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<mods:title id="A29C1DB5FCA4663793FBB53D22A1FEC2">&lt;strong&gt; Phylogeny and revision of &lt;em&gt; Messatoporus &lt;/ em&gt; Cushman (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of sixty five new species &lt;/ strong&gt;</mods:title>
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<heading id="E5778010FF092ED0B8BFFA41FC80B6C1" bold="true" box="[151,799,1470,1496]" fontSize="11" level="2" pageId="151" pageNumber="152" reason="4">
<taxonomicName id="79804CFFFF092ED0B8BFFA41FC80B6C1" ID-CoL="73BGH" authority="Kasparyan &amp; Ruiz 2005" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[151,799,1470,1496]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF092ED0B8BFFA41FC80B6C1" bold="true" box="[151,799,1470,1496]" pageId="151" pageNumber="152">
<emphasis id="8CF4EB6EFF092ED0B8BFFA41FE75B6C1" bold="true" box="[151,490,1470,1496]" italics="true" pageId="151" pageNumber="152">Messatoporus mesosternalis</emphasis>
<bibRefCitation id="DA114A8DFF092ED0B9DAFA41FC80B6C1" author="Kasparyan, D. R. &amp; Ruiz, E. C." box="[498,799,1470,1496]" pageId="151" pageNumber="152" pagination="105 - 122" refId="ref210720" refString="Kasparyan, D. R. &amp; Ruiz, E. C. (2005) A review of the North American species of Messatoporus Cushman (Hymenoptera: Ichneumonidae, Cryptini). Zoosystematica Rossica, 14 (1), 105 - 122." type="journal article" year="2005">Kasparyan &amp; Ruíz 2005</bibRefCitation>
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<paragraph id="BE3F377CFF092ED0B8BFFA1EFEE4B6E0" blockId="151.[151,799,1470,1529]" box="[151,379,1505,1529]" pageId="151" pageNumber="152">
(
<figureCitation id="26BB2BF9FF092ED0B8B7FA1EFE9BB6E0" box="[159,260,1505,1529]" captionStart="FIGURES 101–105" captionStartId="247.[151,258,1976,1997]" captionTargetBox="[151,1436,181,1969]" captionTargetId="figure-52@247.[151,1436,181,1969]" captionTargetPageId="247" captionText="FIGURES 101–105. Lateral view. 101, M. basiflavus*. 102, M. occidentalis*. 103, M. mesosternalis. 104, M. orientalis. 105, M. semiaurantigaster." figureDoi="http://doi.org/10.5281/zenodo.5262169" httpUri="https://zenodo.org/record/5262169/files/figure.png" pageId="151" pageNumber="152">Figs 103</figureCitation>
,
<figureCitation id="26BB2BF9FF092ED0B927FA1EFEA4B6E0" box="[271,315,1505,1529]" captionStart="FIGURES 236–241" captionStartId="268.[151,258,1967,1988]" captionTargetBox="[151,1436,193,1946]" captionTargetId="figure-56@268.[151,1436,193,1946]" captionTargetPageId="268" captionText="FIGURES 236–241. Hind leg. 236, M. mesonotator. 237, M. arcuatus. 238, M. keraiopetalus. 239, M. mesosternalis. 240, M. albiaterartus*. 241, M. versicolor*." figureDoi="http://doi.org/10.5281/zenodo.5262211" httpUri="https://zenodo.org/record/5262211/files/figure.png" pageId="151" pageNumber="152">239</figureCitation>
,
<figureCitation id="26BB2BF9FF092ED0B96FFA1EFEEDB6E0" box="[327,370,1505,1529]" captionStart="FIGURES 345–352" captionStartId="280.[151,258,1969,1990]" captionTargetBox="[151,1398,195,1954]" captionTargetId="figure-67@280.[151,1398,193,1954]" captionTargetPageId="280" captionText="FIGURES 345–352. Distribution maps for species of Messatoporus. 345, M. mesosternalis. 346, M. nigriangulatus. 347, M. nigriscapus. 348, M. nigriscutus. 349, M. occidentalis. 350, M. opacus. 351, M. orientalis. 352, M. paeneater." figureDoi="http://doi.org/10.5281/zenodo.5262235" httpUri="https://zenodo.org/record/5262235/files/figure.png" pageId="151" pageNumber="152">345</figureCitation>
)
</paragraph>
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<subSubSection id="F69A64F7FF092ED0B8BFF9DDFABDB58D" pageId="151" pageNumber="152" type="reference_group">
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<treatmentCitationGroup id="9E901052FF092ED0B8BFF9DDFEA5B56C" pageId="151" pageNumber="152">
<emphasis id="8CF4EB6EFF092ED0B8BFF9DDFF3EB521" box="[151,161,1570,1592]" italics="true" pageId="151" pageNumber="152">*</emphasis>
<treatmentCitation id="3F21116DFF092ED0B889F9DDFC9CB521" author="Kasparyan, D. R. &amp; Ruiz, E. C." box="[161,771,1570,1592]" page="105" pageId="151" pageNumber="152" year="2005">
<taxonomicName id="79804CFFFF092ED0B889F9DDFC9CB521" ID-CoL="73BGH" authority="Kasparyan &amp; Ruiz 2005: 105" authorityName=": Kasparyan &amp; Ruiz" authorityPageNumber="105" authorityYear="2008" box="[161,771,1570,1592]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF092ED0B889F9DDFE5AB521" box="[161,453,1570,1592]" italics="true" pageId="151" pageNumber="152">Messatoporus mesosternalis</emphasis>
<materialsCitation id="0EE83D21FF092ED0B9E6F9DCFC9CB521" box="[462,771,1570,1592]" collectingDate="2000-10-07" collectingDateMax="2000-10-14" collectingDateMin="2000-10-07" collectionCode="UATM" collectorName="D. R. Kasparyan." country="Mexico" elevation="900" location="Alta Cima" municipality="Gomes Farias" pageId="151" pageNumber="152" specimenCount="1" specimenCount-female="1" stateProvince="Tamaulipas" typeStatus="holotype">
<bibRefCitation id="DA114A8DFF092ED0B9E6F9DCFC9CB521" author="Kasparyan, D. R. &amp; Ruiz, E. C." box="[462,771,1570,1592]" pageId="151" pageNumber="152" pagination="105 - 122" refId="ref210720" refString="Kasparyan, D. R. &amp; Ruiz, E. C. (2005) A review of the North American species of Messatoporus Cushman (Hymenoptera: Ichneumonidae, Cryptini). Zoosystematica Rossica, 14 (1), 105 - 122." type="journal article" year="2005">Kasparyan &amp; Ruíz 2005: 105</bibRefCitation>
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, 118. Description, distribution, keyed.
<typeStatus id="613B89DEFF092ED0BCB1F9DDFB67B521" box="[1177,1272,1570,1592]" pageId="151" pageNumber="152" type="holotype">Holotype</typeStatus>
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(UATM, not examined). Type data:
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,
<collectingRegion id="7C44F99EFF092ED0BA00F9BEFD00B54E" box="[552,671,1601,1623]" country="Mexico" name="Tamaulipas" pageId="151" pageNumber="152">Tamaulipas</collectingRegion>
,
<collectingMunicipality id="5E5BAD06FF092ED0BA99F9BEFCD9B54E" box="[689,838,1601,1623]" pageId="151" pageNumber="152">Gómes Farías</collectingMunicipality>
,
<location id="BB5F61A7FF092ED0BB70F9BEFC54B54E" LSID="urn:lsid:plazi:treatment:3629866AFF092EDEB8BFFA41FCEFB7DD:BB5F61A7FF092ED0BB70F9BEFC54B54E" box="[856,971,1601,1623]" country="Mexico" municipality="Gomes Farias" name="Alta Cima" pageId="151" pageNumber="152" stateProvince="Tamaulipas">Alta Cima</location>
,
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<elevation id="35ADD04FFF092ED0BBF5F9BEFBB8B54E" box="[989,1063,1601,1623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="151" pageNumber="152" unit="m" value="900.0">900 m</elevation>
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,
<collectingMethod id="67C14F6BFF092ED0BC12F9BEFB5DB54E" box="[1082,1218,1601,1623]" pageId="151" pageNumber="152">Malaise trap</collectingMethod>
,
<date id="CA3E11BCFF092ED0BCFCF9BEFAC5B54E" box="[1236,1370,1601,1623]" pageId="151" pageNumber="152" value="2000-10-07" valueMax="2000-10-14" valueMin="2000-10-07">
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,
<collectorName id="137552AAFF092ED0BD44F9BDFEA5B56C" pageId="151" pageNumber="152">D.R. Kasparyan.</collectorName>
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</paragraph>
<paragraph id="BE3F377CFF092ED0B8BFF980FABDB58D" blockId="151.[151,1437,1570,1685]" box="[151,1314,1662,1685]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="9E901052FF092ED0B8BFF980FABDB58D" box="[151,1314,1662,1685]" pageId="151" pageNumber="152">
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<taxonomicName id="79804CFFFF092ED0B8BFF980FD71B58D" ID-CoL="73BGH" authority=": Kasparyan &amp; Ruiz 2008: 308" authorityName=": Kasparyan &amp; Ruiz" authorityPageNumber="308" authorityYear="2008" box="[151,750,1662,1685]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF092ED0B8BFF980FE29B58C" box="[151,438,1663,1685]" italics="true" pageId="151" pageNumber="152">Messatoporus mesosternalis</emphasis>
:
<bibRefCitation id="DA114A8DFF092ED0B9ECF980FD71B58D" author="Kasparyan, D. R. &amp; Ruiz, E. C." box="[452,750,1662,1684]" pageId="151" pageNumber="152" refId="ref210766" refString="Kasparyan, D. R. &amp; Ruiz, E. C. (2008) Cryptini de Mexico (Hymenoptera: Ichneumonidae: Cryptinae) Parte II. Serie Avispas parasiticas de plagas de plagas y otros insectos. Universidad Autonoma de Tamaulipas, Victoria, Mexico, 377 pp." type="book" year="2008">Kasparyan &amp; Ruíz 2008: 308</bibRefCitation>
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, 330–331, 373. Description, distribution, figure, keyed.
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</paragraph>
</subSubSection>
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<paragraph id="BE3F377CFF092ED0B8BFF944FDBFB4EB" blockId="151.[151,1436,1722,2035]" pageId="151" pageNumber="152">
<emphasis id="8CF4EB6EFF092ED0B8BFF944FF71B5CA" bold="true" box="[151,238,1723,1747]" pageId="151" pageNumber="152">Female</emphasis>
. Fore wing
<quantity id="79789A99FF092ED0B950F945FE7EB5CB" box="[376,481,1722,1746]" metricMagnitude="-3" metricUnit="m" metricValue="7.92" pageId="151" pageNumber="152" unit="mm" value="7.92">7.92 mm</quantity>
. HEAD: Mandible densely covered with long hairs; MLW 1.66; MWW 0.34; dorsal tooth in front view projected upwards, much longer than ventral tooth; ventral tooth somewhat rhomboid. MSM 0.23. Clypeus almost entirely flattened, centrally concave; CHW 2.21; CWW 2.34; apical area medially concave, laterally slightly projected, its margin very sharp, raised. Antenna with 21 flagellomeres; flagellum strongly flattened and compressed on apical 0.3; apex of apical flagellomere about as wide as base, not distinctly tapered. Supra-antennal area shiny, scarcely punctate, not striate near antennal sockets, ventrally and around ocelli slightly concave, medially with a distinct depression but without a distinct line. Occipital carina dorsally acuminate, fading out at distance shorter than basal width of mandible from the hypostomal carina. Temple and gena moderately wide, gena regular (as in
<figureCitation id="26BB2BF9FF092ED0B99AF825FD8CB4EB" box="[434,531,2010,2034]" captionStart="FIGURES 260–271. 260–261" captionStartId="270.[151,258,1798,1819]" captionTargetBox="[151,1436,193,1777]" captionTargetId="figure-105@270.[151,1436,193,1777]" captionTargetPageId="270" captionText="FIGURES 260–271. 260–261, Gena, lateral. 260, M. longicaudis*. 261, M. compressicornis. 262–264, Antenna. 262, M. tibiator. 263, M. compresscornis. 264, M. antennator, male. 265–266, Hind t4. 265, M. paeneater*. 266, M. nigriscutus*. 267–271, Metasoma and ovipositor. 267, M. opacus*. 268, M. discoidalis. 269, M. longicaudis*. 270, M. convexus. 271, M. townesi." figureDoi="http://doi.org/10.5281/zenodo.5262215" httpUri="https://zenodo.org/record/5262215/files/figure.png" pageId="151" pageNumber="152">Fig. 260</figureCitation>
).
</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFFF68FB4BB199" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">THORAX: Pronotum glabrous, shiny, impunctate, with weak and short wrinkles at ventral half of posterior margin; epomia weak, distinct only after diverging from pronotal collar, short, ending far from dorsal margin of pronotum, after diverging from pronotal collar approximately straight. Mesoscutum strongly and uniformly convex, subcircular, 1.11 × as long as wide, glabrate, shiny, impunctate; notaulus reaching about 0.69 of mesoscutum length, weakly impressed, surface over notaulus not wrinkled; scuto-scutellar groove moderately deep, without wrinkles. Subalar ridge moderately projected; epicnemial carina reaching about 0.7 of distance to subalar ridge, more or less uniformly curved; sternaulus moderately strong, wide and shallow at posterior 0.25, strongly sinuous, smooth; scrobe moderately deep; mesopleural suture with weak longitudinal wrinkles along entire length. Mesosternum medially with short transverse wrinkles; median portion of posterior transverse carina of the mesothoracic venter short and almost indistinct, distinctly arched forwards. Transverse sulcus at base of propodeum moderately wide, about 0.56 × as long as anterior area of propodeum, impunctate; metapleuron scarcely punctate, without transverse wrinkles, dorsally moderately pilose, ventrally densely pilose; juxtacoxal carina represented by very short ridges. Fore tibia distinctly swollen. Mesal lobe of t4 with a distinct cluster of stout bristles. Hind coxa with moderately strong punctures separated by more than their own diameter.</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFFD73FBF2B016" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">PROPODEUM: 1.11 × as long as wide, shiny, sparsely pilose; anterior area almost impunctate; spiracle elliptic, SWL 2.83; anterior transverse carina medially straight. Propodeal wrinkles anteriorly faint, posteriorly moderately strong, closely spaced, mostly incomplete and overlapped, with some disperse complete wrinkles, mostly slightly curved or somewhat sinuous; posterior transverse carina entirely absent.</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFFCE4FCD3B748" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">WINGS: Fore wing vein 1-Rs+M distinctly sinuous, with bulla placed on basal 0.25; crossvein 1m-cu more or less uniformly curved, limit between 1m-cu and 1-Rs+M clearly traceable; vein 1M+Rs anteriorly straight, posteriorly curved; fore wing crossvein 1 cu-a basad of 1M+Rs by 0.49 of its own length; vein 2Cua 0.52 × as long as crossvein 2cu-a; bulla of crossvein 2m-cu placed on midlength; cell 1+2Rs small, APH 0.63, subquadratic, distinctly higher than wide, AWH 0.79; crossvein 3r-m fully spectral, 2r-m and 3r-m subparallel, about same length; vein 3-M distinctly longer than 2-M; 4-Rs straight, apically slightly upcurved; 4-M spectral, slightly curved backwards. Hind wing vein 1-M forming straight angle with vein Cua; vein 2-Rs tubular, apically spectral, reaching wing margin; HW1C 1.53; vein Cub distinctly convex on posterior half, forming straight angle with Cua; vein 2-1A reaching 0.87 of distance to posterior wing margin.</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFFBA3FC49B676" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">METASOMA: First tergite short, about 0.31 × as long as T2–8, somewhat depressed, ventrolaterally angled, dorsally sparsely pilose, laterally moderately pilose; T1LW 3.45; T1WW 1.66; spiracle on 0.55 of its length, distinctly prominent; median depression elongate, very shallow; lateral and median posterior depressions absent; ventrolateral carina absent. T2LW 1.18; T2WW 1.92; thyridium about as long as wide, without small circular depression just behind it; T2–8 smooth between punctures, laterally moderately pilose, dorsally T2–3 and 8 glabrate, T4–7 sparsely to moderately pilose. OST 1.32; ovipositor moderately stout, straight, basally cylindric, apically slightly depressed; dorsal valve with four weak ridges; ventral valve apex with 9 teeth, apical teeth progressively more closely spaced; surface anterior to first tooth rugulose.</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFFA84FB99B402" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">
<emphasis id="8CF4EB6EFF062EDFB8EFFA84FE98B68A" box="[199,263,1403,1427]" italics="true" pageId="152" pageNumber="153">Color</emphasis>
. Head, mesosoma and metasoma black and yellow. Head: black; clypeus, mandible except apex and borders, mouthparts, supraclypeal area except small lateral marks and complete orbital band, pale yellow (228,220,141); f5–9 entirely white; f10 almost entirely white, f11–12 partially white. Mesosoma: black; pronotal collar, propleuron ventrally, subcircular spot on mesoscutum, scutellum, axillar carinae, postscutellum, tegula, subalar ridge, central mark on mesopleuron, dorsal division of metapleuron, posterior 0.5 of metapleuron and large trapezoidal marks at posterior area of propodeum, pale yellow; mesal face of fore and mid coxae with large blackish mark, mid coxa with small apical brown spot; all legs apically to first trochanter bright yellow, whitish towards apex; fore and mid femora with longitudinal blackish stripes at posterior face; fore and mid t5 dark brown; hind coxa anteriorly black, posteriorly pale yellow with narrow lateral blackish line; first trochanter ventrally black, second trochanter almost entirely blackish; ventral face and apical 0.15 of dorsal face of hind femur black; extreme base of hind tibia marked with blackish. Metasoma: black; T1–8 with posterior pale yellow stripes, expanding laterally at T2–8; S2–6 brown on basal 0.5, posteriorly pale yellow.
</paragraph>
<paragraph id="BE3F377CFF062EDFB8EFF8D7FB37B4EB" blockId="152.[151,1437,151,2034]" pageId="152" pageNumber="153">
<emphasis id="8CF4EB6EFF062EDFB8EFF8D7FE9BB459" bold="true" box="[199,260,1832,1856]" pageId="152" pageNumber="153">Male</emphasis>
. Not examined. According to
<bibRefCitation id="DA114A8DFF062EDFBA4AF8D7FC3FB426" author="Kasparyan, D. R. &amp; Ruiz, E. C." box="[610,928,1831,1856]" pageId="152" pageNumber="153" refId="ref210766" refString="Kasparyan, D. R. &amp; Ruiz, E. C. (2008) Cryptini de Mexico (Hymenoptera: Ichneumonidae: Cryptinae) Parte II. Serie Avispas parasiticas de plagas de plagas y otros insectos. Universidad Autonoma de Tamaulipas, Victoria, Mexico, 377 pp." type="book" year="2008">Kasparyan and Ruíz (2008)</bibRefCitation>
, similar to female, except for the following: fore wing 5.00–6.00 mm long; antenna with 23–24 flagellomeres; propleuron, fore and mid coxae and trochanters completely whitish; whitish central spot on mesopleuron connected with whitish color at fore coxa; mesepimeron whitish; black portion at posterior area of propodeum weak, not connected with anterior black portion; fore and mid femora not ventrally black; hind femur reddish, with apical 0.2 black; hind tibia yellowish with apical 0.35 blackish; tibial spurs and basal 0.4 of t1 blackish; T2 about 3.0 × as long as maximum width.
</paragraph>
<paragraph id="BE3F377CFF072EDEB8EFFF68FDEDB226" blockId="153.[151,1436,151,1220]" pageId="153" pageNumber="154">
<emphasis id="8CF4EB6EFF072EDEB8EFFF68FEA8B3A9" bold="true" box="[199,311,151,176]" pageId="153" pageNumber="154">Variation</emphasis>
. Fore wing
<quantity id="79789A99FF072EDEB9F8FF68FDE7B3B6" box="[464,632,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="8.725" metricValueMax="9.53" metricValueMin="7.92" pageId="153" pageNumber="154" unit="mm" value="8.725" valueMax="9.53" valueMin="7.92">7.92–9.53 mm</quantity>
long. Specimen from Fortín de las
<collectingCountry id="C69777ECFF072EDEBC06FF68FBEBB3B6" box="[1070,1140,151,175]" name="Indonesia" pageId="153" pageNumber="154">Flores</collectingCountry>
with yellowish spots on mesosternum ventrally to sternaulus and median anterior portion of transverse sulcus of propodeum, without narrow blackish stripe at hind coxa, and with ventrolateral yellowish mark on apical 0.4 of T1. Specimen from Puntarenas with longitudinal black mark on propodeum very narrow anteriorly. posterior face of hind coxa sometimes without narrow blackish stripe.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FF072EDEB8EFFEB4FB8AB0F5" pageId="153" pageNumber="154" type="discussion">
<paragraph id="BE3F377CFF072EDEB8EFFEB4FBD8B146" blockId="153.[151,1436,151,1220]" pageId="153" pageNumber="154">
<emphasis id="8CF4EB6EFF072EDEB8EFFEB4FED7B27D" bold="true" box="[199,328,331,356]" pageId="153" pageNumber="154">Comments</emphasis>
.
<taxonomicName id="79804CFFFF072EDEB97EFEB2FD0BB27D" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[342,660,332,356]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF072EDEB97EFEB2FD0BB27D" box="[342,660,332,356]" italics="true" pageId="153" pageNumber="154">Messatoporus mesosternalis</emphasis>
</taxonomicName>
is part of a group of five very close species, all very similar although not forming a strictly monophyletic group (see Analysis 2). Along with the recovered synapomorphies, species of that group are generally similar in size and share features like body very shiny and scarcely punctate to almost impunctate; pronotum mostly black, yellow or whitish only at collar, but not at dorsal margin (which is unusual; species with yellow collar usually do also have the dorsal margin of pronotum marked with yellowish); mesosternum entirely black; hind tibia entirely bright yellow or blackish only on apical 0.2; T1 short, 0.31–0.38 × as long as T2–8, with posterior yellow or whitish stripe, anteriorly entirely black except in
<taxonomicName id="79804CFFFF072EDEBCB3FDDAFAACB125" authority="Santos &amp; Aguiar, 2013" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[1179,1331,548,572]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="basiflavus" status="sp. nov.">
<emphasis id="8CF4EB6EFF072EDEBCB3FDDAFB2DB125" box="[1179,1202,549,572]" italics="true" pageId="153" pageNumber="154">M</emphasis>
.
<emphasis id="8CF4EB6EFF072EDEBCE9FDDBFAACB125" box="[1217,1331,548,572]" italics="true" pageId="153" pageNumber="154">basiflavus</emphasis>
</taxonomicName>
<emphasis id="8CF4EB6EFF072EDEBD12FDDBFA0BB125" bold="true" box="[1338,1428,548,572]" pageId="153" pageNumber="154">
<taxonomicNameLabel id="97C75615FF072EDEBD12FDDBFA0BB125" box="[1338,1428,548,572]" pageId="153" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; and ovipositor dorsal valve with four ridges. Most body proportions are also similar.
</paragraph>
<paragraph id="BE3F377CFF072EDEB8EFFD93FB8AB0F5" blockId="153.[151,1436,151,1220]" pageId="153" pageNumber="154">
Among these species,
<taxonomicName id="79804CFFFF072EDEB9E3FD92FD11B19D" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[459,654,620,644]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF072EDEB9E3FD92FE7DB19D" box="[459,482,621,644]" italics="true" pageId="153" pageNumber="154">M</emphasis>
.
<emphasis id="8CF4EB6EFF072EDEB9DCFD93FD11B19D" box="[500,654,620,644]" italics="true" pageId="153" pageNumber="154">mesosternalis</emphasis>
</taxonomicName>
can be readily identified by its unique color pattern at hind femur, which is mostly black, dorsally bright yellow on basal 0.7.
<taxonomicName id="79804CFFFF072EDEBB12FD6EFBF8B1B1" authority="Santos &amp; Aguiar, 2013" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[826,1127,656,680]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="8CF4EB6EFF072EDEBB12FD6EFBF8B1B1" box="[826,1127,656,680]" italics="true" pageId="153" pageNumber="154">Messatoporus occidentalis</emphasis>
</taxonomicName>
<emphasis id="8CF4EB6EFF072EDEBC47FD6FFB55B1B1" bold="true" box="[1135,1226,656,680]" pageId="153" pageNumber="154">
<taxonomicNameLabel id="97C75615FF072EDEBC47FD6FFB55B1B1" box="[1135,1226,656,680]" pageId="153" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="79804CFFFF072EDEBD2CFD6EFA04B1B1" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[1284,1435,656,680]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="basiflavus">
<emphasis id="8CF4EB6EFF072EDEBD2CFD6EFA84B1B1" box="[1284,1307,657,680]" italics="true" pageId="153" pageNumber="154">M</emphasis>
.
<emphasis id="8CF4EB6EFF072EDEBD02FD6FFA04B1B1" box="[1322,1435,656,680]" italics="true" pageId="153" pageNumber="154">basiflavus</emphasis>
</taxonomicName>
can be readily differentiated from
<taxonomicName id="79804CFFFF072EDEBA0BFD4AFD7AB1D5" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[547,741,692,716]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF072EDEBA0BFD4AFD7AB1D5" box="[547,741,692,716]" italics="true" pageId="153" pageNumber="154">M. mesosternalis</emphasis>
</taxonomicName>
by the mostly or entirely orange hind coxa, trochanters and femur, and by having the yellow mark on propodeum distinctly arc-shaped (vs. posterior area of propodeum with distinctly separated yellow marks, black area T-shaped). In
<taxonomicName id="79804CFFFF072EDEBB46FD02FBF9B00D" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[878,1126,764,788]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="semiaurantigaster">
<emphasis id="8CF4EB6EFF072EDEBB46FD02FC1AB00D" box="[878,901,765,788]" italics="true" pageId="153" pageNumber="154">M</emphasis>
.
<emphasis id="8CF4EB6EFF072EDEBBB3FD02FBF9B00D" box="[923,1126,765,788]" italics="true" pageId="153" pageNumber="154">semiaurantigaster</emphasis>
</taxonomicName>
, the yellow mark on the propodeum is arc-shaped, but very narrow medially; it can be promptly isolated from
<taxonomicName id="79804CFFFF072EDEBC78FCDEFA90B021" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[1104,1295,800,824]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF072EDEBC78FCDEFA90B021" box="[1104,1295,800,824]" italics="true" pageId="153" pageNumber="154">M. mesosternalis</emphasis>
</taxonomicName>
by the T4–8 entirely bright orange, orbital band shortly interrupted at dorsal margin of eye and hind tibia blackish on apical 0.2.
<taxonomicName id="79804CFFFF072EDEB8BFFC96FE39B099" authority="Santos &amp; Aguiar, 2013" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[151,422,872,896]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="8CF4EB6EFF072EDEB8BFFC96FE39B099" box="[151,422,872,896]" italics="true" pageId="153" pageNumber="154">Messatoporus orientalis</emphasis>
</taxonomicName>
<emphasis id="8CF4EB6EFF072EDEB985FC97FD98B099" bold="true" box="[429,519,872,896]" pageId="153" pageNumber="154">
<taxonomicNameLabel id="97C75615FF072EDEB985FC97FD98B099" box="[429,519,872,896]" pageId="153" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the most similar to
<taxonomicName id="79804CFFFF072EDEBB28FC96FC20B099" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[768,959,872,896]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mesosternalis">
<emphasis id="8CF4EB6EFF072EDEBB28FC96FC20B099" box="[768,959,872,896]" italics="true" pageId="153" pageNumber="154">M. mesosternalis</emphasis>
</taxonomicName>
, from which it can be separated by having orbital band incomplete, hind femur entirely dark brown or blackish; crossvein 3r-m absent (vs. present, though spectral); T2 with circular depressions just posteriorly to thyridium (vs. without depressions); thyridium longer than wide (vs. about as wide as high); and yellow marks on propodeum smaller.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FF072EDEB8EFFC08FE3CB716" box="[199,419,1015,1040]" pageId="153" pageNumber="154" type="biology_ecology">
<paragraph id="BE3F377CFF072EDEB8EFFC08FE3CB716" blockId="153.[151,1436,151,1220]" box="[199,419,1015,1040]" pageId="153" pageNumber="154">
<emphasis id="8CF4EB6EFF072EDEB8EFFC08FE80B709" bold="true" box="[199,287,1015,1040]" pageId="153" pageNumber="154">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FF072EDEB8EFFBE4FC49B786" pageId="153" pageNumber="154" type="materials_examined">
<paragraph id="BE3F377CFF072EDEB8EFFBE4FC49B786" blockId="153.[151,1436,151,1220]" pageId="153" pageNumber="154">
<emphasis id="8CF4EB6EFF072EDEB8EFFBE4FE30B72D" bold="true" box="[199,431,1051,1076]" pageId="153" pageNumber="154">Material examined</emphasis>
.
<specimenCount id="A886FCF5FF072EDEB9EAFBE3FE6EB72A" box="[450,497,1052,1076]" pageId="153" pageNumber="154" type="female">3 ♀</specimenCount>
.
<materialsCitation id="0EE83D21FF072EDEBA32FBE3FB28B74E" collectingDate="1968-07-12" collectorName="Planta de la Cerveceria &amp; Ing. Daniel Rabago Res &amp; Jr" country="Mexico" county="Fortin" elevation="838" location="Sumidero" municipality="Flores" pageId="153" pageNumber="154" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave">
<specimenCount id="A886FCF5FF072EDEBA32FBE3FDD6B72A" box="[538,585,1052,1076]" pageId="153" pageNumber="154" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFF072EDEBAB1FBE3FC95B72D" box="[665,778,1052,1076]" name="Mexico" pageId="153" pageNumber="154">MEXICO</collectingCountry>
,
<collectingRegion id="7C44F99EFF072EDEBB33FBE3FC1EB72A" box="[795,897,1052,1075]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="153" pageNumber="154">Veracruz</collectingRegion>
,
<collectingCounty id="575E4FF0FF072EDEBBB9FBE3FC49B72D" box="[913,982,1052,1076]" pageId="153" pageNumber="154">Fortin</collectingCounty>
de las
<collectingMunicipality id="5E5BAD06FF072EDEBC1CFBE3FBE2B72D" box="[1076,1149,1052,1076]" pageId="153" pageNumber="154">Flores</collectingMunicipality>
,
<location id="BB5F61A7FF072EDEBCA6FBE3FB61B72D" LSID="urn:lsid:plazi:treatment:3629866AFF092EDEB8BFFA41FCEFB7DD:BB5F61A7FF072EDEBCA6FBE3FB61B72D" box="[1166,1278,1052,1076]" country="Mexico" county="Fortin" municipality="Flores" name="Sumidero" pageId="153" pageNumber="154" stateProvince="Veracruz de Ignacio de la Llave">Sumidero</location>
,
<collectorName id="137552AAFF072EDEBD26FBE3FE8CB74E" pageId="153" pageNumber="154">Planta de la Cerveceria</collectorName>
,
<collectorName id="137552AAFF072EDEB90BFBBFFDA0B74E" box="[291,575,1087,1112]" pageId="153" pageNumber="154">Ing. Daniel Rabago Res</collectorName>
,
<quantity id="79789A99FF072EDEBA67FBC0FD4AB74E" box="[591,725,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="8.375" metricValueMax="9.15" metricValueMin="7.6" pageId="153" pageNumber="154" unit="m" value="837.5" valueMax="915.0" valueMin="760.0">
<elevation id="35ADD04FFF072EDEBA67FBC0FD4AB74E" box="[591,725,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="8.375" metricValueMax="9.15" metricValueMin="7.6" pageId="153" pageNumber="154" unit="m" value="837.5" valueMax="915.0" valueMin="760.0">760–915 m</elevation>
</quantity>
,
<date id="CA3E11BCFF072EDEBACDFBC0FCEFB74E" box="[741,880,1087,1111]" pageId="153" pageNumber="154" value="1968-07-12">
<collectingDate id="DA7AE854FF072EDEBACDFBC0FCEFB74E" box="[741,880,1087,1111]" pageId="153" pageNumber="154" value="1968-07-12">12.VII.1968</collectingDate>
</date>
,
<collectingMethod id="67C14F6BFF072EDEBBA9FBC0FB8DB74E" box="[897,1042,1087,1111]" pageId="153" pageNumber="154">Malaise trap</collectingMethod>
, HVWeens
<collectorName id="137552AAFF072EDEBC89FBBFFB28B74E" box="[1185,1207,1088,1111]" pageId="153" pageNumber="154">Jr</collectorName>
</materialsCitation>
.
<materialsCitation id="0EE83D21FF072EDEBCEEFBC0FA89B765" collectingDate="1981-07-23" collectingDateMax="1981-07-26" collectingDateMin="1981-07-23" collectionCode="FSCA" collectorName="Prov. Piedras Blancas &amp; Jr." country="Costa Rica" location="Puntarenas" pageId="153" pageNumber="154" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<specimenCount id="A886FCF5FF072EDEBCEEFBC0FB6AB74E" box="[1222,1269,1087,1111]" pageId="153" pageNumber="154" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFF072EDEBD6AFBC0FF42B765" name="Costa Rica" pageId="153" pageNumber="154">COSTA RICA</collectingCountry>
,
<collectingRegion id="7C44F99EFF072EDEB8C6FB9BFEF3B762" box="[238,364,1124,1147]" country="Costa Rica" name="Puntarenas" pageId="153" pageNumber="154">Puntarenas</collectingRegion>
,
<collectorName id="137552AAFF072EDEB955FB9BFD1FB765" box="[381,640,1124,1148]" pageId="153" pageNumber="154">Prov. Piedras Blancas</collectorName>
,
<date id="CA3E11BCFF072EDEBABAFB9BFCD8B765" box="[658,839,1124,1148]" pageId="153" pageNumber="154" value="1981-07-23" valueMax="1981-07-26" valueMin="1981-07-23">
<collectingDate id="DA7AE854FF072EDEBABAFB9BFCD8B765" box="[658,839,1124,1148]" pageId="153" pageNumber="154" value="1981-07-23" valueMax="1981-07-26" valueMin="1981-07-23">23–26.VII.1981</collectingDate>
</date>
, HVWeens
<collectorName id="137552AAFF072EDEBBF1FB9BFC6CB762" box="[985,1011,1124,1147]" pageId="153" pageNumber="154">Jr.</collectorName>
&amp; GBEdwards (
<collectionCode id="D891AFB9FF072EDEBCEAFB9BFA91B765" box="[1218,1294,1124,1148]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="153" pageNumber="154" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0EE83D21FF072EDEBD0FFB9BFC4DB786" collectingDate="1987-07-07" collectionCode="AEIC" collectorName="Pena Blanca" country="Costa Rica" elevation="700" location="Alajuela" pageId="153" pageNumber="154" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela">
<specimenCount id="A886FCF5FF072EDEBD0FFB9BFAC7B762" box="[1319,1368,1124,1148]" pageId="153" pageNumber="154" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFF072EDEB8BFFB78FEA2B786" box="[151,317,1159,1183]" name="Costa Rica" pageId="153" pageNumber="154">COSTA RICA</collectingCountry>
,
<collectingRegion id="7C44F99EFF072EDEB960FB78FE35B786" box="[328,426,1159,1183]" country="Costa Rica" name="Alajuela" pageId="153" pageNumber="154">Alajuela</collectingRegion>
,
<collectorName id="137552AAFF072EDEB99DFB77FDDCB786" box="[437,579,1159,1183]" pageId="153" pageNumber="154">Pena Blanca</collectorName>
,
<quantity id="79789A99FF072EDEBA66FB78FD08B786" box="[590,663,1159,1183]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="153" pageNumber="154" unit="m" value="700.0">
<elevation id="35ADD04FFF072EDEBA66FB78FD08B786" box="[590,663,1159,1183]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="153" pageNumber="154" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="CA3E11BCFF072EDEBA8BFB78FCBFB786" box="[675,800,1159,1183]" pageId="153" pageNumber="154" value="1987-07-07">
<collectingDate id="DA7AE854FF072EDEBA8BFB78FCBFB786" box="[675,800,1159,1183]" pageId="153" pageNumber="154" value="1987-07-07">7.VII.1987</collectingDate>
</date>
, ECruz (
<collectionCode id="D891AFB9FF072EDEBBAAFB78FC55B786" box="[898,970,1159,1183]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" lsid="urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="153" pageNumber="154">AEIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FF072EDEB8EFFB54FCEFB7DD" box="[199,880,1195,1220]" pageId="153" pageNumber="154" type="distribution">
<paragraph id="BE3F377CFF072EDEB8EFFB54FCEFB7DD" blockId="153.[151,1436,151,1220]" box="[199,880,1195,1220]" pageId="153" pageNumber="154">
<emphasis id="8CF4EB6EFF072EDEB8EFFB54FEC6B7DD" bold="true" box="[199,345,1195,1220]" pageId="153" pageNumber="154">Distribution</emphasis>
. Northeastern Mexico to
<collectingCountry id="C69777ECFF072EDEBA50FB53FD6DB7DD" box="[632,754,1196,1220]" name="Costa Rica" pageId="153" pageNumber="154">Costa Rica</collectingCountry>
(
<figureCitation id="26BB2BF9FF072EDEBB29FB53FCFBB7DD" box="[769,868,1196,1220]" captionStart="FIGURES 345–352" captionStartId="280.[151,258,1969,1990]" captionTargetBox="[151,1398,195,1954]" captionTargetId="figure-67@280.[151,1398,193,1954]" captionTargetPageId="280" captionText="FIGURES 345–352. Distribution maps for species of Messatoporus. 345, M. mesosternalis. 346, M. nigriangulatus. 347, M. nigriscapus. 348, M. nigriscutus. 349, M. occidentalis. 350, M. opacus. 351, M. orientalis. 352, M. paeneater." figureDoi="http://doi.org/10.5281/zenodo.5262235" httpUri="https://zenodo.org/record/5262235/files/figure.png" pageId="153" pageNumber="154">Fig. 345</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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