188 lines
32 KiB
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188 lines
32 KiB
XML
<document ID-CLB-Dataset="21976" ID-DOI="http://dx.doi.org/10.3897/zse.96.50360" ID-GBIF-Dataset="d57080cc-b609-47ae-90ad-f26b430471aa" ID-Pensoft-Pub="1860-0743-1-205" ID-Pensoft-UUID="05A8F88FEA125CD184DD0B44874DB41F" ID-ZooBank="3195954C4A454EC4AE4020B158D75C5E" ModsDocID="1860-0743-1-205" checkinTime="1590026314960" checkinUser="pensoft" docAuthor="Colavite, Jessica, Windsor, Amanda M. & Santana, William" docDate="2020" docId="EA66951E647958F19BC1DAAEBE4A41C9" docLanguage="en" docName="ZoosystEvol 96(1): 205-216" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.50360" docTitle="Pohleus heptacanthus gen. nov. et 2020, comb. nov." docType="treatment" docVersion="5" id="05A8F88FEA125CD184DD0B44874DB41F" lastPageNumber="205" masterDocId="05A8F88FEA125CD184DD0B44874DB41F" masterDocTitle="A new genus for Pericera septemspinosa Stimpson, 1871 and Pericera heptacantha Bell, 1836 (Crustacea, Brachyura, Majoidea), based on morphology and molecular data" masterLastPageNumber="216" masterPageNumber="205" pageNumber="205" updateTime="1732809766295" updateUser="ExternalLinkService">
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<mods:title id="0513974D0892F1A7FC9C1ABB069B4AD2">A new genus for Pericera septemspinosa Stimpson, 1871 and Pericera heptacantha Bell, 1836 (Crustacea, Brachyura, Majoidea), based on morphology and molecular data</mods:title>
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<mods:namePart id="C902CE281112F545AA92625F5586070D">Colavite, Jessica</mods:namePart>
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<mods:affiliation id="56D2329A64CCA0462617DFDE11982563">Departamento de Zoologia, Instituto de Biociencias, Universidade Estadual Paulista " Julio de Mesquita Filho " (UNESP), Botucatu, SP 18618 - 970, Brazil & Laboratory of Systematic Zoology, Centro Universitario Sagrado Coracao (UNISAGRADO), Rua Irma Arminda, 10 - 50, Jd. Brazil, Bauru, SP 17011 - 160, Brazil</mods:affiliation>
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<mods:namePart id="6E4AB3D3E856EF6D9CCA7E79CB59EC0F">Windsor, Amanda M.</mods:namePart>
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<mods:affiliation id="A3BA1CAC1335BE943DF6C229B907334F">Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Museum Support Center, 4210 Silver Hill Road, Suitland, MD 20746 - 2863, USA</mods:affiliation>
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<mods:namePart id="E8FFF26C44AE581354A9DEB244946240">Santana, William</mods:namePart>
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<mods:affiliation id="D2115519A38690D1EFF9C0A19CBBD179">Departamento de Zoologia, Instituto de Biociencias, Universidade Estadual Paulista " Julio de Mesquita Filho " (UNESP), Botucatu, SP 18618 - 970, Brazil & Laboratory of Systematic Zoology, Centro Universitario Sagrado Coracao (UNISAGRADO), Rua Irma Arminda, 10 - 50, Jd. Brazil, Bauru, SP 17011 - 160, Brazil</mods:affiliation>
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<mods:nameIdentifier id="9F3F5C0E1F6350FDED927D2BD07C4A5E" type="email">willsantana@gmail.com</mods:nameIdentifier>
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<mods:title id="B9A451A95004993152089170C724EFFC">Zoosystematics and Evolution</mods:title>
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<treatment id="EA66951E647958F19BC1DAAEBE4A41C9" ID-GBIF-Taxon="164244857" LSID="urn:lsid:plazi:treatment:EA66951E647958F19BC1DAAEBE4A41C9" httpUri="http://treatment.plazi.org/id/EA66951E647958F19BC1DAAEBE4A41C9" lastPageNumber="205" pageId="0" pageNumber="205" scope_family="Epialtidae" scope_infraOrder="Brachyura" scope_subFamily="Pisinae" scope_superFamily="Majoidea">
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<subSubSection id="F63F45F007AEDCC5B3416D831DC0D1A4" pageId="0" pageNumber="205" type="nomenclature">
