169 lines
20 KiB
XML
169 lines
20 KiB
XML
<document id="D11CA3F3D98EAB343F86973DC4CAB477" ID-CLB-Dataset="29151" ID-DOI="10.11646/zootaxa.4422.1.5" ID-GBIF-Dataset="ea160d3a-63f4-4a24-b4b1-042ef1508b5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455505" ID-ZooBank="D167DDAD-17D0-4F35-9873-85B4BC7E8FEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539184953362" checkinUser="plazi" docAuthor="Jaschhof, Mathias & Spungis, Voldemars" docDate="2018" docId="9E1D954CFFEC9660FF51FF1DD948FE5E" docLanguage="en" docName="zootaxa.4422.1.5.pdf" docOrigin="Zootaxa 4422 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dicerura formosa Mamaev 1998" docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="6224ED34FFE5966AFFC6FF8ADA0CFFEC" masterDocTitle="Towards reliable identification of male Dicerura: descriptions of three new and seven poorly known species in the Palearctic region (Diptera: Cecidomyiidae, Porricondylinae)" masterLastPageNumber="103" masterPageNumber="85" pageNumber="94" updateTime="1698573658146" updateUser="ExternalLinkService">
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<mods:title id="3FB92AD3ECA9A093DB65104047F5A0C5">Towards reliable identification of male Dicerura: descriptions of three new and seven poorly known species in the Palearctic region (Diptera: Cecidomyiidae, Porricondylinae)</mods:title>
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<mods:namePart id="321E79F2D3D9F5AA3672038810F0A309">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="A8CFD7FCCE7E0D46B5F0E7DF0715B3FE">Spungis, Voldemars</mods:namePart>
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<mods:title id="E205A7BA6855576FFD756E0ECEB19D9F">Zootaxa</mods:title>
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<mods:date id="826308A91900EE2EE9651F823DC2DE98">2018</mods:date>
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<treatment id="9E1D954CFFEC9660FF51FF1DD948FE5E" ID-DOI="http://doi.org/10.5281/zenodo.6492313" ID-GBIF-Taxon="148671183" ID-Zenodo-Dep="6492313" LSID="urn:lsid:plazi:treatment:9E1D954CFFEC9660FF51FF1DD948FE5E" httpUri="http://treatment.plazi.org/id/9E1D954CFFEC9660FF51FF1DD948FE5E" lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94">
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<subSubSection id="5EAE77D1FFEC9663FF51FF1DDB19FF3F" pageId="9" pageNumber="94" type="nomenclature">
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<paragraph id="160B245AFFEC9663FF51FF1DD827FF5E" blockId="9.[151,555,151,211]" box="[151,555,151,178]" pageId="9" pageNumber="94">
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<heading id="4D439336FFEC9663FF51FF1DD827FF5E" bold="true" box="[151,555,151,178]" fontSize="11" level="1" pageId="9" pageNumber="94" reason="1">
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<taxonomicName id="D1B45FD9FFEC9663FF51FF1DD827FF5E" ID-CoL="35JW7" authority="Mamaev, 1998" authorityName="Mamaev" authorityYear="1998" box="[151,555,151,178]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FF51FF1DD827FF5E" bold="true" box="[151,555,151,178]" pageId="9" pageNumber="94">
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<emphasis id="24C0F848FFEC9663FF51FF1DDB62FF5D" bold="true" box="[151,366,151,177]" italics="true" pageId="9" pageNumber="94">Dicerura formosa</emphasis>
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<bibRefCitation id="722559ABFFEC9663FEB0FF12D827FF5E" box="[374,555,152,178]" pageId="9" pageNumber="94" refId="ref11327" refString="Mamaev, B. M. (1998) New species of gall midges of various taxa (Diptera, Cecidomyiidae). All-RuSSian InStitute of ContinuouS Education in ForeStry, PuShkino, MoScow Region, 13, 1 - 10." type="journal article">Mamaev, 1998</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="160B245AFFEC9663FF51FF30DB19FF3F" blockId="9.[151,555,151,211]" box="[151,277,186,211]" pageId="9" pageNumber="94">
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<figureCitation id="8E8F38DFFFEC9663FF51FF30DB19FF3F" box="[151,277,186,211]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Figs 16–18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5EAE77D1FFEC9663FF51FE89D9CFFE6B" pageId="9" pageNumber="94" type="description">
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<paragraph id="160B245AFFEC9663FF51FE89D9CFFE6B" blockId="9.[151,1436,258,391]" pageId="9" pageNumber="94">
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We identified our specimens from sketches made by one of us (MJ) while examining the
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<typeStatus id="C90F9AF8FFEC9663FBBDFE88DED1FEF7" box="[1147,1245,258,283]" pageId="9" pageNumber="94" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="D1B45FD9FFEC9663FAC7FE89DF73FEF7" authorityName="Mamaev" authorityYear="1998" box="[1281,1407,259,283]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FAC7FE89DF73FEF7" box="[1281,1407,259,283]" italics="true" pageId="9" pageNumber="94">D. formosa</emphasis>
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</taxonomicName>
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in 2012. Mamaev’s (1998) description of this species, whose only illustration was a schematic drawing of the gonostylus, is insufficient for specific identification.
