260 lines
40 KiB
XML
260 lines
40 KiB
XML
<document id="E973B89E1F77BE242C803738B2E36A35" ID-DOI="10.15298/rusentj.31.4.15" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596379676" checkinUser="felipe" docAuthor="Golovatch, S. I." docDate="2022" docId="75733D00FFA8D42CDE01FEB5050CFD85" docLanguage="en" docName="RussEntomolJ.31.4.432-435.pdf" docOrigin="Russian Entomological Journal 31 (4)" docSource="http://dx.doi.org/10.15298/rusentj.31.4.15" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Epanerchodus telnovi Golovatch 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="435" masterDocId="894A4578FFA9D42FDF13FFC40426FFF2" masterDocTitle="A new species of the millipede genus Epanerchodus from western Nepal (Diplopoda: Polydesmida: Polydesmidae)" masterLastPageNumber="435" masterPageNumber="432" pageNumber="433" updateTime="1722637417502" updateUser="ExternalLinkService">
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<mods:titleInfo id="50203FFADDE1067457193B481816408F">
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<mods:title id="54F9697892C9380682DD5D77F1106B2C">A new species of the millipede genus Epanerchodus from western Nepal (Diplopoda: Polydesmida: Polydesmidae)</mods:title>
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<mods:name id="7495EEE5FC5A95A31463FF984F9FB5AF" type="personal">
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<mods:namePart id="205E42A48BEC3C5EEFFBC698C94F1F33">Golovatch, S. I.</mods:namePart>
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<mods:title id="9472D9665CDD3D2C7F1CFC2E20345FFD">Russian Entomological Journal</mods:title>
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<mods:part id="A1CB8B7303492C66D75F9D8DACEDD310">
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<mods:date id="25BE33F091A96E6E86972755E448D4A1">2022</mods:date>
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<mods:number id="1C42A73A6E688F9EAB0A6E64DA2F96CD">2022-12-31</mods:number>
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<mods:number id="090B05214CA94E9A56C6533F9FD219C5">31</mods:number>
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<mods:start id="9E24E24129697DC928BCA08DE08CCDDA">432</mods:start>
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<mods:identifier id="9413E19465CE401F165B5BEC7D5E74FF" type="DOI">10.15298/rusentj.31.4.15</mods:identifier>
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<mods:identifier id="1999406C5762D107C4DF7B20A720927C" type="ISSN">0132-8069</mods:identifier>
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<mods:identifier id="2263184D0EF63F1619B02A6C0E738ABC" type="Zenodo-Dep">13179562</mods:identifier>
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<treatment id="75733D00FFA8D42CDE01FEB5050CFD85" LSID="urn:lsid:plazi:treatment:75733D00FFA8D42CDE01FEB5050CFD85" httpUri="http://treatment.plazi.org/id/75733D00FFA8D42CDE01FEB5050CFD85" lastPageId="3" lastPageNumber="435" pageId="1" pageNumber="433">
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<paragraph id="FD658C16FFA8D42EDE01FEB50697FE7B" blockId="1.[189,775,369,551]" box="[274,689,369,393]" pageId="1" pageNumber="433">
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<heading id="A62D3B7AFFA8D42EDE01FEB50697FE7B" box="[274,689,369,393]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="433" reason="8">
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<taxonomicName id="3ADAF795FFA8D42EDE01FEB50655FE7B" ID-CoL="CGMDW" authority="Golovatch" authorityName="Golovatch" authorityYear="2022" box="[274,627,369,393]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="433" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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<emphasis id="CFAE5004FFA8D42EDE01FEB505DBFE7B" box="[274,509,369,393]" italics="true" pageId="1" pageNumber="433">Epanerchodus telnovi</emphasis>
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Golovatch
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</taxonomicName>
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<emphasis id="CFAE5004FFA8D42EDD6FFEB50697FE7B" bold="true" box="[636,689,369,393]" pageId="1" pageNumber="433">
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<taxonomicNameLabel id="D49DED7FFFA8D42EDD6FFEB50697FE7B" box="[636,689,369,393]" pageId="1" pageNumber="433" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B5C0DF9DFFA8D42EDEBCFE4A0139FEA3" pageId="1" pageNumber="433" type="description">
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<paragraph id="FD658C16FFA8D42EDEBCFE4A0518FDD4" blockId="1.[189,775,369,551]" pageId="1" pageNumber="433">
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<materialsCitation id="4DB2864BFFA8D42EDEBCFE4A0518FDD4" collectingDate="2022-06-17" collectingDateMax="2022-06-18" collectingDateMin="2022-06-17" collectionCode="ZMUM" collectorName="D. Telnov" country="Nepal" elevation="3150" latitude="29.90639" location="Humla Distr." longLatPrecision="20" longitude="81.91861" municipality="Western" pageId="1" pageNumber="433" specimenCount="1" specimenCount-male="1" stateProvince="Karnali" typeStatus="holotype">
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<figureCitation id="65E19093FFA8D42EDEBCFE4A0636FE54" box="[431,528,398,422]" captionStart-0="Figs 1–3" captionStart-1="Figs 4–5" captionStartId-0="1.[226,263,1966,1985]" captionStartId-1="2.[226,263,1966,1985]" captionTargetBox-0="[189,1399,576,1954]" captionTargetBox-1="[230,1373,1153,1944]" captionTargetId-0="figure-205@1.[189,1399,576,1954]" captionTargetId-1="figure-586@2.[230,1374,1153,1944]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." captionText-1="Figs 4–5. Epanerchodus telnovi sp.n., ♂ holotype, right gonopod, mesal and lateral views, respectively. Scale bar: 0.2 mm. Abbreviations explained in text. Рис. 4–5. Epanerchodus telnovi sp.n., голотип ♂, правый гонопод, соответственно иЗнутри и сбоку. МасШтаб: 0,2 мм. ОбЪЯснение обоЗначений в тексте." figureDoi-0="http://doi.org/10.5281/zenodo.13179564" figureDoi-1="http://doi.org/10.5281/zenodo.13179566" httpUri-0="https://zenodo.org/record/13179564/files/figure.png" httpUri-1="https://zenodo.org/record/13179566/files/figure.png" pageId="1" pageNumber="433">Figs 1–5</figureCitation>
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.
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<typeStatus id="226132B4FFA8D42EDFF1FE770572FE34" box="[226,340,435,454]" pageId="1" pageNumber="433" type="holotype">HOLOTYPE</typeStatus>
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<specimenCount id="EBDC479FFFA8D42EDE4EFE720557FE34" box="[349,369,438,454]" count="1" pageId="1" pageNumber="433" type="male">♂</specimenCount>
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(somewhat macerated and broken into three pieces) (
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<collectionCode id="9BCB14D3FFA8D42EDE14FE0F0569FE2C" box="[263,335,459,478]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="433">ZMUM</collectionCode>
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),
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<collectingMunicipality id="1D01166CFFA8D42EDE73FE08058FFE2D" box="[352,425,460,479]" pageId="1" pageNumber="433">Western</collectingMunicipality>
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<collectingCountry id="85CDCC86FFA8D42EDEBCFE0F05C0FE2C" box="[431,486,459,478]" name="Nepal" pageId="1" pageNumber="433">Nepal</collectingCountry>
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,
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<collectingRegion id="3F1E42F4FFA8D42EDEE3FE0F064DFE2D" box="[496,619,459,479]" country="Nepal" name="Karnali" pageId="1" pageNumber="433">Karnali Prov.</collectingRegion>
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,
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<location id="F805DACDFFA8D42EDD60FE0F06CCFE2C" LSID="urn:lsid:plazi:treatment:75733D00FFA8D42CDE01FEB5050CFD85:F805DACDFFA8D42EDD60FE0F06CCFE2C" box="[627,746,459,478]" country="Nepal" latitude="29.90639" longLatPrecision="20" longitude="81.91861" municipality="Western" name="Humla Distr." pageId="1" pageNumber="433" stateProvince="Karnali">Humla Distr.</location>
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, ca
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<locationDeviation id="ECD79534FFA8D42EDFAEFE2705B9FE04" box="[189,415,483,502]" location="Simikot" pageId="1" pageNumber="433">
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<quantity id="3A2221F3FFA8D42EDFAEFE270530FE04" box="[189,278,483,502]" metricMagnitude="4" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="1" pageNumber="433" unit="km" value="12.5" valueMax="13.0" valueMin="12.0">12–13 km</quantity>
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SE of Simikot
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</locationDeviation>
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,
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<geoCoordinate id="98EEEAD1FFA8D42EDEBAFE200638FE04" box="[425,542,483,503]" degrees="29" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="433" precision="15" seconds="23" value="29.90639">N 29°54´23´´</geoCoordinate>
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–29°54´00´´,
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<geoCoordinate id="98EEEAD1FFA8D42EDD87FE2006DFFE04" box="[660,761,483,503]" degrees="81" direction="east" minutes="55" orientation="longitude" pageId="1" pageNumber="433" precision="15" seconds="7" value="81.91861">E 81°55´7´´</geoCoordinate>
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– 81°55´11´´,
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<quantity id="3A2221F3FFA8D42EDE35FE3F0587FDFD" box="[294,417,507,527]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.31" metricValueMin="2.9899999999999998" pageId="1" pageNumber="433" unit="m" value="3150.0" valueMax="3310.0" valueMin="2990.0">
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<elevation id="76F76B25FFA8D42EDE35FE3F0587FDFD" box="[294,417,507,527]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.31" metricValueMin="2.9899999999999998" pageId="1" pageNumber="433" unit="m" value="3150.0" valueMax="3310.0" valueMin="2990.0">2990–3310 m</elevation>
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</quantity>
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, disturbed mixed forest,
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||
<date id="8964AAD6FFA8D42EDD6FFE3F0725FDFC" box="[636,771,507,526]" pageId="1" pageNumber="433" value="2022-06-17" valueMax="2022-06-18" valueMin="2022-06-17">
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<collectingDate id="9920533EFFA8D42EDD6FFE3F0725FDFC" box="[636,771,507,526]" pageId="1" pageNumber="433" value="2022-06-17" valueMax="2022-06-18" valueMin="2022-06-17">17–18.VI.2022</collectingDate>
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||
</date>
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||
,
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<collectorName id="502FE9C0FFA8D42EDFAEFDD00530FDD4" box="[189,278,531,551]" pageId="1" pageNumber="433">D. Telnov</collectorName>
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leg.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="FD658C16FFA8D42EDC41FEFF0139FEA3" blockId="1.[812,1399,315,551]" box="[850,1311,315,337]" pageId="1" pageNumber="433">NAME. Honours Dmitry Telnov, the collector.</paragraph>
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</subSubSection>
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<subSubSection id="B5C0DF9DFFA8D42EDC41FE9200FDFE03" pageId="1" pageNumber="433" type="diagnosis">
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<paragraph id="FD658C16FFA8D42EDC41FE9200FDFE03" blockId="1.[812,1399,315,551]" pageId="1" pageNumber="433">
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DIAGNOSIS. Differs from congeners by the relatively small size (width ca 2.0 mm), coupled with the head being narrower than the collum, the absence of sphaerotrichomes, lateral bulges on
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<specimenCount id="EBDC479FFFA8D42EDCF2FE6D07D1FE49" box="[993,1015,425,443]" count="1" pageId="1" pageNumber="433" type="male">♂</specimenCount>
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prefemora, and an exomere, and the shapes of the endomere (
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<emphasis id="CFAE5004FFA8D42EDB35FE050018FE24" bold="true" box="[1062,1086,449,470]" pageId="1" pageNumber="433">en</emphasis>
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) and its single, basal, postfemoral process (
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<emphasis id="CFAE5004FFA8D42EDCA0FE1F07EEFE03" bold="true" box="[947,968,475,497]" pageId="1" pageNumber="433">pf</emphasis>
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). See also Remarks below.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5C0DF9DFFA8D42DDC41FE320156FC69" lastPageId="2" lastPageNumber="434" pageId="1" pageNumber="433" type="description">
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<paragraph id="FD658C16FFA8D42DDC41FE320538FE74" blockId="1.[812,1399,315,551]" lastBlockId="2.[189,776,315,1111]" lastPageId="2" lastPageNumber="434" pageId="1" pageNumber="433">
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DESCRIPTION. Length ca
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<quantity id="3A2221F3FFA8D42EDB78FE320093FDF9" box="[1131,1205,502,523]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="433" unit="mm" value="14.0">14 mm</quantity>
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, width of midbody pro- and metazonae 1.0 and 2.0 mm, respectively (
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<specimenCount id="EBDC479FFFA8D42EDA0CFDD70113FDD7" box="[1311,1333,531,549]" count="1" pageId="1" pageNumber="433" type="male">♂</specimenCount>
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). Colouration in alcohol uniformly brown with mostly lighter sterna and brown-yellow legs (
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<figureCitation id="65E19093FFABD42DDD13FE920678FE99" box="[512,606,342,364]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 1–3</figureCitation>
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). Body with 20 segments.
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</paragraph>
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||
<caption id="A9A5DC9EFFA8D42EDFF1F86A078BF7FF" ID-DOI="http://doi.org/10.5281/zenodo.13179564" ID-Zenodo-Dep="13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="1" pageNumber="433" startId="1.[226,263,1966,1985]" targetBox="[189,1399,576,1954]" targetPageId="1" targetType="figure">
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||
<paragraph id="FD658C16FFA8D42EDFF1F86A06C4F82B" blockId="1.[189,1399,1966,2061]" pageId="1" pageNumber="433">
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||
Figs 1–3.
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<taxonomicName id="3ADAF795FFA8D42EDE28F86A05DEF833" authorityName="Golovatch" authorityYear="2022" box="[315,504,1966,1985]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="433" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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<emphasis id="CFAE5004FFA8D42EDE28F86A05DEF833" box="[315,504,1966,1985]" italics="true" pageId="1" pageNumber="433">Epanerchodus telnovi</emphasis>
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</taxonomicName>
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<emphasis id="CFAE5004FFA8D42EDEECF86A060CF833" bold="true" box="[511,554,1966,1985]" pageId="1" pageNumber="433">
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||
<taxonomicNameLabel id="D49DED7FFFA8D42EDEECF86A060CF833" box="[511,554,1966,1985]" pageId="1" pageNumber="433" rank="species">sp.n.</taxonomicNameLabel>
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||
</emphasis>
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||
, ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale.
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||
</paragraph>
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||
<paragraph id="FD658C16FFA8D42EDFF1F826078BF7FF" blockId="1.[189,1399,1966,2061]" pageId="1" pageNumber="433">
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Рис. 1–3.
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<taxonomicName id="3ADAF795FFA8D42EDE29F82505D0F806" authorityName="Golovatch" authorityYear="2022" box="[314,502,2017,2036]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="433" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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||
<emphasis id="CFAE5004FFA8D42EDE29F82505D0F806" box="[314,502,2017,2036]" italics="true" pageId="1" pageNumber="433">Epanerchodus telnovi</emphasis>
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||
</taxonomicName>
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||
<emphasis id="CFAE5004FFA8D42EDEE8F8250600F806" bold="true" box="[507,550,2017,2036]" pageId="1" pageNumber="433">
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||
<taxonomicNameLabel id="D49DED7FFFA8D42EDEE8F8250600F806" box="[507,550,2017,2036]" pageId="1" pageNumber="433" rank="species">sp.n.</taxonomicNameLabel>
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||
</emphasis>
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||
, голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба.
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</paragraph>
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</caption>
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<paragraph id="FD658C16FFABD42DDFF1FE4806A5FE02" blockId="2.[189,776,315,1111]" pageId="2" pageNumber="434">
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Tegument mostly dull, texture very delicately shagreened. Head pilose nearly throughout, with squarish genae. Antennae and most legs broken off. Interantennal isthmus about twice as large as diameter of antennal socket (
|
||
<figureCitation id="65E19093FFABD42DDD33FE1F0651FE02" box="[544,631,475,497]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD658C16FFABD42DDFF1FE32011DFD84" blockId="2.[189,776,315,1111]" lastBlockId="2.[812,1399,315,1111]" pageId="2" pageNumber="434">
|
||
In width, head <collum <segment 3=4 <2 <5=15, thereafter body gradually tapering towards telson (
|
||
<figureCitation id="65E19093FFABD42DDDB0FDD506DDFDD4" box="[675,763,529,551]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 1–3</figureCitation>
|
||
). Paraterga strongly developed, set high (at about upper ¼ of midbody height), starting with collum, dorsum very faintly convex; paraterga mostly weakly upturned above dorsum. Anterior shoulders and sides of paraterga mostly straight to only slightly rounded. Collum crescent-shaped, each side with only one lateral incision near midway (
|
||
<figureCitation id="65E19093FFABD42DDD85FD7506DCFD34" box="[662,762,689,711]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 1, 3</figureCitation>
|
||
). Caudolateral corner of paraterga spiniform and pointed, postcollum ones extending increasingly past rear tergal margin starting with paraterga 4, especially clearly so on rings 17 and 18. All poreless paraterga with three, all pore-bearing ones with four, minute incisions at lateral margin. Front margins of metaterga narrowly bordered and forming distinct shoulders. Pore formula normal, ozopores small, but evident, dorsal, located in front of posteriormost marginal indentation. Metatergal sculpture typical, starting with collum, poorly developed, but readily visible, with three transverse rows of typical (= polydesmid), setigerous, polygonal bosses (
|
||
<figureCitation id="65E19093FFABD42DDDACFC1204ECFBF4" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 1– 3</figureCitation>
|
||
), anterior row being the most obliterate and caudal row the most evident (up to rough wrinkles on ring 18). Tergal setae very short, simple, mostly obliterated. Stricture between pro- and metazona wide, shallow and nearly smooth. Limbus very thin, microdenticulate. Pleurosternal carinae absent. Epiproct rather short, conical, pre-apical lateral papillae inconspicuous. Hypoproct semi-circular; caudal, paramedian, setigerous papillae very prominent and well separated teeth (
|
||
<figureCitation id="65E19093FFABD42DDA5CFE4F071CFE49" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Fig. 2</figureCitation>
|
||
). Sterna without modifications, nearly flat, densely setose. Legs (
|
||
<specimenCount id="EBDC479FFFABD42DDC7BFE070758FE27" box="[872,894,451,469]" count="1" pageId="2" pageNumber="434" type="male">♂</specimenCount>
|
||
) generally rather long and slender, apparently slightly incrassate (
|
||
<figureCitation id="65E19093FFABD42DDCA3FE1F0021FE02" box="[944,1031,475,497]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 1–2</figureCitation>
|
||
), ca 1.7–1.8 times as long as midbody height, densely setose, almost all setae simple, very poorly branching setae with minute, distal, side branchlets being observed only on slender prefemora, the latter devoid of lateral bulges; sphaerotrichomes absent. In length, tarsus = femur> prefemur> coxa = postfemur = tibia (
|
||
<figureCitation id="65E19093FFABD42DDBE0FDA50108FD84" box="[1267,1326,609,631]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD658C16FFABD42DDC41FDBF0156FC69" blockId="2.[812,1399,315,1111]" pageId="2" pageNumber="434">
|
||
Gonopods (
|
||
<figureCitation id="65E19093FFABD42DDCD7FDBF0022FD62" box="[964,1028,635,657]" captionStart="Figs 1–3" captionStartId="1.[226,263,1966,1985]" captionTargetBox="[189,1399,576,1954]" captionTargetId="figure-205@1.[189,1399,576,1954]" captionTargetPageId="1" captionText="Figs 1–3. Epanerchodus telnovi sp.n., ♂ holotype: 1–2 — habitus, dorsal and ventral views, respectively; 3 — anterior part of body, anterodorsal view. Pictures by K. Makarov, not taken to scale. Рис. 1–3. Epanerchodus telnovi sp.n., голотип ♂: 1–2 — обЩий вид, соответственно сверху и сниЗу; 3 — переднЯЯ часть головы, одновременно спереди и сверху. Фотографии К. Макарова, снЯты беЗ масШтаба." figureDoi="http://doi.org/10.5281/zenodo.13179564" httpUri="https://zenodo.org/record/13179564/files/figure.png" pageId="2" pageNumber="434">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="65E19093FFABD42DDB03FDBF0013FD63" box="[1040,1077,635,657]" captionStart="Figs 4–5" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[230,1373,1153,1944]" captionTargetId="figure-586@2.[230,1374,1153,1944]" captionTargetPageId="2" captionText="Figs 4–5. Epanerchodus telnovi sp.n., ♂ holotype, right gonopod, mesal and lateral views, respectively. Scale bar: 0.2 mm. Abbreviations explained in text. Рис. 4–5. Epanerchodus telnovi sp.n., голотип ♂, правый гонопод, соответственно иЗнутри и сбоку. МасШтаб: 0,2 мм. ОбЪЯснение обоЗначений в тексте." figureDoi="http://doi.org/10.5281/zenodo.13179566" httpUri="https://zenodo.org/record/13179566/files/figure.png" pageId="2" pageNumber="434">4–5</figureCitation>
|
||
) with large, subquadrate coxites (
|
||
<emphasis id="CFAE5004FFABD42DDC27FD52076DFD59" bold="true" box="[820,843,662,683]" pageId="2" pageNumber="434">cx</emphasis>
|
||
) strongly fused medially at base and carrying a few long setae ventrally; a long, simple and unciform cannula, as usual. Telopodite elongated, stout, subfalcate only distally, prefemorite (= densely setose part) more than half as long as entire telopodite; seminal groove running mesally over most of its extent, only distally moving frontally to recurve first laterad and then a little basad at base of a longer, subunciform and quite simple endomere (
|
||
<emphasis id="CFAE5004FFABD42DDB2CFC950071FC94" bold="true" box="[1087,1111,849,870]" pageId="2" pageNumber="434">en</emphasis>
|
||
); an exomere absent; a round accessory seminal chamber hidden inside at base of a single, stout and apically unequally bifid postfemoral process (
|
||
<emphasis id="CFAE5004FFABD42DDA5EFC420144FC6E" bold="true" box="[1357,1378,902,924]" pageId="2" pageNumber="434">pf</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5C0DF9DFFABD42CDC41FC65050CFD85" lastPageId="3" lastPageNumber="435" pageId="2" pageNumber="434" type="discussion">
|
||
<paragraph id="FD658C16FFABD42CDC41FC650580FE99" blockId="2.[812,1399,315,1111]" lastBlockId="3.[189,775,315,364]" lastPageId="3" lastPageNumber="435" pageId="2" pageNumber="434">
|
||
REMARKS. The new species differs readily from all hitherto known Himalayan congeners primarily by the single, undivided postfemoral process of the gonopod [
|
||
<bibRefCitation id="994BF1E7FFABD42DDA1AFC120779FBF4" author="Golovatch S. I." pageId="2" pageNumber="434" pagination="345 - 369" refId="ref1947" refString="Golovatch S. I. 1986. Diplopoda from the Nepal Himalayas: Polydesmidae, Fuhrmannodesmidae // Senckenbergiana biologica. Bd. 66. H. 4 / 6. S. 345 - 369." type="journal article" year="1986">Golovatch, 1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="994BF1E7FFABD42DDC79FC3507BAFBF4" author="Golovatch S. I." box="[874,924,1009,1030]" pageId="2" pageNumber="434" pagination="203 - 217" refId="ref1983" refString="Golovatch S. I. 1987. Diplopoda from the Nepal Himalayas. Opisotretidae, additional Polydesmidae and Fuhrmannodesmidae // Courier Forschungsinstitut Senckenberg. Bd. 93. S. 203 - 217." type="book chapter" year="1987">1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="994BF1E7FFABD42DDCBBFC3507FBFBF4" author="Golovatch S. I." box="[936,989,1009,1030]" pageId="2" pageNumber="434" pagination="237 - 252" refId="ref2017" refString="Golovatch S. I. 1990. Diplopoda from the Nepal Himalayas. Several additional Polydesmidae and Fuhrmannodesmidae (Polydesmida) // Spixiana. Bd. 13. H. 3. S. 237 - 252." type="journal article" year="1990">1990</bibRefCitation>
|
||
], in this respect rather resembling several species from
|
||
<collectingCountry id="85CDCC86FFABD42DDCBCFBCF07CDFBD3" box="[943,1003,1035,1057]" name="China" pageId="2" pageNumber="434">China</collectingCountry>
|
||
, e.g.
|
||
<taxonomicName id="3ADAF795FFABD42DDB33FBC80155FBD2" authority="Liu et Golovatch, 2018" authorityName="Liu et Golovatch" authorityYear="2018" box="[1056,1395,1035,1057]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="coniger">
|
||
<emphasis id="CFAE5004FFABD42DDB33FBC800A0FBD3" box="[1056,1158,1036,1057]" italics="true" pageId="2" pageNumber="434">E. coniger</emphasis>
|
||
<bibRefCitation id="994BF1E7FFABD42DDB9EFBCF0155FBD2" author="Liu Weixin & Golovatch S. I." box="[1165,1395,1035,1057]" pageId="2" pageNumber="434" pagination="53 - 84" refId="ref2240" refString="Liu Weixin, Golovatch S. I. 2018. The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae) // Zootaxa. Vol. 4459. No. 1. P. 53 - 84." type="journal article" year="2018">Liu et Golovatch, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3ADAF795FFABD42DDC3FFBE30050FBC9" authority="Liu et Golovatch, 2018" authorityName="Liu et Golovatch" authorityYear="2018" box="[812,1142,1062,1084]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis id="CFAE5004FFABD42DDC3FFBE307ACFBCE" box="[812,906,1063,1084]" italics="true" pageId="2" pageNumber="434">E. parvus</emphasis>
|
||
<bibRefCitation id="994BF1E7FFABD42DDC82FBE20050FBC9" author="Liu Weixin & Golovatch S. I." box="[913,1142,1062,1084]" pageId="2" pageNumber="434" pagination="53 - 84" refId="ref2240" refString="Liu Weixin, Golovatch S. I. 2018. The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae) // Zootaxa. Vol. 4459. No. 1. P. 53 - 84." type="journal article" year="2018">Liu et Golovatch, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3ADAF795FFABD42DDB92FBE307BAFBA4" authority="Liu et Golovatch, 2018" authorityName="Liu et Golovatch" authorityYear="2018" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="gladiatus">
|
||
<emphasis id="CFAE5004FFABD42DDB92FBE300D0FBC9" box="[1153,1270,1062,1084]" italics="true" pageId="2" pageNumber="434">E. gladiatus</emphasis>
|
||
<bibRefCitation id="994BF1E7FFABD42DDBEEFBE207BAFBA4" author="Liu Weixin & Golovatch S. I." pageId="2" pageNumber="434" pagination="53 - 84" refId="ref2240" refString="Liu Weixin, Golovatch S. I. 2018. The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae) // Zootaxa. Vol. 4459. No. 1. P. 53 - 84." type="journal article" year="2018">Liu et Golovatch, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and some others, in which the single postfemoral process is relatively small, sometimes even barely discernible [
|
||
<bibRefCitation id="994BF1E7FFAAD42CDFD1FE9205BAFE99" author="Liu Weixin & Golovatch S. I." box="[194,412,342,364]" pageId="3" pageNumber="435" pagination="53 - 84" refId="ref2240" refString="Liu Weixin, Golovatch S. I. 2018. The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae) // Zootaxa. Vol. 4459. No. 1. P. 53 - 84." type="journal article" year="2018">Liu, Golovatch, 2018</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<caption id="A9A5DC9EFFABD42DDFF1F86A05AFF7FF" ID-DOI="http://doi.org/10.5281/zenodo.13179566" ID-Zenodo-Dep="13179566" httpUri="https://zenodo.org/record/13179566/files/figure.png" pageId="2" pageNumber="434" startId="2.[226,263,1966,1985]" targetBox="[230,1373,1153,1944]" targetPageId="2" targetType="figure">
|
||
<paragraph id="FD658C16FFABD42DDFF1F86A0571F82B" blockId="2.[189,1398,1965,2061]" pageId="2" pageNumber="434">
|
||
Figs 4–5.
|
||
<taxonomicName id="3ADAF795FFABD42DDE26F86A05D7F833" authorityName="Golovatch" authorityYear="2022" box="[309,497,1966,1985]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
|
||
<emphasis id="CFAE5004FFABD42DDE26F86A05D7F833" box="[309,497,1966,1985]" italics="true" pageId="2" pageNumber="434">Epanerchodus telnovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFAE5004FFABD42DDEE6F86A0606F833" bold="true" box="[501,544,1966,1985]" pageId="2" pageNumber="434">
|
||
<taxonomicNameLabel id="D49DED7FFFABD42DDEE6F86A0606F833" box="[501,544,1966,1985]" pageId="2" pageNumber="434" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype, right gonopod, mesal and lateral views, respectively. Scale bar: 0.2 mm. Abbreviations explained in text.
|
||
</paragraph>
|
||
<paragraph id="FD658C16FFABD42DDFF1F82605AFF7FF" blockId="2.[189,1398,1965,2061]" pageId="2" pageNumber="434">
|
||
Рис. 4–5.
|
||
<taxonomicName id="3ADAF795FFABD42DDE24F82505D5F806" authorityName="Golovatch" authorityYear="2022" box="[311,499,2017,2036]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
|
||
<emphasis id="CFAE5004FFABD42DDE24F82505D5F806" box="[311,499,2017,2036]" italics="true" pageId="2" pageNumber="434">Epanerchodus telnovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFAE5004FFABD42DDEE4F8250604F806" bold="true" box="[503,546,2017,2036]" pageId="2" pageNumber="434">
|
||
<taxonomicNameLabel id="D49DED7FFFABD42DDEE4F8250604F806" box="[503,546,2017,2036]" pageId="2" pageNumber="434" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, голотип ♂, правый гонопод, соответственно иЗнутри и сбоку. МасШтаб: 0,2 мм. ОбЪЯснение обоЗначений в тексте.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FD658C16FFAAD42CDFF1FE4F050CFD85" blockId="3.[189,775,395,631]" pageId="3" pageNumber="435">
|
||
<emphasis id="CFAE5004FFAAD42CDFF1FE4F058FFE53" bold="true" box="[226,425,395,417]" pageId="3" pageNumber="435">Acknowledgements</emphasis>
|
||
.
|
||
<materialsCitation id="4DB2864BFFAAD42CDEABFE480500FD85" collectionCode="ZMUM" collectorName="Special & Kirill Makarov & The & Presidium & Programme No." country="United Kingdom" location="Russian Academy of Sciences" pageId="3" pageNumber="435" specimenCount="1" stateProvince="Moscow" typeStatus="holotype">
|
||
I am most grateful to Dmitry Telnov (Natural History Museum, London,
|
||
<collectingCountry id="85CDCC86FFAAD42CDD44FE620726FE4E" box="[599,768,422,444]" name="United Kingdom" pageId="3" pageNumber="435">United Kingdom</collectingCountry>
|
||
) for the provision of the
|
||
<typeStatus id="226132B4FFAAD42CDEB3FE0505D3FE25" box="[416,501,449,471]" pageId="3" pageNumber="435" type="holotype">holotype</typeStatus>
|
||
of the new species for study and the donation to the
|
||
<collectionCode id="9BCB14D3FFAAD42CDEB3FE1805CDFE03" box="[416,491,476,497]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="3" pageNumber="435">ZMUM</collectionCode>
|
||
collection.
|
||
<collectorName id="502FE9C0FFAAD42CDD4FFE1F0685FE03" box="[604,675,475,497]" pageId="3" pageNumber="435">Special</collectorName>
|
||
thanks go to
|
||
<collectorName id="502FE9C0FFAAD42CDFC4FE32054DFDFE" box="[215,363,502,524]" pageId="3" pageNumber="435">Kirill Makarov</collectorName>
|
||
(
|
||
<collectingRegion id="3F1E42F4FFAAD42CDE69FE3205F7FDF9" box="[378,465,502,523]" country="Russia" name="Moscow" pageId="3" pageNumber="435">Moscow</collectingRegion>
|
||
,
|
||
<collectingCountry id="85CDCC86FFAAD42CDECFFE320638FDFE" box="[476,542,502,524]" name="Russia" pageId="3" pageNumber="435">Russia</collectingCountry>
|
||
) who so skillfully took all pictures.
|
||
<collectorName id="502FE9C0FFAAD42CDE26FDD5057CFDD5" box="[309,346,529,551]" pageId="3" pageNumber="435">The</collectorName>
|
||
author was partly supported by the
|
||
<collectorName id="502FE9C0FFAAD42CDDA8FDD504FAFDB3" pageId="3" pageNumber="435">Presidium</collectorName>
|
||
of the
|
||
<location id="F805DACDFFAAD42CDE0CFDEF0663FDB3" LSID="urn:lsid:plazi:treatment:75733D00FFA8D42CDE01FEB5050CFD85:F805DACDFFAAD42CDE0CFDEF0663FDB3" box="[287,581,555,577]" country="United Kingdom" name="Russian Academy of Sciences" pageId="3" pageNumber="435" stateProvince="Moscow">Russian Academy of Sciences</location>
|
||
,
|
||
<collectorName id="502FE9C0FFAAD42CDD5DFDE806CFFDB3" box="[590,745,556,577]" pageId="3" pageNumber="435">Programme No.</collectorName>
|
||
41 “Biodiversity of Natural Systems and Biological Resources of
|
||
<collectingCountry id="85CDCC86FFAAD42CDFCBFDA5053BFD85" box="[216,285,609,631]" name="Russia" pageId="3" pageNumber="435">Russia</collectingCountry>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |