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<mods:title id="10E5DA477648933A37A880A2EC1851A0">Reappraisal of species attributed to Halicarcinus White, 1846 (Crustacea: Decapoda: Brachyura: Hymenosomatidae) with diagnosis of four new genera and one new species from New Ireland, Papua New Guinea</mods:title>
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<taxonomicName id="3A944D2D050FFD4DFF22FC0BFD07FBBC" ID-CoL="3JBMF" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[151,713,1023,1049]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFF22FC0BFD07FBBC" bold="true" box="[151,713,1023,1049]" pageId="9" pageNumber="489">
<emphasis id="CFE0EABC050FFD4DFF22FC0BFEB1FBBC" bold="true" box="[151,383,1023,1049]" italics="true" pageId="9" pageNumber="489">Halicarcinus quoyi</emphasis>
(H.
<bibRefCitation id="99054B5F050FFD4DFE00FC0BFD0FFBBC" author="Milne" box="[437,705,1023,1049]" pageId="9" pageNumber="513" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles (ser. 3), 20, 163 - 228, pls 1 - 11. Avaliable from: http: // biodiversitylibrary. org / page / 35466171 (Accessed 21 Mar. 2016)" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
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(
<figureCitation id="65AF2A2B050FFD4DFF2AFBD6FF11FB9E" box="[159,223,1058,1083]" captionStart="FIGURE 1" captionStartId="7.[151,250,1881,1903]" captionTargetBox="[218,1375,1012,1862]" captionTargetPageId="7" captionText="FIGURE 1. Halicarcinus cookii (Filhol, 1885): a c, lectotype of Hymenicus cookii Filhol, 1885, MNHN IU- 2014 - 7143 (B 32572), (female, 6.1 mm). Halicarcinus planatus (Fabricius, 1775): d, holotype of Elamena mexicana (H. Milne Edwards, 1853), MNHN IU- 2000 - 655 (B 655), (male, 7.3 mm). Halicarcinus quoyi (H. Milne Edwards, 1853): e, lectotype of Elamena quoyi H. Milne Edwards, 1853, MNHN- 2000 - 667 (B 667), (male without rostrum, 6.4 mm); f, paralectotype, MNHN- 2000 - 32609 (B 32609), (sex unknown, dry fragments)." httpUri="https://zenodo.org/record/258479/files/figure.png" pageId="9" pageNumber="489">Fig. 1</figureCitation>
e, f)
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<emphasis id="CFE0EABC050FFD4DFF22FB9EFF3EFB25" box="[151,240,1130,1152]" italics="true" pageId="9" pageNumber="489">Elamene</emphasis>
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[sic]
<taxonomicName id="3A944D2D050FFD4DFE9CFB9FFD58FB25" authority="H. Milne Edwards, 1853: 223" authorityName="H. Milne Edwards" authorityPageNumber="223" authorityYear="1853" box="[297,662,1130,1152]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFE9CFB9FFEAFFB25" box="[297,353,1131,1152]" italics="true" pageId="9" pageNumber="489">quoyi</emphasis>
H.
<treatmentCitation id="7C3510BF050FFD4DFE33FB9EFD58FB25" author="Milne" box="[390,662,1130,1152]" page="223" pageId="9" pageNumber="489" year="1853">
<bibRefCitation id="99054B5F050FFD4DFE33FB9EFDAAFB25" author="Milne" box="[390,612,1130,1152]" pageId="9" pageNumber="513" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles (ser. 3), 20, 163 - 228, pls 1 - 11. Avaliable from: http: // biodiversitylibrary. org / page / 35466171 (Accessed 21 Mar. 2016)" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
: 223
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, pl. 11 fig. 3.
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<treatmentCitationGroup id="DD841180050FFD4DFF22FB7DFDE4FB3A" box="[151,554,1161,1183]" pageId="9" pageNumber="489">
<taxonomicName id="3A944D2D050FFD4DFF22FB7DFDE8FB3A" ID-CoL="7PJ4L" authority="Huttoni Filhol, 1885b: 46" authorityName="Huttoni Filhol" authorityPageNumber="46" authorityYear="1885" box="[151,550,1161,1183]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="CFE0EABC050FFD4DFF22FB7DFEA3FB3A" box="[151,365,1161,1183]" italics="true" pageId="9" pageNumber="489">Halicarcinus Huttoni</emphasis>
<treatmentCitation id="7C3510BF050FFD4DFEC1FB7DFDE8FB3A" author="Filhol" box="[372,550,1161,1183]" page="46" pageId="9" pageNumber="489" year="1885">
<bibRefCitation id="99054B5F050FFD4DFEC1FB7DFDCFFB3A" author="Filhol" box="[372,513,1161,1183]" pageId="9" pageNumber="512" refString="Filhol, H. (1885 b) Description d'une nouvelle espece de Crustace appartenant au genre Halicarcinus. Bulletin de la Societe Philomathique de Paris, ser. 7, 9, 45 - 46. Avaliable from: http: // biodiversitylibrary. org / page / 31945639 (Accessed 21 Mar. 2016)" type="journal article" year="1885" yearSuffix="b">Filhol, 1885b</bibRefCitation>
: 46
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.
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<treatmentCitationGroup id="DD841180050FFD4DFF22FB53FD0CFB18" box="[151,706,1191,1213]" pageId="9" pageNumber="489">
<emphasis id="CFE0EABC050FFD4DFF22FB53FE40FB18" box="[151,398,1191,1213]" italics="true" pageId="9" pageNumber="489">
<taxonomicName id="3A944D2D050FFD4DFF22FB53FED4FB18" ID-CoL="7PJ4L" box="[151,282,1191,1213]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="genus">Halicarcinus</taxonomicName>
planatus—
</emphasis>
<treatmentCitation id="7C3510BF050FFD4DFE3BFB53FDA0FB18" author="Richardson" box="[398,622,1191,1213]" page="68" pageId="9" pageNumber="489" year="1949">
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: 68
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, fig. 48.
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</paragraph>
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<treatmentCitationGroup id="DD841180050FFD4DFF22FB32FCF7FB79" box="[151,825,1222,1244]" pageId="9" pageNumber="489">
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<emphasis id="CFE0EABC050FFD4DFF22FB32FE5DFB79" box="[151,403,1222,1244]" italics="true" pageId="9" pageNumber="489">Halicarcinus innominata</emphasis>
<treatmentCitation id="7C3510BF050FFD4DFE2FFB32FD43FB79" author="Richardson" box="[410,653,1222,1244]" page="130" pageId="9" pageNumber="489" year="1949">
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: 130
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[
<emphasis id="CFE0EABC050FFD4DFD2EFB33FCE4FB79" box="[667,810,1222,1244]" italics="true" pageId="9" pageNumber="489">nomen nudum</emphasis>
].
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<paragraph id="FD2B36AE050FFD4DFF22FB11FC81FABC" blockId="9.[151,1437,1130,1305]" pageId="9" pageNumber="489">
<treatmentCitationGroup id="DD841180050FFD4DFF22FB11FC81FABC" pageId="9" pageNumber="489">
<emphasis id="CFE0EABC050FFD4DFF22FB11FE76FB5E" box="[151,440,1253,1275]" italics="true" pageId="9" pageNumber="489">
<taxonomicName id="3A944D2D050FFD4DFF22FB11FED4FB5E" ID-CoL="7PJ4L" box="[151,282,1253,1275]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="genus">Halicarcinus</taxonomicName>
innominatus—
</emphasis>
<treatmentCitation id="7C3510BF050FFD4DFE02FB11FDBCFB5E" author="Melrose" box="[439,626,1253,1275]" page="26" pageId="9" pageNumber="489" year="1975">
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: 26
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34, figs 511, pl. 1 (for extensive synonymy).
<emphasis id="CFE0EABC050FFD4DFBD1FB12FBB4FB5E" box="[1124,1146,1254,1275]" italics="true" pageId="9" pageNumber="489"></emphasis>
<treatmentCitation id="7C3510BF050FFD4DFBCEFB12FAF6FB5E" author="McLay" box="[1147,1336,1253,1275]" page="362" pageId="9" pageNumber="489" year="1988">
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: 362
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365, fig. 77.
<emphasis id="CFE0EABC050FFD4DFF53FAF0FF32FABC" box="[230,252,1284,1305]" italics="true" pageId="9" pageNumber="489"></emphasis>
<treatmentCitation id="7C3510BF050FFD4DFF48FAF0FE68FABC" author="Poore" box="[253,422,1283,1305]" page="394" pageId="9" pageNumber="489" year="2004">
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: 394
</treatmentCitation>
, fig. 120c.
<emphasis id="CFE0EABC050FFD4DFDA4FAF0FDE9FABC" box="[529,551,1284,1305]" italics="true" pageId="9" pageNumber="489"></emphasis>
<treatmentCitation id="7C3510BF050FFD4DFD9DFAF7FD2AFABC" author="Webber" box="[552,740,1283,1305]" page="185" pageId="9" pageNumber="489" year="2010">
<bibRefCitation id="99054B5F050FFD4DFD9DFAF7FD7DFABC" author="Webber" box="[552,691,1283,1305]" pageId="9" pageNumber="514" refString="Webber, W. R., Fenwick, G. D., Bradford-Grieve, J. M., Eagar, S. H., Buckeridge, J. S., Poore, G. C. B., Dawson, E. W., Watling, L., Jones, J. B., Wells, J. B. J., Bruce, N. L., Ahyong, S. T., Larsen, K., Chapman, M. A., Olesen, J., Ho, J. S., Green, J. D., Shiel, R. J., Rocha, C. E. F., Lorz, A. - N., Bird, G. J. &amp; Charleston, W. A. (2010) Chapter 8. Phylum Arthropoda. Subphylum Crustacea. Shrimps, crabs, lobsters, barnacles, slaters, and kin. In: Gordon, D. P. (Ed.) New Zealand Inventory of Biodiversity. 2. Kingdom Animalia. Canterbury University Press, Christchurch, pp. 98 - 232." type="journal article" year="2010">Webber 2010</bibRefCitation>
: 185
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, 186, 226.
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</paragraph>
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<emphasis id="CFE0EABC050FFD4DFF22FAB2FEB3FAFA" bold="true" box="[151,381,1350,1375]" pageId="9" pageNumber="489">Material examined</emphasis>
.
<typeStatus id="222F880C050FFD4DFE3AFAB3FE31FAFA" box="[399,511,1351,1375]" pageId="9" pageNumber="489" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="3A944D2D050FFD4DFD99FAB2FBC8FAFA" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[556,1030,1350,1375]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFD99FAB2FD16FAFB" box="[556,728,1350,1375]" italics="true" pageId="9" pageNumber="489">Elamena quoyi</emphasis>
H.
<bibRefCitation id="99054B5F050FFD4DFCBCFAB2FBC8FAFA" author="Milne" box="[777,1030,1350,1375]" pageId="9" pageNumber="513" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles (ser. 3), 20, 163 - 228, pls 1 - 11. Avaliable from: http: // biodiversitylibrary. org / page / 35466171 (Accessed 21 Mar. 2016)" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
, herein selected.
<collectingCountry id="8583763E050FFD4DFB67FAB2FA52FAFA" box="[1234,1436,1350,1375]" name="New Zealand" pageId="9" pageNumber="489">Nouvelle-Zélande</collectingCountry>
(
<collectingCountry id="8583763E050FFD4DFF2BFA9FFEF3FA26" box="[158,317,1386,1411]" name="New Zealand" pageId="9" pageNumber="489">New Zealand</collectingCountry>
; mislabelled
<collectingCountry id="8583763E050FFD4DFE62FA9EFD73FA26" box="[471,701,1386,1411]" name="New Caledonia" pageId="9" pageNumber="489">Nouvelle-Calédonie</collectingCountry>
), MNHN-2000-667 (B667), (male,
<quantity id="3A6C9B4B050FFD4DFBEFFA9EFB7BFA27" box="[1114,1205,1386,1411]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="489" unit="mm" value="6.4">6.4 mm</quantity>
, dry).
<typeStatus id="222F880C050FFD4DFAB1FA9EFA52FA26" box="[1284,1436,1386,1411]" pageId="9" pageNumber="489" type="paralectotype">Paralectotype</typeStatus>
fragments of
<taxonomicName id="3A944D2D050FFD4DFE99FA7AFE1BFA03" box="[300,469,1422,1447]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFE99FA7AFE1BFA03" box="[300,469,1422,1447]" italics="true" pageId="9" pageNumber="489">Elamena quoyi</emphasis>
</taxonomicName>
. Same locality, MNHN-2000-32609 (B32609), (sex unknown, dry).
</paragraph>
<paragraph id="FD2B36AE050FFD4DFF72FA47FEB9FA4A" blockId="9.[151,1437,1350,2023]" pageId="9" pageNumber="489">
<typeStatus id="222F880C050FFD4DFF72FA47FEEDFA6E" box="[199,291,1459,1483]" pageId="9" pageNumber="489" type="syntype">Syntype</typeStatus>
of
<taxonomicName id="3A944D2D050FFD4DFEEEFA46FD3DFA6E" authority="Filhol, 1885" authorityName="Filhol" authorityYear="1885" box="[347,755,1458,1483]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="huttoni">
<emphasis id="CFE0EABC050FFD4DFEEEFA46FD82FA6E" box="[347,588,1458,1483]" italics="true" pageId="9" pageNumber="489">Halicarcinus huttoni</emphasis>
<bibRefCitation id="99054B5F050FFD4DFDE8FA46FD3DFA6E" author="Filhol" box="[605,755,1458,1483]" lastPageId="9" lastPageNumber="512" pageId="9" pageNumber="511" refString="Filhol, H. (1885 a) Descriptions de nouvelles especes de Crustaces appartenant au genre Hymenicus, provenant de la Nouvelle- Zelande. Bulletin de la Societe Philomathique de Paris, ser. 7, 9, 43 - 44. Avaliable from: http: // biodiversitylibrary. org / page / 31945637 (Accessed 21 Mar. 2016)" type="journal article" year="1885">Filhol, 1885</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="8583763E050FFD4DFCB2FA47FC64FA6E" box="[775,938,1458,1483]" name="New Zealand" pageId="9" pageNumber="489">New Zealand</collectingCountry>
, Port Chambers, MNHN IU-2000-32613 (B32613), (female).
</paragraph>
<paragraph id="FD2B36AE050FFD4DFF72FA0EFE4EF992" blockId="9.[151,1437,1350,2023]" pageId="9" pageNumber="489">
Other material.
<collectingCountry id="8583763E050FFD4DFEC9FA0FFDD9F9B6" box="[380,535,1530,1555]" name="New Zealand" pageId="9" pageNumber="489">New Zealand</collectingCountry>
, Otago coast, NMV J7372 (ovigerous female,
<quantity id="3A6C9B4B050FFD4DFB85FA0FFB57F9B7" box="[1072,1177,1531,1555]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="9" pageNumber="489" unit="mm" value="12.1">12.1 mm</quantity>
;
<specimenCount id="EB92FD27050FFD4DFB12FA0FFB37F9B6" box="[1191,1273,1530,1555]" pageId="9" pageNumber="489" type="male">1 male</specimenCount>
); NMV J7036 (males and females).
</paragraph>
</subSubSection>
<subSubSection id="B58E6525050FFD4DFF72F9B6FA92F9FE" box="[199,1372,1602,1627]" pageId="9" pageNumber="489" type="distribution">
<paragraph id="FD2B36AE050FFD4DFF72F9B6FA92F9FE" blockId="9.[151,1437,1350,2023]" box="[199,1372,1602,1627]" pageId="9" pageNumber="489">
<emphasis id="CFE0EABC050FFD4DFF72F9B6FE97F9FE" bold="true" box="[199,345,1602,1627]" pageId="9" pageNumber="489">Distribution</emphasis>
.
<collectingCountry id="8583763E050FFD4DFED2F9B7FE33F9FE" box="[359,509,1602,1627]" name="New Zealand" pageId="9" pageNumber="489">New Zealand</collectingCountry>
except subantarctic islands; introduced to Tasmania,
<collectingCountry id="8583763E050FFD4DFBE6F9B6FB74F9FE" box="[1107,1210,1602,1627]" name="Australia" pageId="9" pageNumber="489">Australia</collectingCountry>
(
<bibRefCitation id="99054B5F050FFD4DFB7CF9B7FA9EF9FE" author="Poore" box="[1225,1360,1603,1627]" pageId="9" pageNumber="513" refString="Poore, G. C. B. (2004) Marine Decapod Crustacea of Southern Australia. A Guide to Identification (with chapter on Stomatopoda by Shane Ahyong). CSIRO Publishing, Melbourne, 574 pp." type="book" year="2004">Poore 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B58E6525050FFD4EFF72F992FC51FF56" lastPageId="10" lastPageNumber="490" pageId="9" pageNumber="489" type="discussion">
<paragraph id="FD2B36AE050FFD4DFF72F992FAE2F843" blockId="9.[151,1437,1350,2023]" pageId="9" pageNumber="489">
<emphasis id="CFE0EABC050FFD4DFF72F992FEFAF9DA" bold="true" box="[199,308,1638,1663]" pageId="9" pageNumber="489">Remarks</emphasis>
. The two dry specimens of the
<typeStatus id="222F880C050FFD4DFD75F993FD3EF9DA" box="[704,752,1639,1663]" pageId="9" pageNumber="489">type</typeStatus>
series of
<taxonomicName id="3A944D2D050FFD4DFCDAF992FA9AF9DA" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[879,1364,1638,1663]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFCDAF992FBD3F9DB" box="[879,1053,1638,1663]" italics="true" pageId="9" pageNumber="489">Elamena quoyi</emphasis>
H.
<bibRefCitation id="99054B5F050FFD4DFBE6F992FA9AF9DA" author="Milne" box="[1107,1364,1638,1663]" pageId="9" pageNumber="513" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles (ser. 3), 20, 163 - 228, pls 1 - 11. Avaliable from: http: // biodiversitylibrary. org / page / 35466171 (Accessed 21 Mar. 2016)" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
, were examined and a
<typeStatus id="222F880C050FFD4DFEECF97EFE0EF906" box="[345,448,1674,1699]" pageId="9" pageNumber="489" type="lectotype">lectotype</typeStatus>
is selected. H. Milne Edwards schematic drawing showed a low and unilobate-like rostrum, which lead
<bibRefCitation id="99054B5F050FFD4DFE34F95AFDDAF962" author="Tesch" box="[385,532,1710,1735]" pageId="9" pageNumber="514" refString="Tesch, J. J. (1918) Decapoda F. (Decapoda Brachyura continued) Hymenosomatidae, Retropilumnidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls 1 - 6. Avaliable from: http: // biodiversitylibrary. org / page / 2037493 (Accessed 21 Mar. 2016)" type="journal article" year="1918">Tesch (1918)</bibRefCitation>
and
<bibRefCitation id="99054B5F050FFD4DFDF8F95AFD35F962" author="Melrose" box="[589,763,1710,1735]" pageId="9" pageNumber="513" refString="Melrose, M. J. (1975) The marine fauna of New Zealand: family Hymenosomatidae (Crustacea, Decapoda, Brachyura). New Zealand Oceanographic Institute Memoir, 34, 1 - 123." type="journal article" year="1975">Melrose (1975)</bibRefCitation>
to erroneously consider this poorly known species a junior synonym of
<taxonomicName id="3A944D2D050FFD4DFE98F926FD2DF94E" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[301,739,1746,1771]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CFE0EABC050FFD4DFE98F926FDF0F94E" box="[301,574,1746,1771]" italics="true" pageId="9" pageNumber="489">Halicarcinus pubescens</emphasis>
(
<bibRefCitation id="99054B5F050FFD4DFDE7F927FD15F94E" author="Dana" box="[594,731,1746,1771]" pageId="9" pageNumber="511" refString="Dana, J. D. (1851) Conspectus crustaceorum quae in orbis terrarum circumnavigatione, Carolo Wilkes e classe reipublicae foederatae duce, lexit e descripsit. Proceedings of the Academy of Natural Science of Philadelphia, 5, 247 - 254. Avaliable from: http: // biodiversitylibrary. org / page / 26301482 (Accessed 21 Mar. 2016)" type="journal article" year="1851">Dana, 1851</bibRefCitation>
)
</taxonomicName>
(now placed in
<taxonomicName id="3A944D2D050FFD4DFC19F926FB20F94E" authority="Lucas, 1980" authorityName="Lucas" authorityYear="1980" box="[940,1262,1746,1771]" class="Malacostraca" family="Hymenosomatidae" genus="Neohymenicus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="CFE0EABC050FFD4DFC19F926FB81F94E" box="[940,1103,1746,1771]" italics="true" pageId="9" pageNumber="489">Neohymenicus</emphasis>
<bibRefCitation id="99054B5F050FFD4DFBE9F927FB20F94E" author="Lucas" box="[1116,1262,1747,1771]" pageId="9" pageNumber="513" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33, 148 - 247." type="journal article" year="1980">Lucas, 1980</bibRefCitation>
</taxonomicName>
). Melrose also included it as a possible synonym of
<emphasis id="CFE0EABC050FFD4DFDF6F903FD37F8AB" box="[579,761,1783,1806]" italics="true" pageId="9" pageNumber="489">
<taxonomicName id="3A944D2D050FFD4DFDF6F903FD3BF8AB" box="[579,757,1783,1806]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="innominatus">H. innominatus</taxonomicName>
.
</emphasis>
The
<typeStatus id="222F880C050FFD4DFC80F902FC52F8AA" box="[821,924,1782,1807]" pageId="9" pageNumber="489" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="3A944D2D050FFD4DFC70F903FBEAF8AB" box="[965,1060,1783,1806]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFC70F903FBEAF8AB" box="[965,1060,1783,1806]" italics="true" pageId="9" pageNumber="489">E. quoyi</emphasis>
</taxonomicName>
(fig. 1e) actually has a short but sharp rostrum and lateral pseudorostral lobes. As
<bibRefCitation id="99054B5F050FFD4DFD72F8EEFC6FF896" author="Melrose" box="[711,929,1818,1843]" pageId="9" pageNumber="513" refString="Melrose, M. J. (1975) The marine fauna of New Zealand: family Hymenosomatidae (Crustacea, Decapoda, Brachyura). New Zealand Oceanographic Institute Memoir, 34, 1 - 123." type="journal article" year="1975">Melrose (1975: 26)</bibRefCitation>
noted,
<taxonomicName id="3A944D2D050FFD4DFC40F8EEFF1CF8F2" authority="Richardson, 1949" authorityName="Richardson" authorityYear="1949" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="innominata">
<emphasis id="CFE0EABC050FFD4DFC40F8EEFAC0F897" box="[1013,1294,1818,1843]" italics="true" pageId="9" pageNumber="489">Halicarcinus innominata</emphasis>
<bibRefCitation id="99054B5F050FFD4DFAA0F8EEFF1CF8F2" author="Richardson" lastPageId="9" lastPageNumber="514" pageId="9" pageNumber="513" refString="Richardson, L. R. (1949 a) A guide to the Oxyrhyncha, Oxystoma and lesser crabs. Tuatara, 2, 58 - 69. Avaliable from: http: // nzetc. victoria. ac. nz / tm / scholarly / tei-Bio 02 Tuat 02 - t 1 - body-d 2. html (Accessed 21 Mar. 2016)" type="journal article" year="1949">Richardson, 1949</bibRefCitation>
</taxonomicName>
, was erected without designation of
<typeStatus id="222F880C050FFD4DFDC3F8CBFD7FF8F2" box="[630,689,1855,1879]" pageId="9" pageNumber="489">types</typeStatus>
or material of any kind or of a
<typeStatus id="222F880C050FFD4DFB95F8CBFB9EF8F2" box="[1056,1104,1855,1879]" pageId="9" pageNumber="489">type</typeStatus>
locality (contravening ICZN Article 61). She corrected the spelling of the species name, nominated a
<typeStatus id="222F880C050FFD4DFC42F897FBE9F8DE" box="[1015,1063,1891,1915]" pageId="9" pageNumber="489">type</typeStatus>
locality but was unable to find specimens identified by Richardson … in the Dominion Museum or Victoria University of Wellington Zoology Department Collections. Richardson also failed to designate a
<typeStatus id="222F880C050FFD4DFCDEF85EFC03F866" box="[875,973,1962,1987]" pageId="9" pageNumber="489" type="holotype">holotype</typeStatus>
and although the name has been widely used in
<collectingCountry id="8583763E050FFD4DFF5BF83BFE4AF842" box="[238,388,1998,2023]" name="New Zealand" pageId="9" pageNumber="489">New Zealand</collectingCountry>
the species remains a
<emphasis id="CFE0EABC050FFD4DFDCAF83BFCD3F842" box="[639,797,1998,2023]" italics="true" pageId="9" pageNumber="489">nomen nudum</emphasis>
. It is here treated as conspecific with
<taxonomicName id="3A944D2D050FFD4DFB71F83BFAEBF843" box="[1220,1317,1999,2022]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050FFD4DFB71F83BFAEBF843" box="[1220,1317,1999,2022]" italics="true" pageId="9" pageNumber="489">H. quoyi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FD2B36AE050CFD4EFF72FF63FC51FF56" blockId="10.[151,1436,151,243]" pageId="10" pageNumber="490">
<bibRefCitation id="99054B5F050CFD4EFF72FF63FEBAFF15" author="Melrose" box="[199,372,151,176]" pageId="10" pageNumber="513" refString="Melrose, M. J. (1975) The marine fauna of New Zealand: family Hymenosomatidae (Crustacea, Decapoda, Brachyura). New Zealand Oceanographic Institute Memoir, 34, 1 - 123." type="journal article" year="1975">Melrose (1975)</bibRefCitation>
included
<taxonomicName id="3A944D2D050CFD4EFE57FF6DFD9AFF15" box="[482,596,151,176]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="490" phylum="Arthropoda" rank="species" species="huttoni">
<emphasis id="CFE0EABC050CFD4EFE57FF6DFD9AFF15" box="[482,596,151,176]" italics="true" pageId="10" pageNumber="490">H. huttoni</emphasis>
</taxonomicName>
in the synonymy of both
<taxonomicName id="3A944D2D050CFD4EFCC6FF6DFC10FF15" box="[883,990,153,176]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="490" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="CFE0EABC050CFD4EFCC6FF6DFC10FF15" box="[883,990,153,176]" italics="true" pageId="10" pageNumber="490">H. ovatus</emphasis>
</taxonomicName>
and
<taxonomicName id="3A944D2D050CFD4EFBA1FF6DFB0FFF15" box="[1044,1217,153,176]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="490" phylum="Arthropoda" rank="species" species="innominatus">
<emphasis id="CFE0EABC050CFD4EFBA1FF6DFB0FFF15" box="[1044,1217,153,176]" italics="true" pageId="10" pageNumber="490">H. innominatus</emphasis>
</taxonomicName>
(now
<taxonomicName id="3A944D2D050CFD4EFABDFF6DFAA7FF15" box="[1288,1385,153,176]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="490" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="CFE0EABC050CFD4EFABDFF6DFAA7FF15" box="[1288,1385,153,176]" italics="true" pageId="10" pageNumber="490">H. quoyi</emphasis>
</taxonomicName>
). Its
<typeStatus id="222F880C050CFD4EFF22FF4EFF09FF77" box="[151,199,186,210]" pageId="10" pageNumber="490">type</typeStatus>
locality in
<collectingCountry id="8583763E050CFD4EFEF8FF4EFE2BFF77" box="[333,485,185,210]" name="New Zealand" pageId="10" pageNumber="490">New Zealand</collectingCountry>
and overall morphology lead to the conclusion that it is a synonym of the latter, a
<collectingCountry id="8583763E050CFD4EFF22FF2FFEE3FF56" box="[151,301,218,243]" name="New Zealand" pageId="10" pageNumber="490">New Zealand</collectingCountry>
endemic rather than the former, an Australian endemic.
</paragraph>
</subSubSection>
</treatment>
</document>