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<taxonomicName id="D1B45FD9FFEC9663FD23FEC1D9A2FE8F" authorityName="Mamaev" authorityYear="1998" box="[741,942,331,355]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FD23FEC1D9A2FE8F" box="[741,942,331,355]" italics="true" pageId="9" pageNumber="94">Dicerura formosa</emphasis>
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</taxonomicName>
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, described originally from southern Siberia, is newly reported here from two additional northern European countries.
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</paragraph>
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</subSubSection>
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<caption id="42CB74D2FFEC9663FF51F90EDE9FF955" httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94" startId="9.[151,264,1668,1690]" targetBox="[195,1387,532,1626]" targetPageId="9">
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<paragraph id="160B245AFFEC9663FF51F90EDE9FF955" blockId="9.[151,1435,1668,1721]" pageId="9" pageNumber="94">
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<emphasis id="24C0F848FFEC9663FF51F90EDB5AF976" bold="true" box="[151,342,1668,1690]" pageId="9" pageNumber="94">FIGURES 16–18.</emphasis>
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<taxonomicName id="D1B45FD9FFEC9663FE98F90FD81BF976" authorityName="Mamaev" authorityYear="1998" box="[350,535,1668,1690]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FE98F90FD81BF976" box="[350,535,1668,1690]" italics="true" pageId="9" pageNumber="94">Dicerura formoSa</emphasis>
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</taxonomicName>
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, male, from Sweden.
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<emphasis id="24C0F848FFEC9663FD39F90ED92CF976" bold="true" box="[767,800,1668,1690]" pageId="9" pageNumber="94">16:</emphasis>
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Genitalia, ventral.
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<emphasis id="24C0F848FFEC9663FC2CF90EDE00F976" bold="true" box="[1002,1036,1668,1690]" pageId="9" pageNumber="94">17:</emphasis>
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Ninth tergite, dorsal.
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<emphasis id="24C0F848FFEC9663FB35F90EDF18F976" bold="true" box="[1267,1300,1668,1690]" pageId="9" pageNumber="94">18:</emphasis>
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Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis.
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</paragraph>
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</caption>
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<subSubSection id="5EAE77D1FFEC9660FF01F96ADEF0FEF3" lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94" type="diagnosis">
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<paragraph id="160B245AFFEC9663FF01F96ADF90F915" blockId="9.[151,1436,1760,2009]" box="[199,1436,1760,1785]" pageId="9" pageNumber="94">
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<emphasis id="24C0F848FFEC9663FF01F96ADB4DF915" bold="true" box="[199,321,1760,1785]" pageId="9" pageNumber="94">Diagnosis.</emphasis>
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Characters to differentiate
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<taxonomicName id="D1B45FD9FFEC9663FDB1F96BD8FAF914" authorityName="Mamaev" authorityYear="1998" box="[631,758,1760,1784]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FDB1F96BD8FAF914" box="[631,758,1760,1784]" italics="true" pageId="9" pageNumber="94">D. formosa</emphasis>
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</taxonomicName>
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from the other species of the
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<taxonomicName id="D1B45FD9FFEC9663FB8CF96ADEAAF914" authorityName="Mamaev" authorityYear="1998" box="[1098,1190,1760,1784]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEC9663FB8CF96ADEAAF914" box="[1098,1190,1760,1784]" italics="true" pageId="9" pageNumber="94">formosa</emphasis>
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</taxonomicName>
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group are as follows.
