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<mods:title id="2E65ECFAD5023FFC09695F6FF7AFB9DB">The genus Cephaloleia Chevrolat, 1836 (Coleoptera, Chrysomelidae, Cassidinae)</mods:title>
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<mods:namePart id="74E0E53D06ED3F3F5E3ADFB615441D40">Staines, Charles L.</mods:namePart>
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<treatment id="9ECD31172F62BE8D3D6A3A01D4EA08E8" ID-GBIF-Taxon="152055253" LSID="urn:lsid:plazi:treatment:9ECD31172F62BE8D3D6A3A01D4EA08E8" httpUri="http://treatment.plazi.org/id/9ECD31172F62BE8D3D6A3A01D4EA08E8" lastPageId="187" lastPageNumber="187" pageId="186" pageNumber="187" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Cassidinae">
<subSubSection id="44F4A798E4075981F391CC79695D6A05" pageId="186" pageNumber="187" type="multiple">
<paragraph id="F053255BBF398C7CEB7DA785CA820055" pageId="186" pageNumber="187">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
</subSubSection>
<subSubSection id="64CCD75E472FB5D0658DF2EFC4A26F3D" pageId="186" pageNumber="187" type="nomenclature">
<paragraph id="2B8C5D74D59EE4D59929425302B8AD1B" pageId="186" pageNumber="187">
<taxonomicName id="9B6E592DC9B1D35C6F898AB40CFE0426" ID-CoL="SDF7" LSID="http://species-id.net/wiki/Cephaloleia_metallescens" authority="Baly, 1885" authorityName="Baly" authorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia metallescens" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="metallescens">Cephaloleia metallescens Baly, 1885</taxonomicName>
Fig. 188
</paragraph>
</subSubSection>
<subSubSection id="032AF4D0C4A946B3930110F91B532E31" pageId="186" pageNumber="187" type="reference_group">
<paragraph id="4686BEEB4D577D0D8CC41C368FF59835" pageId="186" pageNumber="187">
<taxonomicName id="B82D83D35E7B040104F85FB029AC77CC" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia metallescens" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="metallescens">Cephaloleia metallescens</taxonomicName>
<bibRefCitation id="FAE736E2AF6309B9B2D8C2E37D5348AF" author="Baly, JS" editor="Godman, FD" journalOrPublisher="London" pageId="284" pageNumber="285" title="Biologia CentraliAmericana, Zoology, Insecta, Coleoptera, Phytophaga" year="1885">Baly 1885</bibRefCitation>
: 25.
<bibRefCitation id="87A5076ED5131FB3611381DC02DBCB21" author="Weise, J" journalOrPublisher="W. Junk" pageId="297" pageNumber="298" pagination="1 - 94" title="Coleopterorum Catalogus, Chrysomelidae: Hispinae" year="1911 a">Weise 1911a</bibRefCitation>
: 8 (catalog),
<bibRefCitation id="16E9860C3BA3FD88616DC84942828F11" author="Weise, J" editor="Wytsman, P" journalOrPublisher="Genera Insectorum, Brussels" pageId="297" pageNumber="298" pagination="1 - 123" title="Coleoptera Phytophaga fam. Chrysomelidae, subfam. Hispinae." volume="fasc. 125" year="1911 b">1911b</bibRefCitation>
: 13 (catalog);
<bibRefCitation id="6BDAE4AA688AB7BD64926DDFF576E1E0" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="284" pageNumber="285" pagination="551 - 763" title="Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1946">Blackwelder 1946</bibRefCitation>
: 719 (catalog);
<bibRefCitation id="5617AE7AAAE197AE2B1CD10CDEF2EBB3" author="Papp, CS" journalOrPublisher="Portugaliae Acta Biologica (B)" pageId="291" pageNumber="292" pagination="1 - 147" title="The Hispinae of America. 3 rd Contribution for promoting the scientifical results of the International Hylean Amazon Institute in Manaos, Brazil." volume="4" year="1953">Papp 1953</bibRefCitation>
: 19 (catalog);
<bibRefCitation id="FF2F94B69E96018EC4CB0D0562CE0959" author="Uhmann, E" journalOrPublisher="W. Junk, Gravenhage" pageId="296" pageNumber="297" pagination="1 - 153" title="Coleopterorum Catalogus. Supplementa. Chrysomelidae: Hispinae, Hispinae Americanae" year="1957 a">Uhmann 1957a</bibRefCitation>
: 22 (catalog);
<bibRefCitation id="D8DA2D312798157752F608221E81AFCC" author="Wilcox, JA" journalOrPublisher="Biological Research Institute of America, Latham, NY" pageId="297" pageNumber="298" title="Checklist of the beetles of Canada, United States, Mexico, Central America and the West Indies" year="1983">Wilcox 1983</bibRefCitation>
: 137 (catalog);
<bibRefCitation id="C0E4BFC2C9F0F88F832D1794099C6639" pageId="186" pageNumber="187">Staines 1996</bibRefCitation>
: 45 (Central America species),
<bibRefCitation id="2FC27F8283D9F342370D356AE2AEA3AC" author="Staines, CL" editor="Cox, ML" journalOrPublisher="Backhuys Publishers, Leiden" pageId="293" pageNumber="294" pagination="239 - 246" title="Possible mimetic complexes in Central American Cephaloleia (Coleoptera: Chrysomelidae: Hispinae)." volumeTitle="Advances in Chrysomelidae Biology" year="1999">1999</bibRefCitation>
: 242 (mimicry),
<bibRefCitation id="86E8CCD98EC0D1930E4E05EEEDAA9D42" author="Staines, CL" journalOrPublisher="ZooKeys" pageId="293" pageNumber="294" pagination="45 - 65" title="Hispines (Chrysomelidae, Cassidinae) of La Selva Biological Station, Costa Rica." url="10.3897/zookeys.157.1338" volume="157" year="2011">2011</bibRefCitation>
: 50 (faunal list);
<bibRefCitation id="D3CC5D68AC3E327AC7FE294E48B951F8" author="Staines, CL" journalOrPublisher="Beitraege zur Entomologie" pageId="293" pageNumber="294" pagination="489 - 530" title="Joseph Sugar Baly: The man and his entomological works." volume="49" year="1999">Staines and Staines 1999</bibRefCitation>
: 524 (Baly species list);
<bibRefCitation id="9C9E66BAE407B35CAFF596A75C7F6108" author="McKenna, DD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="290" pageNumber="291" pagination="117 - 131" title="Molecular phylogenetics and evolution of host plant use in the Neotropical rolled leaf ' hispine' beetle genus Cephaloleia (Chevrolat) (Coleoptera: Cassidinae)." url="10.1016/j.ympev.2005.06.011" volume="37" year="2005">McKenna and Farrell 2005</bibRefCitation>
: 119 (phylogeny),
<bibRefCitation id="8BB428DCD7CA24938D7BFD552CD777E0" author="McKenna, DD" journalOrPublisher="Proceedings of the National Academy of Sciences" pageId="290" pageNumber="291" pagination="10947 - 10951" title="Tropical forests are both evolutionary cradles and museums of leaf beetle diversity." url="10.1073/pnas.0602712103" volume="103" year="2006">2006</bibRefCitation>
: 10949 (phylogeny).
</paragraph>
<paragraph id="0EA975B31208340DC82269A18F0564A8" pageId="186" pageNumber="187">
<taxonomicName id="3E361237F4DB3AFEB7D5A9C08A551A5E" genus="Cephalolia" lsidName="Cephalolia metallescens" pageId="186" pageNumber="187" rank="species" species="metallescens">Cephalolia metallescens</taxonomicName>
Baly.
<bibRefCitation id="A3B50E7A2DB41BA75519302568353385" author="Donckier, H" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="286" pageNumber="287" pagination="540 - 615" title="Catalogue systematique des Hispides." volume="68" year="1899">Donckier 1899</bibRefCitation>
: 550 (catalog);
<bibRefCitation id="B1607184D30E5C2BBE85702638C3C06B" author="Uhmann, E" journalOrPublisher="Arbeiten ueber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem" pageId="295" pageNumber="296" pagination="93 - 102" title="Der Borstenkegel und das Pygidium bei Cephalolia-Arten. (Coleoptera: Chrysomelidae). 96. Beitrag zur Kenntnis der Hispinen." volume="9" year="1942">Uhmann 1942</bibRefCitation>
: 94 (noted).
</paragraph>
<paragraph id="0241D6079111EB83486CF1FBD0C46DFF" pageId="186" pageNumber="187">
<taxonomicName id="B6D6D4E08A2E3D7C0BB99B0A8621523D" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia metalescens" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="metalescens">Cephaloleia metalescens</taxonomicName>
Baly.
<bibRefCitation id="836A5A71D0A8AACD2905E7892FC8EB2E" author="Meskins, C" journalOrPublisher="International Journal of Tropical Insect Science" pageId="290" pageNumber="291" pagination="159 - 171" title="A comparison of hispine beetles (Coleoptera: Chrysomelidae) associated with three orders of monocot host plants in lowland Panama." volume="27" year="2008">Meskins et al. 2008</bibRefCitation>
: 163 (misspelling, host plants).
</paragraph>
</subSubSection>
<subSubSection id="452043D09E1A424334D912782E081D7D" pageId="186" pageNumber="187" type="description">
<paragraph id="32B5E0C4ADEA77FC822F68E880643F9E" pageId="186" pageNumber="187">Description.</paragraph>
<paragraph id="FDF3379C5FB25FBB4017459E2DD17E87" pageId="186" pageNumber="187">
Broadly oblong-ovate; small; subdepressed; metallic blue; pronotum with lateral margin paler; venter and legs yellowish-red; antenna with antennomeres 1 and apical
<normalizedToken id="B8DB10E19D28888FD8DD3EE97D611098" originalValue="½">1/2</normalizedToken>
of 11 reddish. Head: vertex punctate, medial carina present; frons not projecting; depressed between eyes. Antenna:
<normalizedToken id="65C4E603C7D2E0CEA6130FA41189C0DC" originalValue="½">1/2</normalizedToken>
body length; slender; antennomeres 1-3 elongate; 1 and 3 subequal in length; 2 slightly shorter; 4-10 transverse, subequal in length; 11 2
<normalizedToken id="971520C0C5DCB105A867113631F17DD3" originalValue="×">x</normalizedToken>
length of 10, pointed at apex; 1-3 punctate with scattered setae; 4-11 setose. Pronotum: nearly twice as wide as long; lateral margin straight then rounding to anterior angle, canaliculate; anterior angle rounded, not produced; posterior angle acute; anterior margin emarginate behind head; disc transversely subconvex, depressed on each side; surface densely punctate; basal impression absent; pronotal length 0.6-0.7 mm; pronotal width 0.7-1.1 mm. Scutellum: pentagonal, impunctate. Elytron: lateral margin slightly curved, smooth, margined; apex rounded; sutural angle without tooth; humerus slightly produced, callus extends on base to scutellum; slightly constricted behind humerus; disc subconvex; moderately punctate-striate, interspaces sulcate; puncture rows obsolete at apex; elytral length 2.1-2.3 mm; elytral width 1.4-1.6 mm. Venter: pro- and mesosterna punctate; metasternum impunctate medially, punctate laterally; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete; last sternite with apical margin emarginate medially in male, rounded, entire in female. Leg: slender; punctate, each puncture with pale seta; tibia with fringe of setae on inner margin of apex. Total length: 3.1-3.3 mm.
</paragraph>
<caption id="7AD827DB869F5F4C9FA2720284722905" pageId="186" pageNumber="187">
<paragraph id="A177C8AD0AC0860176F4E967F5C2AE9D" pageId="186" pageNumber="187">
Figures 188-196. Habitus. 188
<taxonomicName id="FF84A03EC14CF63B168D3BA894305F02" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia metallescens" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="metallescens">Cephaloleia metallescens</taxonomicName>
189
<taxonomicName id="EE9990DF20DF0E0F45CD0C1377373725" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nana" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nana">Cephaloleia nana</taxonomicName>
sp. n. 190
<taxonomicName id="243098FAB6A03BFE9B9E69AF8AE448D2" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia neglecta" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="neglecta">Cephaloleia neglecta</taxonomicName>
191
<taxonomicName id="704EC2E3816BACB7A862FDD18FCE89E4" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nevermanni" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nevermanni">Cephaloleia nevermanni</taxonomicName>
192
<taxonomicName id="BFE3E17897DD3EFFA60B4E09508A180D" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nigriceps" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nigriceps">Cephaloleia nigriceps</taxonomicName>
193
<taxonomicName id="B7DFF694BE656A9BA3658E5D518C3E34" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nigricornis" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nigricornis">Cephaloleia nigricornis</taxonomicName>
194
<taxonomicName id="7DFB605BD23F07585FA0C0DF807FC861" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nigrithorax" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nigrithorax">Cephaloleia nigrithorax</taxonomicName>
195
<taxonomicName id="84342A542E6321264AFF1F94A80D7343" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nigropicta" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nigropicta">Cephaloleia nigropicta</taxonomicName>
196
<taxonomicName id="6BFD1F5A6F6C464761C70DACAD268448" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia nitida" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="nitida">Cephaloleia nitida</taxonomicName>
. Scale bars equal 3 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="561F79623B72FC8CD65C0D13DAABD748" pageId="186" pageNumber="187" type="diagnosis">
<paragraph id="34584C15F2D84AAFBF927555C099EEC5" pageId="186" pageNumber="187">Diagnosis.</paragraph>
<paragraph id="F1A207396F0A82BB4036E8D06BA5D3C0" pageId="186" pageNumber="187">This is a very distinctive species with the flattened elongate body, the smooth rounded elytral apical margin, the smooth lateral margin of the pronotum, the lack of additional puncture rows on the elytra, the smooth lateral margins of the elytra, the lack of a declivity beginning at puncture row 7 on the elytra, and the unicolorous metallic blue dorsum.</paragraph>
</subSubSection>
<subSubSection id="8635ADF63434BA8DDCC5C544FA4BD21F" pageId="186" pageNumber="187" type="host plant">
<paragraph id="4C3D9FE6BAEE9F5EF45D0CB0D4515EA8" pageId="186" pageNumber="187">Host plant.</paragraph>
<paragraph id="2A3E1105B6D5917491DE838309BBB396" pageId="186" pageNumber="187">
<taxonomicName id="68368D667F1F34D10A4101259E3DF0B6" class="Liliopsida" family="Arecaceae" genus="Bactris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bactris major" order="Arecales" pageId="186" pageNumber="187" phylum="Tracheophyta" rank="species" species="major">Bactris major</taxonomicName>
Jacq.,
<taxonomicName id="446725C8FE1EE4081D232FD4C615CC23" class="Liliopsida" family="Arecaceae" genus="Chamaedorea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chamaedorea wendlandiana" order="Arecales" pageId="186" pageNumber="187" phylum="Tracheophyta" rank="species" species="wendlandiana">Chamaedorea wendlandiana</taxonomicName>
Hemsl. (
<taxonomicName id="9CE321FACCBDB21C8C8038CA59602092" genus="Arecaceae" lsidName="Arecaceae" pageId="186" pageNumber="187" rank="genus">Arecaceae</taxonomicName>
) (
<bibRefCitation id="42BC56155EEAE87660A919CC02E1E62F" author="Meskins, C" journalOrPublisher="International Journal of Tropical Insect Science" pageId="290" pageNumber="291" pagination="159 - 171" title="A comparison of hispine beetles (Coleoptera: Chrysomelidae) associated with three orders of monocot host plants in lowland Panama." volume="27" year="2008">Meskins et al. 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CBF5778765FA2DEF4D6AA0F7EF3F6314" pageId="186" pageNumber="187" type="distribution">
<paragraph id="F9250EF5CDFAC3EEED604BF001AFA04D" pageId="186" pageNumber="187">Distribution.</paragraph>
<paragraph id="03C3E3EEFAFD869BC88F875FD31C7079" pageId="186" pageNumber="187">Costa Rica, Guatemala, Nicaragua, Panama.</paragraph>
</subSubSection>
<subSubSection id="6A36FF762B579D205B1C0BC800BC94F4" pageId="186" pageNumber="187" type="type material examined">
<paragraph id="0E8FA26323CF070A914854A98F1BCEA0" pageId="186" pageNumber="187">Type material examined.</paragraph>
<paragraph id="BCF6CC8B8AC573B26D9C6EFE669DE739" pageId="186" pageNumber="187">Holotype: Type H. T. [white disk with red border]/ Guatemala, Vera Paz, San Juan, Champion/ B. C. A., Col. VI, 2. Cephaloleia metallescens, Baly/ Cephaloleia/ Cephalolia metallescens Baly, Guatemala [blue handwritten label] (BMNH).</paragraph>
</subSubSection>
<subSubSection id="6ED531CC974710BECE429C0A2E6E0044" lastPageId="187" lastPageNumber="188" pageId="186" pageNumber="187" type="specimens examined">
<paragraph id="DFAAB9E3F9263CBC869B857A0AEDB48B" pageId="186" pageNumber="187">Specimens examined.</paragraph>
<paragraph id="4896E8181EF04AC64205FFE00481D934" lastPageId="187" lastPageNumber="188" pageId="186" pageNumber="187">
COSTA RICA: Alajuela- Brasilia, 3 April 1988 (MUCR). Cartago- Turrialba (DEI, USNM); Turrialba, Tayutic, Grano de Oro, 1100-1200 m (INBIO). Guanacaste-
<normalizedToken id="6EA8F2A24304BA3A7BBFACBBA911C55C" originalValue="Estación">Estacion</normalizedToken>
Pitila, 9 km S Santa Cecilia, 600-700 m (INBIO); La Cruz, 9 km S Santa Cecilia, 600-700 m (INBIO).
<normalizedToken id="DB830345533ED10FA99DA2CD6EB9F55C" originalValue="Limón-">Limon-</normalizedToken>
Est. Cuatro Equinas, 0 m, P.N. Tortuguero November 1991 (INBIO);
<normalizedToken id="390F27B3FC40198BB69376EA1EE415D7" originalValue="Pococí">Pococi</normalizedToken>
, 30 km N Cariari, 100-200 m (INBIO); Pque Nal Corcovado, Est Sirena, 0-100 m (INBIO). NICARAGUA:
<normalizedToken id="79EA6865B9824057D1EF1E0C816916C0" originalValue="Río">Rio</normalizedToken>
San Juan- Refugio Bartola, 16 km ESE El Castillo, 26 April 1993 (USNM). PANAMA:
<normalizedToken id="1C96BD580EEE6C80D7FC8D670D3AB8CA" originalValue="Chiriquí-">Chiriqui-</normalizedToken>
Reserva Fortuna, Continental Divide Trail, 25 May 1993 (CDFA).
<pageBreakToken id="579CC7C23EE8C443E4D498771BF2EB69" pageId="187" pageNumber="188" start="start">
<normalizedToken id="6CA8F97C6ACC9C8557CA4567E5192976" originalValue="Colón-">Colon-</normalizedToken>
</pageBreakToken>
5 mi NW Gamboa, 29 September 1969 (CMNC); Paraiso, 26 March 1911, 18 April 1911, 6 April 1911, 4 May 1911, 20 May 1911, 17 April 1911, 28 March 1911, 21 April 1911, 14 April 1911 (USNM).
<normalizedToken id="9FDDB568AE0799358D7A8166B7B82F4B" originalValue="Colón-">Colon-</normalizedToken>
Parq. Nac. Soberania, Pipeline road, 23 May 1993 (EGRC).
<normalizedToken id="51644C8E9648374B981123C37269ACDA" originalValue="Panamá-">Panama-</normalizedToken>
Barro Colorado Is., 7 January 1929 (USNM); Cerro Campana, 800 m, 3 November 1969 (EGRC); Cerro Jefe, 700 m, 19 June, 76 (EGRC); Coco Solito Hosp., 11 December 1971 (EGRC); Llano Carti Rd. at km 9, 18 May 1993 (EGRC); Madden Forest, 14 May 1978 (USNM); Nusagandi area, I. K. U. S. A. Igar, 20 May 1993 (EGRC). Total: 69.
</paragraph>
</subSubSection>
</treatment>
</document>