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<document id="66945A031674A6BD537FB6FEEDB4D156" ID-CLB-Dataset="20238" ID-DOI="10.11646/zootaxa.5115.2.5" ID-GBIF-Dataset="fd2d13c3-71c1-4378-902a-92e3635376f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6352414" ID-ZooBank="784CAC77-93E2-4305-BADB-68C0C9A9BBD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647259178825" checkinUser="plazi" docAuthor="Anderson, Robert S." docDate="2022" docId="03EA5C1DFFE7347BFF0604FDFE7FF828" docLanguage="en" docName="zootaxa.5115.2.5.pdf" docOrigin="Zootaxa 5115 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhodobaenus howdenorum Anderson 2022, new species" docType="treatment" docVersion="7" lastPageNumber="270" masterDocId="FFD32465FFE43478FF91050EFFE7FF87" masterDocTitle="New species of Rhodobaenus LeConte, 1876 (Coleoptera: Curculionidae: Dryophthorinae) from Central and South America" masterLastPageNumber="273" masterPageNumber="267" pageNumber="270" updateTime="1699317997003" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6F67DD10B8165A2F8808952A6690D6CF">New species of Rhodobaenus LeConte, 1876 (Coleoptera: Curculionidae: Dryophthorinae) from Central and South America</mods:title>
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<mods:namePart id="4EF1DF3AC2DB8DB90717AEF46389C3AF">Anderson, Robert S.</mods:namePart>
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<mods:date id="1E5BE10D74248BAA613DFA8078BC28CB">2022</mods:date>
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<treatment id="03EA5C1DFFE7347BFF0604FDFE7FF828" ID-DOI="http://doi.org/10.5281/zenodo.6352428" ID-GBIF-Taxon="194020419" ID-Zenodo-Dep="6352428" LSID="urn:lsid:plazi:treatment:03EA5C1DFFE7347BFF0604FDFE7FF828" httpUri="http://treatment.plazi.org/id/03EA5C1DFFE7347BFF0604FDFE7FF828" lastPageNumber="270" pageId="3" pageNumber="270">
<subSubSection id="C359BE80FFE7347BFF0604FDFCE4FD89" box="[151,771,499,526]" pageId="3" pageNumber="270" type="nomenclature">
<paragraph id="8BFCED0BFFE7347BFF0604FDFCE4FD89" blockId="3.[151,771,499,563]" box="[151,771,499,526]" pageId="3" pageNumber="270">
<heading id="D0B45A67FFE7347BFF0604FDFCE4FD89" bold="true" box="[151,771,499,526]" fontSize="11" level="1" pageId="3" pageNumber="270" reason="1">
<emphasis id="B9373119FFE7347BFF0604FDFCE4FD89" bold="true" box="[151,771,499,526]" pageId="3" pageNumber="270">
<taxonomicName id="4C439688FFE7347BFF0604FDFD81FD89" authority="Anderson" authorityName="Anderson" authorityYear="2022" box="[151,614,499,526]" class="Insecta" family="Curculionidae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="howdenorum" status="sp. nov.">
<emphasis id="B9373119FFE7347BFF0604FDFE02FD89" bold="true" box="[151,485,499,526]" italics="true" pageId="3" pageNumber="270">Rhodobaenus howdenorum</emphasis>
Anderson
</taxonomicName>
,
<taxonomicNameLabel id="A2048C62FFE7347BFDE204FAFCE4FD89" box="[627,771,500,526]" pageId="3" pageNumber="270" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C359BE80FFE7347BFF060716FCBEFD44" pageId="3" pageNumber="270" type="description">
<paragraph id="8BFCED0BFFE7347BFF060716FF1EFDB4" blockId="3.[151,771,499,563]" box="[151,249,536,563]" pageId="3" pageNumber="270">
<figureCitation id="1378F18EFFE7347BFF060716FF1EFDB4" box="[151,249,536,563]" captionStart="FIGURES 18" captionStartId="2.[151,264,1614,1638]" captionTargetBox="[151,1436,179,1588]" captionTargetId="figure-30@2.[151,1436,179,1588]" captionTargetPageId="2" captionText="FIGURES 18. Species of Rhodobaenus. 12, Rhodobaenus chamorroae. 1, Dorsal habitus, male. 2, Lateral habitus, male. 34, Rhodobenus flecha. 3, Dorsal habitus, female. 4, Lateral habitus, female. 56, Rhodobenus howdenorum. 5, Dorsal habitus, male. 6, Lateral habitus, male. 78, Rhodobenus schusteri. 7, Dorsal habitus, male. 8, Lateral habitus, male." figureDoi="http://doi.org/10.5281/zenodo.6352416" httpUri="https://zenodo.org/record/6352416/files/figure.png" pageId="3" pageNumber="270">Figs 56</figureCitation>
</paragraph>
<paragraph id="8BFCED0BFFE7347BFF06076EFCBEFD44" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF06076EFE7EFDFD" bold="true" box="[151,409,608,634]" pageId="3" pageNumber="270">Specimens examined:</emphasis>
<materialsCitation id="3B2BE756FFE7347BFE0E076EFDF0FD19" ID-GBIF-Occurrence="3704873305" collectingDate="1976-02-27" collectionCode="CMNC" collectorName="H. Howden &amp; A. Howden" country="Ecuador" location="Rio Palenque" pageId="3" pageNumber="270" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F853A9FFE7347BFE0E076EFDE0FDFD" box="[415,519,608,634]" pageId="3" pageNumber="270" type="holotype">Holotype</typeStatus>
male, labeled
<collectingCountry id="F354AD9BFFE7347BFD3A076EFCD0FDFD" box="[683,823,608,634]" name="Ecuador" pageId="3" pageNumber="270">ECUADOR</collectingCountry>
:
<location id="8E9CBBD0FFE7347BFCD0076EFC3DFDFD" LSID="urn:lsid:plazi:treatment:03EA5C1DFFE7347BFF0604FDFE7FF828:8E9CBBD0FFE7347BFCD0076EFC3DFDFD" box="[833,986,608,634]" country="Ecuador" name="Rio Palenque" pageId="3" pageNumber="270">Rio Palenque</location>
,
<quantity id="4CBB40EEFFE7347BFC74076EFBCCFDFD" box="[997,1067,608,634]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="270" unit="km" value="47.0">47 km</quantity>
<collectorName id="26B688DDFFE7347BFBA3076EFAE4FDFD" box="[1074,1283,608,634]" pageId="3" pageNumber="270">S. Santo Domingo</collectorName>
, 700,
<date id="FFFDCBCBFFE7347BFADE076EFF35FD19" pageId="3" pageNumber="270" value="1976-02-27">
<collectingDate id="EFB93223FFE7347BFADE076EFF35FD19" pageId="3" pageNumber="270" value="1976-02-27">27 Feb 1976</collectingDate>
</date>
,
<collectorName id="26B688DDFFE7347BFF71078BFE42FD19" box="[224,421,644,671]" pageId="3" pageNumber="270">H. &amp; A. Howden</collectorName>
(
<collectionCode id="ED5275CEFFE7347BFE27078AFDE8FD19" box="[438,527,644,670]" pageId="3" pageNumber="270">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2BE756FFE7347BFDB5078BFCB1FD44" ID-GBIF-Occurrence="3704873308" collectingDate="1896-12" collectionCode="MNHN" collectorName="Oberthur" country="Ecuador" location="San Javier de Cachabi" pageId="3" pageNumber="270" specimenCount="1" specimenCount-male="1" stateProvince="Esmeraldas" typeStatus="paratype">
<typeStatus id="54F853A9FFE7347BFDB5078BFD6EFD18" box="[548,649,645,671]" pageId="3" pageNumber="270" type="paratype">Paratype</typeStatus>
,
<collectingCountry id="F354AD9BFFE7347BFD07078AFCC5FD19" box="[662,802,644,670]" name="Ecuador" pageId="3" pageNumber="270">ECUADOR</collectingCountry>
:
<collectingRegion id="498723E9FFE7347BFCBE078AFC54FD19" box="[815,947,644,670]" country="Ecuador" name="Esmeraldas" pageId="3" pageNumber="270">Esmeraldas</collectingRegion>
,
<location id="8E9CBBD0FFE7347BFC51078AFB24FD19" LSID="urn:lsid:plazi:treatment:03EA5C1DFFE7347BFF0604FDFE7FF828:8E9CBBD0FFE7347BFC51078AFB24FD19" box="[960,1219,644,670]" country="Ecuador" name="San Javier de Cachabi" pageId="3" pageNumber="270" stateProvince="Esmeraldas">San Javier de Cachabi</location>
, low c.,
<date id="FFFDCBCBFFE7347BFAB6078BFA7FFD19" box="[1319,1432,644,671]" pageId="3" pageNumber="270" value="1896-12">
<collectingDate id="EFB93223FFE7347BFAB6078BFA7FFD19" box="[1319,1432,644,671]" pageId="3" pageNumber="270" value="1896-12">Dec 1896</collectingDate>
</date>
,
<collectorName id="26B688DDFFE7347BFF0607A6FEF6FD45" box="[151,273,680,706]" pageId="3" pageNumber="270">Rosenberg</collectorName>
,
<collectorName id="26B688DDFFE7347BFE8D07A7FE5AFD45" box="[284,445,680,707]" pageId="3" pageNumber="270">Museum Paris</collectorName>
ex
<collectorName id="26B688DDFFE7347BFE7707A6FD6EFD45" box="[486,649,680,706]" pageId="3" pageNumber="270">Coll. Oberthur</collectorName>
(
<specimenCount id="9D452682FFE7347BFD0A07A6FD0FFD45" box="[667,744,680,706]" pageId="3" pageNumber="270" type="male">1 male</specimenCount>
,
<collectionCode id="ED5275CEFFE7347BFD6207A7FCA9FD44" box="[755,846,681,707]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="270">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359BE80FFE7347BFF5607C2FD0FFB3D" pageId="3" pageNumber="270" type="diagnosis">
<paragraph id="8BFCED0BFFE7347BFF5607C2FC63FCF1" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF5607C2FEA3FD61" bold="true" box="[199,324,716,742]" pageId="3" pageNumber="270">Diagnosis:</emphasis>
Length
<quantity id="4CBB40EEFFE7347BFE3307C2FE1BFD60" box="[418,508,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="270" unit="mm" value="9.5">9.5 mm</quantity>
. Dorsally dark red (excepting black midline and lateral lines of the pronotum) and a short black line along elytral interval 3 at about the midlength of the elytra. Rostrum moderately long, curved, humped at base, red, lacking any dorsal hairs; ventrally with peduncle only very slightly produced, rounded. Pygidium with sparse short yellowish setae not forming distinct apical tuft. Ventral surfaces of femora lacking rows of long fine hairs, tarsus with claw bearing article flat between claws.
</paragraph>
<paragraph id="8BFCED0BFFE7347BFF56068EFB32FBAD" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF56068EFE95FC1D" bold="true" box="[199,370,896,922]" pageId="3" pageNumber="270">Identification:</emphasis>
Despite very close similarity to
<emphasis id="B9373119FFE7347BFD72068FFC9EFC1D" box="[739,889,896,922]" italics="true" pageId="3" pageNumber="270">R. longicollis</emphasis>
Hustache, in
<bibRefCitation id="EFD290FAFFE7347BFB85068EFB57FC1D" author="Vaurie, P." box="[1044,1200,896,922]" pageId="3" pageNumber="270" pagination="1 - 44" refId="ref3955" refString="Vaurie, P. (1980) Revision of Rhodobaenus. Part 1. Species in South America (Coleoptera, Curculionidae, Rhynchophorinae). Bulletin of the American Museum of Natural History, 167 (1), 1 - 44." type="journal article" year="1980">Vaurie (1980)</bibRefCitation>
this species will key to
<taxonomicName id="4C439688FFE7347BFF2206AAFDFBFC38" authority="(Kirsch)" baseAuthorityName="Kirsch" box="[179,540,932,959]" class="Insecta" family="Curculionidae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="nawradii">
<emphasis id="B9373119FFE7347BFF2206AAFE50FC39" box="[179,439,932,958]" italics="true" pageId="3" pageNumber="270">Rhodobaenus nawradii</emphasis>
(Kirsch)
</taxonomicName>
in couplet 13. The two species share the tarsus with claw bearing article flat, not excavate, between claws, basally transversely impressed pronotum and the completely pilose venter of tarsal article 3.
<taxonomicName id="4C439688FFE7347BFF2306E2FE04FB81" authorityName="Anderson" authorityYear="2022" box="[178,483,1004,1030]" class="Insecta" family="Curculionidae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B9373119FFE7347BFF2306E2FE04FB81" box="[178,483,1004,1030]" italics="true" pageId="3" pageNumber="270">Rhodobaenus howdenorum</emphasis>
</taxonomicName>
is a smaller primarily reddish species with some longitudinal black markings whereas
<taxonomicName id="4C439688FFE7347BFF06011FFEFCFBAD" box="[151,283,1040,1066]" class="Insecta" family="Curculionidae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="nawradii">
<emphasis id="B9373119FFE7347BFF06011FFEFCFBAD" box="[151,283,1040,1066]" italics="true" pageId="3" pageNumber="270">R. nawradii</emphasis>
</taxonomicName>
is larger and predominantly grey with two large black velvety-looking elytral marks.
</paragraph>
<paragraph id="8BFCED0BFFE7347BFF56013AFD0FFB3D" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<taxonomicName id="4C439688FFE7347BFF56013AFE1FFBC9" authorityName="Anderson" authorityYear="2022" box="[199,504,1076,1102]" class="Insecta" family="Curculionidae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B9373119FFE7347BFF56013AFE1FFBC9" box="[199,504,1076,1102]" italics="true" pageId="3" pageNumber="270">Rhodobaenus howdenorum</emphasis>
</taxonomicName>
can be separated from
<emphasis id="B9373119FFE7347BFD6B013BFC69FBC9" box="[762,910,1076,1102]" italics="true" pageId="3" pageNumber="270">R. longicollis</emphasis>
by the lack of an excavation between the tarsal claws, a slightly less deep transverse impression across the base of the pronotum, dorsal surface of rostrum lacking hairs, a very slightly ventrally projected gular peduncle in the male, sparsely tomentose pygidium, and lack of a row of long hairs along the ventral surface of each femur.
</paragraph>
</subSubSection>
<subSubSection id="C359BE80FFE7347BFF5601CAFD31F97D" pageId="3" pageNumber="270" type="description">
<paragraph id="8BFCED0BFFE7347BFF5601CAFA9BF935" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF5601CAFEBCFB59" bold="true" box="[199,347,1220,1246]" pageId="3" pageNumber="270">Description:</emphasis>
Length
<quantity id="4CBB40EEFFE7347BFE2A01CAFDF4FB58" box="[443,531,1220,1247]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="270" unit="mm" value="9.5">9.5 mm</quantity>
in male. Cuticle dark red with black longitudinal lines on pronotum, one along midline, the other two lateral, at top of flanks, short black line along elytral interval 3 at about the midlength of the elytra, no whitish spots around punctures. Rostrum humped over scrobe, more strongly arcuate basally, less so apically, moderately long, length slightly shorter than length pronotum, basal area slightly transversely impressed. Gular peduncle slightly produced, rounded, not angulate behind. Antennal scape subcylindrical, about two-thirds length of rostrum, almost twice length funicle, slightly clavate apically. Antennal scape with articles 1 and 2 longer than wide, articles 36 moniliform, width of club subequal to width of apex of rostrum; spongy apex about threefifths length of club. Antennal groove with posterior border separated from anterior margin of eye by about width of apex of scape. Pronotum very slightly longer than width across base, basal area slightly transversely impressed. Elytra slightly wider than pronotum, about twice as long as pronotum, striae with vaguely defined punctures. Prosternum flat in front of coxa. Femora lacking rows of long fine hairs along ventral surfaces, posterior femur with width at apex about twice width at base, tarsus with claw bearing article flat, not excavate, between claws. Pygidium with punctate part short, about as wide as long, somewhat convex, with setae sparse, not forming apical tuft. Abdomen lacking ventral tubercle. Male not dissected although aedeagus is extruded on
<typeStatus id="54F853A9FFE7347BFBAB0396FB78F935" box="[1082,1183,1688,1714]" pageId="3" pageNumber="270" type="holotype">holotype</typeStatus>
, female not known.
</paragraph>
<paragraph id="8BFCED0BFFE7347BFF5603B2FD31F97D" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF5603B2FEA6F951" bold="true" box="[199,321,1724,1750]" pageId="3" pageNumber="270">Variation:</emphasis>
The
<typeStatus id="54F853A9FFE7347BFEEE03B3FE38F950" box="[383,479,1725,1751]" pageId="3" pageNumber="270" type="paratype">paratype</typeStatus>
male has the short black line along elytral interval 3 at about the midlength of the elytra much less well-defined than on the
<typeStatus id="54F853A9FFE7347BFDFD03EEFD35F97D" box="[620,722,1760,1786]" pageId="3" pageNumber="270" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C359BE80FFE7347BFF56020AFE7FF828" pageId="3" pageNumber="270" type="etymology">
<paragraph id="8BFCED0BFFE7347BFF56020AFA63F899" blockId="3.[151,1437,608,1967]" box="[199,1412,1796,1823]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF56020AFEB5F899" bold="true" box="[199,338,1796,1822]" pageId="3" pageNumber="270">Etymology:</emphasis>
This species is named after Dr. Henry and Anne Howden who collected the
<typeStatus id="54F853A9FFE7347BFB38020AFAECF899" box="[1193,1291,1796,1822]" pageId="3" pageNumber="270" type="holotype">holotype</typeStatus>
specimen.
</paragraph>
<paragraph id="8BFCED0BFFE7347BFF560226FE7FF828" blockId="3.[151,1437,608,1967]" pageId="3" pageNumber="270">
<emphasis id="B9373119FFE7347BFF560226FEF4F8C5" bold="true" box="[199,275,1832,1858]" pageId="3" pageNumber="270">Notes:</emphasis>
The
<typeStatus id="54F853A9FFE7347BFEDE0226FE56F8C5" box="[335,433,1832,1858]" pageId="3" pageNumber="270" type="holotype">holotype</typeStatus>
has a small note handwritten by Patricia Vaurie under the specimen “Resembles
<emphasis id="B9373119FFE7347BFADD0226FEEAF8E1" italics="true" pageId="3" pageNumber="270">Rhodo. longicollis</emphasis>
but has no claw excavation, and no white spots. Is specimen like this in Paris Mus. P.
<date id="FFFDCBCBFFE7347BFB5C0243FACEF8E1" box="[1229,1321,1868,1895]" pageId="3" pageNumber="270" value="1981-05">V. 1981</date>
”. This latter specimen, with the following labels, “claw very tiny, one in Howden Coll.,
<emphasis id="B9373119FFE7347BFB8E027EFB50F80D" box="[1055,1207,1904,1930]" italics="true" pageId="3" pageNumber="270">Rhodobaenus</emphasis>
<taxonomicNameLabel id="A2048C62FFE7347BFB50027FFB19F80C" box="[1217,1278,1905,1931]" pageId="3" pageNumber="270" rank="species">n. sp.</taxonomicNameLabel>
Vaurie 80” is included as a
<typeStatus id="54F853A9FFE7347BFEA1029BFE73F828" box="[304,404,1941,1967]" pageId="3" pageNumber="270" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>