129 lines
14 KiB
XML
129 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.252.3588" ID-GBIF-Dataset="a4366e50-a271-44c0-bdef-b52c8cad0b85" ID-PMC="PMC3560839" ID-Pensoft-Pub="1313-2970-252-1" ID-PubMed="23378811" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-252-1" ModsDocOrigin="ZooKeys 252" ModsDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" checkinTime="1451247847790" checkinUser="pensoft" docAuthor="Bond, Jason E." docDate="2012" docId="5304B8EB732CF10BE8A4C2DFE9803C63" docLanguage="en" docName="ZooKeys 252: 1-209" docOrigin="ZooKeys 252" docSource="http://dx.doi.org/10.3897/zookeys.252.3588" docTitle="Aptostichus fornax Bond, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="142" masterDocId="FFAD871BFFD4FF8CFFED1A5CFFE7FFA0" masterDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" masterLastPageNumber="209" masterPageNumber="1" pageNumber="140" updateTime="1668155039594" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bond, Jason E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>252</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>209</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.252.3588</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.252.3588</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-252-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039295" LSID="urn:lsid:zoobank.org:act:50095886-C9E7-4098-84D7-ADB6911C7FCC" httpUri="http://treatment.plazi.org/id/5304B8EB732CF10BE8A4C2DFE9803C63" lastPageId="141" lastPageNumber="142" pageId="139" pageNumber="140">
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<subSubSection pageId="139" pageNumber="140" type="nomenclature">
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<paragraph pageId="139" pageNumber="140">
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<taxonomicName LSID="‘The Furnace Trapdoor Spider’" class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
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<taxonomicNameLabel pageId="139" pageNumber="140">sp. n.</taxonomicNameLabel>
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Figures 298-303Map 33
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</paragraph>
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</subSubSection>
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<subSubSection pageId="139" pageNumber="140" type="types">
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<paragraph pageId="139" pageNumber="140">Types.</paragraph>
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<paragraph pageId="139" pageNumber="140">Male holotype (AP418) from California, Inyo County, Panamint Valley sand dunes, 36.09167, -117.25917 4, 317m, coll. D. Giuliani 17.ii.1972; female paratype (AP578) from type locality, coll. 27.ix.1974; holotype deposited in CAS, paratype in AMNH.</paragraph>
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</subSubSection>
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<subSubSection pageId="139" pageNumber="140" type="etymology">
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<paragraph pageId="139" pageNumber="140">Etymology.</paragraph>
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<paragraph pageId="139" pageNumber="140">The specific epithet is a noun taken in apposition and refers to the hot climate in which this species is found. Fornax is a southern constellation; the name is Latin for furnace. In Roman mythology Fornax is the goddess of the hearth.</paragraph>
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</subSubSection>
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<subSubSection pageId="139" pageNumber="140" type="diagnosis">
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<paragraph pageId="139" pageNumber="140">Diagnosis.</paragraph>
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<paragraph pageId="139" pageNumber="140">
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Males can be distinguished by a palpal tibia with a retrolateral patch of small spines and at least one large, stout spine (Fig. 303, 302) and numerous spines on the retrolateral surface of tibia II;
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus elisabethae" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="elisabethae">Aptostichus elisabethae</taxonomicName>
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lacks the palpal spine and tibia II retrolateral spines. The enlarged palpal spine is shared bya number of
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<taxonomicName class="Actinopterygii" family="Scombridae" genus="Sierra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sierra" order="Perciformes" pageId="139" pageNumber="140" phylum="Chordata" rank="genus">Sierra</taxonomicName>
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species group taxa; however, these species lack the proximal - ventral metatarsus I excavation that differentially characterizes
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
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.
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
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specimens lack the serrated embolus of other
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Simus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simus" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">Simus</taxonomicName>
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group species (
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus simus" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="simus">Aptostichus simus</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus lucerne" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="lucerne">Aptostichus lucerne</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus satleri" order="Araneae" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="satleri">Aptostichus satleri</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="139" pageNumber="140" type="description">
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<paragraph pageId="139" pageNumber="140">Description of male holotype.</paragraph>
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<paragraph pageId="139" pageNumber="140">
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Specimen preparation and condition. Specimen collected from pitfall trap, preserved in 70% EtOH. Coloration faded. Pedipalp, leg I left side removed, stored in vial with specimen. General coloration. Carapace, chelicerae, legs dark red 2.5YR 3/6. Abdomen uniform very pale brown 10YR 7/4, lacks distinct markings (Fig. 298). Cephalothorax. Carapace 4.8 long, 3.96 wide, glabrous, stout black bristles along fringe; surface smooth, pars cephalica elevated. Fringe, posterior margin with black bristles. Foveal groove deep, strongly recurved. Eyes on low mound. AER slightly procurved, PER slightly recurved.
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<normalizedToken originalValue="PME’s">PME's</normalizedToken>
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much smaller in size than
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<normalizedToken originalValue="AME’s">AME's</normalizedToken>
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. Sternum moderately setose, STRl 2.75, STRw 2.23. Posterior sternal sigilla medium in size, elongate, positioned centrally, not contiguous, anterior sigilla pairs small, sub-oval to elongate, marginal. Chelicerae with distinct anterior tooth row comprising 4 teeth, posterior margin with single row of small denticles. Palpal endites, labium, lacks cuspules, LBw 0.77, LBl 0.37. Rastellum consists of 8 stout spines arranged in continuous row. Abdomen. Setose, heavy black setae intermingled with fine black setae. Legs. Leg I: 4.50, 3.64, 2.82, 1.83, 1.50; leg IV: 4.2, 2.36. Light tarsal scopulae on legs I, II. Tarsus I with single, slightly staggered row of 12 trichobothria. Leg I spination pattern illustrated in Figures 299, 300, 302; TSp 17, TSr 12, TSrd 0. Pedipalp. Articles stout, palpal tibia width greater than
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<normalizedToken originalValue="½">1/2</normalizedToken>
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length, with one large spine situated among distinct patch of smaller spines (Figs 301, 302). Palpal bulb short, broad with slight curvature at tip, lacks serrations. PTw 0.92, PTl 1.70, Bl 0.62.
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</paragraph>
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<paragraph pageId="139" pageNumber="140">Variation.Known only from the type specimen.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="141" lastPageNumber="142" pageId="139" pageNumber="140" type="description">
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<paragraph pageId="139" pageNumber="140">Description of female paratype.</paragraph>
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<paragraph lastPageId="141" lastPageNumber="142" pageId="139" pageNumber="140">
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Specimen preparation and condition. Female presumed to have been collected live from burrow, prepared in same manner as male holotype; in very poor condition, abdomen destroyed. Genital plate in poor condition, removed,
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<pageBreakToken pageId="140" pageNumber="141" start="start">cleared</pageBreakToken>
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in trypsin, stored in microvial with specimen. General coloration. Carapace, legs, chelicerae, dark reddish brown 2.5YR 2.5/3. Cephalothorax. Carapace 4.40 long, 3.32 wide, generally glabrous, very sparse fine black setae; generally smooth surface, pars cephalica
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<pageBreakToken pageId="141" pageNumber="142" start="start">moderately</pageBreakToken>
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elevated. Fringe lacks setae. Foveal groove deep, slightly procurved. Eye group slightly elevated on very low mound. AER slightly procurved, PER slightly recurved.
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<normalizedToken originalValue="PME’s">PME's</normalizedToken>
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smaller in diameter than
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<normalizedToken originalValue="AME’s">AME's</normalizedToken>
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. Sternum widest at coxae II/III, moderately setose, STRl 2.45, STRw 1.88. Three pairs of sternal sigilla anterior pairs small in size, oval, marginal; posterior pair moderate in size, elongate, mesially positioned but not contiguous. Chelicerae anterior tooth row comprising 4 teeth with posterior margin denticle patch. Palpal endites with>75 cuspules concentrated at the inner (promargin) posterior heel; labium lacks cuspules, LBw 0.80, LBl 0.63. Rastellum consists of 17 stout spines not positioned on mound; fringe of short spines along distal promargin extending upward from rastellum. Abdomen. Moderately setose. PLS all 3 segments with spigots. Terminal segment 1/2 length of medial segment, 2 enlarged spigots visible at tip. PMS single segment, with spigots, short with rounded terminus. Walking legs. Anterior two pairs noticeably more slender than posterior pairs. Leg I 10.46 long. Tarsus I with 7 trichobothria arranged in staggered row. Legs I, II with light scopulae on tarsus, metatarsus; light scopulae on distal aspect tarsus legs III, IV. PTLs 23, TBs 4. Rudimentary preening comb on retrolateral distal surface, tarsus-metatarsus joint, of metatarsus IV; well developed, wide preening comb on leg III. Spermathecae. 2 short, heavily sclerotized, simple spermathecal bulbs; basal extension small (Fig. 303).
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</paragraph>
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<paragraph pageId="141" pageNumber="142">Variation.Known only from the type specimen.</paragraph>
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<caption pageId="141" pageNumber="142">
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<paragraph pageId="141" pageNumber="142">
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Figures 298-303.
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
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sp. n. 298-302 male holotype (AP418) from Inyo County; scale bars = 1.0mm 298 habitus [806456] 299 retrolateral aspect, leg I [806460] 300 prolateral aspect, leg I [806462] 301 retrolateral aspect, pedipalp [806464] 302 line drawings of leg I spination pattern and pedipalp; retrolateral aspect patella, tibia, metatarsus; prolateral aspect patella and tibia; retrolateral aspect, pedipalp 303 female paratype (AP578), cleared spermathecae [806465]; scale bar = 0.1mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="141" pageNumber="142" type="material examined">
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<paragraph pageId="141" pageNumber="142">Material examined.</paragraph>
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<paragraph pageId="141" pageNumber="142">Known only from the type material.</paragraph>
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</subSubSection>
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<subSubSection pageId="141" pageNumber="142" type="distribution">
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<paragraph pageId="141" pageNumber="142">Distribution and natural history.</paragraph>
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<paragraph pageId="141" pageNumber="142">Known only from the type locality in Inyo County, characterized as Mojave Desert habitat.</paragraph>
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</subSubSection>
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<subSubSection pageId="141" pageNumber="142" type="conservation status">
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<paragraph pageId="141" pageNumber="142">Conservation status.</paragraph>
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<paragraph pageId="141" pageNumber="142">
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The conservation status of
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<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
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is likely to be imperiled given its very restricted distribution in Death Valley National Park and paucity of specimens collected.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="141" pageNumber="142" type="species concept applied">
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<paragraph pageId="141" pageNumber="142">Species concept applied.</paragraph>
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<paragraph pageId="141" pageNumber="142">Morphological.</paragraph>
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</subSubSection>
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</treatment>
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</document> |