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<paragraph id="2B5BEA69AFAF72685C9CCB07EDFF6546" pageId="0" pageNumber="205">
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<taxonomicName id="9AA03829882BDDB42090574839CE612B" LSID="EA66951E-6479-58F1-9BC1-DAAEBE4A41C9" authority="(Bell, 1836) gen. nov. et" authorityName="gen. nov. et" authorityYear="2020" baseAuthorityName="Bell" baseAuthorityYear="1836" class="Malacostraca" family="Majidae" genus="Pohleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pohleus heptacanthus" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacanthus" status="comb. nov.">Pohleus heptacanthus (Bell, 1836) gen. nov. et</taxonomicName>
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<taxonomicNameLabel id="D506E93F03C20B6F7AB1414032267399" pageId="0" pageNumber="205">comb. nov.</taxonomicNameLabel>
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<figureCitation id="C47E7122CEF69FBECAB13E80A9281DD3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pohleus septemspinosus (Stimpson, 1871) gen. nov. et comb. nov. A, B. female (MNHN IU 2013 - 32682). Pohleus heptacanthus (Bell, 1836) gen. nov. et comb. nov.; C, D. female lectotype (OUM 13764). A, C. habitus; B, D. ventral view. Note the spine on merus of the second pereopod (white arrow). Scale bars: 10 mm." figureDoi="10.3897/zse.96.50360.figure5" httpUri="https://binary.pensoft.net/fig/412094" pageId="0" pageNumber="205">Fig. 5C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7F0862587ECF0FA1D9A73E689F5C6F26" pageId="0" pageNumber="205" type="reference_group">
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<paragraph id="91A8394E9F39D73CF0F7ED581BA95EE8" pageId="0" pageNumber="205">
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<taxonomicName id="8FCC3A3F44401B9EBE06758937907A65" authorityName="Bell" authorityYear="1836" class="Malacostraca" family="Majidae" genus="Pericera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericera heptacantha" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacantha">Pericera heptacantha</taxonomicName>
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Bell, 1836: 173 [type locality: Puerto Potrero, Central America, 23.7 m depth; type material: syntypes, 1 male (non-extant), 1 female (OUM 13764)] -
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<bibRefCitation id="10B595C284FF1E7A1B654C5DCD4A4662" DOI="https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" author="Bell, T" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="205" refId="B6" refString="Bell, T, 1836b. IV. Some account of the Crustacea of the coasts of South America, with descriptions of new genera and species: founded principally on the collections obtained by Mr. Cuming and Mr. Miller. Transactions of the Zoological Society of London (Zoology) 2(1): 39-66. [pls 8-13]. Date of publication, November 1836 from note inside NHM volume. https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" title="IV. Some account of the Crustacea of the coasts of South America, with descriptions of new genera and species: founded principally on the collections obtained by Mr. Cuming and Mr. Miller. Transactions of the Zoological Society of London (Zoology) 2 (1): 39 - 66. [pls 8 - 13]. Date of publication, November 1836 from note inside NHM volume." url="https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" year="1836 b">Bell 1836b</bibRefCitation>
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: 61, pl. 12, fig. 6 and 6r-u;
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<bibRefCitation id="2826EE8C5D21D128D33B5BCA4304A626" author="White, A" journalOrPublisher="British Museum, London" pageId="0" pageNumber="205" refId="B47" refString="White, A, 1847. List of the Specimens in the Collection of the British Museum (Vol. 8). British Museum, London" title="List of the Specimens in the Collection of the British Museum (Vol. 8)." year="1847">White 1847</bibRefCitation>
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: 10; A.
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<bibRefCitation id="8BBDFD780541872A8694C95BEFEA76E2" author="Milne-Edwards, A" journalOrPublisher="Milne Edwards, membre de l'Institut. Cinquieme partie. Tome premier. Paris: Imprimerie Nationale. 8 [unnumbered]" pageId="0" pageNumber="205" refId="B31" refString="Milne-Edwards, A, 1873. Etudes sur les xiphosures et les crustaces de la region mexicaine. Mission scientifique au Mexique et dans l'Amerique centrale, ouvrage publie par ordre du Ministre de l'Instruction publique. Recherches zoologiques pour servir a l'histoire de la faune de l'Amerique central et du Mexique, publiees sous la direction de M.H. Milne Edwards, membre de l'Institut. Cinquieme partie. Tome premier. Paris: Imprimerie Nationale. 8 [unnumbered]" title="Etudes sur les xiphosures et les crustaces de la region mexicaine. Mission scientifique au Mexique et dans l'Amerique centrale, ouvrage publie par ordre du Ministre de l'Instruction publique. Recherches zoologiques pour servir a l'histoire de la faune de l'Amerique central et du Mexique, publiees sous la direction de M. H." year="1873">Milne-Edwards 1873</bibRefCitation>
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: 55.
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</paragraph>
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<paragraph id="5103639B56D2D0F3E9B859BE70721029" pageId="0" pageNumber="205">
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<taxonomicName id="28B9F910975F04618E7DA6A4581A1488" class="Malacostraca" family="Majidae" genus="Macrocoeloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocoeloma heptacantha" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacantha">Macrocoeloma heptacantha</taxonomicName>
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-
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<bibRefCitation id="BFF5D760D8BCD9ADB11FE3D1563B3359" author="Miers, EJ" journalOrPublisher="Her Majesty Stationery Office, London, Edinburgh and Dublin" pageId="0" pageNumber="205" refId="B29" refString="Miers, EJ, 1886. Miers, E. J., 1886. Part II. - Report on the Brachyura collected by H.M.S. Challenger during the years 1873-76. Report on the Scientific Results of the Voyage of HMS Challenger during the years 1873-1876 under the command of Captain George S Nares, RN, FRS and the late Captain Frank Tourle Thomson, RN prepared under the Superintendence of the late Sir C Wyville Thomson, Knt, FRS &c Regius Professor of Natural History in the University of Edinburgh Director of the civilian scientific staff on board and now of John Murray, LLD, PhD, &c one of the naturalists of the Expedition Zoology, Published by Order of Her Majesty's Government (Vol 17). Her Majesty Stationery Office, London, Edinburgh and Dublin" title="Miers, E. J., 1886. Part II. - Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of HMS Challenger during the years 1873 - 1876 under the command of Captain George S Nares, RN, FRS and the late Captain Frank Tourle Thomson, RN prepared under the Superintendence of the late Sir C Wyville Thomson, Knt, FRS & c Regius Professor of Natural History in the University of Edinburgh Director of the civilian scientific staff on board and now of John Murray, LLD, PhD, & c one of the naturalists of the Expedition Zoology, Published by Order of Her Majesty's Government (Vol 17)." year="1886">Miers 1886</bibRefCitation>
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: 79, 81.
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</paragraph>
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<paragraph id="F898F3FE15D55F8D469B99334ADA4F17" pageId="0" pageNumber="205">
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<taxonomicName id="F53EDBD497B2D4C72D00988D3C507893" baseAuthorityName="Bell" baseAuthorityYear="1835" class="Malacostraca" family="Majidae" genus="Macrocoeloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocoeloma heptacanthum" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacanthum">Macrocoeloma heptacanthum</taxonomicName>
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-
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<bibRefCitation id="8C4325D9767BB13B8CF76807F91623C2" DOI="https://doi.org/10.5479/si.00963801.21-1162.567" author="Rathbun, MJ" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="205" pagination="567 - 616" refId="B37" refString="Rathbun, MJ, 1898a. The Brachyura collected by the U.S. Fish Commission steamer Albatross on the voyage from Norfolk, Virginia, to San Francisco, California, 1887-1888. Proceedings of the United States National Museum 21: 567 - 616, DOI: https://doi.org/10.5479/si.00963801.21-1162.567" title="The Brachyura collected by the U. S. Fish Commission steamer Albatross on the voyage from Norfolk, Virginia, to San Francisco, California, 1887 - 1888." url="https://doi.org/10.5479/si.00963801.21-1162.567" volume="21" year="1898 a">Rathbun 1898a</bibRefCitation>
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: 576;
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<bibRefCitation id="C022C54F46D53C5D92458F5C065DEB77" DOI="https://doi.org/10.5479/si.03629236.129.i" author="Rathbun, MJ" journalOrPublisher="Bulletin of the United States National Museum" pageId="0" pageNumber="205" pagination="1 - 613" refId="B40" refString="Rathbun, MJ, 1925. The spider crabs of America. Bulletin of the United States National Museum 129: 1 - 613, DOI: https://doi.org/10.5479/si.03629236.129.i" title="The spider crabs of America." url="https://doi.org/10.5479/si.03629236.129.i" volume="129" year="1925">1925</bibRefCitation>
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: 473, pl. 173, fig. 1; pl. 269, fig. 8-11, text-figs 133, 134;
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<bibRefCitation id="86D68D27B90DBFBC0B9FFFE756880A9B" author="Garth, JS" journalOrPublisher="Allan Hancock Pacific Expeditions" pageId="0" pageNumber="205" pagination="501 - 854" refId="B18" refString="Garth, JS, 1958. Brachyura of the Pacific coast of America. Oxyrhyncha. Tables and Plates. Allan Hancock Pacific Expeditions 21 (2): 501 - 854" title="Brachyura of the Pacific coast of America. Oxyrhyncha. Tables and Plates." volume="21" year="1958">Garth 1958</bibRefCitation>
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: 415;
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<bibRefCitation id="BAA41A0E0688E42EA66F25EDC6327A76" DOI="https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" author="Di Mauro, Jr AA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="205" pagination="155 - 182" refId="B16" refString="Di Mauro, Jr AA, 1982. Rediscovery of Professor Thomas Bell's type Crustacea (Brachyura) in the dry crustacean collection of the Zoological Collections, University Museum, Oxford. Zoological Journal of the Linnean Society 76 (2): 155 - 182, DOI: https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" title="Rediscovery of Professor Thomas Bell's type Crustacea (Brachyura) in the dry crustacean collection of the Zoological Collections, University Museum, Oxford." url="https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" volume="76" year="1982">Di Mauro 1982</bibRefCitation>
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: 170;
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<bibRefCitation id="C0047EF5836D80968594A6B8D41AE6AC" author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="205" pagination="1 - 286" refId="B34" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
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: 119.
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</paragraph>
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</subSubSection>
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<subSubSection id="8CFA0BA071713278F0289582E64E9025" pageId="0" pageNumber="205" type="lectotype">
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<paragraph id="633029BADB9FBA01BEEE278265292145" pageId="0" pageNumber="205">Lectotype</paragraph>
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<paragraph id="F7735457FBFCD24FE5D2E9582085A768" pageId="0" pageNumber="205">(Here designated).</paragraph>
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<paragraph id="90C2EC2D2D1CC70DFA28DA19C8F0B1D5" pageId="0" pageNumber="205">
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Costa Rica • 1 female, cl: 35 mm, cw: 43 mm; Central America, Puerto Potrero, 23.7 m depth; H Cumming leg. (OUM 13764) (Fig.
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<figureCitation id="2019F10ED31458AF41963AC003D3AA8C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pohleus septemspinosus (Stimpson, 1871) gen. nov. et comb. nov. A, B. female (MNHN IU 2013 - 32682). Pohleus heptacanthus (Bell, 1836) gen. nov. et comb. nov.; C, D. female lectotype (OUM 13764). A, C. habitus; B, D. ventral view. Note the spine on merus of the second pereopod (white arrow). Scale bars: 10 mm." figureDoi="10.3897/zse.96.50360.figure5" httpUri="https://binary.pensoft.net/fig/412094" pageId="0" pageNumber="205">5C, D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="719D775BF84398F1A8CA52C6C6C4D9BA" pageId="0" pageNumber="205" type="materials_examined">
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<paragraph id="5EE1067393EE494DFC6FE51D338328AD" pageId="0" pageNumber="205">Material examined.</paragraph>
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<paragraph id="5950EB6A91ADEDA1E20992DA2359A58B" pageId="0" pageNumber="205">
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Mexico • 1 juv. female; Off Cape San Lucas, R/V Albatross, stn 2829,
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<geoCoordinate id="8B85A3AFF7F28ACEAF560C3A3780EB2E" degrees="22" direction="north" minutes="52" orientation="latitude" precision="15" seconds="00" value="22.866667">22°52'00"N</geoCoordinate>
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,
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<geoCoordinate id="128F74D1721792640AA56080C9A6452C" degrees="109" direction="west" minutes="55" orientation="longitude" precision="15" seconds="00" value="-109.916664">109°55'00"W</geoCoordinate>
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, RKY leg., 56.6 m depth, 74.1 °F; 01 May 1888, M J Rathbun det. (USNM 21933 illustrated). Panama • 1 juv. female; Panama Bay, R/V Albatross, stn 2798,
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<geoCoordinate id="4A1AD728FBE3F449882C8359272F5E4F" degrees="8" direction="north" minutes="10" orientation="latitude" precision="15" seconds="30" value="8.175">8°10'30"N</geoCoordinate>
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,
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<geoCoordinate id="2384B766A533C098DD1E788E8AEE2E84" degrees="78" direction="west" minutes="50" orientation="longitude" precision="15" seconds="30" value="-78.84167">78°50'30"W</geoCoordinate>
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, 114.6 m depth; 05 Mar1888, M J Rathbun det. (USNM 21932).
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</paragraph>
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</subSubSection>
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<subSubSection id="CEEB93E400EDEC0207A5E746E75DF226" pageId="0" pageNumber="205" type="diagnosis">
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<paragraph id="7CB78921B94DDB3B2712812B75C749E2" pageId="0" pageNumber="205">Diagnosis.</paragraph>
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<paragraph id="06542E103546F9409B9A3D90C7BE26D9" pageId="0" pageNumber="205">Rostrum width less than one-third of interorbital length, bifurcated, base elongated, fused, diverging abruptly forming a Y-shape, ending in acute tip. Pleonal somite II with one spine or tubercle. Merus of second pereopod smooth.</paragraph>
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</subSubSection>
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<subSubSection id="9A3B6A2DB11178AB40B4E8A6EA9F80AB" pageId="0" pageNumber="205" type="description">
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<paragraph id="6E0256C0349CE610EC104C81D5374334" pageId="0" pageNumber="205">Description based on female lectotype</paragraph>
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<paragraph id="FD7C46D9879393FA2790015E06579120" pageId="0" pageNumber="205">
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(male characters modified from
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<bibRefCitation id="8797E2231666F8E7105561549B9AC14E" DOI="https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" author="Bell, T" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="205" refId="B6" refString="Bell, T, 1836b. IV. Some account of the Crustacea of the coasts of South America, with descriptions of new genera and species: founded principally on the collections obtained by Mr. Cuming and Mr. Miller. Transactions of the Zoological Society of London (Zoology) 2(1): 39-66. [pls 8-13]. Date of publication, November 1836 from note inside NHM volume. https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" title="IV. Some account of the Crustacea of the coasts of South America, with descriptions of new genera and species: founded principally on the collections obtained by Mr. Cuming and Mr. Miller. Transactions of the Zoological Society of London (Zoology) 2 (1): 39 - 66. [pls 8 - 13]. Date of publication, November 1836 from note inside NHM volume." url="https://doi.org/10.1111/j.1469-7998.1839.tb00007.x" year="1836 b">Bell 1836b</bibRefCitation>
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). Cephalothorax and appendages slightly covered with short, velvet-like pubescence. Carapace subglobose wider than long, convex, with long lines of hooked and simple setae in all regions. Rostrum short, less than one-third of interorbital length, bifurcated, base elongated, fused, diverging abruptly forming a Y-shape, ending in acute tips. Interorbital region slightly depressed medially. Hepatic region broad. One metagastric spine. Four long, strong, conical lateral spines (two in each branchial region), in line with cardiac spine. One short intestinal spine. Orbital region very prominent, eyes completely protected when retracted, ocular peduncle visible when not retracted. Pre-orbital spine directed upwards, slightly curved on tip, longer than post-orbital spine, ventral margin of pre-orbital spine with small crenulation; post-orbital spine curved upwards.
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</paragraph>
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<paragraph id="908A444A03FABBCAD117AB4FFB97CA92" pageId="0" pageNumber="205">Antennular fossae wider than long, margins smooth. Interantennular septum elongate, laterally compressed, forming distinct ventrally-directed lobe. First and second antennal articles fused to epistome, with suture between antenna and epistome visible, antennal gland opening near suture line. Basal article of antenna with two spines, not visible in dorsal view: proximal spine smallest. Antennal flagella longer than rostral spines, behind rostrum in dorsal view.</paragraph>
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<paragraph id="D816C8F7BCD46BFA09383B7B9BE5D5E9" pageId="0" pageNumber="205">Epistome anterior margin narrower than antennular fossae, smooth, posterior margin slightly depressed. Buccal field sub-rectangular, narrower at posterior edge with one acute spine in anterolateral angle aligned with antennal spines. Third maxillipeds covering buccal frame when closed, leaving a small gap between ischia. Exopod long, nearly reaching distal margin of merus. Pterygostomial region subtriangular, slightly inflated, separated from subhepatic region by marked groove, with one long, strong spine slightly curved upwards on medial margin, visible in dorsal view.</paragraph>
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<paragraph id="81ED6AA661D2E989752EA18840C8FB32" pageId="0" pageNumber="205">Male chelipeds equal, longer than pereopods; covered with sparse granulation, unarmed. Dactylus arched in adult males, leaving small gap between fingers, distinctly shorter than palm. Cutting edges with sub-equal teeth in distal half, one distinct proximal tooth in larger males; distal half with light brown colour in fixed specimens. Pereopods short, slender, cylindrical. P2 longest; P3-P5 progressively decreasing in length. Dactylus slightly curved, covered with short setae.</paragraph>
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<paragraph id="257F60D449E05369583A4755516CC0FB" pageId="0" pageNumber="205">Female chelipeds equal, longer than pereopods, slender and smooth. Dactylus arched in adult, shorter than palm, sub-equal teeth in distal half. Pereopods, slender, cylindrical. P2 longest, P3-P5 progressively decreasing in length. Dactylus slightly curved, shorter than propodus, smooth ventrally, with corneous tips.</paragraph>
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<paragraph id="19BDF84CAB29E8E27A874CFB80EBD3FB" pageId="0" pageNumber="205">Male thoracic sternites I-IV fused, broadly triangular, smooth, anterior half declivous in ventral view. Telson fully fitted to cavity, anterior margin smooth.</paragraph>
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<paragraph id="989F769335B3AC320A35394B19A08F3B" pageId="0" pageNumber="205">Female pleonal somites I-VI markedly arched, telson free, transversally oval, with a row of setae on margin and one small spine in first somite. Male pleon rather prominent, pleonal somites I-VI, telson free, somite II with a mesial tubercle. Somite III with slight elevations. Somite VI longest, with a mesial tubercle and a small projection each side.</paragraph>
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</subSubSection>
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<subSubSection id="456C37B20A79BC6E50DA2025AD7D5A1D" pageId="0" pageNumber="205" type="colour in life">
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<paragraph id="381B1BE403564CE7FA7CEA50CB5A95AC" pageId="0" pageNumber="205">Colour in life.</paragraph>
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<paragraph id="EF4AC8E9B16AD8B49E1F7CA79E7ACEF2" pageId="0" pageNumber="205">Light brown, covered with darker hair, first pair of pereopods reddish (Bell 1936b).</paragraph>
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</subSubSection>
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<subSubSection id="71A538FDD0C1B12A536096CA0AC73037" pageId="0" pageNumber="205" type="type locality">
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<paragraph id="76B68FBE31684E5D63B463B7ED6DB770" pageId="0" pageNumber="205">Type locality.</paragraph>
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<paragraph id="A67033743F39D8EB9140E0D8A6E68C3B" pageId="0" pageNumber="205">Costa Rica, Central America, Puerto Potrero, in sand at a depth of 23.7 m.</paragraph>
