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28 KiB
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210 lines
28 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.362.6300" ID-PMC="PMC3867105" ID-Pensoft-Pub="1313-2970-362-29" ID-Pensoft-UUID="EA63FFD35254FFFC603C7A5E8D08FFA0" ID-PubMed="24363599" ID-Zenodo-Dep="578109" ID-ZooBank="569812E982B54BCF84E0D4C43AEE8273" ModsDocID="1313-2970-362-29" checkinTime="1451246566598" checkinUser="pensoft" docAuthor="Erwin, Terry L. & Amundson, Lauren M." docDate="2013" docId="99EAD918EF7D577882248AD3FCB06941" docLanguage="en" docName="ZooKeys 362: 29-54" docOrigin="ZooKeys 362" docPubDate="2013-12-13" docSource="http://dx.doi.org/10.3897/zookeys.362.6300" docTitle="Pseudomorpha (Pseudomorpha) santarita Erwin & Amundson 2013, sp. n." docType="treatment" docUuid="2CF69165-EC7A-49E4-8E75-3FC288BAB924" docUuidSource="ZooBank" docVersion="3" id="EA63FFD35254FFFC603C7A5E8D08FFA0" lastPageNumber="43" masterDocId="EA63FFD35254FFFC603C7A5E8D08FFA0" masterDocTitle="Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group" masterLastPageNumber="54" masterPageNumber="29" pageNumber="42" updateTime="1668157171424" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erwin, Terry L.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Amundson, Lauren M.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group Summer Intern, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-12-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>362</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.362.6300</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.362.6300</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-362-29</mods:identifier>
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<mods:identifier type="ZooBank">569812E982B54BCF84E0D4C43AEE8273</mods:identifier>
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<mods:identifier type="Pensoft-UUID">EA63FFD35254FFFC603C7A5E8D08FFA0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578109</mods:identifier>
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</mods:mods>
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<subSection lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42" type="accounts of taxa">
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<treatment ID-GBIF-Taxon="182235301" LSID="urn:lsid:zoobank.org:act:2CF69165-EC7A-49E4-8E75-3FC288BAB924" httpUri="http://treatment.plazi.org/id/99EAD918EF7D577882248AD3FCB06941" lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
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<subSubSection pageId="13" pageNumber="42" type="nomenclature">
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<paragraph pageId="13" pageNumber="42">
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<taxonomicName LSID="http://zoobank.org/2CF69165-EC7A-49E4-8E75-3FC288BAB924" authority="Erwin & Amundson" authorityName="Erwin & Amundson" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pseudomorpha (Pseudomorpha) santarita" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="santarita" status="sp. n." subGenus="Pseudomorpha">Pseudomorpha (Pseudomorpha) santarita Erwin & Amundson</taxonomicName>
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<taxonomicNameLabel pageId="13" pageNumber="42">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="13" pageNumber="42">Figures 6</figureCitation>
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="13" pageNumber="42">, 12</figureCitation>
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="13" pageNumber="42">, 16</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="13" pageNumber="42">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="42" type="holotype">
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<paragraph pageId="13" pageNumber="42">Holotype.</paragraph>
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<paragraph pageId="13" pageNumber="42">
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<emphasis bold="true" pageId="13" pageNumber="42">USA:</emphasis>
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Arizona,
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<emphasis italics="true" pageId="13" pageNumber="42">Pima County</emphasis>
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, Santa Rita Ranch,
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<geoCoordinate degrees="31.946" direction="north" orientation="latitude" precision="55" value="31.946">31.946°N</geoCoordinate>
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,
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<geoCoordinate degrees="110.758" direction="west" orientation="longitude" precision="55" value="-110.758">110.758°W</geoCoordinate>
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, 1080m, July 1978 (R. Lenczy) (NMNH: ADP110388, male). Paratypes are listed below; see other specimens examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="42" type="derivation of specific epithet">
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<paragraph pageId="13" pageNumber="42">Derivation of specific epithet.</paragraph>
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<paragraph pageId="13" pageNumber="42">
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The epithet "
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<emphasis italics="true" pageId="13" pageNumber="42">santarita</emphasis>
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" is a singular feminine noun used in apposition and is based on the name of the upland range, Santa Rita Mountains in the Coronado National Forest, which includes the type locality of this species. This area was once the home of the indigenous peoples, Papago.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="42" type="proposed english vernacular name">
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<paragraph pageId="13" pageNumber="42">Proposed english vernacular name.</paragraph>
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<paragraph pageId="13" pageNumber="42">Santa Rita False-form beetle.</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="42" type="diagnosis">
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<paragraph pageId="13" pageNumber="42">Diagnosis.</paragraph>
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<paragraph pageId="13" pageNumber="42">
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With the attributes of the species group as described above and color tawny (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="13" pageNumber="42">Fig. 6</figureCitation>
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), color tone of dorsum uniform; form broad and stout; head with preapical lobe prominent, slightly exceeding gena-eye line; pronotum (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="13" pageNumber="42">Fig. 6</figureCitation>
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) slightly wider at base than elytra across humeri; elytron slightly tapered from humerus to narrower truncated apex and with a bare trace of costae, interneurs of finely impressed zig-zag striae, intervals micropunctate, setae very short, fine, and wide-spaced.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42" type="description">
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<paragraph pageId="13" pageNumber="42">Description.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
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(
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="13" pageNumber="42">Figs 6</figureCitation>
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,
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="13" pageNumber="42">12</figureCitation>
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,
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="13" pageNumber="42">16</figureCitation>
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)
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<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Tables with measurements and ratios for Pseudomorpha santarita sp. n.; all measurements are in millimeters." httpUri="http://table.plazi.org/id/5A598BCA8A3B400AD20272A923C53425" pageId="13" pageNumber="42" tableUuid="5A598BCA8A3B400AD20272A923C53425">Table 6</tableCitation>
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.
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<emphasis italics="true" pageId="14" pageNumber="43">
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<pageBreakToken pageId="14" pageNumber="43" start="start">Size</pageBreakToken>
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</emphasis>
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: Medium for genus, ABL = 5.3 to 5.7 mm, SBL = 5.1 to 5.7 mm, TW = 4.4 to 4.5 mm. Preocular lobe-eye ratio (L/L): 0.551 to 0.693. Pronotum ratio (L/W): 0.225 to 0.286. Elytron ratio (L/W): 1.545 to 1.681.
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<emphasis italics="true" pageId="14" pageNumber="43">Color</emphasis>
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: Head, pronotum and elytra tawny, their lateral margins somewhat diaphanous rufous, appendages flavotestaceous.
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<emphasis italics="true" pageId="14" pageNumber="43">Luster</emphasis>
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: Dorsal surface moderately alutaceous, moderately matte.
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<emphasis italics="true" pageId="14" pageNumber="43">Microsculpture</emphasis>
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: Dorsal surface with very fine flat isodiametric sculpticells.
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<emphasis bold="true" pageId="14" pageNumber="43">Head:</emphasis>
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(
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="14" pageNumber="43">Fig. 12</figureCitation>
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) Eye setiferous. Clypeal suture effaced at middle. Frons and vertex with very sparse micropunctulate, setigerous pores with very short setae widely scattered, no transverse line of setae present. Occiput medial to hind margin of eye without small group of coarse setiferous pores.
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<emphasis bold="true" pageId="14" pageNumber="43">Prothorax:</emphasis>
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Pronotum (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="14" pageNumber="43">Fig. 6</figureCitation>
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) moderately convex with broad explanate sides, wider than long, without fringe of long stout setae along lateral margin although present at both anterior and hind angles, fringe of short setae present along anterior and posterior margins; anterior margin bead effaced medially, posterior margin discolored but not beaded; disk with longitudinal pigmented line.
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<emphasis bold="true" pageId="14" pageNumber="43">Pterothorax:</emphasis>
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Elytron (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="14" pageNumber="43">Fig. 6</figureCitation>
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) with interval micropunctate, setigerous pores very widely spaced, finely impressed; interneurs very finely zig-zag striate. Metepisternum longer than wide, surface sparsely setiferous anteriorly. Metasternum sparsely setiferous medially. Metathoracic wing fully developed.
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<emphasis bold="true" pageId="14" pageNumber="43">Abdomen:</emphasis>
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Sternum III broadly and shallowly incised medially. All sterna sparsely setiferous, IV broadly and densely so medially; male with dense patches of setae medially on sterna V and VI, their width coequal to the combined length of posterior basitarsomere plus tarsomere 2.
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<emphasis bold="true" pageId="14" pageNumber="43">Male genitalia:</emphasis>
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(
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="14" pageNumber="43">Fig. 16</figureCitation>
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) Phallobase crested; phalloshaft arched nearly 90°, then straight and depressed in lateral aspect to phalloapex; phalloapex narrowed both in lateral and dorsal aspects to acute tip. Parameres in ventral aspect with left shorter than right and somewhat narrower, distal margins of both narrowly rounded, asetose.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="43" type="dispersal potential">
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<paragraph pageId="14" pageNumber="43">Dispersal potential.</paragraph>
|
||
<paragraph pageId="14" pageNumber="43">These beetles are macropterous and have been recorded at lights, hence capable of flight; they are swift and agile runners. Accordingly, the species may be expected to be more broadly distributed across a wider geographical range than current records indicate.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="43" type="way of life">
|
||
<paragraph pageId="14" pageNumber="43">Way of life.</paragraph>
|
||
<paragraph pageId="14" pageNumber="43">
|
||
Adults are likely found in ant nests and the surrounding vicinity; females are ovoviviparous (
|
||
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Journal of Natural History" pageId="16" pageNumber="45" pagination="1079 - 1086" publicationUrl="10.1080/00222938500770681" refId="B18" refString="Liebherr, JK, Kavanaugh, DH, 1985. Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera). Journal of Natural History 19: 1079 - 1086, 10.1080/00222938500770681" title="Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera)." url="10.1080/00222938500770681" volume="19" year="1985">Liebherr and Kavanaugh 1985</bibRefCitation>
|
||
); larvae are ant nest inquilines (
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopterists Bulletin" pageId="16" pageNumber="45" pagination="53 - 68" refId="B8" refString="Erwin, TL, 1981. A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants. The Coleopterists Bulletin 35 (1): 53 - 68" title="A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants." volume="35" year="1981">Erwin 1981</bibRefCitation>
|
||
). Members of
|
||
<taxonomicName authorityName="Erwin & Amundson" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="santarita">
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Pseudomorpha santarita</emphasis>
|
||
</taxonomicName>
|
||
occur at midland and upland altitudes in between the Sonoran and Chihuahuan Deserts in the riparian vegetation zones with Sycamore (
|
||
<taxonomicName class="Dicotyledoneae" family="Platanaceae" genus="Platanus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="14" pageNumber="43" phylum="Angiospermae" rank="species" species="occidentalis">
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Platanus occidentalis</emphasis>
|
||
</taxonomicName>
|
||
L.) and Cottonwood (
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Populus Fremontii</emphasis>
|
||
S. Wats.) and desert scrub on the slopes. Adults are active in July -August, very hot months in this area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="43" type="other specimens examined">
|
||
<paragraph pageId="14" pageNumber="43">Other specimens examined.</paragraph>
|
||
<paragraph pageId="14" pageNumber="43">
|
||
<emphasis bold="true" pageId="14" pageNumber="43">USA</emphasis>
|
||
: Arizona,
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Santa Cruz County</emphasis>
|
||
, Pajarito Mountains,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
Blanca Canyon,
|
||
<geoCoordinate degrees="31.386" direction="north" orientation="latitude" precision="55" value="31.386">31.386°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="111.093" direction="west" orientation="longitude" precision="55" value="-111.093">111.093°W</geoCoordinate>
|
||
, 1191m, 2 July 1980 (S. McCleve) (UATC: ADP110817, male paratype);
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Graham County</emphasis>
|
||
, Galiuro Mountains north, Ash Creek,
|
||
<geoCoordinate degrees="32.514" direction="north" orientation="latitude" precision="55" value="32.514">32.514°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="110.139" direction="west" orientation="longitude" precision="55" value="-110.139">110.139°W</geoCoordinate>
|
||
, 1400m, 16-17 August 1982 (D.R. Maddison, G.E. Ball & S. McCleve) (DRMC: ADP110591, male paratype);
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Pima County</emphasis>
|
||
, Santa Rita Mountains, Madera Canyon,
|
||
<geoCoordinate degrees="31.724" direction="north" orientation="latitude" precision="55" value="31.724">31.724°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="110.880" direction="west" orientation="longitude" precision="55" value="-110.88">110.880°W</geoCoordinate>
|
||
, 1487m, 11 July 1963 (V.L. Vesterby) (UCDC: ADP111898, female paratype), Madera Canyon, 1499m,
|
||
<geoCoordinate degrees="31.724" direction="north" orientation="latitude" precision="55" value="31.724">31.724°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="110.880" direction="west" orientation="longitude" precision="55" value="-110.88">110.880°W</geoCoordinate>
|
||
, 8 July 1970 (K. Stephan) (FSCA: ADP112570, male paratype). New Mexico,
|
||
<emphasis italics="true" pageId="14" pageNumber="43">Hidalgo County</emphasis>
|
||
, Animas Mountains, Double Adobe Creek,
|
||
<geoCoordinate degrees="31.614" direction="north" orientation="latitude" precision="55" value="31.614">31.614°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="108.779" direction="west" orientation="longitude" precision="55" value="-108.779">108.779°W</geoCoordinate>
|
||
, 1755m, 11 July 1981 (S. McCleve) (UATC: ADP110861, male paratype).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="43" type="geographic distribution">
|
||
<paragraph pageId="14" pageNumber="43">Geographic distribution.</paragraph>
|
||
<paragraph pageId="14" pageNumber="43">
|
||
(
|
||
<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="14" pageNumber="43">Fig. 18</figureCitation>
|
||
). This species is currently known from Arizona and New Mexico.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |