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<document ID-DOI="http://dx.doi.org/10.3897/jhr.70.34206" ID-GBIF-Dataset="2be0b95e-7f6b-4817-b733-c6e2a4ebd224" ID-Pensoft-Pub="1314-2607-70-41" ID-Pensoft-UUID="043FFFE5D662FFB1FF8BFF88FF94FF84" ID-ZBK="B8F3FB3E723749E79AA71AAC9EB4C787" ID-Zenodo-Dep="3269963" ID-ZooBank="B8F3FB3E723749E79AA71AAC9EB4C787" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2607-70-41" ModsDocOrigin="Journal of Hymenoptera Research 70" ModsDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" checkinTime="1561791832741" checkinUser="pensoft" docAuthor="Kimsey, Lynn S." docDate="2019" docId="8C7CC50F9DC194A6F0E0AE6456987056" docLanguage="en" docName="JourHymenoptRes 70: 41-64" docOrigin="Journal of Hymenoptera Research 70" docPubDate="2019-06-28" docSource="http://dx.doi.org/10.3897/jhr.70.34206" docTitle="Cladobethylus darlingi Kimsey 2019, sp. nov." docType="treatment" docUuid="71BED8B5-7E81-4C25-AA13-4E850E01A370" docUuidSource="ZooBank" docVersion="5" id="043FFFE5D662FFB1FF8BFF88FF94FF84" lastPageNumber="41" masterDocId="043FFFE5D662FFB1FF8BFF88FF94FF84" masterDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="41" updateTime="1678751192564" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimsey, Lynn S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology and Nematology, University of California, Davis, California, 95616, USA</mods:affiliation>
<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>64</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.70.34206</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.70.34206</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-70-41</mods:identifier>
<mods:identifier type="ZooBank">B8F3FB3E723749E79AA71AAC9EB4C787</mods:identifier>
<mods:identifier type="Pensoft-UUID">043FFFE5D662FFB1FF8BFF88FF94FF84</mods:identifier>
<mods:identifier type="Zenodo-Dep">3269963</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158471894" LSID="urn:lsid:zoobank.org:act:71BED8B5-7E81-4C25-AA13-4E850E01A370" httpUri="http://treatment.plazi.org/id/8C7CC50F9DC194A6F0E0AE6456987056" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
<taxonomicName LSID="http://zoobank.org/71BED8B5-7E81-4C25-AA13-4E850E01A370" authority="Kimsey" authorityName="Kimsey" authorityYear="2019" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus darlingi" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="darlingi" status="sp. nov.">Cladobethylus darlingi Kimsey</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">
<figureCitation captionStart="Figures 1219" captionStartId="F3" captionText="Figures 12 - 19. 12, 13, 16, 17 Male Cladobethylus 14, 15, 18, 19 Female Cladobethylus 12, 14, 16, 18 Lateral view 13, 15, 17, 19 dorsal view." figureDoi="10.3897/jhr.70.34206.figures12-19" httpUri="https://binary.pensoft.net/fig/313610" pageId="0" pageNumber="41">Figs 12-15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">
Males of this species most closely resemble those of
<taxonomicName lsidName="Cl. cruciger" pageId="0" pageNumber="41" rank="species" species="cruciger">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. cruciger</emphasis>
</taxonomicName>
as discussed under that species. It can be distinguished from
<taxonomicName lsidName="Cl. cruciger" pageId="0" pageNumber="41" rank="species" species="cruciger">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. cruciger</emphasis>
</taxonomicName>
and other male
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
by the combination of the malar space 3.5 midocellar diameters wide or wider, interocular distance more than twice eye width in front view, flagellomere I less than 3.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, flagellomere II less than 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, and flagellomere XI more than 6.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Male description.</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis italics="true" pageId="0" pageNumber="41">Body</emphasis>
(Figs
<figureCitation captionStart="Figures 1219" captionStartId="F3" captionText="Figures 12 - 19. 12, 13, 16, 17 Male Cladobethylus 14, 15, 18, 19 Female Cladobethylus 12, 14, 16, 18 Lateral view 13, 15, 17, 19 dorsal view." figureDoi="10.3897/jhr.70.34206.figures12-19" httpUri="https://binary.pensoft.net/fig/313610" pageId="0" pageNumber="41">12</figureCitation>
,
<figureCitation captionStart="Figures 1219" captionStartId="F3" captionText="Figures 12 - 19. 12, 13, 16, 17 Male Cladobethylus 14, 15, 18, 19 Female Cladobethylus 12, 14, 16, 18 Lateral view 13, 15, 17, 19 dorsal view." figureDoi="10.3897/jhr.70.34206.figures12-19" httpUri="https://binary.pensoft.net/fig/313610" pageId="0" pageNumber="41">13</figureCitation>
). Length 3.0-5.0 mm.
<emphasis italics="true" pageId="0" pageNumber="41">Head</emphasis>
. Frons with punctures 1-2 puncture diameters apart; scapal basin primarily smooth with narrow strip of cross-ridges on either side of broad medial stripe and scattered punctures laterally along eye margins; clypeus long, projecting ventrally, projection apicomedially irregularly truncate; subantennal distance 1.2 MOD long; malar space 3.8 midocellar diameters wide; head about as long as wide; interocular distance 2.2
<normalizedToken originalValue="×">x</normalizedToken>
eye width in front view; midocellus 2.4 diameters from ocular margin; ocelli arranged in nearly equilateral triangle; hindocelli separated from ocular margin by 1.0-1.2 diameters; flagellomere I 3.5-3.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, setae length 0.3
<normalizedToken originalValue="×">x</normalizedToken>
flagellar width; flagellomere II 2.3-2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomere XI 6.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; apical flagellomeres asymmetrically slightly bulging medially.
<emphasis italics="true" pageId="0" pageNumber="41">Mesosoma</emphasis>
. Pronotum slightly shorter than scutum; mesopleural punctures 0.5 puncture diameters apart; metapleuron polished with 1-3 irregular transverse ridges; propodeum laterally polished, with subrectangular posteromedial enclosure.
<emphasis italics="true" pageId="0" pageNumber="41">Metasoma</emphasis>
. Tergum I with medial transverse zone of widely separated tiny punctures, otherwise impunctate; tergum II highly polished, with tiny punctures 2-10 puncture diameters apart, with impunctate longitudinal medial band and posterior margin; terga III-IV with tiny punctures, 1-3 puncture diameters apart, posterior margins impunctate.
<emphasis italics="true" pageId="0" pageNumber="41">Color</emphasis>
. Body black, except head vertex with purplish highlights and dorsum of pronotum, scutum, scutellum and metanotal triangle with metallic blue highlights; metasomal tergum I reddish basally, apically black, often with bluish highlights, terga II-V black, often with bluish highlights; legs including coxae yellow; antenna orange to light brown; wing membrane light brown-tinted, with dark brown veins.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Female description.</paragraph>
<paragraph pageId="0" pageNumber="41">
Body (Figs
<figureCitation captionStart="Figures 1219" captionStartId="F3" captionText="Figures 12 - 19. 12, 13, 16, 17 Male Cladobethylus 14, 15, 18, 19 Female Cladobethylus 12, 14, 16, 18 Lateral view 13, 15, 17, 19 dorsal view." figureDoi="10.3897/jhr.70.34206.figures12-19" httpUri="https://binary.pensoft.net/fig/313610" pageId="0" pageNumber="41">14</figureCitation>
,
<figureCitation captionStart="Figures 1219" captionStartId="F3" captionText="Figures 12 - 19. 12, 13, 16, 17 Male Cladobethylus 14, 15, 18, 19 Female Cladobethylus 12, 14, 16, 18 Lateral view 13, 15, 17, 19 dorsal view." figureDoi="10.3897/jhr.70.34206.figures12-19" httpUri="https://binary.pensoft.net/fig/313610" pageId="0" pageNumber="41">15</figureCitation>
). Length 3.0 mm.
<emphasis italics="true" pageId="0" pageNumber="41">Head</emphasis>
. Frons punctures contiguous to 0.5 puncture diameter apart; scapal basin impunctate, with narrow band of cross-ridges submedially; clypeus short, truncate apicomedially; subantennal distance 0.4 MOD wide; malar space 4 MOD long; head 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, interocular distance 1.1
<normalizedToken originalValue="×">x</normalizedToken>
eye width in front view; postocular distance 0.4 MOD; vertex rounded angulate posterolaterally in dorsal view; midocellus 2.2 MOD from ocular margin; ocelli arranged in isosceles triangle; hindocelli separated from ocular margin by 1 hindocellar diameter; flagellomere I 2.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomere II as long as broad; flagellomere X 1.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad.
<emphasis italics="true" pageId="0" pageNumber="41">Mesosoma</emphasis>
. Mesopleural punctation contiguous; metapleuron polished with metapleural-propodeal suture foveate; propodeum smooth, impunctate below wing.
<emphasis italics="true" pageId="0" pageNumber="41">Metasoma</emphasis>
. Tergum I and II with tiny widely separated punctures, 10 puncture diameters apart or more; terga III and IV punctures 1-3 puncture diameters apart, with impunctate apical rim.
<emphasis italics="true" pageId="0" pageNumber="41">Color</emphasis>
. Body black, without metallic highlights; legs brown; antenna brown; wing membrane light brown tinted, with dark brown veins.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="type material">
<paragraph pageId="0" pageNumber="41">Type material.</paragraph>
<paragraph pageId="0" pageNumber="41">
Holotype male: Indonesia: Borneo,
<emphasis bold="true" pageId="0" pageNumber="41">West Kalimantan</emphasis>
, Gunung Palung Nat. Park, Cabang Panti Res. Sta., 100m, Malaise trap, rainforest alluvial,
<geoCoordinate degrees="1" direction="south" minutes="15" orientation="latitude" precision="925" value="-1.25">1°15'S</geoCoordinate>
,
<geoCoordinate degrees="110" direction="east" minutes="5" orientation="longitude" precision="925" value="110.083336">110°5'E</geoCoordinate>
, 15 Jun-15 Aug, 1991, Darling, Rosichon, Sutrisno, IIS910122 (ROM). Paratypes: 8 males, 2 females; 3 males: same data as holotype; 2 males: same data, except rainforest sandstone, IIS910119; 1 male: rainforest sandstone, light gap, IIS910116; 1 male: 100-400m, rainforest sandstone, IIS910129; 1 male: 29/vi/1991, rainforest alluvial, screen sweep, IIS910165; 1 female: rainforest sandstone, Malaise trap pans, IIS9310134; 1 male: rainforest peat swamp, Malaise trap pans, IIS910132; 1 female: rainforest granite, Malaise trap pans, IIS910115; 1 male: 17-29/vi/1991, rainforest sandstone, Malaise trap, IIS910135 (BME, MZB, ROM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="41">
Borneo. Additional non-type material from Borneo includes three males:
<emphasis bold="true" pageId="0" pageNumber="41">East Kalimantan</emphasis>
, Kac. Pujungan, Kayan-Mentarang Nat. Res., 21/iii/-4/iii/1992,
<geoCoordinate degrees="2" direction="north" minutes="52" orientation="latitude" precision="925" value="2.8666668">2°52'N</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="49" orientation="longitude" precision="925" value="115.816666">115°49'E</geoCoordinate>
, lowland Dipt. Forest, Lalut Birai riparian vegetation 378m, screen sweep, DC Darling, IIS930008; 4 males:
<emphasis bold="true" pageId="0" pageNumber="41">Brunei</emphasis>
, Kuala Belalong, FSC,
<geoCoordinate degrees="4" direction="north" minutes="34" orientation="latitude" precision="925" value="4.5666666">4°34'N</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="7" orientation="longitude" precision="925" value="115.11667">115°7'E</geoCoordinate>
, 18/v/1991, Malaise, N. Mawdsley, BMNH(E)1991-173; 1 male: iii/1991, BMNH(E)1991-173; 1 male: Brunei, Ulu Temburong base camp, 300m, 16/ii-9/iii/1982, M.C. Day; 1 male:
<emphasis bold="true" pageId="0" pageNumber="41">North Borneo</emphasis>
(SE), 19 km n, forest camp, 60 m, 21/xi/1962, K.J. Kuncheria; 1 male: 7/xi/1862; 1 male:
<emphasis bold="true" pageId="0" pageNumber="41">Sarawak</emphasis>
, 4th div., Gn. Mulu, RGS Exp. 17/xi/1923, D. Hollis, BM77-543; 1 male: British N. Borneo, Gomantong Caves, 22-26/xi/1958, TC Maa; 2 males: Borneo, Sarawak, sw Gunung Buda, 64km s Limbang,
<geoCoordinate degrees="4" direction="north" minutes="13" orientation="latitude" precision="925" value="4.2166667">4°13'N</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="56" orientation="longitude" precision="925" value="114.933334">114°56'E</geoCoordinate>
, 22-28/xi/1996, MT, SL Heydon &amp; S Fung; 1 male: 8-15/xi/1996.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="etymology">
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="41">
The species is named after Chris Darling who not only collected this species, but also has collected many other
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
species in Southeast Asia.
</paragraph>
</subSubSection>
</treatment>
</document>