308 lines
46 KiB
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308 lines
46 KiB
XML
<document ID-CLB-Dataset="270987" ID-DOI="http://dx.doi.org/10.3897/zookeys.1181.110628" ID-GBIF-Dataset="0e51071e-1347-4745-8658-fe7e92330e45" ID-PMC="PMC10576206" ID-Pensoft-Pub="1313-2970-1181-201" ID-Pensoft-UUID="CA5BA4745FFD57E19CB7B6E98161293E" ID-PubMed="37841026" ID-ZooBank="AB0B755AC53046CD87F277599B974343" ModsDocID="1313-2970-1181-201" checkinTime="1696648911163" checkinUser="pensoft" docAuthor="Chu, Chang, Li, Shuqiang, Pham, Dinh-Sac & Yao, Zhiyuan" docDate="2023" docId="9E6F30D605C152EABF31E24548C2B168" docLanguage="en" docName="ZooKeys 1181: 201-217" docOrigin="ZooKeys 1181" docPubDate="2023-10-06" docSource="http://dx.doi.org/10.3897/zookeys.1181.110628" docTitle="Utivarachna tamdao Chu & Li 2023, sp. nov." docType="treatment" docUuid="7CC2C53C-5BBB-49BD-A81E-13835F82B7E5" docUuidSource="ZooBank" docVersion="4" id="CA5BA4745FFD57E19CB7B6E98161293E" lastPageNumber="201" masterDocId="CA5BA4745FFD57E19CB7B6E98161293E" masterDocTitle="Three new species of the spider genus Utivarachna Kishida, 1940 (Araneae, Trachelidae) from China and Vietnam" masterLastPageNumber="217" masterPageNumber="201" pageNumber="201" updateTime="1732484100492" updateUser="ExternalLinkService">
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<mods:titleInfo id="6AC7443DC3E5E626D8388B6D29D687C8">
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<mods:title id="C71F7F13648342687EC4509F1C8B3B93">Three new species of the spider genus Utivarachna Kishida, 1940 (Araneae, Trachelidae) from China and Vietnam</mods:title>
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<mods:name id="D58748093BC632C4B22A0546811EAC6A" type="personal">
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<mods:namePart id="A45EDD9EDF81622ACAD20C0338C24778">Chu, Chang</mods:namePart>
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<mods:nameIdentifier id="B1AA7DB6292274B1E026D76A82FA9097" type="ORCID">https://orcid.org/0000-0003-3520-5463</mods:nameIdentifier>
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<mods:affiliation id="14F92D490CD1837CAC0D2DC1EB637D6A">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:roleTerm id="3AA27AB69666214434D169C53F7CA429">Author</mods:roleTerm>
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<mods:namePart id="179C52F88ADD566ACFB6E00545E0A483">Li, Shuqiang</mods:namePart>
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<mods:nameIdentifier id="F12B25A5683E6E6BDDC6C03294342A2D" type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:affiliation id="DFFD9C65FB01316D7115E9260354E022">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier id="68DD348D9F111A61563475A9B2AF0BD4" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:roleTerm id="80839057C4D56F7378D2C77321120B92">Author</mods:roleTerm>
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<mods:namePart id="BBBA6888973C94B1A1C1C297CEAD27E9">Pham, Dinh-Sac</mods:namePart>
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<mods:nameIdentifier id="CACDA5E250900D0550E2738E61D98651" type="ORCID">https://orcid.org/0000-0001-8594-5270</mods:nameIdentifier>
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<mods:affiliation id="5FD77EBBB9FF52176D33B32A475F5015">Vietnam National Museum of Nature (VNMN), Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="4B44DCA3DFD5A2D599D70115213922EA">Yao, Zhiyuan</mods:namePart>
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<mods:nameIdentifier id="B81CB687CBFB1BD27065FDAACD337BE0" type="ORCID">https://orcid.org/0000-0002-1631-0949</mods:nameIdentifier>
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<mods:affiliation id="8A6B99870444362C1FF2EEC6BB481334">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:nameIdentifier id="7A6CFB376BC9FEB2B26F3CDC9B556962" type="email">yaozy@synu.edu.cn</mods:nameIdentifier>
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<mods:title id="5E787C252819225F61FF62785D7BC378">ZooKeys</mods:title>
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<mods:part id="7F1C376097B637CF9EC419D6A0E21697">
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<mods:date id="26EDF52F22197139DE1D8145CDBB4C6E">2023</mods:date>
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<mods:number id="3A07C237B616BCF1D68B0F595C96210C">2023-10-06</mods:number>
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<mods:number id="EA33F0A5B03FADDF3D68F02942B3F6AC">1181</mods:number>
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<mods:classification id="E0737964F426BC5566F2B6A762E477CF">journal article</mods:classification>
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<mods:identifier id="4F9A64B7D5F9003F3759C3BF14DD0A70" type="DOI">http://dx.doi.org/10.3897/zookeys.1181.110628</mods:identifier>
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<treatment id="9E6F30D605C152EABF31E24548C2B168" ID-GBIF-Taxon="213736217" LSID="urn:lsid:zoobank.org:act:7CC2C53C-5BBB-49BD-A81E-13835F82B7E5" httpUri="http://treatment.plazi.org/id/9E6F30D605C152EABF31E24548C2B168" lastPageNumber="201" pageId="0" pageNumber="201" scope_family="Trachelidae" scope_order="Araneae">
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<subSubSection id="E84F3CD252428A18F21D5078DA74BE38" pageId="0" pageNumber="201" type="nomenclature">
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<paragraph id="7A1F2A6E0276BD9395E7496856B9CCAC" pageId="0" pageNumber="201">
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<taxonomicName id="72F7260DFCC1704F50D43C1A885286AD" ID-CoL="C8MQV" LSID="https://zoobank.org/7CC2C53C-5BBB-49BD-A81E-13835F82B7E5" authority="Chu & Li" authorityName="Chu & Li" authorityYear="2023" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Utivarachna tamdao" order="Araneae" pageId="0" pageNumber="201" phylum="Arthropoda" rank="species" species="tamdao" status="sp. nov.">Utivarachna tamdao Chu & Li</taxonomicName>
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<taxonomicNameLabel id="ED57CE19BA66737D432902B3A43C05E1" pageId="0" pageNumber="201">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DF1159B4EAF21B3682CCF52EA6F43F3" pageId="0" pageNumber="201" type="description">
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<paragraph id="8C79C7A8640D42C1336AF05020D2CAD1" pageId="0" pageNumber="201">
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<figureCitation id="8E055420F333D2079CCFAD8DD8B89341" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">Figs 4</figureCitation>
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<figureCitation id="2BFE3BB16921F11770B26260B6DA305A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">, 5</figureCitation>
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<figureCitation id="F6BCD13168199BC767D7A03723437E58" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="55B0939224BF216F83AF2EB20194CB45" pageId="0" pageNumber="201" type="type material">
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<paragraph id="445BAAF179EDEC46EA144CB8D7FE718B" pageId="0" pageNumber="201">Type materials.</paragraph>
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<paragraph id="F5D0349C3F012E8BE6EB77581B09D3B2" pageId="0" pageNumber="201">
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<emphasis id="471BA62971F13A4A9B7BB965878F6ABF" bold="true" italics="true" pageId="0" pageNumber="201">Holotype</emphasis>
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♂ (IZCAS-Ar44630):
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<emphasis id="314EB67EA4D2CD2F67C300642A694DBF" bold="true" pageId="0" pageNumber="201">Vietnam</emphasis>
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: Vinh Phuc Province: Tam Dao National Park, natural forest (
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<geoCoordinate id="E54F0B481B437FBBBCDB85BF471AA31B" degrees="21.4872" direction="north" orientation="latitude" precision="5" value="21.4872">21.4872°N</geoCoordinate>
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,
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<geoCoordinate id="CE2CE7C5C97D521E34D1D202B43662FA" degrees="105.6201" direction="east" orientation="longitude" precision="5" value="105.6201">105.6201°E</geoCoordinate>
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, 870 m a.s.l.), hand caught in leaf litter, 18.IX.2007, leg. Dinh-Sac Pham.
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<emphasis id="B402703E902D746FD828FE8EEAD5281F" bold="true" italics="true" pageId="0" pageNumber="201">Paratypes</emphasis>
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: 1♂ (IZCAS-Ar44631) and 2♀ (IZCAS-Ar44632, 44633), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="CA86C9469B8B76637657AE98C6C56B8D" pageId="0" pageNumber="201" type="etymology">
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<paragraph id="2E2343909544E86E3395E64C54F9935B" pageId="0" pageNumber="201">Etymology.</paragraph>
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<paragraph id="B5DA15F365F6EB30F974ED998262A39B" pageId="0" pageNumber="201">The specific name is named after type locality; noun in apposition.</paragraph>
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</subSubSection>
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||
<subSubSection id="756E76CF3C0F4564E15CBFE0D3F51BE4" pageId="0" pageNumber="201" type="diagnosis">
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<paragraph id="E8F76F30C8FFF8F50E4FA5DDA4626BF8" pageId="0" pageNumber="201">Diagnosis.</paragraph>
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<paragraph id="7CE584A9964F2E78DCC240A55E25F45D" pageId="0" pageNumber="201">
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The new species resembles
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<taxonomicName id="BCBDED065A552A4EEB6793C67B1C5E59" lsidName="U. kinabaluensis" pageId="0" pageNumber="201" rank="species" species="kinabaluensis">
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<emphasis id="E70AEE90FA4FA61FCBB89D16752AAC1E" italics="true" pageId="0" pageNumber="201">U. kinabaluensis</emphasis>
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</taxonomicName>
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Deeleman-Reinhold, 2001 (cf. Figs
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<figureCitation id="BC1BB49D92054B73E4541C6022DF6101" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4</figureCitation>
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-
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<figureCitation id="63411FF78476D727F3A52E2C298C0A5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6</figureCitation>
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and
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<bibRefCitation id="C8A447E6FD9C320F140A2AD7AB882936" DOI="https://doi.org/10.1163/9789004475588" author="Deeleman-Reinhold, CL" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="201" refId="B3" refString="Deeleman-Reinhold, CL, 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, DOI: https://doi.org/10.1163/9789004475588" title="Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae)." url="https://doi.org/10.1163/9789004475588" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
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: 381, figs 593-597), as males have a similar U-shaped sperm duct (SD) (Fig.
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<figureCitation id="EE87758BE06F60A84606BD1E4C799539" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B</figureCitation>
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), the tegulum is widest in anterior part (Fig.
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<figureCitation id="F4D93D566AA02511C2F4E8B05C003DB4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B</figureCitation>
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), the embolus (E) is coiled (Fig.
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<figureCitation id="6B2353BF276EA57FCF3F07F73BA95B8B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4A-C</figureCitation>
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), females have a nearly trapezoidal atrium (A) (Fig.
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<figureCitation id="F1FFCA4D34D766E199B0FB872E53DD19" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A</figureCitation>
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), connecting ducts (CnD) are thin and slender (Fig.
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<figureCitation id="56CD67FDAB8CC8411A9533DCB6BC374F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
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), and fertilization ducts (FD) are laminar (Fig.
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<figureCitation id="4673752DD65C7296BBCAF73D0AEB426E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
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). Males can be distinguished in having the RTA widest at its base (Fig.
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<figureCitation id="AE5E60940A0F2026F0E7E6D74AA06C00" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B, C</figureCitation>
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; vs RTA widest at middle, distally with hook-shaped apophysal claw in
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<taxonomicName id="BE10CCC820BEE74211F8435BE24DD689" lsidName="U. kinabaluensis" pageId="0" pageNumber="201" rank="species" species="kinabaluensis">
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<emphasis id="E51518005AD3B5BFCD064F7217065308" italics="true" pageId="0" pageNumber="201">U. kinabaluensis</emphasis>
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</taxonomicName>
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). Females can be distinguished from
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<taxonomicName id="C6F227731E24F93B5EC9DFB07738DC74" lsidName="U. kinabaluensis" pageId="0" pageNumber="201" rank="species" species="kinabaluensis">
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<emphasis id="8DDCBF8E3C65581BAAFD37A81E01B50D" italics="true" pageId="0" pageNumber="201">U. kinabaluensis</emphasis>
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</taxonomicName>
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in having the copulatory openings (CO) located anteriorly (Fig.
