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<document id="B7EBC12D9C070B12779157533FF0C537" ID="10.11646/zootaxa.4254.4.6SLASH10857" ID-CLB-Dataset="33198" ID-DOI="10.11646/zootaxa.4254.4.6" ID-GBIF-Dataset="da672c83-e552-4115-bccb-144bdea41b05" ID-ISSN="1175-5326" ID-Zenodo-Dep="556070" ID-ZooBank="CE64E8D5-BB9C-4633-92B2-94F6B6889164" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492588944747" checkinUser="plazi" docAuthor="Kottelat, Maurice" docDate="2017" docId="B839F52AAC15FFD652A220671343FC09" docLanguage="en" docName="zootaxa.4254.4.6SLASH10857.pdf" docOrigin="Zootaxa 4254 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speolabeo Kottelat, 2017, new genus" docType="treatment" docVersion="8" lastPageNumber="498" masterDocId="44008D52AC11FFD35235210D117EFF89" masterDocTitle="Speolabeo, a new genus name for the cave fish Bangana musaei (Teleostei: Cyprinidae)" masterLastPageNumber="499" masterPageNumber="493" pageNumber="497" updateTime="1698438967479" updateUser="ExternalLinkService">
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<mods:title id="39408E11CF79C33C5554E8254771C3DB">Speolabeo, a new genus name for the cave fish Bangana musaei (Teleostei: Cyprinidae)</mods:title>
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<mods:name id="B1A9B42C2C2EB280F8F07853A3B95D14" type="personal">
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<mods:namePart id="6A1132B74C04E82E937CD384AF815932">Kottelat, Maurice</mods:namePart>
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<mods:date id="EEDC5582E772380E9908750C8D9BBAEA">2017</mods:date>
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<mods:classification id="A8168E414D906A320A7AE032B6254AD5">journal article</mods:classification>
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<treatment id="B839F52AAC15FFD652A220671343FC09" ID-DOI="http://doi.org/10.5281/zenodo.6017858" ID-GBIF-Taxon="128539725" ID-Zenodo-Dep="6017858" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09" httpUri="http://treatment.plazi.org/id/B839F52AAC15FFD652A220671343FC09" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
<subSubSection id="788A17B7AC15FFD752A2206710DDFE0D" box="[151,419,362,388]" pageId="4" pageNumber="497" type="nomenclature">
<paragraph id="302F443CAC15FFD752A2206710DDFE0D" blockId="4.[151,419,362,388]" box="[151,419,362,388]" pageId="4" pageNumber="497">
<heading id="6B67F350AC15FFD752A2206710DDFE0D" bold="true" box="[151,419,362,388]" fontSize="11" level="1" pageId="4" pageNumber="497" reason="1">
<emphasis id="02E4982EAC15FFD752A2206710DDFE0D" bold="true" box="[151,419,362,388]" pageId="4" pageNumber="497">
<taxonomicName id="F7903FBFAC15FFD752A22067106DFE0D" ID-CoL="644HN" box="[151,275,362,388]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="02E4982EAC15FFD752A22067106DFE0D" bold="true" box="[151,275,362,388]" italics="true" pageId="4" pageNumber="497">Speolabeo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="19D72555AC15FFD75314206710DDFE0D" box="[289,419,362,388]" pageId="4" pageNumber="497" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="788A17B7AC15FFD752A220BC13A9FE4E" box="[151,727,433,455]" pageId="4" pageNumber="497" type="materials_examined">
<paragraph id="302F443CAC15FFD752A220BC13A9FE4E" blockId="4.[151,727,433,455]" box="[151,727,433,455]" pageId="4" pageNumber="497">
<emphasis id="02E4982EAC15FFD752A220BC105CFE4E" bold="true" box="[151,290,433,455]" pageId="4" pageNumber="497">Type species.</emphasis>
<taxonomicName id="F7903FBFAC15FFD7531C20BF13ADFE4E" authority="Kottelat &amp; Steiner, 2011" authorityName="Kottelat &amp; Steiner" authorityYear="2011" box="[297,723,433,455]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC15FFD7531C20BF10ADFE4E" box="[297,467,434,455]" italics="true" pageId="4" pageNumber="497">Bangana musaei</emphasis>
<bibRefCitation id="540139CDAC15FFD753EC20BC13ADFE4E" author="Kottelat" box="[473,723,433,455]" pageId="4" pageNumber="497" refString="Kottelat, M. &amp; Steiner, H. (2011) Bangana musaei, a new cave fish from central Laos (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 21 (4), 313 - 322. [2010]" type="journal article" year="2011">Kottelat &amp; Steiner, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="788A17B7AC15FFD752A220F91017FDF1" pageId="4" pageNumber="497" type="etymology">
<paragraph id="302F443CAC15FFD752A220F91017FDF1" blockId="4.[151,1437,500,2037]" pageId="4" pageNumber="497">
<emphasis id="02E4982EAC15FFD752A220F91060FD84" bold="true" box="[151,286,500,525]" pageId="4" pageNumber="497">Etymology.</emphasis>
The name is formed on the classical Greek noun
<emphasis id="02E4982EAC15FFD7517120F912FAFD84" box="[836,900,500,525]" italics="true" pageId="4" pageNumber="497">σπέΟς</emphasis>
(speos), which means cave, cavern, excavation; and
<emphasis id="02E4982EAC15FFD752F0231A107DFDB9" box="[197,259,535,560]" italics="true" pageId="4" pageNumber="497">labeo</emphasis>
, originally a Latin noun meaning a man with thick lips, which in ichthyology became a genus name, then a vernacular name as well as the base of the family-group name as a subfamily
<taxonomicName id="F7903FBFAC15FFD7560C233115C2FDDC" box="[1081,1212,572,597]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="subFamily" subFamily="Labeoninae">Labeoninae</taxonomicName>
or tribe Labeonini. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="788A17B7AC15FFD752F223891510F971" pageId="4" pageNumber="497" type="diagnosis">
<paragraph id="302F443CAC15FFD752F2238913F4FCFD" blockId="4.[151,1437,500,2037]" pageId="4" pageNumber="497">
<emphasis id="02E4982EAC15FFD752F22389103FFD14" bold="true" box="[199,321,644,669]" pageId="4" pageNumber="497">Diagnosis.</emphasis>
The diagnostic characters of the genus are the same as those of the only included species.
<taxonomicName id="F7903FBFAC15FFD752A223AA101CFD49" box="[151,354,679,704]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC15FFD752A223AA101CFD49" box="[151,354,679,704]" italics="true" pageId="4" pageNumber="497">Speolabeo musaei</emphasis>
</taxonomicName>
is a cave fish showing characters common to many hypogean fishes (and other animals), which are unique among the species earlier placed in
<taxonomicName id="F7903FBFAC15FFD7508723C01269FD6D" box="[690,791,717,740]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="02E4982EAC15FFD7508723C01269FD6D" box="[690,791,717,740]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D72555AC15FFD7512A23C1123DFD6C" box="[799,835,716,741]" pageId="4" pageNumber="497" sensu="lato">s.l.</taxonomicNameLabel>
: absence of eyes and whitish colour (pinkish in life) due to the lack of pigmentation. These characters are cave adaptations that have appeared in parallel in numerous fish species living in caves, in aquifers, in crevices, under stones as well as in deep rapids (
<bibRefCitation id="540139CDAC15FFD756A52219142EFCA4" author="Proudlove" box="[1168,1360,788,813]" pageId="4" pageNumber="497" refString="Proudlove, G. S. (2006) Subterranean fishes of the world. An account of the subterranean (hypogean) fishes described up to 2003 with a bibliography 1541 - 2004. International Society for Subterranean Biology, Moulis, xviii + 300 pp." type="book chapter" year="2006">Proudlove, 2006</bibRefCitation>
). They are useful to identify the genus, but by themselves do not justify the recognition of a distinct genus (see, e.g.,
<bibRefCitation id="540139CDAC15FFD752A22251101CFCFC" author="Greenwood" box="[151,354,860,885]" pageId="4" pageNumber="497" refString="Greenwood, P. H. (1976) A new and eyeless cobitid fish (Pisces, Cypriniformes) from the Zagros Mountains, Iran. Journal of Zoology, 180, 129 - 137." type="journal article" year="1976">Greenwood, 1976</bibRefCitation>
,
<bibRefCitation id="540139CDAC15FFD7535822511303FCFD" author="Banister" box="[365,637,860,885]" pageId="4" pageNumber="497" refString="Banister, K. E. &amp; Bunni, M. K. (1980) A new blind cyprinid fish from Iraq. Bulletin of the British Museum (Natural History), Zoology, 38 (3), 151 - 158." type="journal article" year="1980">Banister &amp; Bunni, 1980</bibRefCitation>
).
</paragraph>
<paragraph id="302F443CAC15FFD752F2228D155FFAAC" blockId="4.[151,1437,500,2037]" pageId="4" pageNumber="497">
However, there are other characters not related to adaptation to life in hypogean habitats that distinguish
<taxonomicName id="F7903FBFAC15FFD752A222A91072FC34" box="[151,268,932,957]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="02E4982EAC15FFD752A222A91072FC34" box="[151,268,932,957]" italics="true" pageId="4" pageNumber="497">Speolabeo</emphasis>
</taxonomicName>
from
<taxonomicName id="F7903FBFAC15FFD7536422A810C8FC35" box="[337,438,933,956]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD7536422A810C8FC35" box="[337,438,933,956]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
,
<taxonomicName id="F7903FBFAC15FFD753F022A9135AFC34" box="[453,548,932,957]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD753F022A9135AFC34" box="[453,548,932,957]" italics="true" pageId="4" pageNumber="497">Altigena</emphasis>
</taxonomicName>
and the
<emphasis id="02E4982EAC15FFD750B022A8125EFC34" box="[645,800,932,957]" italics="true" pageId="4" pageNumber="497">'B.' lemassoni</emphasis>
-group: the dorsal fin has 7–8½ branched dorsal-fin rays (vs. 10–12½); the origin of the pelvic fin is located at the vertical between the first unbranched and first branched dorsal-fin rays (vs. at the vertical through 3rd to 6th branched rays); there are no tubercles on the snout (vs. tubercles present); the edge of the upper jaw is distinct, very narrow, adnate to the upper lip and separated from it by a narrow and very shallow furrow (vs. upper jaw entirely enclosed in upper lip); the distal edge of the dorsal fin is straight or slightly concave (vs. conspicuously concave in
<taxonomicName id="F7903FBFAC15FFD75170255A12DAFBF9" box="[837,932,1111,1136]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD75170255A12DAFBF9" box="[837,932,1111,1136]" italics="true" pageId="4" pageNumber="497">Altigena</emphasis>
</taxonomicName>
and
<taxonomicName id="F7903FBFAC15FFD751E92554153FFBF9" box="[988,1089,1113,1136]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD751E92554153FFBF9" box="[988,1089,1113,1136]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
); 35–37 lateral-line scales (vs.
<quantity id="F768E9D9AC15FFD752A225711183FB1C" box="[151,253,1148,1173]" metricMagnitude="0" metricUnit="m" metricValue="1.1049" metricValueMax="1.1938" metricValueMin="1.016" pageId="4" pageNumber="497" unit="in" value="43.5" valueMax="47.0" valueMin="40.0">40–47 in</quantity>
the
<emphasis id="02E4982EAC15FFD75300257010AAFB1C" box="[309,468,1148,1173]" italics="true" pageId="4" pageNumber="497">'B.' lemassoni</emphasis>
-group,
<quantity id="F768E9D9AC15FFD75005257113E8FB1C" box="[560,662,1148,1173]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.0922" metricValueMin="1.016" pageId="4" pageNumber="497" unit="in" value="41.5" valueMax="43.0" valueMin="40.0">40–43 in</quantity>
<taxonomicName id="F7903FBFAC15FFD7509425701278FB1D" box="[673,774,1149,1172]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD7509425701278FB1D" box="[673,774,1149,1172]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
). The postlabial groove is restricted to the corner of the mouth (vs. complete in
<taxonomicName id="F7903FBFAC15FFD753AA25921080FB31" box="[415,510,1183,1208]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD753AA25921080FB31" box="[415,510,1183,1208]" italics="true" pageId="4" pageNumber="497">Altigena</emphasis>
</taxonomicName>
, medially interrupted in
<taxonomicName id="F7903FBFAC15FFD7512725AC1209FB31" box="[786,887,1185,1208]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD7512725AC1209FB31" box="[786,887,1185,1208]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
and the
<emphasis id="02E4982EAC15FFD751E025AD150EFB31" box="[981,1136,1183,1208]" italics="true" pageId="4" pageNumber="497">'B.' lemassoni</emphasis>
-group); however, in CMK 26632, two small specimens from a separate cave, a very shallow groove is present. This could be geographical variation or related to the smaller size. Also, the smaller specimens were fixed in ethanol, and the soft tissues have collapsed and are not in an optimal condition for the description of soft anatomy.
</paragraph>
<paragraph id="302F443CAC15FFD752F224221510F971" blockId="4.[151,1437,500,2037]" pageId="4" pageNumber="497">
<bibRefCitation id="540139CDAC15FFD752F2242210B5FAC1" author="Zhang" box="[199,459,1327,1352]" pageId="4" pageNumber="497" refString="Zhang, E. &amp; Chen, Y. - Y. (2006) Revised diagnosis of the genus Bangana Hamilton, 1822 (Pisces: Cyprinidae), with taxonomic and nomenclatural notes on the Chinese species. Zootaxa, 1281, 41 - 54." type="journal article" year="2006">Zhang &amp; Chen (2006)</bibRefCitation>
diagnosed
<taxonomicName id="F7903FBFAC15FFD75066243C13C6FAC1" box="[595,696,1329,1352]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="02E4982EAC15FFD75066243C13C6FAC1" box="[595,696,1329,1352]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D72555AC15FFD750F62422139AFAC1" box="[707,740,1327,1352]" pageId="4" pageNumber="497" sensu="lato">s.l.</taxonomicNameLabel>
with, among others, the following characters related to the mouth morphology: the upper jaw is entirely enclosed in the upper lip, whose median part is covered by the rostral fold; the rostral fold is thick, pendulous, separated from the upper lip by a deep groove, not continuous with the lower lip; the upper lip is continuous with the lower lip; the lower lip is separated from the lower jaw by a groove, its anterior margin is free and has numerous papillae on the dorsal surface; the lower jaw is cornified, with a sharp edge. All these structures are present in
<taxonomicName id="F7903FBFAC15FFD7505724E913A9FA74" box="[610,727,1508,1533]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="02E4982EAC15FFD7505724E913A9FA74" box="[610,727,1508,1533]" italics="true" pageId="4" pageNumber="497">Speolabeo</emphasis>
</taxonomicName>
, except that the upper jaw is not enclosed in the upper lip but adnate behind it (a condition overlooked by
<bibRefCitation id="540139CDAC15FFD750AF270A12C6F9A9" author="Kottelat" box="[666,952,1543,1568]" pageId="4" pageNumber="497" refString="Kottelat, M. &amp; Steiner, H. (2011) Bangana musaei, a new cave fish from central Laos (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 21 (4), 313 - 322. [2010]" type="journal article" year="2011">Kottelat &amp; Steiner, 2011</bibRefCitation>
). The rostral fold is present but, unlike in
<taxonomicName id="F7903FBFAC15FFD752A227201182F9CD" box="[151,252,1581,1604]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD752A227201182F9CD" box="[151,252,1581,1604]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
,
<taxonomicName id="F7903FBFAC15FFD7533F27211017F9CC" box="[266,361,1580,1605]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD7533F27211017F9CC" box="[266,361,1580,1605]" italics="true" pageId="4" pageNumber="497">Altigena</emphasis>
</taxonomicName>
and the
<emphasis id="02E4982EAC15FFD753FC2720131AF9CC" box="[457,612,1580,1605]" italics="true" pageId="4" pageNumber="497">'B.' lemassoni</emphasis>
group, it is not thick and fleshy. The upper and lower lips are continuous in
<taxonomicName id="F7903FBFAC15FFD7528127421056F9E1" box="[180,296,1615,1640]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="02E4982EAC15FFD7528127421056F9E1" box="[180,296,1615,1640]" italics="true" pageId="4" pageNumber="497">Speolabeo</emphasis>
</taxonomicName>
. The lower lip is separated from the lower jaw by a shallow groove and leaves most of the lower jaw exposed; the dorsal surface of the lip is narrow, not covered by papillae, and its anterior margin is crenulated. The median part of the lower lip is very thin and cannot easily be distinguished from the skin of the gular area; if it were not for the crenulated edge and the groove separating it from the jaw, the lip would appear as if widely interrupted and restricted to the corners of the mouth. The lower jaw has a cornified but blunt edge.
</paragraph>
</subSubSection>
<subSubSection id="788A17B7AC15FFD652F2260910D2FD21" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497" type="discussion">
<paragraph id="302F443CAC15FFD752F22609145FF801" blockId="4.[151,1437,500,2037]" pageId="4" pageNumber="497">
<emphasis id="02E4982EAC15FFD752F226091045F894" bold="true" box="[199,315,1796,1821]" pageId="4" pageNumber="497">Remarks.</emphasis>
Since its original description,
<taxonomicName id="F7903FBFAC15FFD7509226081269F895" box="[679,791,1797,1820]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC15FFD7509226081269F895" box="[679,791,1797,1820]" italics="true" pageId="4" pageNumber="497">S. musaei</emphasis>
</taxonomicName>
has been observed and collected in other caves in the Khammouan Karst near the type locality (H. Steiner, pers. comm.). These additional specimens do not show differences from the original description except as noted under the diagnosis for the postlabial groove. For the time being it does not seem worthwhile to speculate on the possible phylogenetic relationships of
<taxonomicName id="F7903FBFAC15FFD7569326621464F801" box="[1190,1306,1903,1928]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="02E4982EAC15FFD7569326621464F801" box="[1190,1306,1903,1928]" italics="true" pageId="4" pageNumber="497">Speolabeo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="302F443CAC15FFD652F2269915FFFE94" blockId="4.[151,1437,500,2037]" lastBlockId="5.[151,1437,151,896]" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
<taxonomicName id="F7903FBFAC15FFD752F2269910ECF825" box="[199,402,1940,1965]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC15FFD752F2269910ECF825" box="[199,402,1940,1965]" italics="true" pageId="4" pageNumber="497">Speolabeo musaei</emphasis>
</taxonomicName>
was originally placed in
<taxonomicName id="F7903FBFAC15FFD7509B2698126DF825" box="[686,787,1941,1964]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC15FFD7509B2698126DF825" box="[686,787,1941,1964]" italics="true" pageId="4" pageNumber="497">Bangana</emphasis>
</taxonomicName>
by default, because of the inability to place it in a known genus and the unwillingness to establish a distinct genus for a species whose most salient characters were related to its specialised habitat. However, a number of the characters used in the diagnosis (especially the relative positions of the dorsal and pelvic fins and the number of dorsal-fin rays) cannot be the result of cave adaptations. After the 'dismantling' of
<taxonomicName id="F7903FBFAC14FFD6536421B010C8FF5D" box="[337,438,189,212]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="02E4982EAC14FFD6536421B010C8FF5D" box="[337,438,189,212]" italics="true" pageId="5" pageNumber="498">Bangana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D72555AC14FFD653F421B1109CFF5C" box="[449,482,188,213]" pageId="5" pageNumber="498" sensu="lato">s.l.</taxonomicNameLabel>
, based on external characters
<emphasis id="02E4982EAC14FFD6517621B012BFFF5D" box="[835,961,189,212]" italics="true" pageId="5" pageNumber="498">
'B.'
<taxonomicName id="F7903FBFAC14FFD6514721B012BFFF5D" box="[882,961,189,212]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="musaei">musaei</taxonomicName>
</emphasis>
cannot be placed in any of the resulting genera, or in any other named labeonine genera. It is therefore recognised here as a distinct genus; keeping it in any of the mentioned genera would be just as artificial as its placement earlier in
<taxonomicName id="F7903FBFAC14FFD651C120081527FE95" box="[1012,1113,261,284]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="02E4982EAC14FFD651C120081527FE95" box="[1012,1113,261,284]" italics="true" pageId="5" pageNumber="498">Bangana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D72555AC14FFD65655200915FFFE94" box="[1120,1153,260,285]" pageId="5" pageNumber="498" sensu="lato">s.l.</taxonomicNameLabel>
</paragraph>
<paragraph id="302F443CAC14FFD652F2202510D2FD21" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="498">
In some cases the relationships of a cave species with one or more epigean species are obvious (e.g.
<taxonomicName id="F7903FBFAC14FFD6570020251196FEEC" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="jordani">
<emphasis id="02E4982EAC14FFD6570020251196FEEC" italics="true" pageId="5" pageNumber="498">Astyanax jordani</emphasis>
</taxonomicName>
, see
<bibRefCitation id="540139CDAC14FFD6531320411099FEEC" author="Proudlove" box="[294,487,332,357]" pageId="5" pageNumber="498" refString="Proudlove, G. S. (2006) Subterranean fishes of the world. An account of the subterranean (hypogean) fishes described up to 2003 with a bibliography 1541 - 2004. International Society for Subterranean Biology, Moulis, xviii + 300 pp." type="book chapter" year="2006">Proudlove, 2006</bibRefCitation>
). In other cases there is apparently no closely related epigean species (at least not based on external characters) (e.g.
<taxonomicName id="F7903FBFAC14FFD65015207D1562FE01" authority="Kottelat, 1988" authorityName="Kottelat" authorityYear="1988" box="[544,1052,367,392]" class="Actinopterygii" family="Balitoridae" genus="Cryptotora" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="thamicola">
<emphasis id="02E4982EAC14FFD65015207D126AFE01" box="[544,788,367,392]" italics="true" pageId="5" pageNumber="498">Cryptotora thamicola</emphasis>
,
<bibRefCitation id="540139CDAC14FFD65116206212B9FE01" author="Kottelat" box="[803,967,367,392]" pageId="5" pageNumber="498" refString="Kottelat, M. (1988) Two species of cavefishes from Northern Thailand in the genera Nemacheilus and Homaloptera (Osteichthyes: Homalopteridae). Records of the Australian Museum, 40 (3 - 4), 225 - 231." type="journal article" year="1988">Kottelat, 1988</bibRefCitation>
,
<bibRefCitation id="540139CDAC14FFD651E620621562FE01" author="Kottelat" box="[979,1052,367,392]" pageId="5" pageNumber="498" refString="Kottelat, M. (1998 b) Homaloptera yuwonoi, a new species of hillstream loach from Borneo, with a new generic name for H. thamicola (Teleostei: Balitoridae). Ichthyological Exploration of Freshwaters, 9 (3), 267 - 272." type="journal article" year="1998" yearSuffix="b">1998b</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="F7903FBFAC14FFD6561D207C11AAFE25" authority="Kottelat, 2012" authorityName="Kottelat" authorityYear="2012" class="Actinopterygii" family="Nemacheilidae" genus="Draconectes" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="narinosus">
<emphasis id="02E4982EAC14FFD6561D207C1452FE01" box="[1064,1324,369,392]" italics="true" pageId="5" pageNumber="498">Draconectes narinosus</emphasis>
,
<bibRefCitation id="540139CDAC14FFD6570E206211AAFE25" author="Kottelat" pageId="5" pageNumber="498" refString="Kottelat, M. (2012) Draconectes narinosus, a new genus and species of cave fish from an island of Halong Bay, Vietnam (Teleostei: Nemacheilidae). Revue Suisse de Zoologie, 119 (3), 341 - 349." type="journal article" year="2012">Kottelat, 2012</bibRefCitation>
</taxonomicName>
). Noteworthy is that two other fish species are known from the caves of the
<collectingRegion id="F2548ADEAC14FFD65608209915B0FE24" box="[1085,1230,404,429]" country="Laos" name="Khammouan" pageId="5" pageNumber="498">Khammouan</collectingRegion>
Karst, for both of which it has not been possible to identify close relatives. The possible relationships of
<taxonomicName id="F7903FBFAC14FFD656FD20BA13FFFE7D" authority="Kottelat &amp; Brehier, 1999" authorityName="Kottelat &amp; Brehier" authorityYear="1999" class="Actinopterygii" family="Cyprinidae" genus="Troglocyclocheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="khammouanensis">
<emphasis id="02E4982EAC14FFD656FD20BA1026FE7C" italics="true" pageId="5" pageNumber="498">Troglocyclocheilus khammouanensis</emphasis>
<bibRefCitation id="540139CDAC14FFD6535420D113FFFE7D" author="Kottelat" box="[353,641,476,501]" pageId="5" pageNumber="498" refString="Kottelat, M. &amp; Brehier, F. (1999) Troglocyclocheilus khammouanensis, a new genus and species of cave fish from the Khammouan karst, Laos (Teleostei: Cyprinidae). Revue Suisse de Zoologie, 106 (2), 347 - 359. https: // doi. org / 10.5962 / bhl. part. 80084" type="journal article" year="1999">Kottelat &amp; Bréhier, 1999</bibRefCitation>
</taxonomicName>
are unclear.
<taxonomicName id="F7903FBFAC14FFD6512D20D11413FE7D" authority="Vidthayanon &amp; Jaruthanin, 2002" authorityName="Vidthayanon &amp; Jaruthanin" authorityYear="2002" box="[792,1389,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="kaysonei">
<emphasis id="02E4982EAC14FFD6512D20D11293FE7C" box="[792,1005,476,501]" italics="true" pageId="5" pageNumber="498">Schistura kaysonei</emphasis>
<bibRefCitation id="540139CDAC14FFD651C220D11413FE7D" author="Vidthayanon" box="[1015,1389,476,501]" pageId="5" pageNumber="498" refString="Vidthayanon, C. &amp; Jaruthanin, K. (2002) Schistura kaysonei, (Teleostei: Balitoridae) a new cave fish from the Khammouan karst, Laos PDR. Aqua, Journal of Ichthyology and Aquatic Biology, 6 (1), 17 - 20." type="journal article" year="2002">Vidthayanon &amp; Jaruthanin, 2002</bibRefCitation>
</taxonomicName>
has characters such as a reduced number of caudal-fin rays and a nasal barbel unknown in any
<taxonomicName id="F7903FBFAC14FFD6568120F21460FD91" box="[1204,1310,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="genus">
<emphasis id="02E4982EAC14FFD6568120F21460FD91" box="[1204,1310,511,536]" italics="true" pageId="5" pageNumber="498">Schistura</emphasis>
</taxonomicName>
(and other
<taxonomicName id="F7903FBFAC14FFD652A22329103FFDB4" box="[151,321,548,573]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
) of the Xe Bangfai and adjacent drainages (
<bibRefCitation id="540139CDAC14FFD651052329129CFDB5" author="Kottelat" box="[816,994,548,573]" pageId="5" pageNumber="498" refString="Kottelat, M. (1998 a) Fishes of the Nam Theun and Xe Bangfai basins, Laos, with diagnoses of twenty-two new species (Teleostei: Cyprinidae, Balitoridae, Cobitidae, Coiidae and Odontobutidae). Ichthyological Exploration of Freshwaters, 9 (1), 1 - 128." type="journal article" year="1998" yearSuffix="a">Kottelat, 1998a</bibRefCitation>
,
<bibRefCitation id="540139CDAC14FFD651D823291556FDB5" author="Kottelat" box="[1005,1064,548,572]" pageId="5" pageNumber="498" refString="Kottelat, M. (2000) Diagnoses of a new genus and 64 new species of fishes from Laos (Teleostei: Cyprinidae, Balitoridae, Bagridae, Syngnathidae, Chaudhuriidae and Tetraodontidae). Journal of South Asian Natural History, 5 (1), 37 - 82." type="journal article" year="2000">2000</bibRefCitation>
,
<bibRefCitation id="540139CDAC14FFD6560123291511FDB5" author="Kottelat" box="[1076,1135,548,572]" pageId="5" pageNumber="498" refString="Kottelat, M. (2001) Fishes of Laos. Wildlife Heritage Trust, Colombo, 198 pp." type="book" year="2001">2001</bibRefCitation>
,
<bibRefCitation id="540139CDAC14FFD6564F232915BDFDB4" author="Kottelat" box="[1146,1219,548,573]" pageId="5" pageNumber="498" refString="Kottelat, M. (2016) The fishes of the Nam Theun and Xe Bangfai drainages, Laos. Hydroecologie Appliquee, 19, 271 - 320. https: // doi. org / 10.1051 / hydro / 2015005" type="journal article" year="2016" yearSuffix="b">2016b</bibRefCitation>
, unpubl. obs.). The
<collectingRegion id="F2548ADEAC14FFD652A2234A1056FDE9" box="[151,296,583,608]" country="Laos" name="Khammouan" pageId="5" pageNumber="498">Khammouan</collectingRegion>
Karst is continued across the border eastwards in
<collectingCountry id="488704ACAC14FFD6516D234A12C4FDE9" box="[856,954,583,608]" name="Vietnam" pageId="5" pageNumber="498">Vietnam</collectingCountry>
; this area is poorly known ichthyologically and relatives of
<taxonomicName id="F7903FBFAC14FFD653562361134BFD0D" box="[355,565,620,645]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC14FFD653562361134BFD0D" box="[355,565,620,645]" italics="true" pageId="5" pageNumber="498">Speolabeo musaei</emphasis>
</taxonomicName>
,
<taxonomicName id="F7903FBFAC14FFD650792361128EFD0C" box="[588,1008,620,645]" class="Actinopterygii" family="Cyprinidae" genus="Troglocyclocheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="khammouanensis">
<emphasis id="02E4982EAC14FFD650792361128EFD0C" box="[588,1008,620,645]" italics="true" pageId="5" pageNumber="498">Troglocyclocheilus khammouanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="F7903FBFAC14FFD656032361146FFD0C" box="[1078,1297,620,645]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="kaysonei">
<emphasis id="02E4982EAC14FFD656032361146FFD0C" box="[1078,1297,620,645]" italics="true" pageId="5" pageNumber="498">Schistura kaysonei</emphasis>
</taxonomicName>
may await discovery in this system.
</paragraph>
</subSubSection>
<subSubSection id="788A17B7AC14FFD652F223B91343FC09" pageId="5" pageNumber="498" type="materials_examined">
<paragraph id="302F443CAC14FFD652F223B91343FC09" blockId="5.[151,1437,151,896]" pageId="5" pageNumber="498">
<emphasis id="02E4982EAC14FFD652F223B910CEFD44" bold="true" box="[199,432,692,717]" pageId="5" pageNumber="498">Material examined.</emphasis>
<taxonomicName id="F7903FBFAC14FFD6538323B913FFFD45" box="[438,641,692,717]" class="Actinopterygii" family="Cyprinidae" genus="Speolabeo" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="musaei">
<emphasis id="02E4982EAC14FFD6538323B913FFFD45" box="[438,641,692,717]" italics="true" pageId="5" pageNumber="498">Speolabeo musaei</emphasis>
</taxonomicName>
, CMK 20111, 1 paratype, 70.1 mm SL;
<materialsCitation id="80F84E61AC14FFD6567123B91290FD79" ID-GBIF-Occurrence="1457229208" country="Laos" location="Xe Bangfai" pageId="5" pageNumber="498" specimenCount="1">
CMK 26632, 2, 41.5–57.7 mm SL;
<collectingCountry id="488704ACAC14FFD652FE23D51078FD66" box="[203,262,728,751]" name="Laos" pageId="5" pageNumber="498">Laos</collectingCountry>
:
<location id="354F12E7AC14FFD6532023D510E3FD79" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD6532023D510E3FD79" box="[277,413,727,752]" country="Laos" name="Xe Bangfai" pageId="5" pageNumber="498">Xe Bangfai</location>
, caves near entrance of tunnel.—
<taxonomicName id="F7903FBFAC14FFD6511D23D41290FD79" box="[808,1006,727,752]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="devdevi">
<emphasis id="02E4982EAC14FFD6511D23D41290FD79" box="[808,1006,727,752]" italics="true" pageId="5" pageNumber="498">Bangana devdevi</emphasis>
</taxonomicName>
</materialsCitation>
,
<materialsCitation id="80F84E61AC14FFD6563423D51300FC9D" ID-GBIF-Occurrence="1457229209" country="Thailand" location="Salween River" pageId="5" pageNumber="498" specimenCount="1">
CMK 16257, 1;
<collectingCountry id="488704ACAC14FFD656F323DA1450FD79" box="[1222,1326,727,752]" name="Thailand" pageId="5" pageNumber="498">Thailand</collectingCountry>
:
<location id="354F12E7AC14FFD6570823DA11A8FC9C" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD6570823DA11A8FC9C" country="Thailand" name="Salween River" pageId="5" pageNumber="498">Salween River</location>
.—
<taxonomicName id="F7903FBFAC14FFD652C323F110CCFC9C" box="[246,434,764,789]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="elegans">
<emphasis id="02E4982EAC14FFD652C323F110CCFC9C" box="[246,434,764,789]" italics="true" pageId="5" pageNumber="498">Altigena elegans</emphasis>
</taxonomicName>
, CMK 191864, 4
</materialsCitation>
;
<materialsCitation id="80F84E61AC14FFD650BE23F012ADFC9C" ID-GBIF-Occurrence="1457229210" box="[651,979,764,789]" country="Laos" location="Nam Theun watershed" pageId="5" pageNumber="498" specimenCount="1">
<collectingCountry id="488704ACAC14FFD650BE23F013B8FC9D" box="[651,710,765,788]" name="Laos" pageId="5" pageNumber="498">Laos</collectingCountry>
:
<location id="354F12E7AC14FFD650E423F012ADFC9C" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD650E423F012ADFC9C" box="[721,979,764,789]" country="Laos" name="Nam Theun watershed" pageId="5" pageNumber="498">Nam Theun watershed</location>
</materialsCitation>
.—
<taxonomicName id="F7903FBFAC14FFD651C123F115EAFC9C" box="[1012,1172,764,789]" family="Cyprinidae" genus="Altigena" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="lippa">
<emphasis id="02E4982EAC14FFD651C123F115EAFC9C" box="[1012,1172,764,789]" italics="true" pageId="5" pageNumber="498">Altigena lippa</emphasis>
</taxonomicName>
, CMK 21050, 5;
<materialsCitation id="80F84E61AC14FFD6576B23F012B0FCB1" ID-GBIF-Occurrence="1457229212" country="Laos" location="Nam Ou watershed" pageId="5" pageNumber="498" specimenCount="1">
<collectingCountry id="488704ACAC14FFD6576B23F014E7FC9D" box="[1374,1433,765,788]" name="Laos" pageId="5" pageNumber="498">Laos</collectingCountry>
:
<location id="354F12E7AC14FFD652A2222D10ECFCB1" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD652A2222D10ECFCB1" box="[151,402,799,824]" country="Laos" name="Nam Ou watershed" pageId="5" pageNumber="498">Nam Ou watershed</location>
.—
<emphasis id="02E4982EAC14FFD65387222D135DFCBE" box="[434,547,800,823]" italics="true" pageId="5" pageNumber="498">'Bangana'</emphasis>
aff.
<emphasis id="02E4982EAC14FFD6504222121399FCB1" box="[631,743,799,824]" italics="true" pageId="5" pageNumber="498">lemassoni</emphasis>
, CMK 25784, 1
</materialsCitation>
,
<materialsCitation id="80F84E61AC14FFD651D9222D10A7FCD4" ID-GBIF-Occurrence="1457229214" country="Laos" location="Nam Ma" pageId="5" pageNumber="498" specimenCount="1">
CMK 25804, 1;
<collectingCountry id="488704ACAC14FFD656E1222D1471FCBE" box="[1236,1295,800,823]" name="Laos" pageId="5" pageNumber="498">Laos</collectingCountry>
:
<location id="354F12E7AC14FFD6571C222D14E2FCBE" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD6571C222D14E2FCBE" box="[1321,1436,800,823]" country="Laos" name="Nam Ma" pageId="5" pageNumber="498">Nam Ma</location>
drainage.—
<taxonomicName id="F7903FBFAC14FFD6532F224910A7FCD4" box="[282,473,836,861]" class="Actinopterygii" family="Cyprinidae" genus="Incisilabeo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="behri">
<emphasis id="02E4982EAC14FFD6532F224910A7FCD4" box="[282,473,836,861]" italics="true" pageId="5" pageNumber="498">Incisilabeo behri</emphasis>
</taxonomicName>
</materialsCitation>
,
<materialsCitation id="80F84E61AC14FFD653D222491343FC09" ID-GBIF-Occurrence="1457229216" country="Laos" county="Steiner" location="Mekong" municipality="Kottelat" pageId="5" pageNumber="498" specimenCount="1">
CMK 15882, 1;
<collectingCountry id="488704ACAC14FFD65095224813A5FCD5" box="[672,731,837,860]" name="Laos" pageId="5" pageNumber="498">Laos</collectingCountry>
:
<location id="354F12E7AC14FFD650D32249123DFCD4" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD650D32249123DFCD4" box="[742,835,836,861]" country="Laos" county="Steiner" municipality="Kottelat" name="Mekong" pageId="5" pageNumber="498">Mekong</location>
at
<location id="354F12E7AC14FFD651522249128DFCD4" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD651522249128DFCD4" box="[871,1011,836,861]" country="Laos" county="Steiner" municipality="Kottelat" name="Khone Falls" pageId="5" pageNumber="498">Khone Falls</location>
. Additional material listed in
<location id="354F12E7AC14FFD65777224914E3FCD4" LSID="urn:lsid:plazi:treatment:B839F52AAC15FFD652A220671343FC09:354F12E7AC14FFD65777224914E3FCD4" box="[1346,1437,836,861]" country="Laos" county="Steiner" municipality="Kottelat" name="Kottelat" pageId="5" pageNumber="498">Kottelat</location>
(1998) and
<bibRefCitation id="540139CDAC14FFD6532D226A1346FC09" author="Kottelat" box="[280,568,871,896]" pageId="5" pageNumber="498" refString="Kottelat, M. &amp; Steiner, H. (2011) Bangana musaei, a new cave fish from central Laos (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 21 (4), 313 - 322. [2010]" type="journal article" year="2011">
<collectingMunicipality id="D04BDE46AC14FFD6532D226A100CFC09" box="[280,370,871,896]" pageId="5" pageNumber="498">Kottelat</collectingMunicipality>
&amp;
<collectingCounty id="D94E3CB0AC14FFD653A3226A109BFC09" box="[406,485,871,896]" pageId="5" pageNumber="498">Steiner</collectingCounty>
(2011)
</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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