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</subSubSection>
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<subSubSection id="DE08B78B63093CBA007C7284F897EC26" pageId="0" pageNumber="205" type="distribution">
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<paragraph id="EFCE3B3A7102B4698CF636FF8D938830" pageId="0" pageNumber="205">Geographic distribution.</paragraph>
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<paragraph id="832E7D9F394E258934F3EB6B3B25DAB7" pageId="0" pageNumber="205">
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Eastern Pacific: Mexico (Cape San Lucas), Costa Rica (Guanacaste) and
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<normalizedToken id="C0D39833B535F3F17DD38D1CA1FA49E4" originalValue="Panamá">Panama</normalizedToken>
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(Panama Bay) (Fig.
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<figureCitation id="91070EC01968D5FB52C9803C232FF142" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution Pohleus heptacanthus (Bell, 1836) gen. nov. et comb. nov.; orange circles = distribution based on examined material; green star = type locality and Pohleus septemspinosus (Stimpson, 1871) gen. nov. et comb. nov.; red circles = distribution based on examined material; yellow star = neotype locality." figureDoi="10.3897/zse.96.50360.figure4" httpUri="https://binary.pensoft.net/fig/412093" pageId="0" pageNumber="205">4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="77FAB3541DAFFAF88E2CCD21678461DC" pageId="0" pageNumber="205" type="remarks">
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<paragraph id="CB654733DC16629ACAC72E790AD4A7F4" pageId="0" pageNumber="205">Remarks.</paragraph>
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<paragraph id="CF2828C5B20300F9DCD438B8F0BEF0EB" pageId="0" pageNumber="205">
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Bell (1836) described
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<taxonomicName id="731871231538DACC9D15A789EFEFD3E0" authorityName="Bell" authorityYear="1836" class="Malacostraca" family="Epialtidae" genus="Pericera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericera heptacantha" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacantha">
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<emphasis id="09717DEA78C87A2E779E1865AC2574CD" italics="true" pageId="0" pageNumber="205">Pericera heptacantha</emphasis>
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</taxonomicName>
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, based on two specimens as syntypes, one male and one female. The male syntype is considered lost and the female is deposited in the dry crustacean collection of the Oxford University Museum (OUM 13764) (
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<bibRefCitation id="56D4E4B78FDD70B9FDB6C8BC8AC74F10" DOI="https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" author="Di Mauro, Jr AA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="205" pagination="155 - 182" refId="B16" refString="Di Mauro, Jr AA, 1982. Rediscovery of Professor Thomas Bell's type Crustacea (Brachyura) in the dry crustacean collection of the Zoological Collections, University Museum, Oxford. Zoological Journal of the Linnean Society 76 (2): 155 - 182, DOI: https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" title="Rediscovery of Professor Thomas Bell's type Crustacea (Brachyura) in the dry crustacean collection of the Zoological Collections, University Museum, Oxford." url="https://doi.org/10.1111/j.1096-3642.1982.tb01499.x" volume="76" year="1982">Di Mauro 1982</bibRefCitation>
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). Thus, the female syntype (OUM 13764) is here designated as the lectotype of
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<taxonomicName id="37838CE34FC5E1ABF528187E444103A1" authorityName="Bell" authorityYear="1836" class="Malacostraca" family="Epialtidae" genus="Pericera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericera heptacantha" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacantha">
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<emphasis id="35196C918D8E02BF2ADF686480689C76" italics="true" pageId="0" pageNumber="205">Pericera heptacantha</emphasis>
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</taxonomicName>
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since it is the only specimen from the type series remaining (Fig.