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<figureCitation id="986553C9F1506319A6164A5252905E7E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A</figureCitation>
|
||
; vs posteriorly), the atrium large, occupying about 2/3 of the epigyne length (vs atrium small, occupying about 1/4 of the epigyne length), the copulatory ducts (CD) forming irregular loops and these loops located anteriorly (Fig.
|
||
<figureCitation id="7842ADCDBC89B718ADC979FB4C00E5BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
; vs copulatory ducts regularly coiled and these coils medially located), the bursae (B) located between the copulatory ducts and spermathecae (SP), anterior part strongly constricted and curved, and posterior part three times width of anterior part (Fig.
|
||
<figureCitation id="75F911AADFAF1B4235346C3AC0AA8C37" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
; vs bursae located in anterior of copulatory ducts, base to middle part strongly constricted and curved, posterior part five times the width of the anterior part), and the spermathecae globular (Fig.
|
||
<figureCitation id="C5897D265F94582C26661CEB402C34F6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
; vs gourd-shaped). The new species also resembles
|
||
<taxonomicName id="F0DBEB29EB1F8DEBF5E15D40CA745B99" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="527B09C0BBB45719C022E6523F7D9A24" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
Jin, Yin & Zhang, 2015 (cf. Figs
|
||
<figureCitation id="0647DBB988833C40AD094BD8C0A2BD6B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4</figureCitation>
|
||
-
|
||
<figureCitation id="B4838899CC2A02B5712B8043257F0A2F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6</figureCitation>
|
||
and
|
||
<bibRefCitation id="B12A5806C8D12E51F0DDA9446336D4BF" DOI="https://doi.org/10.11646/zootaxa.4057.4.6" author="Jin, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="201" pagination="569 - 581" refId="B7" refString="Jin, C, Yin, X, Zhang, F, 2015. A new species of the genus Utivarachna Kishida, 1940 (Araneae: Trachelidae) from China, with the first description of the male of U. fabaria Zhao & Peng, 2014 and a redescription of U. gui (Zhu, Song & Kim, 1998). Zootaxa 4057 (4): 569 - 581, DOI: https://doi.org/10.11646/zootaxa.4057.4.6" title="A new species of the genus Utivarachna Kishida, 1940 (Araneae: Trachelidae) from China, with the first description of the male of U. fabaria Zhao & Peng, 2014 and a redescription of U. gui (Zhu, Song & Kim, 1998)." url="https://doi.org/10.11646/zootaxa.4057.4.6" volume="4057" year="2015">Jin et al. 2015</bibRefCitation>
|
||
: 570, figs 1-3) as males have a similar U-shaped sperm duct (Fig.
|
||
<figureCitation id="64C479FE0B328D7875DFD3979FD9AFC3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B</figureCitation>
|
||
), leaf-shaped subtegulum (ST) in ventral view (Fig.
|
||
<figureCitation id="4DF3B12B90F1CBFF7E860A01097D2089" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B</figureCitation>
|
||
), and long RTA (Fig.
|
||
<figureCitation id="11D3BA2452B6C3313EEC4C5EDFFC3406" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B, C</figureCitation>
|
||
), and females have similar copulatory openings located anteriorly (Fig.
|
||
<figureCitation id="FB9B52A929795840162D46542A59685A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A</figureCitation>
|
||
), a large atrium (Fig.
|
||
<figureCitation id="D8A66460A5DF9A3754F7C69DEBC96299" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A</figureCitation>
|
||
), and globular spermathecae (Fig.
|
||
<figureCitation id="E04A0BB89CE327A458EE416A5CC173A8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
). Males can be distinguished by the oval bulb, which is widest in anteriorly (Fig.
|
||
<figureCitation id="2117D104A147E39C2E9AD9622488A267" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B</figureCitation>
|
||
; vs almost square in
|
||
<taxonomicName id="2B3FEEF5C57EF892E3F1E00DCC1572FA" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="BF616EF45D68A8C5A1FD8414D034CC91" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
) and by the curved basal portion of the embolus (Fig.
|
||
<figureCitation id="01992C0DC259AEA4B6205B3ACCAA779B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4B, C</figureCitation>
|
||
; vs oblique in
|
||
<taxonomicName id="6A170A579E29F1489250D8AE9DB9EC4E" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="6AFE68DF1E8623D88BDF5C8CC7D35391" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
). Females can by distinguished by the copulatory openings width/atrium posterior width: 1/6 (Fig.
|
||
<figureCitation id="129655DE160605337CEAA4B163CA81BA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A</figureCitation>
|
||
; vs copulatory openings width/atrium posterior width: 1/2 in
|
||
<taxonomicName id="E37662803EF44E2A59D4E17FBD1D5012" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="2DFD0F4D10D2C9F66FBD4EC47EFA3E68" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
), by the copulatory ducts forming irregular loops (Fig.
|
||
<figureCitation id="BDB158186D918BFBFB7CC2CA97DF0EFC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
; vs copulatory ducts coiled three times around the anterior of connecting duct in
|
||
<taxonomicName id="E7B1EAC63B1D8954D1462839FB844CDF" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="C03F98B6E68C05A268E6C7F0A8C1CB94" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
), and by the bursae anterior part strongly constricted and curved (Fig.
|
||
<figureCitation id="48C61C3283294490F1514BD5EF82CC3C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5B</figureCitation>
|
||
; vs bursae anterior part slightly constricted and curved, posterior end very close to spermathecae in
|
||
<taxonomicName id="90BB5BBDE3B061668E2ECF84E29C7AB6" lsidName="U. lata" pageId="0" pageNumber="201" rank="species" species="lata">
|
||
<emphasis id="3A97E36914734845841ECD81EF67AA16" italics="true" pageId="0" pageNumber="201">U. lata</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="44BDE2C945174813DFADDF72B9F4FA76" doi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201" start="Figure 4" startId="F4">
|
||
<paragraph id="B2FB14634F404A3B7C860F334D50D590" pageId="0" pageNumber="201">
|
||
<emphasis id="8AD14BB900B56C4029B918EC47DE4B59" bold="true" pageId="0" pageNumber="201">Figure 4.</emphasis>
|
||
<taxonomicName id="E9822CE8059654FC735D86E7F2362FE8" authorityName="Chu & Li" authorityYear="2023" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Utivarachna tamdao" order="Araneae" pageId="0" pageNumber="201" phylum="Arthropoda" rank="species" species="tamdao">
|
||
<emphasis id="A6F1904C6D93EF6D667C06E08E4F8D40" italics="true" pageId="0" pageNumber="201">Utivarachna tamdao</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="231BE3C040FDD40BB17FD8DA90607C58" bold="true" pageId="0" pageNumber="201">A-C</emphasis>
|
||
palp
|
||
<emphasis id="1C2F168481640A9CE7EAB5BAB5039874" bold="true" pageId="0" pageNumber="201">A</emphasis>
|
||
prolateral view
|
||
<emphasis id="F1FF942B228CD7A821BF4B78CDA3BE96" bold="true" pageId="0" pageNumber="201">B</emphasis>
|
||
ventral view
|
||
<emphasis id="F8028A0293CCA920C404D4C42B88C108" bold="true" pageId="0" pageNumber="201">C</emphasis>
|
||
retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="688D419368670B889A225E631F9B65EC" doi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201" start="Figure 5" startId="F5">
|
||
<paragraph id="3ED0639B23092A1401653AA25241BBE0" pageId="0" pageNumber="201">
|
||
<emphasis id="3C074FDF16E1CF41249C9155430D8964" bold="true" pageId="0" pageNumber="201">Figure 5.</emphasis>
|
||
<taxonomicName id="B1DF3B77C530794E60F5FC558636CBAC" authorityName="Chu & Li" authorityYear="2023" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Utivarachna tamdao" order="Araneae" pageId="0" pageNumber="201" phylum="Arthropoda" rank="species" species="tamdao">
|
||
<emphasis id="9FF226537F045EF67C05168AB2F6894C" italics="true" pageId="0" pageNumber="201">Utivarachna tamdao</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype female
|
||
<emphasis id="0CEC6507138BD81339748183BC17CA3C" bold="true" pageId="0" pageNumber="201">A</emphasis>
|
||
epigyne, ventral view
|
||
<emphasis id="637E86A2DC0CF1B0D63F58E32E49A0E7" bold="true" pageId="0" pageNumber="201">B</emphasis>
|
||
vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="56D597F504C48DA8B4068A64A509A92D" doi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201" start="Figure 6" startId="F6">
|
||
<paragraph id="0DE96D7688BB4FE4573B9FB400327E6E" pageId="0" pageNumber="201">
|
||
<emphasis id="32AA51EB7476ADF95EDDAF685A8D0241" bold="true" pageId="0" pageNumber="201">Figure 6.</emphasis>
|
||
<taxonomicName id="28C46766B217343E6FAB56BBB89E1FD3" authorityName="Chu & Li" authorityYear="2023" class="Arachnida" family="Trachelidae" genus="Utivarachna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Utivarachna tamdao" order="Araneae" pageId="0" pageNumber="201" phylum="Arthropoda" rank="species" species="tamdao">
|
||
<emphasis id="952D9E518A05C52E199AADE5C01F9277" italics="true" pageId="0" pageNumber="201">Utivarachna tamdao</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="26980CDB0B092E81F5A7287999C6C26E" bold="true" pageId="0" pageNumber="201">A, B</emphasis>
|
||
) and paratype female (
|
||
<emphasis id="E44F10F925A004CB54648B8926D38F2E" bold="true" pageId="0" pageNumber="201">C, D</emphasis>
|
||
)
|
||
<emphasis id="68C24A428EDD2BDDC914EA0FD7505BB5" bold="true" pageId="0" pageNumber="201">A-D</emphasis>
|
||
habitus
|
||
<emphasis id="7548A9CE07BE3E01DFDAF7358AB75EC4" bold="true" pageId="0" pageNumber="201">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="7279A5F72139A4F1D054E71A478B846D" bold="true" pageId="0" pageNumber="201">B</emphasis>
|
||
ventral view
|
||
<emphasis id="EEAA66CB2A4FDF43AD8FEF629EC3011D" bold="true" pageId="0" pageNumber="201">C</emphasis>
|
||
dorsal view
|
||
<emphasis id="52467E13F277A720E03F5A005C9A9838" bold="true" pageId="0" pageNumber="201">D</emphasis>
|
||
ventral view. Scale bars: 1.00 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="306ECEA43288659815CB0FAA0A132C29" pageId="0" pageNumber="201" type="description">
|
||
<paragraph id="447C43F2C29BBD21DCDE107E1AC94116" pageId="0" pageNumber="201">Description.</paragraph>
|
||
<paragraph id="DF4360255F96FCF6BE89CAC1C2CC569E" pageId="0" pageNumber="201">
|
||
<emphasis id="41477422A74CFE81C13AA0C658B76052" bold="true" pageId="0" pageNumber="201">Male.</emphasis>
|
||
Habitus (Fig.
|
||
<figureCitation id="212B8A4C1E7444F9E8FB385ED83B2FC5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6A, B</figureCitation>
|
||
). Total length 4.68. Carapace (Fig.
|
||
<figureCitation id="681CA9EA87D236D476D15F6D91BB566E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6A</figureCitation>
|
||
): length 2.48, width 1.86, deep reddish brown; cervical groove, radial grooves, and fovea indistinct. Eyes (Fig.
|
||
<figureCitation id="FFFF1E799056EFCE5F2A6A7353C8E0B8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6A</figureCitation>
|
||
): AER procurved, PER recurved in dorsal view, PER wider than AER. Eye sizes and interdistances: AME 0.15, ALE 0.13, PME 0.13, PLE 0.14; AME-AME 0.06, AME-ALE 0.12, PME-PME 0.19, PME-PLE 0.27; MOA 0.36 long, anterior width 0.34, posterior width 0.46. Mouthparts (Fig.
|
||
<figureCitation id="6990DB5DD81D7994ADA8F33363924EF5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6B</figureCitation>
|
||
): chelicerae deep reddish brown, with three promarginal (middle largest) and five retromarginal (proximal largest, distal smallest) teeth; endites depressed posteriorly, slightly convergent anteriorly, with dense setae on inner margin; labium nearly trapezoidal, length 0.47, width 0.39. Sternum (Fig.
|
||
<figureCitation id="B2D7E9ACAC85305252C4C6BF699E5C14" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6B</figureCitation>
|
||
) length 1.29, width 0.97, deep reddish-brown, with dark edges, with precoxal triangles and intercoxal extensions, posterior region protruding strongly between coxae IV. Pedicel cylindrical, sclerotized, relatively short, reddish brown. Abdomen (Fig.
|
||
<figureCitation id="508C0B0A5BC9B7BE260BA39A1050754D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6A, B</figureCitation>
|
||
) grey, 2.20 long, 1.75 wide, dorsum with scutum covering more than half of dorsal surface, with four brown central spots; venter with indistinct two lines of spots in the median field. Spinnerets yellow. Legs: anterior legs reddish brown, distinctly thicker than yellowish-brown posterior legs. Leg measurements: I 6.03 (1.83, 0.77, 1.55, 1.18, 0.70); II 4.82 (1.62, 0.71, 0.89, 1.07, 0.53); III 4.37 (1.26, 0.62, 0.89, 1.07, 0.53); IV 5.50 (1.52, 0.67, 1.20, 1.52, 0.59).
|
||
</paragraph>
|
||
<paragraph id="4F5461DCE6CF2C60BE0321AF72224124" pageId="0" pageNumber="201">
|
||
<emphasis id="4BF0FC39FA2C2C1D1CE4BF539D943777" bold="true" italics="true" pageId="0" pageNumber="201">Palp</emphasis>
|
||
(Fig.
|
||
<figureCitation id="5991B6C0EC67B349CBC1B78069FEFBC5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Utivarachna tamdao sp. nov., holotype male A-C palp A prolateral view B ventral view C retrolateral view. Abbreviations: E = embolus, RTA = retrolateral tibial apophysis, SD = sperm duct, ST = subtegulum. Scale bar: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure4" httpUri="https://binary.pensoft.net/fig/917669" pageId="0" pageNumber="201">4A-C</figureCitation>
|
||
): tibia shorter than half of cymbium length; RTA about 0.98 times longer than tibia, distinctly narrow subdistally to distally, with distinct anterior curvature distally. Bulb oval, widest in anterior part; tegulum approximately 1.39 times as long as its maximum width in ventral view; subtegulum (ST) sclerotized, occupying approximately 1/3 of tegulum width in ventral view; sperm duct (SD) distinct, U-shaped in ventral view, extending to base of tegulum. Embolus (E) long, anticlockwise, obliquely coiled twice, coils as wide as maximum width of tegulum; basal portion of embolus lamellar, wide, arising at 2:30
|
||
<normalizedToken id="07CF0C66E65A90E33A8D5BB741163C78" originalValue="o’clock">o'clock</normalizedToken>
|
||
from bulb; terminal portion of embolus filiform, resting in distal cymbial alveolus.
|
||
</paragraph>
|
||
<paragraph id="3B6FC27D019360EF6BA36131CFE8E7FA" pageId="0" pageNumber="201">
|
||
<emphasis id="F39FE76DCE74645D2D7BCCC751F0EE81" bold="true" pageId="0" pageNumber="201">Female.</emphasis>
|
||
Habitus (Fig.
|
||
<figureCitation id="CD1B58D04526C7ED278220967EC87869" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6C, D</figureCitation>
|
||
). As in male except as noted. Total length 5.34. Carapace length 2.70, width 2.02, reddish brown; fovea distinct, dark, and short. Eye (Fig.
|
||
<figureCitation id="216C69FC5C43FF60787D6B3BBC8ECC8D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6C</figureCitation>
|
||
) sizes and interdistances: AME 0.12, ALE 0.13, PME 0.13, PLE 0.13; AME-AME 0.09, AME-ALE 0.12, PME-PME 0.21, PME-PLE 0.25; MOA 0.32 long, anterior width 0.31, posterior width 0.44. Mouthparts (Fig.
|
||
<figureCitation id="96582F216976197F2ED8635553D7AE59" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6D</figureCitation>
|
||
): chelicerae with three promarginal (middle largest) and five retromarginal (proximal largest, distal smallest) teeth. Sternum (Fig.
|
||
<figureCitation id="8C9C89ADE3D4065EFFB3921A29FECFF2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6D</figureCitation>
|
||
) length 1.50, width 1.17, reddish brown. Abdomen (Fig.
|
||
<figureCitation id="63093CFF651C7E76056357090312DE97" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Utivarachna tamdao sp. nov., holotype male (A, B) and paratype female (C, D) A-D habitus A dorsal view B ventral view C dorsal view D ventral view. Scale bars: 1.00 mm." figureDoi="10.3897/zookeys.1181.110628.figure6" httpUri="https://binary.pensoft.net/fig/917671" pageId="0" pageNumber="201">6C, D</figureCitation>
|
||
): length 2.54, width 2.08, dorsum with four central, indistinct, reddish-brown spots; venter without pattern. Spinnerets surrounded by brown rings. Leg measurements: I 5.56 (1.64, 0.75, 1.32, 1.12, 0.73); II 5.24(1.54, 0.69, 1.19, 1.14, 0.68); III 4.41 (1.24, 0.63, 0.90, 1.07, 0.57); IV 5.73 (1.57, 0.66, 1.27, 1.58, 0.65).
|
||
</paragraph>
|
||
<paragraph id="7BB5C564B556963A6B20E45F1BF52369" pageId="0" pageNumber="201">
|
||
<emphasis id="4310309DE88230D621F0ED2FAD750E0F" bold="true" italics="true" pageId="0" pageNumber="201">Epigyne</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2ED694AB1FE6D796BA3AEB5893C4691A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Utivarachna tamdao sp. nov., paratype female A epigyne, ventral view B vulva, dorsal view. Abbreviations: A = atrium, B = bursa, CD = copulatory duct, CnD = connecting duct, CO = copulatory opening, FD = fertilization duct, GP = glandular particles, SP = spermathecae. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1181.110628.figure5" httpUri="https://binary.pensoft.net/fig/917670" pageId="0" pageNumber="201">5A, B</figureCitation>
|
||
): epigynal plate longer than wide, spermathecae distinct and bursae indistinct in ventral view. Atrium (A) large and nearly trapezoidal, occupying about 2/3 of epigyne length, posterior margin wider than anterior margin. Copulatory openings (CO) semicircular, located at anteriorly, separated by about their diameter. Copulatory ducts (CD) long, anterior part wide and posterior part narrow; copulatory ducts convoluted posteriorly, forming irregular loops. Connecting ducts (CnD) thin and slender, located on lateral areas of copulatory openings, separated by more than spermathecae (SP) diameter. Bursae (B) nearly rod-shaped, anterior part strongly constricted and curved, posterior part three times width of anterior part; bursae with several small clusters of glandular particles (GP) on surface of distal margin. Spermathecae globular, separated by less than half of their diameter. Fertilization ducts (FD) laminar, separated from each other by posterior width of atrium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7D4268959CAA0525B8E7F2D98F22E699" pageId="0" pageNumber="201" type="distribution">
|
||
<paragraph id="1BB4725141E78E8D98FDA4A75A1F7648" pageId="0" pageNumber="201">Distribution.</paragraph>
|
||
<paragraph id="DBBBAB6BA719A35AF0C0FC58EC0B742C" pageId="0" pageNumber="201">
|
||
Vietnam (Vinh Phuc, type locality; Fig.
|
||
<figureCitation id="41489C3613910F9FCF2359AE955CC99B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of new Utivarachna species from China and Vietnam 1 = U. linyejiei sp. nov., 2 = U. tamdao sp. nov., 3 = U. zhengguoi sp. nov." figureDoi="10.3897/zookeys.1181.110628.figure10" httpUri="https://binary.pensoft.net/fig/917675" pageId="0" pageNumber="201">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |