonomicName id="3DA236D9FFEDFF82FC23FCD01AB7A939" authorityName="Delamare Debouteville" authorityYear="1951" box="[957,1209,855,881]" class="Entognatha" family="Paronellidae" genus="Dicranocentruga" higherTaxonomySource="CoL" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="nigromaculata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FC23FCD01AB7A939" box="[957,1209,855,881]" italics="true" pageId="13" pageNumber="190">P. nigromaculata pallida</emphasis>
</taxonomicName>
in 1952, and this time states that the taxon is characterized by very sparse punctate blue pigment but with the same morphological characters as the nominal subspecies.
</paragraph>
<paragraph id="FA1D4D5AFFEDFF82FCDDFC5E1B95AE78" blockId="13.[813,1456,663,1073]" pageId="13" pageNumber="190">
<emphasis id="C8D69148FFEDFF82FCDDFC5E1A13A9B8" box="[835,1053,982,1008]" italics="true" pageId="13" pageNumber="190">
<taxonomicName id="3DA236D9FFEDFF82FCDDFC5E1DA6A9B8" baseAuthorityName="Barra" baseAuthorityYear="1969" box="[835,936,982,1008]" class="Collembola" family="Paronellidae" genus="Paronella" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="13" pageNumber="190" phylum="Arthropoda" rank="genus">Paronella</taxonomicName>
n. pallida
</emphasis>
was described on the same page as
<taxonomicName id="3DA236D9FFEDFF82FCB3FC7F1A0DAE58" authorityName="Delamare Debouteville in Delamare Debouteville &amp; Paulian" authorityYear="1952" box="[813,1027,1014,1040]" class="Collembola" family="Paronellidae" genus="Paronella" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="13" pageNumber="190" phylum="Arthropoda" rank="subSpecies" species="penicillata" subSpecies="pallida">
<emphasis id="C8D69148FFEDFF82FCB3FC7F1A0DAE58" box="[813,1027,1014,1040]" italics="true" pageId="13" pageNumber="190">P. penicillata pallida</emphasis>
</taxonomicName>
(see below). If these two taxa turn out to be good species one of them will require a new epithet.
</paragraph>
</subSubSection>
</treatment>
</document>
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