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</paragraph>
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<paragraph id="160B245AFFEC9663FF51F88EDF90F8A8" blockId="9.[151,1436,1760,2009]" pageId="9" pageNumber="94">
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The slightly bent gonostylus, whose apex is clearly tapered, bears large setae posteriorly and clearly smaller setae medially, the latter intermingled with short, dense microtrichia (
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<figureCitation id="8E8F38DFFFEC9663FCA4F8A1D9B8F8A8" box="[866,948,1835,1860]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Fig. 16</figureCitation>
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, ↓1). The ventral gonocoxal emargination is
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</paragraph>
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<paragraph id="160B245AFFEC9663FF51F8C5DF90F859" blockId="9.[151,1436,1760,2009]" pageId="9" pageNumber="94">
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slightly convex basally, reinforced by sclerotization, and bordered by two pairs of gonocoxal processes, of which the ventromedial processes are ovate (
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<figureCitation id="8E8F38DFFFEC9663FD8CF8FCD892F863" box="[586,670,1910,1935]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Fig. 16</figureCitation>
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, ↓2), while the dorsolateral processes are elongate and slightly bent (
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<figureCitation id="8E8F38DFFFEC9663FF59F816DAF8F859" box="[159,244,1948,1973]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Fig. 16</figureCitation>
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, ↓3). The semicircular collar of the tegmen has 5 inconspicuous, sclerotized knobs (
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<figureCitation id="8E8F38DFFFEC9663FB06F816DF1AF859" box="[1216,1302,1948,1973]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Fig. 18</figureCitation>
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). The ninth
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</paragraph>
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<paragraph id="160B245AFFEC9660FF51F84ADEF0FEF3" blockId="9.[151,1436,1760,2009]" lastBlockId="10.[151,1436,131,434]" lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94">
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tergite is unusual in two different aspects. First, the posterior edge has a small, broadly V-shaped incision (
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<figureCitation id="8E8F38DFFFEC9663FA82F84ADF9BF835" box="[1348,1431,1984,2009]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1668,1690]" captionTargetBox="[195,1387,532,1626]" captionTargetId="figure@9.[151,1436,443,1647]" captionTargetPageId="9" captionText="FIGURES 16–18. Dicerura formoSa, male, from Sweden. 16: Genitalia, ventral. 17: Ninth tergite, dorsal. 18: Tegmen and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455517/files/figure.png" pageId="9" pageNumber="94">Fig. 17</figureCitation>
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, ↓4) that is bordered on the inside by subtriangular, densely microtrichose protuberances, and second, the anterior edge is broadly rounded (convex) and reinforced by sclerotization, not straight and faint as in other
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<taxonomicName id="D1B45FD9FFEF9660FB30FF4ADF50FF3B" baseAuthorityName="Jaschhof & Jaschhof" baseAuthorityYear="2013" box="[1270,1372,192,215]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="24C0F848FFEF9660FB30FF4ADF50FF3B" box="[1270,1372,192,215]" italics="true" pageId="10" pageNumber="95">Dicerura</emphasis>
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</taxonomicName>
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. Two non-genitalic characters uncommon in
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<taxonomicName id="D1B45FD9FFEF9660FD8BFF69D8B8FF16" baseAuthorityName="Jaschhof & Jaschhof" baseAuthorityYear="2013" box="[589,692,227,250]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="24C0F848FFEF9660FD8BFF69D8B8FF16" box="[589,692,227,250]" italics="true" pageId="10" pageNumber="95">Dicerura</emphasis>
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</taxonomicName>
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, which were not mentioned in Mamaev’s (1998) description of
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<taxonomicName id="D1B45FD9FFEF9660FA47FF69DAFFFEF3" authorityName="Mamaev" authorityYear="1998" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="24C0F848FFEF9660FA47FF69DAFFFEF3" italics="true" pageId="10" pageNumber="95">D. formosa</emphasis>
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</taxonomicName>
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, are the presence of setae on the anepisternum and the long, sometimes even complete M1+2.
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</paragraph>
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</subSubSection>
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<subSubSection id="5EAE77D1FFEF9660FF01FEA7D948FE5E" pageId="10" pageNumber="95" type="materials_examined">
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||
<paragraph id="160B245AFFEF9660FF01FEA7D948FE5E" blockId="10.[151,1436,131,434]" pageId="10" pageNumber="95">
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||
<emphasis id="24C0F848FFEF9660FF01FEA7DBBEFEAA" bold="true" box="[199,434,301,326]" pageId="10" pageNumber="95">Material examined.</emphasis>
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<collectingCountry id="6EA364CAFFEF9660FE7DFEA7D814FEAA" box="[443,536,301,326]" name="Sweden" pageId="10" pageNumber="95">Sweden</collectingCountry>
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:
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<specimenCount id="00B2EFD3FFEF9660FDE3FEA4D880FEAA" box="[549,652,301,326]" pageId="10" pageNumber="95" type="male">11 males</specimenCount>
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, Lule Lappmark, Jokkmokk, Kaltisbäcken NR, herb-rich old-growth taiga near stream,
|
||
<date id="620A029AFFEF9660FEA2FED8D876FE86" box="[356,634,337,362]" pageId="10" pageNumber="95" value="2016-07-31" valueMax="2016-08-28" valueMin="2016-07-31">31 July–28 August 2016</date>
|
||
, MT, M. & C. Jaschhof (specimens nos CEC1386–CEC
|
||
<quantity id="D14C89BFFFEF9660FB36FEDBDF48FE86" box="[1264,1348,337,362]" metricMagnitude="1" metricUnit="m" metricValue="3.52806" pageId="10" pageNumber="95" unit="in" value="1389.0">1389 in</quantity>
|
||
<collectionCode id="70A5BC9FFFEF9660FA8DFED8DF94FE86" box="[1355,1432,338,362]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="10" pageNumber="95">NHRS</collectionCode>
|
||
, nos CEC1390–CEC
|
||
<quantity id="D14C89BFFFEF9660FEB9FEFCDBDAFE62" box="[383,470,373,398]" metricMagnitude="1" metricUnit="m" metricValue="3.53822" pageId="10" pageNumber="95" unit="in" value="1393.0">1393 in</quantity>
|
||
<collectionCode id="70A5BC9FFFEF9660FE24FEFCD82FFE62" box="[482,547,374,398]" pageId="10" pageNumber="95">SDEI</collectionCode>
|
||
, nos CEC1394–CEC
|
||
<quantity id="D14C89BFFFEF9660FCDCFEFFD97EFE62" box="[794,882,373,398]" metricMagnitude="1" metricUnit="m" metricValue="3.5458399999999997" pageId="10" pageNumber="95" unit="in" value="1396.0">1396 in</quantity>
|
||
<collectionCode id="70A5BC9FFFEF9660FCB8FEFCD9C8FE62" box="[894,964,374,398]" pageId="10" pageNumber="95">IBUL</collectionCode>
|
||
).
|
||
<collectingCountry id="6EA364CAFFEF9660FC1DFEFCDE3CFE61" box="[987,1072,374,397]" name="Estonia" pageId="10" pageNumber="95">
|
||
<emphasis id="24C0F848FFEF9660FC1DFEFCDE3CFE61" box="[987,1072,374,397]" italics="true" pageId="10" pageNumber="95">Estonia</emphasis>
|
||
</collectingCountry>
|
||
: male,
|
||
<collectingRegion id="D470EAB8FFEF9660FB48FEFCDF08FE61" box="[1166,1284,374,397]" country="Estonia" name="Laanemaa" pageId="10" pageNumber="95">Läänemaa</collectingRegion>
|
||
, Kunila near Lihula,
|
||
<date id="620A029AFFEF9660FF28FE13DBA0FE5E" box="[238,428,409,434]" pageId="10" pageNumber="95" value="2010-07-10" valueMax="2010-07-25" valueMin="2010-07-10">10–25 July 2010</date>
|
||
, MT, R. Nikkel &
|
||
<collectionCode id="70A5BC9FFFEF9660FDBBFE10D8C4FE5D" box="[637,712,410,433]" pageId="10" pageNumber="95">EMTP</collectionCode>
|
||
(in
|
||
<collectionCode id="70A5BC9FFFEF9660FD33FE10D934FE5E" box="[757,824,410,434]" country="Estonia" httpUri="http://grbio.org/cool/9bph-kj5p" name="Institute of Zoology and Botany" pageId="10" pageNumber="95">IZBE</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |