480 lines
65 KiB
XML
480 lines
65 KiB
XML
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<mods:title id="1566930A3B3945DC5D09F2B13FB21E20">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:namePart id="22446E9999596145F99285B73577C39A">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="E7660C82ED840B710227DF76316C8FA6">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="34A5E35AFE6451B9B2A9E9577143E235">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="7B7EE8851F588FC64F4B8A6C9B44DB4B">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:namePart id="B6F3660502C2165379EAD5BC796E2CE5">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="3D3AF3D1CE0945CED45F735FF0C3B431">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:namePart id="E20985FDFA8E3FB28E8E5739DAF32D5D">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="4925AE523E5D43AFF3394622664754ED">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="4B775BF0885EDABA0A810C8658E21BAB">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="529D0F0262F2165DB785770010826CBC">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="3B59D14D8803E73E750E668B957FC478">Panha, Somsak</mods:namePart>
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<mods:affiliation id="DAAB0D6E69278192486E9D9D14D8840F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="C6B91E8F677ED7FDFBC38EA853C827FA">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="8B69872D227813524FB986E3CC3FDA3E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<treatment id="8C1BB00DBF89582596040088370AD469" ID-GBIF-Taxon="209102244" LSID="urn:lsid:plazi:treatment:8C1BB00DBF89582596040088370AD469" httpUri="http://treatment.plazi.org/id/8C1BB00DBF89582596040088370AD469" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="A8150C6218296CD85EACBD01E414DCB1" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="21FF06B1F148C3A39A828AC3135DF4A4" pageId="0" pageNumber="1">
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5.
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<taxonomicName id="ADCF939DC5443FC374195A917ADD9C59" ID-CoL="7SZWZ" ID-ENA="1316078" LSID="8C1BB00D-BF89-5825-9604-0088370AD469" authority="Pfeiffer, 1862" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Rhiostoma hainesi Pfeiffer, 1862</taxonomicName>
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</paragraph>
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<subSubSection id="E59BFC250C7E704673A7FC29DB1DE9EE" pageId="0" pageNumber="1" type="description">
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<figureCitation id="50E65CCD47C41AC4E86548D06EB6FCB1" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)." figureDoi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1">Figs 17B, C</figureCitation>
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<figureCitation id="4A7929DE019562E799C9C75B78BDEB1B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">, 18</figureCitation>
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<figureCitation id="85528E6D4583491E8E268BFB47B383E0" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">, 21</figureCitation>
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<figureCitation id="89C571711219716B0C5C52BB4628A1C4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">, 22</figureCitation>
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<figureCitation id="8FBFB0E7EF06424ACE8435DD7BCF1319" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32B</figureCitation>
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</paragraph>
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<subSubSection id="D4AC349700E5C1E74AE1529B3BFF485F" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="F4E28D567757E303D94644D19999E7FC" pageId="0" pageNumber="1">
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<taxonomicName id="0EF7D6DD3A9CF6385D4B3737847F5C69" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Rhiostoma hainesi</taxonomicName>
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Pfeiffer, 1862a: 115, pl. 12, fig. 8. Type locality: Camboja [Cambodia].
|
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<bibRefCitation id="0F460B720486593168143230D432F599" DOI="https://doi.org/10.5962/bhl.title.10621" author="Pfeiffer, L" journalOrPublisher="Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri" pageId="0" pageNumber="1" refId="B136" refString="Pfeiffer, L, 1865. Monographia Pneumonopomorum viventium. Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri, DOI: https://doi.org/10.5962/bhl.title.10621" title="Monographia Pneumonopomorum viventium." url="https://doi.org/10.5962/bhl.title.10621" year="1865">Pfeiffer 1865</bibRefCitation>
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||
: 38, 39.
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<bibRefCitation id="4DE256890621B66C5B70F344BD709810" author="Martens, E von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B91" refString="Martens, E von, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." title="Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." year="1867">Martens 1867</bibRefCitation>
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: 64.
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<bibRefCitation id="BA850CBF170DACA8467123FAE9816602" DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun" pageId="0" pageNumber="1" pagination="1 - 193" refId="B40" refString="Fischer, P, 1891. Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun 4: 1 - 193, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" volume="4" year="1891">Fischer 1891</bibRefCitation>
|
||
: 101.
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||
<bibRefCitation id="D95B2E2D5F644D625ED81BB3492FE03E" author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
|
||
: 178.
|
||
<bibRefCitation id="49BC32074DE63D5BDD3A52D061A59129" author="Fischer, H" editor="Pavie, A" journalOrPublisher="Ernest Leroux, Editeur, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B42" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour. In: Pavie, A, Ed., Mission Pavie, Indo-Chine 1879-1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III. Ernest Leroux, Editeur, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour." volumeTitle="Mission Pavie, Indo-Chine 1879 - 1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
|
||
: 427. Kobelt 1911: 762, pl. 113, fig. 1.
|
||
<bibRefCitation id="978E3EC725B0C6B2D98BCBE75B97E2A6" author="Fischer, P-H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B41" refString="Fischer, P-H, 1973. Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches." volume="90" year="1973">Fischer 1973</bibRefCitation>
|
||
: 47.
|
||
<bibRefCitation id="6801CC143443C393852D416790F66C3C" author="Tarruella, A" journalOrPublisher="Spira" pageId="0" pageNumber="1" pagination="187 - 192" refId="B164" refString="Tarruella, A, Domenech, JL, 2010. Listado taxonomico ilustrado de la familia Cyclophoridae J.E. Gray, 1847 (Mollusca: Gastropoda): Parte 1. El genero Rhiostoma Benson, 1860. Spira 3: 187 - 192" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 1. El genero Rhiostoma Benson, 1860." volume="3" year="2010">
|
||
Tarruella and
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||
<normalizedToken id="DDB83B191669383C654B3FCBEC8E6953" originalValue="Doménech">Domenech</normalizedToken>
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2010
|
||
</bibRefCitation>
|
||
: 189, fig. 1d.
|
||
<bibRefCitation id="6E844907DF76CEB0F96381CFEFEE8F53" author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
|
||
: 35-40, figs 4.7-4.9.
|
||
<bibRefCitation id="ACE45A9D5B21E1DF0B4CB203EE226687" author="Kongim, B" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="13 - 20" refId="B78" refString="Kongim, B, Sutcharit, C, Tongkerd, P, Panha, S, 2013b. Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae). The Raffles Bulletin of Zoology 61: 13 - 20" title="Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae)." volume="61" year="2013 b">Kongim et al. 2013b</bibRefCitation>
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||
: 16, fig. 2e.
|
||
<bibRefCitation id="1FD827773E906E90D8C9E275734BE456" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
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: 28, fig. 6f, g.
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</paragraph>
|
||
<paragraph id="1F8BEF4A621307060BEC811416C57CCA" pageId="0" pageNumber="1">
|
||
<taxonomicName id="69CBDD846545E27124D7B74B4797DCCB" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos hainesii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesii">Pterocyclos hainesii</taxonomicName>
|
||
[sic]-
|
||
<bibRefCitation id="357B85E83C57A8C78AC7F02B7976AD3C" author="Reeve, LA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" refId="B145" refString="Reeve, LA, 1863. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1-5. Lovell Reeve and Co., London." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1 - 5. Lovell Reeve and Co., London." year="1863">Reeve 1863</bibRefCitation>
|
||
:
|
||
<taxonomicName id="83B22AD0C2E1B1FB62DEBFBD465F0766" authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclos</taxonomicName>
|
||
pl. 4, species 19.
|
||
</paragraph>
|
||
<paragraph id="08001C94418DCFEB7A39851E7F114B3B" pageId="0" pageNumber="1">
|
||
<taxonomicName id="838C59D47BC98CCBCF1713A4A378BB9D" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclus</taxonomicName>
|
||
[sic] (
|
||
<taxonomicName id="78A4AD78B9773ECD558C2FBCC03B9FFC" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
|
||
)
|
||
<taxonomicName id="FCA224EE2494496191539D7FB0CEA745" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Rhiostoma hainesi</taxonomicName>
|
||
-
|
||
<bibRefCitation id="846A4A4EFB82536AA86C3C7F9A1B5EAF" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Calcutta, Trustees" pageId="0" pageNumber="1" refId="B114" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Calcutta, Trustees, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
|
||
: 263.
|
||
</paragraph>
|
||
<paragraph id="49859BAA65E8845DDBF546FBF5C5769E" pageId="0" pageNumber="1">
|
||
<taxonomicName id="FAB98394B90621A94B63B03039AA98F1" authorityName="Bartsch" authorityYear="1932" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma smithi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="smithi">Rhiostoma smithi</taxonomicName>
|
||
Bartsch, 1932: 70, 71, fig. 1. Type locality: Khao Sabap, southern Siam [Khao Sabap, Khlung District, Chanthaburi Province, Thailand].
|
||
<bibRefCitation id="EEAACCAAAA8E2D428B5C4FD446A7A9EF" author="Abbott, RT" journalOrPublisher=", Melbourne, Florida" pageId="0" pageNumber="1" refId="B1" refString="Abbott, RT, 1989. Compendium of Landshells. American Malacologists Inc., Melbourne, Florida" title="Compendium of Landshells. American Malacologists Inc." year="1989">Abbott 1989</bibRefCitation>
|
||
: 34, with 2 figures (paratype).
|
||
<bibRefCitation id="1A2E90D3395A00FEAFBE484EBAC23FBC" author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
||
: 11.
|
||
<bibRefCitation id="308CDC43B879AFE556C72F8B06423354" author="Panha, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="1 - 3" refId="B124" refString="Panha, S, Thanamitramanee, P, 1997. Land snails of Plieu National Park, Thailand. The Papustyla 114: 1 - 3" title="Land snails of Plieu National Park, Thailand." volume="114" year="1997">Panha and Thanamitramanee 1997</bibRefCitation>
|
||
: 2.
|
||
<bibRefCitation id="C91C7C6E995822164327B5865E077F37" author="Hemmen, J" journalOrPublisher="La Conchiglia" pageId="0" pageNumber="1" pagination="1 - 26" refId="B62" refString="Hemmen, J, Hemmen, C, Patamakanthin, S, 1999. Rhiostoma (Fam. Cyclophoridae). La Conchiglia 31: 1 - 26" title="Rhiostoma (Fam. Cyclophoridae)." volume="31" year="1999">Hemmen et al. 1999</bibRefCitation>
|
||
: 8, fig. 4.
|
||
<bibRefCitation id="7AEF555A5A73146966AB227B0066DB27" author="Patamakanthin, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B125" refString="Patamakanthin, S, 2001. Rhiostoma from Thailand and Malaysia. Of Sea and Shore 23: 222, 223." title="Rhiostoma from Thailand and Malaysia. Of Sea and Shore 23: 222, 223." year="2001">Patamakanthin 2001</bibRefCitation>
|
||
: 222, 223, fig. m1-m3. New synonym.
|
||
</paragraph>
|
||
<paragraph id="F94D6CE9431EBDAB2D32AF74F953D35A" pageId="0" pageNumber="1">
|
||
<taxonomicName id="F917659F8772A52B6E6CB9C0C6C20F61" authorityName="Salisbury" authorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma tomlini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tomlini">Rhiostoma tomlini</taxonomicName>
|
||
Salisbury, 1949: 41, 42, pl. 3b, figs 3, 4. Type locality: Khao Sabap, Siam [Khao Sabap, Khlung District, Chanthaburi Province, Thailand].
|
||
<bibRefCitation id="3DA6E2A0343FEB4B3B9F3E3502174FFF" author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
||
: 11.
|
||
<bibRefCitation id="018CC43F67C466DB4858944D024D5B1E" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
|
||
: 51, fig. 12d.
|
||
<bibRefCitation id="40535157A2B8C256E58B780AC2F91D21" DOI="https://doi.org/10.1093/mollus/eyac015" author="Chen, Z-Y" journalOrPublisher="Paleobiology" pageId="0" pageNumber="1" refId="B29" refString="Chen, Z-Y, Huang, B, Pall-Gergely, B, 2022. Exceptional among the exceptional: a new species of Stenogyropsis Moellendorff, 1899 (Eupulmonata: Camaenidae), with a review of the free last whorl in terrestrial gastropods. Journal of Molluscan Studies 88(2): eyac015. https://doi.org/10.1093/mollus/eyac015" title="Exceptional among the exceptional: a new species of Stenogyropsis Moellendorff, 1899 (Eupulmonata: Camaenidae), with a review of the free last whorl in terrestrial gastropods. Journal of Molluscan Studies 88 (2): eyac 015." url="https://doi.org/10.1093/mollus/eyac015" year="2022">Chen et al. 2022</bibRefCitation>
|
||
: fig. 1b. New synonym.
|
||
</paragraph>
|
||
<paragraph id="1002BBF6EDF3591D17C280DF48DC1909" pageId="0" pageNumber="1">
|
||
<taxonomicName id="604D87B0018401752C0F43A41EB24A91" class="Gastropoda" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhinostoma</taxonomicName>
|
||
[sic]
|
||
<taxonomicName id="1F6684D11CD761E6460DBF9D17B8C647" authorityName="Habe" authorityYear="1961" class="Gastropoda" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma housei subsp. kirai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="housei" subSpecies="kirai">Rhinostoma housei kirai</taxonomicName>
|
||
Habe, 1965: 128 [in part], pl. 2, fig. 11. Type locality: Chanthaburi, Thailand. New synonym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AC19545544D05296A52B613DDD639D0" pageId="0" pageNumber="1" type="type material">
|
||
<paragraph id="77F41C4E71BBB96CD86487772DB8CBCE" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="6F94EF5E43BC6F4062C58733AB1999ED" pageId="0" pageNumber="1">
|
||
<emphasis id="65E9F54A2829BAB412E2A04EFDCE19C7" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
|
||
NHMUK 20170371 (2 shells; Fig.
|
||
<figureCitation id="8AC3521EF960E6356F467C6C746C50AC" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21A</figureCitation>
|
||
) from Camboja.
|
||
<emphasis id="BD1AD0A7E4DB6FC4552711429325BB6D" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
NMNH 382943 (Fig.
|
||
<figureCitation id="3FD0DD8D39FF336AB00F5B16DE8B28F2" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21B</figureCitation>
|
||
) of
|
||
<taxonomicName id="77C00150F91AAE6F9B0983D2846DE161" authorityName="Bartsch" authorityYear="1932" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma smithi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="smithi">
|
||
<emphasis id="61399E3596247C4ADE9C5D656CD6A3F6" italics="true" pageId="0" pageNumber="1">Rhiostoma smithi</emphasis>
|
||
</taxonomicName>
|
||
Bartsch, 1932, from Kao Sabap, S.E. Siam.
|
||
<emphasis id="E4612A4FDC0C7E8845AEDA22781B514A" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
NMW 1955.158.24924 (Fig.
|
||
<figureCitation id="1647F844A1BB7606B503B9D1B52E9F2A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21C</figureCitation>
|
||
) of
|
||
<taxonomicName id="C1C347BF4CD39DF1F99922E5E99E5038" authorityName="Salisbury" authorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma tomlini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tomlini">
|
||
<emphasis id="ADDF63DC093A7B109D725B334E4691C7" italics="true" pageId="0" pageNumber="1">Rhiostoma tomlini</emphasis>
|
||
</taxonomicName>
|
||
Salisbury, 1949, from Khao Sabap, Thailand,
|
||
<emphasis id="238BB862DD9B4AFEDF4C36C660CF54F0" bold="true" italics="true" pageId="0" pageNumber="1">paratype</emphasis>
|
||
NHMUK 1949.6.7.1 (2 shells; Fig.
|
||
<figureCitation id="E608A4308270E3F5334AEDFE845B082B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21D</figureCitation>
|
||
) from Khao Sabap, Siam, and
|
||
<emphasis id="D2B00B3FED58B0788FA4743DB3C18AA2" bold="true" italics="true" pageId="0" pageNumber="1">paratype</emphasis>
|
||
NHMUK 20170372 (1 juvenile shell) from Khao Sabap, Siam.
|
||
<emphasis id="1F444080EAF1E51B1DED2884B345A6C2" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
NSMT 52242 (Fig.
|
||
<figureCitation id="7C486F343FF168EC1B37B3DEE0E0AEC6" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21E</figureCitation>
|
||
) of
|
||
<taxonomicName id="7BAEEBDA047D64278BB1FC8C91A1C573" authorityName="Habe" authorityYear="1965" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei subsp. kirai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="housei" subSpecies="kirai">
|
||
<emphasis id="2FA38937946BD530F81EA38C52038C04" italics="true" pageId="0" pageNumber="1">Rhiostoma housei kirai</emphasis>
|
||
</taxonomicName>
|
||
Habe, 1965, from Chanthaburi, Thailand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="176A54CF8B72384831C966674B0D0E7B" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="5CD2035079D5D20D573305DB68EE7783" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||
<paragraph id="6C62BBC8B6986A4A7A70687B589D938F" pageId="0" pageNumber="1">
|
||
<materialsCitation id="6B6878955B6F4B0B3FB2D5A8B72389D7" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Chanthaburi" specimenCount="1" stateProvince="Chanthaburi">
|
||
<emphasis id="EC43A5A041A3DE7B0E23534AF9C3082F" bold="true" pageId="0" pageNumber="1">
|
||
<collectingCountry id="1A86ECE0C5EFFA742FFA1FF487421F17" name="Thailand">Thailand</collectingCountry>
|
||
</emphasis>
|
||
: Khao Yai, Muang, Nakornnayok: CUMZ 4458. Khoa Sib Ha Chun, Kaeng Hang Maeo,
|
||
<collectingRegion id="ABCE3238081E5BC11D2E77FD045B2E6B" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10075, 10107. Klong Plub, Khlung,
|
||
<collectingRegion id="990AA247BCA7A5BD6E6A7695505734BD" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10017. Makok Waterfall, Khlung,
|
||
<collectingRegion id="1AE30015AF253EFA8A0075D0FF860CA7" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 3807, 3814, 3920, 3921, 4421, 4814 (Fig.
|
||
<figureCitation id="A6C99EB128B7A3C18AC11F584B8A311A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22B</figureCitation>
|
||
), 4819, 4827, 10102, 10105, 10109. Trok Nong Waterfall, Khlung,
|
||
<collectingRegion id="35302F5EC37F73608B352CEB23C70258" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 4455. Ban Ma Kham, Ma Kham,
|
||
<collectingRegion id="8B5FFA89D8394F8B5D5F6C6C58F6FEFB" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10099. Phrabat Khao Ma Kham, Ma Kham,
|
||
<collectingRegion id="B0532AAE766D183E453965528360009B" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10116. Khlong Narai waterfall, Muang,
|
||
<collectingRegion id="90558D2BC358F4584CA121B876568F82" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10101. Plieu Waterfall,
|
||
<collectingRegion id="1252A7EC479B01090012A7B00607F956" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 3801, 4336, 4338, 4339, 4340, 4380, 4381 (Figs
|
||
<figureCitation id="C09B767AB962CB74351461890EED1197" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21F</figureCitation>
|
||
,
|
||
<figureCitation id="44B407B2F6517905FEE2008FAEE9AE23" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22A</figureCitation>
|
||
), 4393, 4394, 4403, 4404, 4405, 4418, 4422, 4425, 4426, 4428, 4434, 10106, 10194 (Fig.
|
||
<figureCitation id="DB6E49415C0718D9291D09DEE9B72E90" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22D</figureCitation>
|
||
). Si Bha Chan Waterfall, Muang,
|
||
<collectingRegion id="B3E926B3D06F9324B79EDC4C7C1C1988" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 4733. Hin Dat waterfall, Pong Nam Ron,
|
||
<collectingRegion id="12173D0C27711BB47724420622BE1DD7" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10100. Khao Soi Dao Wildlife Breeding Center, Soi Dao,
|
||
<collectingRegion id="066C51B19DC801524C011B0F9E0704F2" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 3855, 3883, 3898, 3917, 4402, 4423, 4457, 4743, 10114, 10117. Wat Khao Sukim, Tha Mai,
|
||
<collectingRegion id="3F29498B6FC4C2CCF08C4D3618038C8A" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 3803, 3815, 3828, 3918, 3919, 4867, 10103, 10108, 10110, 10111, 10118, 10119, 10120. Wat Tham Klong Tip, Tha Mai,
|
||
<collectingRegion id="FEA7B1A1D63AE3196805A260D7AFE354" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10112.
|
||
<collectingRegion id="FCEAD889334471C437E2A5D6B7BFBACB" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
: CUMZ 10113. Khao Chamao-Khao Wong, Khao Chamao,
|
||
<collectingRegion id="3C47A14A281A7E28DD8B6BD5160543BE" country="Thailand" name="Rayong">Rayong</collectingRegion>
|
||
: CUMZ 4456, 4816 (Fig.
|
||
<figureCitation id="2823B905B15232FA194433862A81283F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22C</figureCitation>
|
||
), 4822, 10098. Khao Chamao Waterfall, Khao Chamao,
|
||
<collectingRegion id="00727AAF8A33673B21F1061F40B337EA" country="Thailand" name="Rayong">Rayong</collectingRegion>
|
||
: CUMZ 3897. Klong Pla Kang waterfall, Khao Chamao-Khao Wong, Khao Chamao,
|
||
<collectingRegion id="FD6C6F781DEFC4C4F6A445911AEE523A" country="Thailand" name="Rayong">Rayong</collectingRegion>
|
||
: CUMZ 10104. Wat Chat San Phatthanaram, Bo Rai,
|
||
<collectingRegion id="4214B4F7B55D9A2C9A43ADE4AE657E66" country="Thailand" name="Trat">Trat</collectingRegion>
|
||
: CUMZ 10115
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="1BE36872D338008CF7BED7D71B095BB7" doi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
|
||
<paragraph id="771CB3F1181967DBA5CDFE353F4E1726" pageId="0" pageNumber="1">
|
||
<emphasis id="1E1F111E3EA47F0D01DB74989FE40AFC" bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
|
||
Shell of
|
||
<taxonomicName id="840D4CD24669C77823A9F6CD325ECFE2" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||
<emphasis id="99990F1E4704C6A77803D13202767D5B" italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B7D90CB829D4FAF702953CF189BBB824" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<typeStatus id="6C7694008FB9E50889577CF45CD663EE">syntype</typeStatus>
|
||
NHMUK 20170371 from
|
||
<collectingCountry id="C016E2652C4FE68A434F7AF3360D4EF8" name="Cambodia">Camboja</collectingCountry>
|
||
<emphasis id="51CD6812FA1A614B2AF96A6DB78A6FC2" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<typeStatus id="6F07051688F03C13DE23A7B6CB11536D">holotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="BA0346287C7357CF27059B5A15A26728" genus="Rhiosotma" lsidName="Rhiosotma smithi" pageId="0" pageNumber="1" rank="species" species="smithi">
|
||
<emphasis id="5020159C7083DEDA7B4E88A61E5C9A12" italics="true" pageId="0" pageNumber="1">Rhiosotma smithi</emphasis>
|
||
</taxonomicName>
|
||
Bartsch, 1932" NMNH 382943 from Khao Sabap, southern
|
||
<collectingCountry id="8B87C57BCCA266AEA42996F6B15C27C4" name="Thailand">Siam</collectingCountry>
|
||
<emphasis id="A5561F28AD904B8B5777600741DBD8DB" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
originally labelled as "
|
||
<taxonomicName id="5DB89E042EC55587F255BD0C65A2CEB3" authorityName="Salisbury" authorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma tomlini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tomlini">
|
||
<emphasis id="88FDE294D67491E16BACF3C24BF193AE" italics="true" pageId="0" pageNumber="1">Rhiostoma tomlini</emphasis>
|
||
</taxonomicName>
|
||
Salisbury, 1949" from Khao Sabap,
|
||
<collectingCountry id="04BADA6E17CB232F85A865C166F0948B" name="Thailand">Siam</collectingCountry>
|
||
<emphasis id="8510F16FBDF3DFA5A8D6F00353DA61DD" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<typeStatus id="77A3FD56A5511E3487AD97203C9F5D07">holotype</typeStatus>
|
||
NMW 1955.158.24924 and
|
||
<emphasis id="4A3F5AA0748113F5212266489E271566" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<typeStatus id="5848FD4FE9AE03EE919AC6517B21772C">paratype</typeStatus>
|
||
NHMUK 1949.6.7.1
|
||
<emphasis id="A04C7404116C069C90FD4818C4D3517E" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<typeStatus id="DB503F2E6D0B786ADA83834CB1AB9DA3">holotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="2038756180261A94108704A1C12373B2" authorityName="Habe" authorityYear="1965" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei subsp. kirai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="housei" subSpecies="kirai">
|
||
<emphasis id="04927AC00B65F83513B9A016B3CCE0EC" italics="true" pageId="0" pageNumber="1">Rhiostoma housei kirai</emphasis>
|
||
</taxonomicName>
|
||
Habe, 1965" NSMT 52242 from
|
||
<collectingRegion id="58607702D1F49E54DC6E2E2AA3CF9806" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
,
|
||
<collectingCountry id="7CB846F1CFB1C9AE6FBEC16D088F8E01" name="Thailand">Thailand</collectingCountry>
|
||
<emphasis id="F4FEED1D1228F0AF2E429ED25950D3F6" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
specimen CUMZ 4381 from
|
||
<collectingRegion id="9381E13D3052A7197B9945AA289BC85E" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="37414B299404C632F7069AAA214FBFC1" doi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1" start="Figure 22" startId="F22">
|
||
<paragraph id="9A76A63C7D6917C52E867545745F5805" pageId="0" pageNumber="1">
|
||
<emphasis id="4E715944381367A78B06873EA3EC1366" bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
|
||
Shell of
|
||
<taxonomicName id="ADCC6DD47149349430F7D906448707D7" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||
<emphasis id="ACFB44BBB04E666673AD9177161E6CCC" italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="505B7C158E759588BE32871548EC8B3B" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
specimen CUMZ 4381 from
|
||
<collectingRegion id="8A6B8F2EB54CB26BC163BA279C287175" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
<emphasis id="67A116FBB855DC31E5ADD9553702BB6A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
specimen CUMZ 4814 from Makok Waterfall,
|
||
<collectingRegion id="4B949DA70BEA9A2DB44CE9C87B511C30" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
<emphasis id="07890697C479F996FE665F7F78F36185" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
specimen CUMZ 4816 from Khao Chamao,
|
||
<collectingRegion id="ECD39161E4103B397002100D652D0E91" country="Thailand" name="Rayong">Rayong</collectingRegion>
|
||
<emphasis id="A332AFF37AF463E6F92E8DF0E70A704E" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
specimens CUMZ 10194 from
|
||
<collectingRegion id="B8C027E62A07BB4477848AA5C03A0652" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
shows series of growth from small juvenile (on the left) to nearly adult (on the right).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="7E5A821C1959292755A0F8CF5F33EFB5" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="844B8E53F55FA07327381D99D18EBD17" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="DA6F9EAD83B064C661BD5EEA3A0302DB" pageId="0" pageNumber="1">Shell large, thick, and depressed. Detached whorl longer than aperture width, curved and descending. Apertural lip much expanded on palatal wall. Breathing device usually tubular and its tip often attached to preceding whorl. Shell usually with thick dark brown periostracum.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8878A01E19E64D2453AC4F680C33F8E" pageId="0" pageNumber="1" type="differential diagnosis">
|
||
<paragraph id="0ECBC53548E01686E3CE2A64C7F7EBA5" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||
<paragraph id="487D5B53E4A64824E5FFFA9C80F9AC99" pageId="0" pageNumber="1">
|
||
<taxonomicName id="2CC220BFE25F7828E8566B21A836D0D2" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||
<emphasis id="879986ED57909C20E47A8BA4BFDE8B02" italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="C560DFCF7198F15B67E36B87FD239E54" lsidName="R. rhothonotaphrosa" pageId="0" pageNumber="1" rank="species" species="rhothonotaphrosa">
|
||
<emphasis id="BB3C96847B508B561F80CBB3353EFFA8" italics="true" pageId="0" pageNumber="1">R. rhothonotaphrosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by having a large and stout last whorl, tubular- or notch-shaped breathing device, thickened brownish periostracum, and lip slightly thickened and widely expanded. In contrast,
|
||
<taxonomicName id="16FE2D8895E076196D343B8C9EF88086" lsidName="R. rhothonotaphrosa" pageId="0" pageNumber="1" rank="species" species="rhothonotaphrosa">
|
||
<emphasis id="014A8CD1B581E972E22D63877C62C22C" italics="true" pageId="0" pageNumber="1">R. rhothonotaphrosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has a smaller shell size, slender last whorl, notch-shaped breathing device, white shell colour with brownish zigzag pattern, periostracum thin corneous, and lip thickened, slightly expanded, and multi-layered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C6724B43D5E27740017EB37678040AA9" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="349F460B0E18DFF596E80D9EE200C05F" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="8A6838BE70C012734B412F948B04AE38" pageId="0" pageNumber="1">
|
||
<emphasis id="4DCBE5912A85BB42E7982274D964AC22" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="021E04318432BAB960313A32B547AFF9" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shell large, cW 22.1-27.8 mm, cH 11.3-15.2 mm, thick, and discoidal shape; detached-whorl length 11.0-28.5 mm. Apex acute; spire elevated. Whorls 5 to 6, convex, increasing regularly; suture wide and slightly deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thick corneous with brownish to dark brown colour (without periostracum shell uniformly whitish to yellowish colour); with narrow dark spiral band on periphery. Detached whorl long, ~2
|
||
<normalizedToken id="27D1A6A24DF782B9904413D05BAD9081" originalValue="×">x</normalizedToken>
|
||
longer than apertural width, curved and descending. Peristome circular and double; lip thickened, broadly expanded on palatal wall, and multi-layered. Aperture opened sub-laterally to sub-ventrally. Breathing device typically tubular (incomplete tube and notch shape sometime present), and its tip sometimes attached to preceding whorl; outer lip forms a short to long closed tube and is located just behind apertural lip; inner lip with small hole inside aperture (or deep incision in notch shape). Umbilicus widely opened and deep. Operculum calcareous, cup-shaped, and multispiral (Figs
|
||
<figureCitation id="C7EEAB62687D3B14A351E8F0859616A4" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21</figureCitation>
|
||
,
|
||
<figureCitation id="406F208E45CADA004A1927BF427EB02A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BFFEBFA0182190812CFFB2CE9CE9BE5A" pageId="0" pageNumber="1">
|
||
<emphasis id="097CDEB6DFFFB76DAFE55C45F6ADBCCA" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="827EAA970AB961D173D32634EB571834" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||
.
|
||
</emphasis>
|
||
Teeth arrangement and shape are almost identical to
|
||
<taxonomicName id="860CFF09A51EB8F5D43E8C1B5346AFF9" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="0E8CF9FD599F49F7FEC8A96B75D1F27E" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
. Central tooth with large central cusp and two lateral cusps on each side. Lateral teeth composed of four cusps; central cusp large with blunt tip; two inner cusps with pointed tips; one outer cusp small and dull. Marginal teeth each consisting of three pointed cusps (Fig.
|
||
<figureCitation id="76E1DD7107609C828B14B5DB8B3F3C79" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4C7277DA7B9AC69FAF5C0715BE27C07E" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="B5CD66D3EB4D9C36891291AC036753D8" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="01C2536AB9EA73B099617029014241E3" pageId="0" pageNumber="1">
|
||
The previous records of
|
||
<taxonomicName id="0A1A99C297B2F5C1794C67A7B7D8FF18" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="043359942BEEB14933D8ACBBBA27A6A4" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
were mostly from vague locations in Cambodia and Thailand: Khao Sabap (= Plieu National Park, Chanthaburi) (
|
||
<bibRefCitation id="1C71E0AE913A6B5F99AAF5199971AD59" author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="115 - 117" refId="B134" refString="Pfeiffer, L, 1862a. Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq. Proceedings of the Zoological Society of London 30: 115 - 117" title="Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq." volume="30" year="1862 a">Pfeiffer 1862a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E11E6EAB0B0B9E0BC0C8EB255DF76470" author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" pagination="111 - 128" refId="B53" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia, Kyoto 4: 111 - 128" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
|
||
). The recent study obtains specimens from several localities in Nakhonnayok, Rayong, and Chanthaburi provinces (Fig.
|
||
<figureCitation id="311D58F878C3B49B75C783F8DC74C3BE" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A40171CA9F37C329B7C06D844E33FC40" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="3C32EAFD6A954C28EA7AAB61CF06504F" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="8889EE56F16B80C2EC377B887D219977" pageId="0" pageNumber="1">
|
||
Three nominal taxa have long been confused with
|
||
<emphasis id="E26900270461A37E093780B7A00D31F7" italics="true" pageId="0" pageNumber="1">
|
||
<taxonomicName id="8551723AE47DB7B2180CBC05FBA268DC" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">R. hainesi</taxonomicName>
|
||
:
|
||
<taxonomicName id="65AA352EA2EA8B89FD73E76F7A6A07C6" lsidName="R. smithi" pageId="0" pageNumber="1" rank="species" species="smithi">R. smithi</taxonomicName>
|
||
</emphasis>
|
||
Bartsch, 1932,
|
||
<taxonomicName id="28872051C99D0F038008DAEAAA0F4B90" lsidName="R. tomlini" pageId="0" pageNumber="1" rank="species" species="tomlini">
|
||
<emphasis id="4709BAC5CB5CEA77853880641153EB67" italics="true" pageId="0" pageNumber="1">R. tomlini</emphasis>
|
||
</taxonomicName>
|
||
Salisbury, 1949, and
|
||
<taxonomicName id="52787166C81AA5199A7153989E72B3D9" lsidName="R. housei subsp. kirai" pageId="0" pageNumber="1" rank="subSpecies" species="housei" subSpecies="kirai">
|
||
<emphasis id="EAC132A37032D96976E67B07E1E83EA5" italics="true" pageId="0" pageNumber="1">R. housei kirai</emphasis>
|
||
</taxonomicName>
|
||
Habe, 1965. The first two species were described based only on differences in shell colour and shapes of the breathing device. The type localities of both are from the same location at "Khao Sabap" [= Plieu National Park, Khlung, Chanthaburi, Thailand], and their holotypes are identical with
|
||
<taxonomicName id="ECAEDB3E904FC9F15D4494B4FF2B4FD2" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="88C7A0B14131B749F442540CEBE4F14A" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="DB92A4BA74BABBCD4E43E275208BC5C0" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21A-D</figureCitation>
|
||
). The last taxon,
|
||
<taxonomicName id="12C9A08E2EEC8D7EEF92B87612C50B9F" lsidName="R. housei subsp. kirai" pageId="0" pageNumber="1" rank="subSpecies" species="housei" subSpecies="kirai">
|
||
<emphasis id="D7CD58F8B6D315A15C80718EDE6B7FB3" italics="true" pageId="0" pageNumber="1">R. housei kirai</emphasis>
|
||
</taxonomicName>
|
||
, was nominated as a subspecies of
|
||
<taxonomicName id="7E222A44782CDB74CDE639094070F7F7" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="C5D931520F1EC1C52AAFCB662EB95B3B" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="D3E54A47002AAB64DF26AFD7B368483E" author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" pagination="111 - 128" refId="B53" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia, Kyoto 4: 111 - 128" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
|
||
). However, the holotype (Fig.
|
||
<figureCitation id="63E3100F790FB7E0F18084AFE326851F" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21E</figureCitation>
|
||
) exhibits a long and curved detached whorl, short tubular breathing device, expanded and thick apertural lip, and brownish periostracum (remaining inside umbilical area), which are the unique characters of
|
||
<taxonomicName id="12E1A06E67C098645DD908C6D4F92548" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="4FAA6BB609EDAB2BB53E58616E3FB19C" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the type locality was Chanthaburi Province, which is within the distribution range of
|
||
<taxonomicName id="3F45FB3EE95657323EE62F8B6F09E176" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="6D75B66EB9BF0A4BDDC4D849A68FCB09" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, we recognise these three nominal taxa as junior subjective synonyms of
|
||
<taxonomicName id="1B012D1CC6DD77645A37AFEC2315B5F8" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="D80DE787D438344AB03F34DED3227B14" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="89B46E120D866E73168C18F70CDDA699" pageId="0" pageNumber="1">
|
||
The three variations in breathing device shape were reported in
|
||
<bibRefCitation id="B146D8F65669D93B3E742D50F968F082" author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan (2001</bibRefCitation>
|
||
: fig. 5.2d-f): tubeless, incomplete tube, and complete tube. In this study,
|
||
<taxonomicName id="E98C0A8C0ED75FAED59B2215D659C2F3" lsidName="R. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
||
<emphasis id="710A83C7E730024BCE6F55FF558E23FF" italics="true" pageId="0" pageNumber="1">R. hainesi</emphasis>
|
||
</taxonomicName>
|
||
from Chanthaburi populations also exhibit sympatrically occurring morphs. The typical and more abundant morph has a short to long tubular breathing device (Figs
|
||
<figureCitation id="9F64E52FF3BB13CE487CCC9986CA877B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shell of Rhiostoma hainesi A syntype NHMUK 20170371 from Camboja B holotype of " Rhiosotma smithi Bartsch, 1932 " NMNH 382943 from Khao Sabap, southern Siam C, D originally labelled as " Rhiostoma tomlini Salisbury, 1949 " from Khao Sabap, Siam C holotype NMW 1955.158.24924 and D paratype NHMUK 1949.6.7.1 E holotype of " Rhiostoma housei kirai Habe, 1965 " NSMT 52242 from Chanthaburi, Thailand F specimen CUMZ 4381 from Chanthaburi." figureDoi="10.3897/zookeys.1142.90097.figure21" httpUri="https://binary.pensoft.net/fig/799879" pageId="0" pageNumber="1">21F</figureCitation>
|
||
,
|
||
<figureCitation id="5C83F61FD202462434D2E7ABB72DDCA2" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22A</figureCitation>
|
||
), and the less abundant morph has a notch-shaped breathing device (Fig.
|
||
<figureCitation id="C9A8E5D8F40162FE08DD2E48F27F1AD9" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Shell of Rhiostoma hainesi A specimen CUMZ 4381 from Chanthaburi B specimen CUMZ 4814 from Makok Waterfall, Chanthaburi C specimen CUMZ 4816 from Khao Chamao, Rayong D specimens CUMZ 10194 from Chanthaburi shows series of growth from small juvenile (on the left) to nearly adult (on the right)." figureDoi="10.3897/zookeys.1142.90097.figure22" httpUri="https://binary.pensoft.net/fig/799880" pageId="0" pageNumber="1">22C</figureCitation>
|
||
). Specimens with these three distinct types of breathing devices were sampled for the COI analysis, and the results suggest that these are conspecific (Fig.
|
||
<figureCitation id="B4E6130AEE056810259615B1F316DEDE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum likelihood (ML) tree showing the relationships among Rhiostoma species based on 660 bp of the mitochondrial COI gene sequence. The major nodes are indicated by numbers as the Maximum likelihood (ML) bootstrap values and Bayesian posterior probabilities of Bayesian inference (BI), respectively. Nodes with sufficient support by both ML and BI are indicated by black circles, and nodes with partial support by only ML are indicated by grey circles, and only BI by white circles. Newly described species in this study are highlighted in blue." figureDoi="10.3897/zookeys.1142.90097.figure3" httpUri="https://binary.pensoft.net/fig/799861" pageId="0" pageNumber="1">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |