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<document ID-DOI="http://doi.org/10.5281/zenodo.4335244" ID-GBIF-Dataset="6790c5bd-2f80-4161-bb6f-2330f95620ed" ID-GBIF-Taxon="164845040" ID-Zenodo-Dep="4335244" approvalRequired="13" approvalRequired_for_document="1" approvalRequired_for_textStreams="10" approvalRequired_for_treatments="2" checkinTime="1593596629447" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2020" docId="75207B3EB00DFFF7A23DF983FF6F113B" docLanguage="en" docName="arachnology.18.4.368-386.pdf" docOrigin="Arachnology 18 (4)" docSource="http://dx.doi.org/10.13156/arac.2020.18.4.368" docStyle="DocumentStyle{}" docTitle="Gorbiscape agelenoides Zamani &amp; Marusik 2020, comb. n." docType="treatment" docVersion="4" lastPageId="15" lastPageNumber="383" masterDocId="89190346B003FFF8A101FFBEFF801909" masterDocTitle="A review of Agelenini (Araneae: Agelenidae: Ageleninae) of Iran and Tajikistan, with descriptions of four new genera" masterLastPageNumber="386" masterPageNumber="368" pageId="14" pageNumber="382" updateTime="1632024453243" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Agelenini (Araneae: Agelenidae: Ageleninae) of Iran and Tajikistan, with descriptions of four new genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arachnology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>18</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>368</mods:start>
<mods:end>386</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.13156/arac.2020.18.4.368</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.13156/arac.2020.18.4.368</mods:identifier>
<mods:identifier type="GBIF-Dataset">6790c5bd-2f80-4161-bb6f-2330f95620ed</mods:identifier>
<mods:identifier type="Zenodo-Dep">3925217</mods:identifier>
<mods:identifier type="ZooBank">CC0827D2-6390-43DC-A02D-D538246814B9</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4335244" ID-GBIF-Taxon="164845040" ID-Zenodo-Dep="4335244" LSID="urn:lsid:plazi:treatment:75207B3EB00DFFF7A23DF983FF6F113B" httpUri="http://treatment.plazi.org/id/75207B3EB00DFFF7A23DF983FF6F113B" lastPageId="15" lastPageNumber="383" pageId="14" pageNumber="382">
<subSubSection box="[828,1478,1596,1621]" pageId="14" pageNumber="382" type="nomenclature">
<paragraph blockId="14.[828,1478,1596,1654]" box="[828,1478,1596,1621]" pageId="14" pageNumber="382">
<taxonomicName authority="(Walckenaer, 1841)" authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" box="[828,1341,1596,1620]" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis bold="true" box="[828,1341,1596,1620]" pageId="14" pageNumber="382">
<emphasis bold="true" box="[828,1097,1597,1620]" italics="true" pageId="14" pageNumber="382">Gorbiscape agelenoides</emphasis>
(
<bibRefCitation author="WALCKENAER, C. A." box="[1120,1333,1596,1620]" journalOrPublisher="Paris: Roret" pageId="14" pageNumber="382" pagination="1 - 549" refId="ref12762" refString="WALCKENAER, C. A. 1841: Histoire naturelle des Insectes. AptXres, tome B. Paris: Roret: 1 - 549." title="Histoire naturelle des Insectes. AptXres, tome B" type="book chapter" year="1841">Walckenaer, 1841</bibRefCitation>
)
</emphasis>
</taxonomicName>
<heading bold="true" box="[1372,1478,1597,1621]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="382" reason="2">
<emphasis bold="true" box="[1372,1478,1597,1621]" pageId="14" pageNumber="382">
<taxonomicNameLabel box="[1372,1478,1597,1621]" pageId="14" pageNumber="382">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[828,924,1632,1654]" pageId="14" pageNumber="382" type="description">
<paragraph blockId="14.[828,1478,1596,1654]" box="[828,924,1632,1654]" pageId="14" pageNumber="382">
(
<figureCitation box="[832,918,1632,1654]" captionStart="Fig" captionStartId="5.[125,156,721,740]" captionTargetBox="[148,733,152,698]" captionTargetId="figure@5.[125,760,152,698]" captionTargetPageId="5" captionText="Fig. 4: Bulb, ventral view. A, B Agelescape cf. livida, from Israel;C Asiascape parthica sp. n.; D Gorbiscape gorbachevi sp. n.; E G. agelenoides. A after Levy (1996), E after de Blauwe (1980), both with modification. Scale bar = 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.3925225" httpUri="https://zenodo.org/record/3925225/files/figure.png" pageId="14" pageNumber="382">Fig. 4E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="382" type="reference_group">
<paragraph blockId="14.[828,1463,1692,1866]" box="[828,1322,1692,1714]" pageId="14" pageNumber="382">
<treatmentCitationGroup box="[828,1322,1692,1714]" pageId="14" pageNumber="382">
<treatmentCitation author="WALCKENAER, C. A." box="[828,1277,1692,1714]" page="454" pageId="14" pageNumber="382" year="1841">
<taxonomicName authority="Walckenaer 1841: 454" authorityName="Walckenaer" authorityPageNumber="454" authorityYear="1841" box="[828,1277,1692,1714]" class="Arachnida" family="Pisauridae" genus="Dolomedes" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis box="[828,1057,1692,1713]" italics="true" pageId="14" pageNumber="382">Dolomedes agelenoides</emphasis>
<bibRefCitation author="WALCKENAER, C. A." box="[1062,1277,1695,1714]" journalOrPublisher="Paris: Roret" pageId="14" pageNumber="382" pagination="1 - 549" refId="ref12762" refString="WALCKENAER, C. A. 1841: Histoire naturelle des Insectes. AptXres, tome B. Paris: Roret: 1 - 549." title="Histoire naturelle des Insectes. AptXres, tome B" type="book chapter" year="1841">Walckenaer 1841: 454</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(D).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="14.[828,1463,1692,1866]" pageId="14" pageNumber="382">
<treatmentCitationGroup pageId="14" pageNumber="382">
<treatmentCitation author="SIMON, E." box="[828,1229,1723,1744]" page="1012" pageId="14" pageNumber="382" year="1937">
<taxonomicName authority=": Simon 1937: 1012" authorityName="Simon" authorityPageNumber="1012" authorityYear="1937" box="[828,1229,1723,1744]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis box="[828,1030,1723,1744]" italics="true" pageId="14" pageNumber="382">Agelena agelenoides</emphasis>
:
<bibRefCitation author="SIMON, E." box="[1045,1229,1725,1744]" journalOrPublisher="Paris: Roret" pageId="14" pageNumber="382" pagination="979 - 1298" refId="ref12722" refString="SIMON, E. 1937: Les arachnides de France. Synopsis generale et catalogue des espXces franYaises de lIordre des Araneae. Tome VI. 5 e et derniere partie. Paris: Roret: 979 - 1298." title="Les arachnides de France. Synopsis generale et catalogue des espXces franYaises de lIordre des Araneae. Tome VI. 5 e et derniere partie" type="book chapter" year="1937">Simon 1937: 1012</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 1042, figs.
<date box="[1355,1463,1725,1744]" pageId="14" pageNumber="382" value="1564" valueMax="1565">15641565</date>
(
<specimenCount box="[895,917,1756,1775]" pageId="14" pageNumber="382"></specimenCount>
<specimenCount box="[917,933,1756,1775]" pageId="14" pageNumber="382"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="14.[828,1463,1692,1866]" box="[828,1379,1784,1805]" pageId="14" pageNumber="382">
<treatmentCitationGroup box="[828,1379,1784,1805]" pageId="14" pageNumber="382">
<treatmentCitation author="DE BLAUWE, R." box="[828,1220,1784,1805]" page="5" pageId="14" pageNumber="382" year="1980">
<taxonomicName authority=": de Blauwe 1980: 5" authorityName=": de Blauwe" authorityPageNumber="5" authorityYear="1980" box="[828,1220,1784,1805]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis box="[828,1027,1784,1805]" italics="true" pageId="14" pageNumber="382">Agelena agelenoides</emphasis>
:
<bibRefCitation author="DE BLAUWE, R." box="[1040,1220,1786,1805]" journalOrPublisher="Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique" pageId="14" pageNumber="382" pagination="1 - 28" part="52" refId="ref12131" refString="DE BLAUWE, R. 1980: Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie). Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique 52 (11): 1 - 28." title="Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie)" type="journal article" year="1980">de Blauwe 1980: 5</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 37 (
<specimenCount box="[1328,1350,1786,1805]" pageId="14" pageNumber="382"></specimenCount>
<specimenCount box="[1350,1366,1786,1805]" pageId="14" pageNumber="382"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="14.[828,1463,1692,1866]" pageId="14" pageNumber="382">
For the complete list of taxonomic entries see
<bibRefCitation author="WORLD SPIDER CATALOG" pageId="14" pageNumber="382" publicationUrl="http://wsc.nmbe.ch" refId="ref12788" refString="WORLD SPIDER CATALOG 2020: World spider catalog, version BA. C. Bern: Natural History Museum, online at http: // wsc. nmbe. ch" title="World spider catalog, version BA. C. Bern: Natural History Museum, online at" type="url" year="2020">World Spider Catalog (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="383" pageId="14" pageNumber="382" type="discussion">
<paragraph blockId="14.[828,1463,1907,2100]" lastBlockId="15.[125,762,1561,2098]" lastPageId="15" lastPageNumber="383" pageId="14" pageNumber="382">
<emphasis box="[858,951,1907,1931]" italics="true" pageId="14" pageNumber="382">Remarks</emphasis>
: This species is tentatively placed in this genus because of the similarity of the male palp with that of the generotype. This species is clearly different from
<taxonomicName authorityName=": Kovblyuk &amp; Kastrygina" authorityYear="2011" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="species" species="labyrinthica">
<emphasis italics="true" pageId="14" pageNumber="382">Agelena labyrinthica</emphasis>
</taxonomicName>
, the
<typeStatus box="[1013,1059,2012,2032]" pageId="14" pageNumber="382">type</typeStatus>
of its genus, and therefore cannot be placed in
<taxonomicName authorityName=": Lehtinen" authorityYear="1967" box="[943,1030,2042,2066]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis box="[943,1030,2042,2066]" italics="true" pageId="14" pageNumber="382">Agelena</emphasis>
</taxonomicName>
. Perhaps, it would belong to its own genus, but as we are reluctant to erect monotypic genera, for now, we transfer it to
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[352,476,1561,1585]" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[352,476,1561,1585]" italics="true" pageId="15" pageNumber="383">Gorbiscape</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[482,553,1565,1585]" pageId="15" pageNumber="383" rank="genus">gen. n.</taxonomicNameLabel>
and leave the question open for future phylogenetic studies.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925245" ID-Zenodo-Dep="3925245" httpUri="https://zenodo.org/record/3925245/files/figure.png" pageId="15" pageNumber="383" startId="15.[125,156,1431,1450]" targetBox="[126,1461,151,1406]" targetPageId="15">
<paragraph blockId="15.[125,1463,1427,1500]" pageId="15" pageNumber="383">
Fig. 14: Epigynes.
<emphasis bold="true" box="[291,310,1427,1451]" pageId="15" pageNumber="383">A</emphasis>
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[317,519,1431,1450]" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="gorbachevi" status="sp. n.">
<emphasis box="[317,519,1431,1450]" italics="true" pageId="15" pageNumber="383">Gorbiscape gorbachevi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[526,575,1432,1451]" pageId="15" pageNumber="383" rank="species">sp. n.</taxonomicNameLabel>
, intact, ventral view;
<emphasis bold="true" box="[765,783,1428,1452]" pageId="15" pageNumber="383">B</emphasis>
same, macerated, dorsal view;
<emphasis bold="true" box="[1059,1112,1427,1452]" pageId="15" pageNumber="383">C, D</emphasis>
same, posterior views;
<emphasis bold="true" box="[1320,1369,1428,1452]" pageId="15" pageNumber="383">E, F</emphasis>
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Asiascape" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="parthica" status="sp. n.">
<emphasis italics="true" pageId="15" pageNumber="383">Asiascape parthica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[277,325,1456,1475]" pageId="15" pageNumber="383" rank="species">sp. n.</taxonomicNameLabel>
, intact, ventral views;
<emphasis bold="true" box="[520,541,1451,1475]" pageId="15" pageNumber="383">G</emphasis>
same, macerated, dorsal view;
<emphasis bold="true" box="[811,832,1452,1476]" pageId="15" pageNumber="383">H</emphasis>
same, posterior view;
<emphasis bold="true" box="[1027,1037,1452,1476]" pageId="15" pageNumber="383">I</emphasis>
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[1043,1196,1455,1474]" class="Arachnida" family="Agelenidae" genus="Persilena" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="sengleti" status="sp. n.">
<emphasis box="[1043,1196,1455,1474]" italics="true" pageId="15" pageNumber="383">Persilena sengleti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1203,1251,1456,1475]" pageId="15" pageNumber="383" rank="species">sp. n.</taxonomicNameLabel>
, posterior view;
<emphasis bold="true" box="[1394,1407,1452,1476]" pageId="15" pageNumber="383">D</emphasis>
same, intact, ventral view;
<emphasis bold="true" box="[375,396,1476,1500]" pageId="15" pageNumber="383">K</emphasis>
same, antero-ventral view;
<emphasis bold="true" box="[633,651,1476,1500]" pageId="15" pageNumber="383">L</emphasis>
same, dorsal view. Scale bars = 0.2 mm. Abbreviation: Cd = copulatory duct.
</paragraph>
</caption>
<subSubSection pageId="15" pageNumber="383" type="diagnosis">
<paragraph blockId="15.[125,762,1561,2098]" pageId="15" pageNumber="383">
<emphasis box="[155,263,1631,1654]" italics="true" pageId="15" pageNumber="383">Diagnosis</emphasis>
: The male of
<taxonomicName authorityName=": de Blauwe" authorityYear="1980" box="[419,581,1629,1653]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis box="[419,581,1629,1653]" italics="true" pageId="15" pageNumber="383">G. agelenoides</emphasis>
</taxonomicName>
differs from the
<typeStatus box="[125,171,1667,1687]" pageId="15" pageNumber="383">type</typeStatus>
species by the broader base of the conductor (cf.
<figureCitation captionStart="Fig" captionStartId="5.[125,156,721,740]" captionTargetBox="[148,733,152,698]" captionTargetId="figure@5.[125,760,152,698]" captionTargetPageId="5" captionText="Fig. 4: Bulb, ventral view. A, B Agelescape cf. livida, from Israel;C Asiascape parthica sp. n.; D Gorbiscape gorbachevi sp. n.; E G. agelenoides. A after Levy (1996), E after de Blauwe (1980), both with modification. Scale bar = 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.3925225" httpUri="https://zenodo.org/record/3925225/files/figure.png" pageId="15" pageNumber="383">Figs. 4E and 4D</figureCitation>
), the rounded tip of the patellar apophysis (
<taxonomicName pageId="15" pageNumber="383" rank="variety" variety="sharply">
<emphasis box="[743,759,1699,1722]" italics="true" pageId="15" pageNumber="383">v.</emphasis>
sharply
</taxonomicName>
pointed), and not subdivided tip of the embolus (
<taxonomicName pageId="15" pageNumber="383" rank="variety" variety="subdivided">
<emphasis box="[743,759,1733,1756]" italics="true" pageId="15" pageNumber="383">v.</emphasis>
subdivided
</taxonomicName>
). Females of two congeners differ by the absence (
<taxonomicName authorityName=": de Blauwe" authorityYear="1980" box="[226,388,1800,1824]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis box="[226,388,1800,1824]" italics="true" pageId="15" pageNumber="383">G. agelenoides</emphasis>
</taxonomicName>
)
<taxonomicName box="[405,523,1802,1825]" pageId="15" pageNumber="383" rank="variety" variety="presence">
<emphasis box="[405,421,1802,1825]" italics="true" pageId="15" pageNumber="383">v.</emphasis>
presence
</taxonomicName>
of the scape (
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="species" species="gorbachevi" status="sp. n.">
<emphasis italics="true" pageId="15" pageNumber="383">G. gorbachevi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[219,285,1837,1858]" pageId="15" pageNumber="383" rank="species">sp. n.</taxonomicNameLabel>
). This species is also similar to
<taxonomicName authorityName=": Lehtinen" authorityYear="1967" box="[665,752,1834,1858]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis box="[665,752,1834,1858]" italics="true" pageId="15" pageNumber="383">Agelena</emphasis>
</taxonomicName>
<emphasis box="[125,248,1870,1893]" italics="true" pageId="15" pageNumber="383">canariensis</emphasis>
by the shape of the embolus and very broad conductor but differs by having a distinct and wide base of the conductor, which is lacking in the other species.
</paragraph>
</subSubSection>
<subSubSection box="[155,538,1973,1996]" pageId="15" pageNumber="383" type="description">
<paragraph blockId="15.[125,762,1561,2098]" box="[155,538,1973,1996]" pageId="15" pageNumber="383">
<emphasis box="[155,281,1973,1996]" italics="true" pageId="15" pageNumber="383">Description</emphasis>
: See
<bibRefCitation author="DE BLAUWE, R." box="[340,534,1974,1995]" journalOrPublisher="Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique" pageId="15" pageNumber="383" pagination="1 - 28" part="52" refId="ref12131" refString="DE BLAUWE, R. 1980: Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie). Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique 52 (11): 1 - 28." title="Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie)" type="journal article" year="1980">de Blauwe (1980)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="383" type="distribution">
<paragraph blockId="15.[125,762,1561,2098]" pageId="15" pageNumber="383">
<emphasis box="[155,285,2006,2030]" italics="true" pageId="15" pageNumber="383">Distribution</emphasis>
: This species is known from the western Mediterranean, from the Iberian Peninsula to
<collectingCountry box="[605,654,2043,2064]" name="Italy" pageId="15" pageNumber="383">Italy</collectingCountry>
(
<bibRefCitation author="HELSDINGEN, P. VAN" pageId="15" pageNumber="383" publicationUrl="http://www.faunaeur.org" refId="ref12270" refString="HELSDINGEN, P. VAN 2020: Fauna Europaea: Araneae. Fauna Europaea, online at: http: // www. faunaeur. org" title="Fauna Europaea: Araneae. Fauna Europaea, online at" type="url" year="2020">Helsdingen 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>