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<figureCitation id="0C4091F60984C75D985FCA1DF3DE7650" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pohleus septemspinosus (Stimpson, 1871) gen. nov. et comb. nov. A, B. female (MNHN IU 2013 - 32682). Pohleus heptacanthus (Bell, 1836) gen. nov. et comb. nov.; C, D. female lectotype (OUM 13764). A, C. habitus; B, D. ventral view. Note the spine on merus of the second pereopod (white arrow). Scale bars: 10 mm." figureDoi="10.3897/zse.96.50360.figure5" httpUri="https://binary.pensoft.net/fig/412094" pageId="0" pageNumber="205">5C, D</figureCitation>
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). The locality Puerto Potrero in Central America probably refers to the Puerto Potrero, Guanacaste, in Costa Rica.
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<bibRefCitation id="B2382A203085DE411E27A453FB868348" DOI="https://doi.org/10.5479/si.03629236.166.i" author="Rathbun, MJ" journalOrPublisher="Bulletin of the United States National Museum" pageId="0" pageNumber="205" pagination="1 - 278" refId="B41" refString="Rathbun, MJ, 1937. The oxystomatous and allied crabs of America. Bulletin of the United States National Museum 166: 1 - 278, DOI: https://doi.org/10.5479/si.03629236.166.i" title="The oxystomatous and allied crabs of America." url="https://doi.org/10.5479/si.03629236.166.i" volume="166" year="1937">Rathbun (1937</bibRefCitation>
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: 136) examining
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<taxonomicName id="0F484A07FFF5D3BD1EBF2959E2A7B467" authorityName="Bell" authorityYear="1855" class="Malacostraca" family="Leucosiidae" genus="Lithadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithadia cumingii" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="cumingii">
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<emphasis id="9D9C46DCDA0E5E26A706D9A06880F0AE" italics="true" pageId="0" pageNumber="205">Lithadia cumingii</emphasis>
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</taxonomicName>
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Bell, 1855, a species described from the same locality by
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<bibRefCitation id="27D3F0378DC45080BE12631122BF9D90" DOI="https://doi.org/10.1111/j.1096-3642.1852.tb00464.x" author="Bell, T" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="205" pagination="277 - 314" refId="B7" refString="Bell, T, 1855. XXXI. Horae carcinologicae, or notices of crustacea. I. A monograph of the Leucosiidae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species. Transactions of the Linnean Society of London 21 (3): 277 - 314, DOI: https://doi.org/10.1111/j.1096-3642.1852.tb00464.x" title="XXXI. Horae carcinologicae, or notices of crustacea. I. A monograph of the Leucosiidae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species." url="https://doi.org/10.1111/j.1096-3642.1852.tb00464.x" volume="21" year="1855">Bell (1855)</bibRefCitation>
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, referred to the type locality as from Potrero, Costa Rica, the same case as for
|
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<taxonomicName id="CD6CDACB1E486EA55533BC5D7F2ACD50" authorityName="gen. nov. et" authorityYear="2020" baseAuthorityName="Bell" baseAuthorityYear="1836" class="Malacostraca" family="Epialtidae" genus="Pohleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pohleus heptacanthus" order="Decapoda" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="heptacanthus">
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<emphasis id="3369F11A638C9867834BFFD7772D30D9" italics="true" pageId="0" pageNumber="205">Pohleus heptacanthus</emphasis>
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</taxonomicName>
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gen. nov. et comb. nov.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |