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<document id="F2EB550206DBEE82DA0DDECDAD80605D" ID-DOI="10.11646/zootaxa.3949.2.9" ID-GBIF-Dataset="14f1d596-9bad-4ea5-b6d7-a9fa8a0ef4a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="233332" ID-ZooBank="6EE53509-2ED0-4466-986B-A9DA84413CB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045809853" checkinUser="plazi" docAuthor="Silva, Rodet Rodriguez" docDate="2015" docId="AC2787ECF334B064FF7C5CF61DE1A07B" docLanguage="en" docName="zt03949p296.pdf" docOrigin="Zootaxa 3949 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rivulus berovidesi Silva, 2015, new species" docType="treatment" docVersion="7" lastPageNumber="294" masterDocId="501EFF94F335B061FFEB5F1B1C72A407" masterDocTitle="Rivulus berovidesi, a new killifish species (Teleostei: Rivulidae) from western Cuba" masterLastPageNumber="296" masterPageNumber="289" pageNumber="290" updateTime="1698576811506" updateUser="plazi">
<mods:mods id="8237139F8E60F1BC6ADF4DD91BE67AB3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D72410E2003845D275FE8A855F187BF9">Rivulus berovidesi, a new killifish species (Teleostei: Rivulidae) from western Cuba</mods:title>
</mods:titleInfo>
<mods:name id="6B0C6206100F89FBB19FEF96540CB7EF" type="personal">
<mods:role id="0EA3C087C54753EFCEFA36E10E57C9AF">
<mods:roleTerm id="288A015158FE4B77558F8625BB04AC8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB212C6EE824FC63B692956F10BAB7E6">Silva, Rodet Rodriguez</mods:namePart>
</mods:name>
<mods:typeOfResource id="5CB7D7F60BDAF02621A80DC2C132021F">text</mods:typeOfResource>
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<mods:title id="AFE5D3568BEF4F1254790E418DF91367">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DC689AC1530C6710C93810E4CE90F235">
<mods:date id="FA8F2511786BE9B765D229C826264CA5">2015</mods:date>
<mods:detail id="4E2D0E434621048323CA5819BA84819E" type="volume">
<mods:number id="D75A9AD6513D4A1F2B3E4F9EEA98CAF1">3949</mods:number>
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<mods:classification id="06B457DB3C09D5A3284E1B9626C743CE">journal article</mods:classification>
<mods:identifier id="AE2E2FA5FEE6629EA8D82FD407633542" type="DOI">10.11646/zootaxa.3949.2.9</mods:identifier>
<mods:identifier id="8868B4225D6A8BCF30322141569E08F8" type="GBIF-Dataset">14f1d596-9bad-4ea5-b6d7-a9fa8a0ef4a0</mods:identifier>
<mods:identifier id="56C87960F96DE45024F83A603048DD4B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D3A026BC893A3496C39265E7D9AD4A94" type="Zenodo-Dep">233332</mods:identifier>
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</mods:mods>
<treatment id="AC2787ECF334B064FF7C5CF61DE1A07B" ID-DOI="http://doi.org/10.5281/zenodo.5662820" ID-GBIF-Taxon="119610587" ID-Zenodo-Dep="5662820" LSID="urn:lsid:plazi:treatment:AC2787ECF334B064FF7C5CF61DE1A07B" httpUri="http://treatment.plazi.org/id/AC2787ECF334B064FF7C5CF61DE1A07B" lastPageId="5" lastPageNumber="294" pageId="1" pageNumber="290">
<subSubSection id="6C946571F334B060FF7C5CF61D62A02E" pageId="1" pageNumber="290" type="nomenclature">
<paragraph id="243136FAF334B060FF7C5CF61E63A000" blockId="1.[151,529,1005,1065]" box="[151,529,1005,1031]" pageId="1" pageNumber="290">
<heading id="7F798196F334B060FF7C5CF61E63A000" bold="true" box="[151,529,1005,1031]" fontSize="11" level="1" pageId="1" pageNumber="290" reason="1">
<emphasis id="16FAEAE8F334B060FF7C5CF61E63A000" bold="true" box="[151,529,1005,1031]" pageId="1" pageNumber="290">
<taxonomicName id="E38E4D79F334B060FF7C5CF61D06A000" ID-CoL="6WWS9" box="[151,372,1005,1031]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F334B060FF7C5CF61D06A000" bold="true" box="[151,372,1005,1031]" italics="true" pageId="1" pageNumber="290">Rivulus berovidesi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="0DC95793F334B060FE695CF61E63A000" box="[386,529,1005,1031]" pageId="1" pageNumber="290" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="243136FAF334B060FF7C5B0B1D62A02E" blockId="1.[151,529,1005,1065]" box="[151,272,1040,1065]" pageId="1" pageNumber="290">
(
<figureCitation id="BCB52A7FF334B060FF745B0B1D7AA02E" box="[159,264,1040,1065]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="2.[151,249,1903,1925]" captionStartId-1="3.[151,249,877,899]" captionStartId-2="3.[151,249,1970,1992]" captionStartId-3="4.[151,249,1020,1042]" captionStartId-4="4.[151,249,1969,1991]" captionTargetBox-0="[155,1436,1127,1881]" captionTargetBox-1="[151,1398,151,840]" captionTargetBox-2="[328,1231,966,1865]" captionTargetBox-3="[166,1417,176,990]" captionTargetBox-4="[166,1417,1137,1930]" captionTargetId-0="figure@2.[151,1436,1127,1881]" captionTargetId-1="figure@3.[151,1399,150,842]" captionTargetId-2="figure@3.[265,1287,964,1935]" captionTargetId-3="figure@4.[166,1418,99,993]" captionTargetId-4="figure@4.[166,1418,1136,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="FIGURE 2. Outline drawings of the frontal scalation patterns of Rivulus from Cuba. A: Rivulus berovidesi sp. n. with a d - type pattern and B: Rivulus cylindraceus with an e - type pattern. The horizontal broken line indicates the pair of scales with their margins fully exposed." captionText-1="FIGURE 3. Rivulus berovidesi sp. n., life coloration in both sexes in specimens kept in captivity for three months after collection in the field." captionText-2="FIGURE 4. Rivulus berovidesi sp. n., life coloration in males (above) and females (below) in specimens just after collection in the field." captionText-3="FIGURE 5. Rivulus berovidesi sp. n., paratype, female, CZACC- 9.85, 37.2 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba." captionText-4="FIGURE 6. Rivulus berovidesi sp. n., holotype, male, CZACC- 9.83, 37.3 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba." httpUri-0="https://zenodo.org/record/233334/files/figure.png" httpUri-1="https://zenodo.org/record/233335/files/figure.png" httpUri-2="https://zenodo.org/record/233336/files/figure.png" httpUri-3="https://zenodo.org/record/233337/files/figure.png" httpUri-4="https://zenodo.org/record/233338/files/figure.png" pageId="1" pageNumber="290">Figs. 2–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6C946571F334B060FF7C5B4F1F0CA141" pageId="1" pageNumber="290" type="materials_examined">
<paragraph id="243136FAF334B060FF7C5B4F1FF2A095" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="290">
<emphasis id="16FAEAE8F334B060FF7C5B4F1D79A06A" bold="true" box="[151,267,1108,1133]" pageId="1" pageNumber="290">
<typeStatus id="FB358858F334B060FF7C5B4F1D74A06A" box="[151,262,1108,1133]" pageId="1" pageNumber="290" type="holotype">Holotype</typeStatus>
.
</emphasis>
CZACC-9.83, adult male,
<quantity id="E3769B1FF334B060FDAD5B4F1EDFA06B" box="[582,685,1108,1132]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="1" pageNumber="290" unit="mm" value="37.3">37.3 mm</quantity>
<collectionCode id="429FAE3FF334B060FD5C5B4F1EA8A06B" box="[695,730,1108,1132]" pageId="1" pageNumber="290">SL</collectionCode>
, Sierra de Cajalbana, Pinar del Rio province,
<collectingCountry id="5C99766AF334B060FB135B4F1947A06A" box="[1272,1333,1108,1133]" name="Cuba" pageId="1" pageNumber="290">Cuba</collectingCountry>
, 220 46’ 31.7’’N, -830 26’ 37.7’’W,
<date id="5030103AF334B060FE215B621E20A095" box="[458,594,1145,1170]" pageId="1" pageNumber="290" value="1926-02">26 February</date>
, 2014. R. Rodriguez Silva.
</paragraph>
<paragraph id="243136FAF334B060FF2C5B861F0CA141" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="290">
<emphasis id="16FAEAE8F334B060FF2C5B861D36A0B1" bold="true" box="[199,324,1181,1206]" pageId="1" pageNumber="290">
<typeStatus id="FB358858F334B060FF2C5B861D32A0B1" box="[199,320,1181,1206]" pageId="1" pageNumber="290" type="paratype">Paratypes</typeStatus>
.
</emphasis>
All from Sierra de Cajalbana, Pinar del Rio province,
<collectingCountry id="5C99766AF334B060FC455B861F9EA0B1" box="[942,1004,1181,1206]" name="Cuba" pageId="1" pageNumber="290">Cuba</collectingCountry>
. Same date and collector as
<typeStatus id="FB358858F334B060FADF5B8619EBA0B1" box="[1332,1433,1181,1206]" pageId="1" pageNumber="290" type="holotype">holotype</typeStatus>
. CZACC-9.84, three males,
<quantity id="E3769B1FF334B060FE235BD91E19A0DE" box="[456,619,1218,1242]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" metricValueMax="3.47" metricValueMin="3.11" pageId="1" pageNumber="290" unit="mm" value="32.9" valueMax="34.7" valueMin="31.1">31.1–34.7 mm</quantity>
<collectionCode id="429FAE3FF334B060FD9A5BD91EE6A0DD" box="[625,660,1218,1242]" pageId="1" pageNumber="290">SL</collectionCode>
; CZACC-9.85, three females,
<quantity id="E3769B1FF334B060FC0D5BDA18FCA0DE" box="[998,1166,1217,1242]" metricMagnitude="-2" metricUnit="m" metricValue="3.7099999999999995" metricValueMax="3.79" metricValueMin="3.63" pageId="1" pageNumber="290" unit="mm" value="37.099999999999994" valueMax="37.9" valueMin="36.3">36.3–37.9 mm</quantity>
<collectionCode id="429FAE3FF334B060FB7E5BD918CBA0DD" box="[1173,1209,1218,1242]" pageId="1" pageNumber="290">SL</collectionCode>
;
<collectionCode id="429FAE3FF334B060FB2F5BDA188FA0DD" box="[1220,1277,1217,1242]" httpUri="http://grbio.org/cool/hsk3-f0jp" name="Museo Felipe Poey" pageId="1" pageNumber="290">MFP</collectionCode>
18.00570, two males
<quantity id="E3769B1FF334B060FF095BFD1DFBA0FA" box="[226,393,1254,1278]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.28" metricValueMin="3.22" pageId="1" pageNumber="290" unit="mm" value="32.5" valueMax="32.8" valueMin="32.2">32.2–32.8 mm</quantity>
<collectionCode id="429FAE3FF334B060FE785BFD1DC6A0F9" box="[403,436,1254,1278]" pageId="1" pageNumber="290">SL</collectionCode>
and two females
<quantity id="E3769B1FF334B060FD6D5BFD1F5FA0FA" box="[646,813,1254,1278]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.4299999999999997" metricValueMin="3.37" pageId="1" pageNumber="290" unit="mm" value="34.0" valueMax="34.3" valueMin="33.7">33.7–34.3 mm</quantity>
<collectionCode id="429FAE3FF334B060FCDC5BFD1F28A0F9" box="[823,858,1254,1278]" pageId="1" pageNumber="290">SL</collectionCode>
;
<collectionCode id="429FAE3FF334B060FC815BFD1FB2A0FA" LSID="urn:lsid:biocol.org:col:34988" box="[874,960,1254,1277]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="290">MNHN</collectionCode>
1457 one male
<quantity id="E3769B1FF334B060FB7F5BFD188DA0FA" box="[1172,1279,1254,1278]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="1" pageNumber="290" unit="mm" value="31.3">31.3 mm</quantity>
<collectionCode id="429FAE3FF334B060FAE65BFD195CA0F9" box="[1293,1326,1254,1278]" pageId="1" pageNumber="290">SL</collectionCode>
and two females
<quantity id="E3769B1FF334B060FF165A121DE6A126" box="[253,404,1289,1314]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" metricValueMax="3.56" metricValueMin="3.1" pageId="1" pageNumber="290" unit="mm" value="33.3" valueMax="35.6" valueMin="31.0">31–35.6 mm</quantity>
<collectionCode id="429FAE3FF334B060FE485A111DB5A125" box="[419,455,1290,1314]" pageId="1" pageNumber="290">SL</collectionCode>
; MZC 171656 and 171657 two males
<quantity id="E3769B1FF334B060FC445A11182AA126" box="[943,1112,1290,1314]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" metricValueMax="3.79" metricValueMin="3.27" pageId="1" pageNumber="290" unit="mm" value="35.3" valueMax="37.9" valueMin="32.7">32.7–37.9 mm</quantity>
<collectionCode id="429FAE3FF334B060FB8D5A1118F5A125" box="[1126,1159,1290,1314]" pageId="1" pageNumber="290">SL</collectionCode>
respectively and MZC 171658 and 171659 two females
<quantity id="E3769B1FF334B060FDF25A361ECDA142" box="[537,703,1325,1350]" metricMagnitude="-2" metricUnit="m" metricValue="3.245" metricValueMax="3.35" metricValueMin="3.1399999999999997" pageId="1" pageNumber="290" unit="mm" value="32.45" valueMax="33.5" valueMin="31.4">31.4–33.5 mm</quantity>
<collectionCode id="429FAE3FF334B060FD205A351E9EA141" box="[715,748,1326,1350]" pageId="1" pageNumber="290">SL</collectionCode>
respectively.
</paragraph>
</subSubSection>
<subSubSection id="6C946571F334B060FF2C5A4A1E8FA219" pageId="1" pageNumber="290" type="diagnosis">
<paragraph id="243136FAF334B060FF2C5A4A1E8FA219" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="290">
<emphasis id="16FAEAE8F334B060FF2C5A4A1D33A16D" bold="true" box="[199,321,1361,1386]" pageId="1" pageNumber="290">Diagnosis.</emphasis>
Both males and females of
<taxonomicName id="E38E4D79F334B060FD4D5A4A1F08A16E" box="[678,890,1361,1385]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F334B060FD4D5A4A1F08A16E" box="[678,890,1361,1385]" italics="true" pageId="1" pageNumber="290">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F334B060FC625A491FCCA16E" box="[905,958,1362,1385]" pageId="1" pageNumber="290" rank="species">sp.n.</taxonomicNameLabel>
are readily distinguished from
<taxonomicName id="E38E4D79F334B060FAA25A4A1D57A18A" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="cylindraceus">
<emphasis id="16FAEAE8F334B060FAA25A4A1D57A18A" italics="true" pageId="1" pageNumber="290">Rivulus cylindraceus</emphasis>
</taxonomicName>
by the presence of a dark lateral band situated longitudinally along the lateral line, extended from the posterior margin of the eye to the base of the caudal fin (versus a lack of this dark lateral band in
<taxonomicName id="E38E4D79F334B060FF7C5AA61DF3A1D2" box="[151,385,1469,1493]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="cylindraceus">
<emphasis id="16FAEAE8F334B060FF7C5AA61DF3A1D2" box="[151,385,1469,1493]" italics="true" pageId="1" pageNumber="290">Rivulus cylindraceus</emphasis>
</taxonomicName>
). Dorsal fin slightly shifted to the caudal peduncle in relation to the origin of anal fin (
<quantity id="E3769B1FF334B060FA995AA51CB6A1FD" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="290" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
) when compared to
<taxonomicName id="E38E4D79F334B060FE5B5AFA1EE9A1FE" box="[432,667,1505,1529]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="cylindraceus">
<emphasis id="16FAEAE8F334B060FE5B5AFA1EE9A1FE" box="[432,667,1505,1529]" italics="true" pageId="1" pageNumber="290">Rivulus cylindraceus</emphasis>
</taxonomicName>
. Frontal scalation pattern in
<taxonomicName id="E38E4D79F334B060FBEA5AFA18A6A1FE" box="[1025,1236,1505,1529]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F334B060FBEA5AFA18A6A1FE" box="[1025,1236,1505,1529]" italics="true" pageId="1" pageNumber="290">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F334B060FB0B5AFA195AA1FD" box="[1248,1320,1505,1530]" pageId="1" pageNumber="290" rank="species">
<emphasis id="16FAEAE8F334B060FB0B5AFA195AA1FD" bold="true" box="[1248,1320,1505,1530]" pageId="1" pageNumber="290">sp. n.</emphasis>
</taxonomicNameLabel>
is
<emphasis id="16FAEAE8F334B060FABE5AFA1911A1FE" box="[1365,1379,1505,1529]" italics="true" pageId="1" pageNumber="290">d</emphasis>
-
<typeStatus id="FB358858F334B060FA815AF919EEA1FD" box="[1386,1436,1506,1530]" pageId="1" pageNumber="290">type</typeStatus>
versus
<emphasis id="16FAEAE8F334B060FF02591D1C87A21A" box="[233,245,1542,1565]" italics="true" pageId="1" pageNumber="290">e</emphasis>
-
<typeStatus id="FB358858F334B060FF16591D1D5DA219" box="[253,303,1542,1566]" pageId="1" pageNumber="290">type</typeStatus>
pattern in
<taxonomicName id="E38E4D79F334B060FE58591E1ED6A21A" box="[435,676,1541,1565]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="cylindraceus">
<emphasis id="16FAEAE8F334B060FE58591E1ED6A21A" box="[435,676,1541,1565]" italics="true" pageId="1" pageNumber="290">Rivulus cylindraceus</emphasis>
</taxonomicName>
(
<figureCitation id="BCB52A7FF334B060FD58591E1E84A219" box="[691,758,1541,1566]" captionStart="FIGURE 2" captionStartId="2.[151,249,1903,1925]" captionTargetBox="[155,1436,1127,1881]" captionTargetId="figure@2.[151,1436,1127,1881]" captionTargetPageId="2" captionText="FIGURE 2. Outline drawings of the frontal scalation patterns of Rivulus from Cuba. A: Rivulus berovidesi sp. n. with a d - type pattern and B: Rivulus cylindraceus with an e - type pattern. The horizontal broken line indicates the pair of scales with their margins fully exposed." httpUri="https://zenodo.org/record/233334/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6C946571F334B064FF2C593218BBA610" lastPageId="5" lastPageNumber="294" pageId="1" pageNumber="290" type="description">
<paragraph id="243136FAF334B060FF2C59321879A31C" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="290">
<emphasis id="16FAEAE8F334B060FF2C59321D25A245" bold="true" box="[199,343,1577,1602]" pageId="1" pageNumber="290">Description.</emphasis>
Morphometric and meristic data for
<typeStatus id="FB358858F334B060FCE759321F1FA245" box="[780,877,1577,1602]" pageId="1" pageNumber="290" type="holotype">holotype</typeStatus>
and 22
<typeStatus id="FB358858F334B060FC385931184FA245" box="[979,1085,1578,1602]" pageId="1" pageNumber="290" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="690C0341F334B060FAFB5932191FA245" box="[1296,1389,1577,1602]" captionStart="TABLE 1" captionStartId="2.[151,237,152,175]" captionTargetBox="[159,1342,266,629]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data for holotype and paratypes of Rivulus berovidesi sp. n. Standard length (SL) and head length (HL) are expressed in millimeters; all other measurements are expressed as percentage of SL except for eye diameter and snout length, which are given as percentage of HL." httpUri="http://table.plazi.org/id/70F16672F337B063FF7C5F831FEDA4E8" pageId="1" pageNumber="290" tableUuid="70F16672F337B063FF7C5F831FEDA4E8">Tables 1</tableCitation>
and
<tableCitation id="690C0341F334B060FF7C59551CDAA261" box="[151,168,1614,1638]" captionStart="TABLE 2" captionStartId="2.[151,237,693,716]" captionText="TABLE 2. Meristic data for holotype and paratypes of Rivulus berovidesi sp. n. Values in parentheses indicate number of specimens examined with that count." pageId="1" pageNumber="290">2</tableCitation>
. Largest examined male
<quantity id="E3769B1FF334B060FE5559551E53A262" box="[446,545,1614,1638]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="1" pageNumber="290" unit="mm" value="37.9">37.9 mm</quantity>
SL; largest examined female 38.0 mm SL. Body subcylindrical anteriorly and compressed posteriorly. Dorsal profile slightly convex from the snout to the posterior base of dorsal fin. Ventral profile slightly convex from the lower jaw to the anal fin origin. Both paired and impaired fins rounded and without filaments. Dorsal fin origin above base of the 5th or 6th anal fin ray. Pectoral fins inserted behind to the posterior margin of opercles. Pelvic fins are the smallest, reaching anus. Dorsal fin rays 6–9, caudal fin rays 15–19, anal fin rays 8–11, pelvic fin rays 4–6 and pectoral fin rays 11–14.
</paragraph>
<paragraph id="243136FAF334B060FF2C583D1F60A3F4" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="290">
Body and head entirely scaled with cycloid scales. Longitudinal series of scales 34–37, transverse series of scales 9, pre-dorsal scales 21–25. Frontal scalation is
<emphasis id="16FAEAE8F334B060FCF758501F58A364" box="[796,810,1867,1891]" italics="true" pageId="1" pageNumber="290">d</emphasis>
-
<typeStatus id="FB358858F334B060FCDA58501F11A364" box="[817,867,1867,1891]" pageId="1" pageNumber="290">type</typeStatus>
pattern. Males of
<taxonomicName id="E38E4D79F334B060FBAD58501956A364" box="[1094,1316,1867,1891]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F334B060FBAD58501956A364" box="[1094,1316,1867,1891]" italics="true" pageId="1" pageNumber="290">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F334B060FADE58501918A365" box="[1333,1386,1867,1890]" pageId="1" pageNumber="290" rank="species">sp.n.</taxonomicNameLabel>
are yellow and orange ventrally, whereas females are less colored.
<figureCitation id="BCB52A7FF334B060FC4D5875187FA380" box="[934,1037,1902,1927]" captionStart="FIGURE 3" captionStartId="3.[151,249,877,899]" captionTargetBox="[151,1398,151,840]" captionTargetId="figure@3.[151,1399,150,842]" captionTargetPageId="3" captionText="FIGURE 3. Rivulus berovidesi sp. n., life coloration in both sexes in specimens kept in captivity for three months after collection in the field." httpUri="https://zenodo.org/record/233335/files/figure.png" pageId="1" pageNumber="290">Figure 3</figureCitation>
shows sexual dimorphism in this species, which includes differences in color of body and fins as well as females characterized by a distinctive ocellated caudal spot. Females have a diffuse ocellated caudal spot which can extend from the posterior base of the dorsal fin to the upper margin of the caudal fin.
</paragraph>
<caption id="70F16672F337B063FF7C5F831FEDA4E8" ID-Table-UUID="70F16672F337B063FF7C5F831FEDA4E8" httpUri="http://table.plazi.org/id/70F16672F337B063FF7C5F831FEDA4E8" pageId="2" pageNumber="291" targetBox="[159,1342,266,629]" targetIsTable="true" targetPageId="2">
<paragraph id="243136FAF337B063FF7C5F831FEDA4E8" blockId="2.[151,1436,151,239]" pageId="2" pageNumber="291">
<emphasis id="16FAEAE8F337B063FF7C5F831D75A4A8" bold="true" box="[151,263,152,175]" pageId="2" pageNumber="291">TABLE 1</emphasis>
. Morphometric data for holotype and paratypes of
<taxonomicName id="E38E4D79F337B063FCBC5F8C1853A4A8" box="[855,1057,151,175]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F337B063FCBC5F8C1853A4A8" box="[855,1057,151,175]" italics="true" pageId="2" pageNumber="291">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F337B063FBC55F831803A4A8" box="[1070,1137,152,175]" pageId="2" pageNumber="291" rank="species">
<emphasis id="16FAEAE8F337B063FBC55F831803A4A8" bold="true" box="[1070,1137,152,175]" pageId="2" pageNumber="291">sp. n.</emphasis>
</taxonomicNameLabel>
Standard length (SL) and head length (HL) are expressed in millimeters; all other measurements are expressed as percentage of SL except for eye diameter and snout length, which are given as percentage of HL.
</paragraph>
</caption>
<paragraph id="243136FAF337B063FDC85E11194CA672" pageId="2" pageNumber="291">
<table id="568EC45AF3374F9EFF745E11194CA672" box="[159,1342,266,629]" gridcols="6" gridrows="10" pageId="2" pageNumber="291">
<tr id="9ABE34B8F3374F9EFF745E11194CA527" box="[159,1342,266,288]" gridrow="0" pageId="2" pageNumber="291" rowspan-0="1" rowspan-4="1" rowspan-5="1">
<th id="D96F5DC4F3374F9EFDC85E111EF3A527" box="[547,641,266,288]" gridcol="1" gridrow="0" pageId="2" pageNumber="291">Holotype</th>
<th id="D96F5DC4F3374F9EFD1A5E111F79A527" box="[753,779,266,288]" gridcol="2" gridrow="0" pageId="2" pageNumber="291">N</th>
<th id="D96F5DC4F3374F9EFC815E111FBEA527" box="[874,972,266,288]" gridcol="3" gridrow="0" pageId="2" pageNumber="291">Paratypes</th>
</tr>
<tr id="9ABE34B8F3374F9EFF745E29194CA54F" box="[159,1342,306,328]" gridrow="1" pageId="2" pageNumber="291" rowspan-1="1" rowspan-2="1">
<th id="D96F5DC4F3374F9EFF745E291DE3A54F" box="[159,401,306,328]" gridcol="0" gridrow="1" pageId="2" pageNumber="291">Morphometric variables</th>
<td id="D96F5DC4F3374F9EFC815E291FBEA54F" box="[874,972,306,328]" gridcol="3" gridrow="1" pageId="2" pageNumber="291">Mean</td>
<td id="D96F5DC4F3374F9EFBD95E2918BFA54F" box="[1074,1229,306,328]" gridcol="4" gridrow="1" pageId="2" pageNumber="291">Min Max</td>
<td id="D96F5DC4F3374F9EFAF55E29194CA54F" box="[1310,1342,306,328]" gridcol="5" gridrow="1" pageId="2" pageNumber="291">SD</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745E42194CA568" box="[159,1342,345,367]" gridrow="2" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745E421DE3A568" box="[159,401,345,367]" gridcol="0" gridrow="2" pageId="2" pageNumber="291">Standard length (mm)</th>
<td id="D96F5DC4F3374F9EFDC85E421EF3A568" box="[547,641,345,367]" gridcol="1" gridrow="2" pageId="2" pageNumber="291">37.3</td>
<td id="D96F5DC4F3374F9EFD1A5E421F79A568" box="[753,779,345,367]" gridcol="2" gridrow="2" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815E421FBEA568" box="[874,972,345,367]" gridcol="3" gridrow="2" pageId="2" pageNumber="291">33.8</td>
<td id="D96F5DC4F3374F9EFBD95E4218BFA568" box="[1074,1229,345,367]" gridcol="4" gridrow="2" pageId="2" pageNumber="291">28.9 37.9</td>
<td id="D96F5DC4F3374F9EFAF55E42194CA568" box="[1310,1342,345,367]" gridcol="5" gridrow="2" pageId="2" pageNumber="291">2.5</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745E64194CA592" box="[159,1342,383,405]" gridrow="3" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745E641DE3A592" box="[159,401,383,405]" gridcol="0" gridrow="3" pageId="2" pageNumber="291">Pre-dorsal length</th>
<td id="D96F5DC4F3374F9EFDC85E641EF3A592" box="[547,641,383,405]" gridcol="1" gridrow="3" pageId="2" pageNumber="291">68.0</td>
<td id="D96F5DC4F3374F9EFD1A5E641F79A592" box="[753,779,383,405]" gridcol="2" gridrow="3" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815E641FBEA592" box="[874,972,383,405]" gridcol="3" gridrow="3" pageId="2" pageNumber="291">70.1</td>
<td id="D96F5DC4F3374F9EFBD95E6418BFA592" box="[1074,1229,383,405]" gridcol="4" gridrow="3" pageId="2" pageNumber="291">67.3 71.5</td>
<td id="D96F5DC4F3374F9EFAF55E64194CA592" box="[1310,1342,383,405]" gridcol="5" gridrow="3" pageId="2" pageNumber="291">1.6</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745EBF194CA5BD" box="[159,1342,420,442]" gridrow="4" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745EBF1DE3A5BD" box="[159,401,420,442]" gridcol="0" gridrow="4" pageId="2" pageNumber="291">Pre-anal length</th>
<td id="D96F5DC4F3374F9EFDC85EBF1EF3A5BD" box="[547,641,420,442]" gridcol="1" gridrow="4" pageId="2" pageNumber="291">60.9</td>
<td id="D96F5DC4F3374F9EFD1A5EBF1F79A5BD" box="[753,779,420,442]" gridcol="2" gridrow="4" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815EBF1FBEA5BD" box="[874,972,420,442]" gridcol="3" gridrow="4" pageId="2" pageNumber="291">61.9</td>
<td id="D96F5DC4F3374F9EFBD95EBF18BFA5BD" box="[1074,1229,420,442]" gridcol="4" gridrow="4" pageId="2" pageNumber="291">59.5 65.3</td>
<td id="D96F5DC4F3374F9EFAF55EBF194CA5BD" box="[1310,1342,420,442]" gridcol="5" gridrow="4" pageId="2" pageNumber="291">1.2</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745ED0194CA5E6" box="[159,1342,459,481]" gridrow="5" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745ED01DE3A5E6" box="[159,401,459,481]" gridcol="0" gridrow="5" pageId="2" pageNumber="291">Interdorsal-anal length</th>
<td id="D96F5DC4F3374F9EFDC85ED01EF3A5E6" box="[547,641,459,481]" gridcol="1" gridrow="5" pageId="2" pageNumber="291">7.1</td>
<td id="D96F5DC4F3374F9EFD1A5ED01F79A5E6" box="[753,779,459,481]" gridcol="2" gridrow="5" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815ED01FBEA5E6" box="[874,972,459,481]" gridcol="3" gridrow="5" pageId="2" pageNumber="291">8.2</td>
<td id="D96F5DC4F3374F9EFBD95ED018BFA5E6" box="[1074,1229,459,481]" gridcol="4" gridrow="5" pageId="2" pageNumber="291">6.8 10.6</td>
<td id="D96F5DC4F3374F9EFAF55ED0194CA5E6" box="[1310,1342,459,481]" gridcol="5" gridrow="5" pageId="2" pageNumber="291">1.1</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745EEA194CA600" box="[159,1342,497,519]" gridrow="6" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745EEA1DE3A600" box="[159,401,497,519]" gridcol="0" gridrow="6" pageId="2" pageNumber="291">Caudal peduncle depth</th>
<td id="D96F5DC4F3374F9EFDC85EEA1EF3A600" box="[547,641,497,519]" gridcol="1" gridrow="6" pageId="2" pageNumber="291">13.7</td>
<td id="D96F5DC4F3374F9EFD1A5EEA1F79A600" box="[753,779,497,519]" gridcol="2" gridrow="6" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815EEA1FBEA600" box="[874,972,497,519]" gridcol="3" gridrow="6" pageId="2" pageNumber="291">12.7</td>
<td id="D96F5DC4F3374F9EFBD95EEA18BFA600" box="[1074,1229,497,519]" gridcol="4" gridrow="6" pageId="2" pageNumber="291">11.1 14.4</td>
<td id="D96F5DC4F3374F9EFAF55EEA194CA600" box="[1310,1342,497,519]" gridcol="5" gridrow="6" pageId="2" pageNumber="291">0.9</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745D0E194CA62C" box="[159,1342,533,555]" gridrow="7" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745D0E1DE3A62C" box="[159,401,533,555]" gridcol="0" gridrow="7" pageId="2" pageNumber="291">Head length (mm)</th>
<td id="D96F5DC4F3374F9EFDC85D0E1EF3A62C" box="[547,641,533,555]" gridcol="1" gridrow="7" pageId="2" pageNumber="291">10.4</td>
<td id="D96F5DC4F3374F9EFD1A5D0E1F79A62C" box="[753,779,533,555]" gridcol="2" gridrow="7" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815D0E1FBEA62C" box="[874,972,533,555]" gridcol="3" gridrow="7" pageId="2" pageNumber="291">9.0</td>
<td id="D96F5DC4F3374F9EFBD95D0E18BFA62C" box="[1074,1229,533,555]" gridcol="4" gridrow="7" pageId="2" pageNumber="291">8.1 10.4</td>
<td id="D96F5DC4F3374F9EFAF55D0E194CA62C" box="[1310,1342,533,555]" gridcol="5" gridrow="7" pageId="2" pageNumber="291">0.6</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745D21194CA657" box="[159,1342,570,592]" gridrow="8" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745D211DE3A657" box="[159,401,570,592]" gridcol="0" gridrow="8" pageId="2" pageNumber="291">Snout length</th>
<td id="D96F5DC4F3374F9EFDC85D211EF3A657" box="[547,641,570,592]" gridcol="1" gridrow="8" pageId="2" pageNumber="291">28.1</td>
<td id="D96F5DC4F3374F9EFD1A5D211F79A657" box="[753,779,570,592]" gridcol="2" gridrow="8" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815D211FBEA657" box="[874,972,570,592]" gridcol="3" gridrow="8" pageId="2" pageNumber="291">30.1</td>
<td id="D96F5DC4F3374F9EFBD95D2118BFA657" box="[1074,1229,570,592]" gridcol="4" gridrow="8" pageId="2" pageNumber="291">25.9 33.5</td>
<td id="D96F5DC4F3374F9EFAF55D21194CA657" box="[1310,1342,570,592]" gridcol="5" gridrow="8" pageId="2" pageNumber="291">2.0</td>
</tr>
<tr id="9ABE34B8F3374F9EFF745D44194CA672" box="[159,1342,607,629]" gridrow="9" pageId="2" pageNumber="291">
<th id="D96F5DC4F3374F9EFF745D441DE3A672" box="[159,401,607,629]" gridcol="0" gridrow="9" pageId="2" pageNumber="291">Eye diameter</th>
<td id="D96F5DC4F3374F9EFDC85D441EF3A672" box="[547,641,607,629]" gridcol="1" gridrow="9" pageId="2" pageNumber="291">23.7</td>
<td id="D96F5DC4F3374F9EFD1A5D441F79A672" box="[753,779,607,629]" gridcol="2" gridrow="9" pageId="2" pageNumber="291">23</td>
<td id="D96F5DC4F3374F9EFC815D441FBEA672" box="[874,972,607,629]" gridcol="3" gridrow="9" pageId="2" pageNumber="291">26.6</td>
<td id="D96F5DC4F3374F9EFBD95D4418BFA672" box="[1074,1229,607,629]" gridcol="4" gridrow="9" pageId="2" pageNumber="291">23.7 30.0</td>
<td id="D96F5DC4F3374F9EFAF55D44194CA672" box="[1310,1342,607,629]" gridcol="5" gridrow="9" pageId="2" pageNumber="291">1.8</td>
</tr>
</table>
</paragraph>
<caption id="70F16672F337B063FF7C5DAE1E52A6EB" pageId="2" pageNumber="291">
<paragraph id="243136FAF337B063FF7C5DAE1E52A6EB" blockId="2.[151,1436,692,748]" pageId="2" pageNumber="291">
<emphasis id="16FAEAE8F337B063FF7C5DAE1D72A6CB" bold="true" box="[151,256,693,716]" pageId="2" pageNumber="291">TABLE 2</emphasis>
. Meristic data for holotype and paratypes of
<taxonomicName id="E38E4D79F337B063FD315DAF1FEEA6CB" box="[730,924,692,716]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F337B063FD315DAF1FEEA6CB" box="[730,924,692,716]" italics="true" pageId="2" pageNumber="291">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F337B063FC495DAE1FAFA6CB" box="[930,989,693,716]" pageId="2" pageNumber="291" rank="species">
<emphasis id="16FAEAE8F337B063FC495DAE1FAFA6CB" bold="true" box="[930,989,693,716]" pageId="2" pageNumber="291">sp. n.</emphasis>
</taxonomicNameLabel>
Values in parentheses indicate number of specimens examined with that count.
</paragraph>
</caption>
<paragraph id="243136FAF337B063FF745C131E9EA719" blockId="2.[159,1354,776,1058]" box="[159,748,776,798]" pageId="2" pageNumber="291">
Counts
<typeStatus id="FB358858F337B063FE9D5C131DA6A719" box="[374,468,776,798]" pageId="2" pageNumber="291" type="holotype">Holotype</typeStatus>
<typeStatus id="FB358858F337B063FDD15C121EEEA719" box="[570,668,777,798]" pageId="2" pageNumber="291" type="paratype">Paratypes</typeStatus>
(N=23)
</paragraph>
<paragraph id="243136FAF337B063FF745C351FBAA7FA" blockId="2.[159,1354,776,1058]" pageId="2" pageNumber="291">Pre-dorsal scales 23 21 (4) 22 (4) 23 (6) 24 (6) 25 (3) Lateral scales 37 34 (4) 35 (7) 36 (7) 37 (5) Dorsal rays 7 6 (4) 7 (9) 8 (8) 9 (2) Caudal rays 19 15 (1) 16 (2) 17 (9) 18 (7) 19 (4) Anal rays 10 8 (2) 9 (12) 10 (8) 11 (1) Pelvic rays 5 4 (4) 5 (10) 6 (9)</paragraph>
<paragraph id="243136FAF337B063FF745B1718FCA025" blockId="2.[159,1354,776,1058]" box="[159,1166,1036,1058]" pageId="2" pageNumber="291">Pectoral rays 11 11 (10) 12 (9) 13 (2) 14 (2)</paragraph>
<caption id="70F16672F337B063FF7C58741DC0A3C5" httpUri="https://zenodo.org/record/233334/files/figure.png" pageId="2" pageNumber="291" targetBox="[155,1436,1127,1881]" targetPageId="2">
<paragraph id="243136FAF337B063FF7C58741DC0A3C5" blockId="2.[151,1436,1903,1986]" pageId="2" pageNumber="291">
<emphasis id="16FAEAE8F337B063FF7C58741D61A383" bold="true" box="[151,275,1903,1925]" pageId="2" pageNumber="291">FIGURE 2.</emphasis>
Outline drawings of the frontal scalation patterns of
<taxonomicName id="E38E4D79F337B063FCDC58741FF0A382" box="[823,898,1903,1925]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="genus">
<emphasis id="16FAEAE8F337B063FCDC58741FF0A382" box="[823,898,1903,1925]" italics="true" pageId="2" pageNumber="291">Rivulus</emphasis>
</taxonomicName>
from Cuba. A:
<taxonomicName id="E38E4D79F337B063FBC358741899A382" box="[1064,1259,1903,1925]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F337B063FBC358741899A382" box="[1064,1259,1903,1925]" italics="true" pageId="2" pageNumber="291">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F337B063FB1F58741941A383" box="[1268,1331,1903,1924]" pageId="2" pageNumber="291" rank="species">
<emphasis id="16FAEAE8F337B063FB1F58741941A383" bold="true" box="[1268,1331,1903,1924]" pageId="2" pageNumber="291">sp. n.</emphasis>
</taxonomicNameLabel>
with a
<emphasis id="16FAEAE8F337B063FA6C587419E1A382" box="[1415,1427,1903,1925]" italics="true" pageId="2" pageNumber="291">d</emphasis>
- type pattern and B:
<taxonomicName id="E38E4D79F337B063FE8758961E33A3A4" box="[364,577,1933,1955]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="cylindraceus">
<emphasis id="16FAEAE8F337B063FE8758961E33A3A4" box="[364,577,1933,1955]" italics="true" pageId="2" pageNumber="291">Rivulus cylindraceus</emphasis>
</taxonomicName>
with an
<emphasis id="16FAEAE8F337B063FD4F58951EDDA3A4" box="[676,687,1934,1955]" italics="true" pageId="2" pageNumber="291">e</emphasis>
-type pattern. The horizontal broken line indicates the pair of scales with their margins fully exposed.
</paragraph>
</caption>
<caption id="70F16672F336B062FF7C5C761D06A7A5" httpUri="https://zenodo.org/record/233335/files/figure.png" pageId="3" pageNumber="292" targetBox="[151,1398,151,840]" targetPageId="3">
<paragraph id="243136FAF336B062FF7C5C761D06A7A5" blockId="3.[151,1436,877,930]" pageId="3" pageNumber="292">
<emphasis id="16FAEAE8F336B062FF7C5C761D67A785" bold="true" box="[151,277,877,899]" pageId="3" pageNumber="292">FIGURE 3.</emphasis>
<taxonomicName id="E38E4D79F336B062FECB5C761DACA784" box="[288,478,877,899]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="292" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F336B062FECB5C761DACA784" box="[288,478,877,899]" italics="true" pageId="3" pageNumber="292">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F336B062FE025C751E5EA784" box="[489,556,878,899]" pageId="3" pageNumber="292" rank="species">
<emphasis id="16FAEAE8F336B062FE025C751E5EA784" bold="true" box="[489,556,878,899]" pageId="3" pageNumber="292">sp. n.</emphasis>
</taxonomicNameLabel>
, life coloration in both sexes in specimens kept in captivity for three months after collection in the field.
</paragraph>
</caption>
<caption id="70F16672F336B062FF7C58A91D06A3E0" httpUri="https://zenodo.org/record/233336/files/figure.png" pageId="3" pageNumber="292" targetBox="[328,1231,966,1865]" targetPageId="3">
<paragraph id="243136FAF336B062FF7C58A91D06A3E0" blockId="3.[151,1436,1970,2023]" pageId="3" pageNumber="292">
<emphasis id="16FAEAE8F336B062FF7C58A91D68A3C0" bold="true" box="[151,282,1970,1992]" pageId="3" pageNumber="292">FIGURE 4.</emphasis>
<taxonomicName id="E38E4D79F336B062FEC358A91D98A3CF" box="[296,490,1970,1992]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="292" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F336B062FEC358A91D98A3CF" box="[296,490,1970,1992]" italics="true" pageId="3" pageNumber="292">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F336B062FE1258A81E4FA3CF" box="[505,573,1971,1992]" pageId="3" pageNumber="292" rank="species">
<emphasis id="16FAEAE8F336B062FE1258A81E4FA3CF" bold="true" box="[505,573,1971,1992]" pageId="3" pageNumber="292">sp. n.</emphasis>
</taxonomicNameLabel>
, life coloration in males (above) and females (below) in specimens just after collection in the field.
</paragraph>
</caption>
<caption id="70F16672F331B065FF7C5CE71DFAA036" httpUri="https://zenodo.org/record/233337/files/figure.png" pageId="4" pageNumber="293" targetBox="[166,1417,176,990]" targetPageId="4">
<paragraph id="243136FAF331B065FF7C5CE71DFAA036" blockId="4.[151,1435,1020,1073]" pageId="4" pageNumber="293">
<emphasis id="16FAEAE8F331B065FF7C5CE71D63A015" bold="true" box="[151,273,1020,1042]" pageId="4" pageNumber="293">FIGURE 5.</emphasis>
<taxonomicName id="E38E4D79F331B065FEFC5CE71DA3A015" box="[279,465,1020,1042]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F331B065FEFC5CE71DA3A015" box="[279,465,1020,1042]" italics="true" pageId="4" pageNumber="293">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F331B065FE3C5CE61E66A015" box="[471,532,1021,1042]" pageId="4" pageNumber="293" rank="species">
<emphasis id="16FAEAE8F331B065FE3C5CE61E66A015" bold="true" box="[471,532,1021,1042]" pageId="4" pageNumber="293">sp. n.</emphasis>
</taxonomicNameLabel>
, paratype, female, CZACC-9.85, 37.2 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba.
</paragraph>
</caption>
<caption id="70F16672F331B065FF7C58AA1DFAA3E2" httpUri="https://zenodo.org/record/233338/files/figure.png" pageId="4" pageNumber="293" targetBox="[166,1417,1137,1930]" targetPageId="4">
<paragraph id="243136FAF331B065FF7C58AA1DFAA3E2" blockId="4.[151,1435,1969,2021]" pageId="4" pageNumber="293">
<emphasis id="16FAEAE8F331B065FF7C58AA1D61A3C1" bold="true" box="[151,275,1969,1991]" pageId="4" pageNumber="293">FIGURE 6.</emphasis>
<taxonomicName id="E38E4D79F331B065FEF058AA1DA4A3C0" box="[283,470,1969,1991]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F331B065FEF058AA1DA4A3C0" box="[283,470,1969,1991]" italics="true" pageId="4" pageNumber="293">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F331B065FE3558AA1E69A3C1" box="[478,539,1969,1990]" pageId="4" pageNumber="293" rank="species">
<emphasis id="16FAEAE8F331B065FE3558AA1E69A3C1" bold="true" box="[478,539,1969,1990]" pageId="4" pageNumber="293">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, male, CZACC-9.83, 37.3 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba.
</paragraph>
</caption>
<paragraph id="243136FAF330B064FF2C5F8C18A0A563" blockId="5.[151,1437,151,1148]" pageId="5" pageNumber="294">
<emphasis id="16FAEAE8F330B064FF2C5F8C1D2CA4B7" bold="true" box="[199,350,151,176]" pageId="5" pageNumber="294">Color in life.</emphasis>
Males (
<figureCitation id="BCB52A7FF330B064FE515F8C1D8DA4B7" box="[442,511,151,176]" captionStart="FIGURE 4" captionStartId="3.[151,249,1970,1992]" captionTargetBox="[328,1231,966,1865]" captionTargetId="figure@3.[265,1287,964,1935]" captionTargetPageId="3" captionText="FIGURE 4. Rivulus berovidesi sp. n., life coloration in males (above) and females (below) in specimens just after collection in the field." httpUri="https://zenodo.org/record/233336/files/figure.png" pageId="5" pageNumber="294">Fig. 4</figureCitation>
): Body coloration is greenish or olive-green dorsally. There is a wide dark lateral band extended from the posterior margin of the eye to the base of the caudal fin. Ventral ground coloration is yellowish with orange spots reaching the anal fin base. An iridescent blue spot is present behind the margin of opercle in both sexes. Males have an iridescent olive- green spot on the opercle too. Dorsal fin is greenish, caudal fin is slightly greenish on its base and more transparent to the edges. Anal fin is spotted in orange in their base and yellow or yellow- green to the ends. Pelvic fins are yellow and pectoral fins are transparent.
</paragraph>
<paragraph id="243136FAF330B064FF2C5E7418BBA610" blockId="5.[151,1437,151,1148]" pageId="5" pageNumber="294">
Females (
<figureCitation id="BCB52A7FF330B064FEDC5E741D0DA58F" box="[311,383,367,392]" captionStart="FIGURE 4" captionStartId="3.[151,249,1970,1992]" captionTargetBox="[328,1231,966,1865]" captionTargetId="figure@3.[265,1287,964,1935]" captionTargetPageId="3" captionText="FIGURE 4. Rivulus berovidesi sp. n., life coloration in males (above) and females (below) in specimens just after collection in the field." httpUri="https://zenodo.org/record/233336/files/figure.png" pageId="5" pageNumber="294">Fig. 4</figureCitation>
): Body coloration less striking when compared to males. General body coloration brown, darker dorsally than ventrally. Ventral coloration is paler with some small yellowish spots. There is a narrow dark lateral band extended from the posterior margin of the eye to the base of the caudal. Diffuse ocellated caudal spot present which can extends from the posterior base of dorsal fin to the superior margin of caudal fin and sometimes along the caudal fin base too (
<figureCitation id="BCB52A7FF330B064FD8C5EE41ED8A61F" box="[615,682,511,536]" captionStart="FIGURE 4" captionStartId="3.[151,249,1970,1992]" captionTargetBox="[328,1231,966,1865]" captionTargetId="figure@3.[265,1287,964,1935]" captionTargetPageId="3" captionText="FIGURE 4. Rivulus berovidesi sp. n., life coloration in males (above) and females (below) in specimens just after collection in the field." httpUri="https://zenodo.org/record/233336/files/figure.png" pageId="5" pageNumber="294">Fig. 4</figureCitation>
and
<figureCitation id="BCB52A7FF330B064FD035EE41E8AA61F" box="[744,760,511,536]" captionStart="FIGURE 5" captionStartId="4.[151,249,1020,1042]" captionTargetBox="[166,1417,176,990]" captionTargetId="figure@4.[166,1418,99,993]" captionTargetPageId="4" captionText="FIGURE 5. Rivulus berovidesi sp. n., paratype, female, CZACC- 9.85, 37.2 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba." httpUri="https://zenodo.org/record/233337/files/figure.png" pageId="5" pageNumber="294">5</figureCitation>
). All fins slightly greenish or transparent.
</paragraph>
</subSubSection>
<subSubSection id="6C946571F330B064FF2C5D3F1D7DA6F7" pageId="5" pageNumber="294" type="materials_examined">
<paragraph id="243136FAF330B064FF2C5D3F1D7DA6F7" blockId="5.[151,1437,151,1148]" pageId="5" pageNumber="294">
<emphasis id="16FAEAE8F330B064FF2C5D3F1E44A63A" bold="true" box="[199,566,548,573]" pageId="5" pageNumber="294">Color in preserved specimens.</emphasis>
Preserved specimens with the typical dark lateral band from the posterior margin of the eye to the base of caudal fin in both sexes, although slightly paler than living specimens. Ventral region of the body with a yellowish homogeneous coloration while dorsal region is brownish-gray. Males with a reticulated pattern on scales above lateral dark band (
<figureCitation id="BCB52A7FF330B064FCF45D941F14A6AF" box="[799,870,655,680]" captionStart="FIGURE 6" captionStartId="4.[151,249,1969,1991]" captionTargetBox="[166,1417,1137,1930]" captionTargetId="figure@4.[166,1418,1136,1932]" captionTargetPageId="4" captionText="FIGURE 6. Rivulus berovidesi sp. n., holotype, male, CZACC- 9.83, 37.3 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba." httpUri="https://zenodo.org/record/233338/files/figure.png" pageId="5" pageNumber="294">Fig. 6</figureCitation>
) while females with the diffuse ocellated caudal spot extended from the end of dorsal fin to the superior margin of caudal fin (
<figureCitation id="BCB52A7FF330B064FBD25DAF180DA6CA" box="[1081,1151,692,717]" captionStart="FIGURE 5" captionStartId="4.[151,249,1020,1042]" captionTargetBox="[166,1417,176,990]" captionTargetId="figure@4.[166,1418,99,993]" captionTargetPageId="4" captionText="FIGURE 5. Rivulus berovidesi sp. n., paratype, female, CZACC- 9.85, 37.2 mm SL, small stream in Sierra de Cajalbana, Pinar del Rio province, Cuba." httpUri="https://zenodo.org/record/233337/files/figure.png" pageId="5" pageNumber="294">Fig. 5</figureCitation>
). All fins with a whitish coloration.
</paragraph>
</subSubSection>
<subSubSection id="6C946571F330B064FF2C5DE71893A75A" pageId="5" pageNumber="294" type="etymology">
<paragraph id="243136FAF330B064FF2C5DE71893A75A" blockId="5.[151,1437,151,1148]" pageId="5" pageNumber="294">
<emphasis id="16FAEAE8F330B064FF2C5DE71D3EA712" bold="true" box="[199,332,764,789]" pageId="5" pageNumber="294">Etymology.</emphasis>
The specific name of this new species is given in honor to the Professor Vicente Berovides Alvarez, professor at the Faculty of Biology, Universidad de la Habana,
<collectingCountry id="5C99766AF330B064FBE95C04184FA73F" box="[1026,1085,799,824]" name="Cuba" pageId="5" pageNumber="294">Cuba</collectingCountry>
in recognition of his life-long dedication and contribution to train several generations of new researchers in biological sciences.
</paragraph>
</subSubSection>
<subSubSection id="6C946571F330B064FF2C5C7C1DE1A07B" pageId="5" pageNumber="294" type="distribution">
<paragraph id="243136FAF330B064FF2C5C7C1DE1A07B" blockId="5.[151,1437,151,1148]" pageId="5" pageNumber="294">
<emphasis id="16FAEAE8F330B064FF2C5C7C1D89A787" bold="true" box="[199,507,871,896]" pageId="5" pageNumber="294">Distribution and habitat.</emphasis>
<taxonomicName id="E38E4D79F330B064FDE35C731EA8A787" box="[520,730,872,896]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F330B064FDE35C731EA8A787" box="[520,730,872,896]" italics="true" pageId="5" pageNumber="294">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F330B064FD035C7C1F5CA787" box="[744,814,871,896]" pageId="5" pageNumber="294" rank="species">
<emphasis id="16FAEAE8F330B064FD035C7C1F5CA787" bold="true" box="[744,814,871,896]" pageId="5" pageNumber="294">sp. n.</emphasis>
</taxonomicNameLabel>
is known only from northwest mountain system in Pinar del Rio province. Particularly, in Sierra de Cajalbana (
<typeStatus id="FB358858F330B064FCB55C971FFDA7A3" box="[862,911,908,932]" pageId="5" pageNumber="294">type</typeStatus>
locality) and Rio Camarones. Both localities are situated in northwestern
<collectingCountry id="5C99766AF330B064FE325CB41E66A7CF" box="[473,532,943,968]" name="Cuba" pageId="5" pageNumber="294">Cuba</collectingCountry>
(
<figureCitation id="BCB52A7FF330B064FDCD5CB41E1BA7CF" box="[550,617,943,968]" captionStart="FIGURE 1" captionStartId="1.[151,249,883,905]" captionTargetBox="[153,1433,375,860]" captionTargetId="figure@1.[153,1433,375,860]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Rivulus species from Cuba. Red spots represent distribution of Rivulus cylindraceus and red triangles represents distribution of Rivulus berovidesi sp. n., 1 Sierra de Cajalbana (type locality); 2 Rio Camarones." httpUri="https://zenodo.org/record/233333/files/figure.png" pageId="5" pageNumber="294">Fig. 1</figureCitation>
). This species inhabits small mountain streams with a depth of
<quantity id="E3769B1FF330B064FABD5CAB1CCBA7EB" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="5" pageNumber="294" unit="cm" value="50.0" valueMax="70.0" valueMin="30.0">30–70 cm</quantity>
approximately and bottom substrate basically composed by stones, sand and dead leaves (
<figureCitation id="BCB52A7FF330B064FB375CCF1957A7EB" box="[1244,1317,980,1005]" captionStart="FIGURE 7" captionStartId="5.[151,249,1976,1998]" captionTargetBox="[313,1273,1234,1954]" captionTargetId="figure@5.[313,1273,1234,1954]" captionTargetPageId="5" captionText="FIGURE 7. Type locality of Rivulus berovidesi sp. n. Small stream at Sierra de Cajalbana, Pinar del Rio province, Cuba" httpUri="https://zenodo.org/record/233339/files/figure.png" pageId="5" pageNumber="294">Fig. 7</figureCitation>
). There is not aquatic vegetation in the area. The streams are clear with
<collectingCountry id="5C99766AF330B064FC735CE31FC8A008" box="[920,954,1016,1039]" name="Philippines" pageId="5" pageNumber="294">pH</collectingCountry>
7.5 and water temperature 24°C. Other freshwater fish collected together with
<taxonomicName id="E38E4D79F330B064FD615B071F10A033" box="[650,866,1052,1076]" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="berovidesi" status="sp. nov.">
<emphasis id="16FAEAE8F330B064FD615B071F10A033" box="[650,866,1052,1076]" italics="true" pageId="5" pageNumber="294">Rivulus berovidesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC95793F330B064FC9F5B071FB0A032" box="[884,962,1052,1077]" pageId="5" pageNumber="294" rank="species">
<emphasis id="16FAEAE8F330B064FC9F5B071FB0A032" bold="true" box="[884,962,1052,1077]" pageId="5" pageNumber="294">sp. n.</emphasis>
</taxonomicNameLabel>
were:
<taxonomicName id="E38E4D79F330B064FBCF5B07197AA033" box="[1060,1288,1052,1076]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="punctata">
<emphasis id="16FAEAE8F330B064FBCF5B07197AA033" box="[1060,1288,1052,1076]" italics="true" pageId="5" pageNumber="294">Gambusia punctata</emphasis>
</taxonomicName>
,
<taxonomicName id="E38E4D79F330B064FACB5B071D7FA050" class="Actinopterygii" family="Poeciliidae" genus="Girardinus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="uninotatus">
<emphasis id="16FAEAE8F330B064FACB5B071D7FA050" italics="true" pageId="5" pageNumber="294">Girardinus uninotatus</emphasis>
</taxonomicName>
,
<taxonomicName id="E38E4D79F330B064FECF5B5B1D8BA05F" box="[292,505,1088,1112]" class="Actinopterygii" family="Poeciliidae" genus="Girardinus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="creolus">
<emphasis id="16FAEAE8F330B064FECF5B5B1D8BA05F" box="[292,505,1088,1112]" italics="true" pageId="5" pageNumber="294">Girardinus creolus</emphasis>
</taxonomicName>
,
<taxonomicName id="E38E4D79F330B064FDE15B5B1F59A05F" box="[522,811,1088,1112]" class="Actinopterygii" family="Poeciliidae" genus="Girardinus" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="microdactylus">
<emphasis id="16FAEAE8F330B064FDE15B5B1F59A05F" box="[522,811,1088,1112]" italics="true" pageId="5" pageNumber="294">Girardinus microdactylus</emphasis>
</taxonomicName>
,
<taxonomicName id="E38E4D79F330B064FCD75B5B183AA05F" box="[828,1096,1088,1112]" class="Actinopterygii" family="Cichlidae" genus="Nandopsis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="tetracanthus">
<emphasis id="16FAEAE8F330B064FCD75B5B183AA05F" box="[828,1096,1088,1112]" italics="true" pageId="5" pageNumber="294">Nandopsis tetracanthus</emphasis>
</taxonomicName>
,
<taxonomicName id="E38E4D79F330B064FBB35B5B191AA05F" box="[1112,1384,1088,1112]" class="Actinopterygii" family="Mugilidae" genus="Agonostomus" kingdom="Animalia" order="Mugiliformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="monticola">
<emphasis id="16FAEAE8F330B064FBB35B5B191AA05F" box="[1112,1384,1088,1112]" italics="true" pageId="5" pageNumber="294">Agonostomus monticola</emphasis>
</taxonomicName>
and
<taxonomicName id="E38E4D79F330B064FF7C5B7F1DFEA07B" box="[151,396,1124,1148]" class="Actinopterygii" family="Eleotridae" genus="Gobiomorus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="dormitor">
<emphasis id="16FAEAE8F330B064FF7C5B7F1DFEA07B" box="[151,396,1124,1148]" italics="true" pageId="5" pageNumber="294">Gobiomorus dormitor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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