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<document id="FA58A4014865C52FE3DD0051C7C3C48F" ID-CLB-Dataset="58033" ID-DOI="10.5852/ejt.2023.882.2177" ID-GBIF-Dataset="57d1d191-2bf5-4229-a0f5-9ab472a114a6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177581" ID-ZooBank="67C909E4-C700-4F8D-B8CE-5FD9B2C5D549" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1690201644379" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Weyer, Dieter" docDate="2023" docId="03EA5C14CA18855DFDE2FE50FAFD80BD" docLanguage="en" docName="ejt-882_1-230.pdf" docOrigin="European Journal of Taxonomy 882" docSource="http://dx.doi.org/10.5852/ejt.2023.882.2177" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Prionoceratinae Hyatt 1884" docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="FFD3246CCA09854FFFCEFF90FFB2846D" masterDocTitle="The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)" masterLastPageNumber="230" masterPageNumber="1" pageNumber="18" updateTime="1699545181523" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D83BCA7DAB74049D939AED365899A472">The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)</mods:title>
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<mods:namePart id="6DA88C0131EE75019812A16F2F971F47">Korn, Dieter</mods:namePart>
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<mods:affiliation id="35DE8E61AFEDF9196D5020C8012D321F">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:affiliation id="F1D11399075BD42D4470DDDA323A94AB">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<treatment id="03EA5C14CA18855DFDE2FE50FAFD80BD" ID-DOI="http://doi.org/10.5281/zenodo.8184800" ID-GBIF-Taxon="212420326" ID-Zenodo-Dep="8184800" LSID="urn:lsid:plazi:treatment:03EA5C14CA18855DFDE2FE50FAFD80BD" httpUri="http://treatment.plazi.org/id/03EA5C14CA18855DFDE2FE50FAFD80BD" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="C359BE89CA18855EFDE2FE50FBBA85B6" box="[556,1032,448,475]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BFCED02CA18855EFDE2FE50FBBA85B6" blockId="17.[556,1032,448,475]" box="[556,1032,448,475]" pageId="17" pageNumber="18">
Subfamily
<taxonomicName id="4C439681CA18855EFD63FE50FBBA85B6" authority="Hyatt, 1884" authorityName="Hyatt" authorityYear="1884" box="[685,1032,448,475]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">
<emphasis id="B9373110CA18855EFD63FE50FCC785B7" bold="true" box="[685,885,448,474]" pageId="17" pageNumber="18">Prionoceratinae</emphasis>
Hyatt, 1884
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA18855EFF73FD93FB868671" box="[189,1076,514,541]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BFCED02CA18855EFF73FD93FB868671" blockId="17.[189,1076,514,541]" box="[189,1076,514,541]" pageId="17" pageNumber="18">
[nom. transl.
<bibRefCitation id="EFD290F3CA18855EFE97FD92FD7D8671" author="Bartzsch K. &amp; Weyer D." box="[345,719,514,541]" pageId="17" pageNumber="18" pagination="130 - 142" refId="ref110993" refString="Bartzsch K. &amp; Weyer D. 1988 a. Die unterkarbonische Ammonoidea-Subfamilia Karagandoceratinae. Freiberger Forschungs-Hefte, C 419: 130 - 142." type="journal article" year="1988">Bartzsch &amp; Weyer (1988a: 136)</bibRefCitation>
, pro Prionocerae Hyatt, 1884]
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA18855EFF73FDD4FD248695" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BFCED02CA18855EFF73FDD4FE848633" blockId="17.[189,310,580,606]" box="[189,310,580,606]" pageId="17" pageNumber="18">
<heading id="D0B45A6ECA18855EFF73FDD4FE848633" bold="true" box="[189,310,580,606]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9373110CA18855EFF73FDD4FE848633" bold="true" box="[189,310,580,606]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA18855EFF73FDE3FD248695" blockId="17.[189,1400,627,760]" pageId="17" pageNumber="18">
Subfamily of the family
<taxonomicName id="4C439681CA18855EFE2DFDE3FD2986E0" authorityName="Hyatt" authorityYear="1884" box="[483,667,627,653]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="family">Prionoceratidae</taxonomicName>
with the sutural formula E A L U I, adventive lobe V-shaped or lanceolate, pointed. Conch in the juvenile stage usually subinvolute, rarely subevolute; adult stage involute or subinvolute. Coiling rate usually very low or low (WER = 1.401.75). Shell ornament with fine to coarse growth lines, without ribs.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA18855EFF73FCB0FB0587B9" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BFCED02CA18855EFF73FCB0FE3A8757" blockId="17.[189,392,800,826]" box="[189,392,800,826]" pageId="17" pageNumber="18">
<heading id="D0B45A6ECA18855EFF73FCB0FE3A8757" bold="true" box="[189,392,800,826]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9373110CA18855EFF73FCB0FE3A8757" bold="true" box="[189,392,800,826]" pageId="17" pageNumber="18">Included genera</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA18855EFF73FCDFFB0587B9" blockId="17.[189,1398,847,980]" pageId="17" pageNumber="18">
<taxonomicName id="4C439681CA18855EFF73FCDFFE408704" authority="Hyatt, 1884" authorityName="Hyatt" authorityYear="1884" box="[189,498,847,873]" class="Cephalopoda" family="Prionoceratidae" genus="Prionoceras" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFF73FCDFFEFC8705" box="[189,334,847,872]" italics="true" pageId="17" pageNumber="18">Prionoceras</emphasis>
Hyatt, 1884
</taxonomicName>
;
<taxonomicName id="4C439681CA18855EFDC8FCDFFCD38704" authority="Cossmann 1900" authorityName="Cossmann" authorityYear="1900" box="[518,865,847,873]" class="Cephalopoda" genus="Haugiceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFDC8FCDFFD3D8705" box="[518,655,847,872]" italics="true" pageId="17" pageNumber="18">Haugiceras</emphasis>
Cossmann 1900
</taxonomicName>
[synonym of
<taxonomicName id="4C439681CA18855EFBD5FCDFFB1E8705" authorityName="Hyatt" authorityYear="1884" box="[1051,1196,847,872]" class="Cephalopoda" family="Prionoceratidae" genus="Prionoceras" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFBD5FCDFFB1E8705" box="[1051,1196,847,872]" italics="true" pageId="17" pageNumber="18">Prionoceras</emphasis>
</taxonomicName>
];
<taxonomicName id="4C439681CA18855EFB02FCDFFECF87E1" authority="Wedekind, 1913" authorityName="Wedekind" authorityYear="1913" class="Cephalopoda" family="Prionoceratidae" genus="Postprolobites" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFB02FCDFFAC48704" box="[1228,1398,847,873]" italics="true" pageId="17" pageNumber="18">Postprolobites</emphasis>
Wedekind, 1913
</taxonomicName>
[synonym of
<taxonomicName id="4C439681CA18855EFDEDFCE3FD0187E1" authorityName="Hyatt" authorityYear="1884" box="[547,691,883,908]" class="Cephalopoda" family="Prionoceratidae" genus="Prionoceras" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFDEDFCE3FD0187E1" box="[547,691,883,908]" italics="true" pageId="17" pageNumber="18">Prionoceras</emphasis>
</taxonomicName>
];
<taxonomicName id="4C439681CA18855EFD03FCE2FB3387E1" authority="Schindewolf, 1924" authorityName="Schindewolf" authorityYear="1924" box="[717,1153,882,908]" class="Cephalopoda" family="Prionoceratidae" genus="Paragattendorfia" kingdom="Animalia" order="Goniatitina" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFD03FCE2FC2587E1" box="[717,919,882,908]" italics="true" pageId="17" pageNumber="18">Paragattendorfia</emphasis>
<bibRefCitation id="EFD290F3CA18855EFC51FCE2FB3387E1" author="Schindewolf O. H." box="[927,1153,882,908]" pageId="17" pageNumber="18" pagination="95 - 113" refId="ref117232" refString="Schindewolf O. H. 1924. Bermerkungen zur Stratigraphie und Ammoneenfauna des Saalfelder Oberdevons. Senckenbergiana 6: 95 - 113." type="journal article" year="1924">Schindewolf, 1924</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA18855EFB40FCE3FF4A87DD" authority="Weyer, 1972" authorityName="Weyer" authorityYear="1972" class="Cephalopoda" family="Prionoceratidae" genus="Cunitoceras" higherTaxonomySource="CoL" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFB40FCE3FAAF87E0" box="[1166,1309,883,909]" italics="true" pageId="17" pageNumber="18">Cunitoceras</emphasis>
<bibRefCitation id="EFD290F3CA18855EFAEAFCE3FF4A87DD" author="Weyer D." pageId="17" pageNumber="18" pagination="325 - 350" refId="ref118335" refString="Weyer D. 1972. Zum Alter der Ammonoideen-Faunen des Marshall-Sandsteins (Unterkarbon; Michigan, USA). Berichte der deutschen Gesellschaft der geologischen Wissenschaften, A, Geologie, Palaontologie 17: 325 - 350." type="journal article" year="1972">Weyer, 1972</bibRefCitation>
</taxonomicName>
[problematic genus];
<taxonomicName id="4C439681CA18855EFDC6FC07FCE687DD" authority="Korn, 1988" authorityName="Korn" authorityYear="1988" box="[520,852,918,944]" class="Cephalopoda" family="Prionoceratidae" genus="Mimimitoceras" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFDC6FC07FD0887DD" box="[520,698,919,944]" italics="true" pageId="17" pageNumber="18">Mimimitoceras</emphasis>
Korn, 1988
</taxonomicName>
;
<taxonomicName id="4C439681CA18855EFCABFC06FB0787DD" authority="Korn, 1993" authorityName="Korn" authorityYear="1993" box="[869,1205,918,944]" class="Cephalopoda" family="Prionoceratidae" genus="Globimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFCABFC06FBAE87DD" box="[869,1052,918,944]" italics="true" pageId="17" pageNumber="18">Globimitoceras</emphasis>
<bibRefCitation id="EFD290F3CA18855EFBE6FC06FB0787DD" author="Korn D." box="[1064,1205,918,944]" pageId="17" pageNumber="18" pagination="581 - 593" refId="ref114371" refString="Korn D. 1993. The ammonoid faunal change near the Devonian - Carboniferous boundary. Annales de la Societe geologique de Belgique 115: 581 - 593." type="journal article" year="1993">Korn, 1993</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA18855EFB08FC07FEE087BE" authority="Becker 1996" authorityName="Becker" authorityYear="1996" class="Cephalopoda" genus="Rectimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFB08FC07FAC487DD" box="[1222,1398,919,944]" italics="true" pageId="17" pageNumber="18">Rectimitoceras</emphasis>
<bibRefCitation id="EFD290F3CA18855EFF73FC29FEE087BE" author="Becker R. T." box="[189,338,953,979]" pageId="17" pageNumber="18" pagination="19 - 45" refId="ref111308" refString="Becker R. T. 1996. New faunal records and holostratigraphic correlation of the Hasselbachtal D / C- Boundary Auxiliary Stratotype (Germany). Annales de la Societe geologique de Belgique 117: 19 - 45." type="journal article" year="1996">Becker 1996</bibRefCitation>
</taxonomicName>
[synonym of
<taxonomicName id="4C439681CA18855EFE38FC2AFD1A87BE" authorityName="Korn" authorityYear="1988" box="[502,680,954,979]" class="Cephalopoda" family="Prionoceratidae" genus="Mimimitoceras" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFE38FC2AFD1A87BE" box="[502,680,954,979]" italics="true" pageId="17" pageNumber="18">Mimimitoceras</emphasis>
</taxonomicName>
];
<taxonomicName id="4C439681CA18855EFD0FFC2AFB0187B9" authority="Ebbighausen &amp; Bockwinkel, 2007" authorityName="Ebbighausen &amp; Bockwinkel" authorityYear="2007" box="[705,1203,953,980]" class="Cephalopoda" family="Prionoceratidae" genus="Kornia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitina" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFD0FFC2AFCA687BE" box="[705,788,954,979]" italics="true" pageId="17" pageNumber="18">Kornia</emphasis>
<bibRefCitation id="EFD290F3CA18855EFCD5FC29FB0187B9" author="Ebbighausen V. &amp; Bockwinkel J." box="[795,1203,953,980]" pageId="17" pageNumber="18" pagination="125 - 163" refId="ref112667" refString="Ebbighausen V. &amp; Bockwinkel J. 2007. Tournaisian (Early Carboniferous / Mississippian) ammonoids from the Ma'der Basin (Anti-Atlas, Morocco). Fossil Record 10: 125 - 163. https: // doi. org / 10.1002 / mmng. 200700003" type="journal article" year="2007">Ebbighausen &amp; Bockwinkel, 2007</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA18855EFF73FC6BFCB58360" pageId="17" pageNumber="18" type="description">
<paragraph id="8BFCED02CA18855EFF73FC6BFEEA8078" blockId="17.[189,344,1019,1045]" box="[189,344,1019,1045]" pageId="17" pageNumber="18">
<heading id="D0B45A6ECA18855EFF73FC6BFEEA8078" bold="true" box="[189,344,1019,1045]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9373110CA18855EFF73FC6BFEEA8078" bold="true" box="[189,344,1019,1045]" pageId="17" pageNumber="18">Morphology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA18855EFF73FBBAFC6A8177" blockId="17.[189,1399,1066,1306]" pageId="17" pageNumber="18">
The species of the subfamily
<taxonomicName id="4C439681CA18855EFDD7FBBAFD638029" authorityName="Hyatt" authorityYear="1884" box="[537,721,1066,1092]" class="Cephalopoda" family="Prionoceratinae" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
are characterised by a simple conch geometry; the conch is sometimes discoidal but more often pachyconic to globular with a nearly closed or closed umbilicus and almost always a very low or low coiling rate. This morphology is shared by the two main genera
<taxonomicName id="4C439681CA18855EFF71FB06FEE280C2" authorityName="Hyatt" authorityYear="1884" box="[191,336,1174,1199]" class="Cephalopoda" family="Prionoceratidae" genus="Prionoceras" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFF71FB06FEE280C2" box="[191,336,1174,1199]" italics="true" pageId="17" pageNumber="18">Prionoceras</emphasis>
</taxonomicName>
and
<taxonomicName id="4C439681CA18855EFE40FB06FDF280C2" authorityName="Korn" authorityYear="1988" box="[398,576,1174,1199]" class="Cephalopoda" family="Prionoceratidae" genus="Mimimitoceras" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFE40FB06FDF280C2" box="[398,576,1174,1199]" italics="true" pageId="17" pageNumber="18">Mimimitoceras</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="EFD290F3CA18855EFD5EFB06FCEF80C2" author="Korn D. &amp; Bockwinkel J. &amp; Ebbighausen V." box="[656,861,1173,1200]" pageId="17" pageNumber="18" pagination="167 - 204" refId="ref115642" refString="Korn D., Bockwinkel J. &amp; Ebbighausen V. 2014. Middle Famennian (Late Devonian) ammonoids from the Anti-Atlas of Morocco. 1. Prionoceras. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 272: 167 - 204. https: // doi. org / 10.1127 / 0077 - 7749 / 2014 / 0405" type="journal article" year="2014">
Korn
<emphasis id="B9373110CA18855EFD19FB06FCA780C2" box="[727,789,1173,1199]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2014
</bibRefCitation>
, 2015). Another common feature of the two genera are the shell constrictions, which are present in all species at least in the early stage of ontogeny. Almost all species have a shell ornament consisting only of growth lines; these almost always have a convex course on the flank and form a broad, shallow ventral sinus.
</paragraph>
<paragraph id="8BFCED02CA18855EFF73FAD2FAC581AB" blockId="17.[188,1401,1346,1479]" pageId="17" pageNumber="18">
The suture line also shows little variation. It consists of the basic elements E A L U I; the shape of the external and adventive lobe varies between species. Usually, species with a discoidal conch show a narrower, sometimes lanceolate external lobe, while in globular species it is often broader and V-shaped. A deviation from this rule, however, is
<taxonomicName id="4C439681CA18855EFD94FA3CFCBF81AB" authorityName="Korn" authorityYear="1993" box="[602,781,1452,1478]" class="Cephalopoda" family="Prionoceratidae" genus="Globimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA18855EFD94FA3CFCBF81AB" box="[602,781,1452,1478]" italics="true" pageId="17" pageNumber="18">Globimitoceras</emphasis>
</taxonomicName>
with a globular conch but a very narrow external lobe.
</paragraph>
<paragraph id="8BFCED02CA18855EFF73FA7EFE858265" blockId="17.[189,311,1518,1544]" box="[189,311,1518,1544]" pageId="17" pageNumber="18">
<heading id="D0B45A6ECA18855EFF73FA7EFE858265" bold="true" box="[189,311,1518,1544]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9373110CA18855EFF73FA7EFE858265" bold="true" box="[189,311,1518,1544]" pageId="17" pageNumber="18">Ontogeny</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA18855EFF73F98DFCB58360" blockId="17.[188,1400,1565,1805]" pageId="17" pageNumber="18">
The species of the subfamily underwent a comparatively simple ontogenetic development; this means that the morphology of juveniles and adults does not differ markedly. Conch allometry is usually weakly developed; the ontogenetic trajectories (
<bibRefCitation id="EFD290F3CA18855EFD5EF9F4FCA38213" author="Korn D." box="[656,785,1636,1662]" pageId="17" pageNumber="18" pagination="501 - 514" refId="ref114700" refString="Korn D. 2012. Quantification of ontogenetic allometry in ammonoids. Evolution &amp; Development 14: 501 - 514. https: // doi. org / 10.1111 / ede. 12003" type="journal article" year="2012">Korn 2012</bibRefCitation>
) are often monophasic with a simple decrease of the ww/dm ratio, for example. These simple ontogenetic trajectories are mainly caused by the small width of the umbilicus in the juvenile stage. Therefore, the whorl profiles are rather similar at all growth stages and range from C-shaped to horseshoe-shaped. Only the trend towards more slender conchs that is present in most Palaeozoic ammonoids is evident.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA18855DFF73F8A5FB598709" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BFCED02CA18855EFF73F8A5FE8D8322" blockId="17.[189,319,1845,1871]" box="[189,319,1845,1871]" pageId="17" pageNumber="18">
<heading id="D0B45A6ECA18855EFF73F8A5FE8D8322" bold="true" box="[189,319,1845,1871]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9373110CA18855EFF73F8A5FE8D8322" bold="true" box="[189,319,1845,1871]" pageId="17" pageNumber="18">Phylogeny</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA18855DFF73F8F3FB578507" blockId="17.[189,1399,1891,2024]" lastBlockId="18.[189,1399,265,363]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
The
<taxonomicName id="4C439681CA18855EFF39F8F3FE1D8310" authorityName="Hyatt" authorityYear="1884" box="[247,431,1891,1917]" class="Cephalopoda" family="Prionoceratinae" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="116" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
are the ancestral subfamily of the family
<taxonomicName id="4C439681CA18855EFC71F8F3FBC58310" authorityName="Hyatt" authorityYear="1884" box="[959,1143,1891,1917]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitida" pageId="17" pageNumber="18" phylum="Mollusca" rank="family">Prionoceratidae</taxonomicName>
and thus of all post Devonian ammonoids. The prionoceratids of the Famennian formed only one previously known side branch, the subfamily
<taxonomicName id="4C439681CA18855EFDC5F83BFDC983A8" box="[523,635,1963,1989]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="subFamily" subFamily="Balviinae">Balviinae</taxonomicName>
with its paedomorphic conchs (
<bibRefCitation id="EFD290F3CA18855EFBC8F83BFB2B83A8" author="Korn D." box="[1030,1177,1963,1989]" pageId="17" pageNumber="18" pagination="21 - 36" refId="ref114266" refString="Korn D. 1992 a. Heterochrony in the evolution of Late Devonian Ammonoids. Acta Palaeontologica Polonica 37: 21 - 36." type="journal article" year="1992">Korn 1992a</bibRefCitation>
,
<bibRefCitation id="EFD290F3CA18855EFB64F83BFB4683AB" author="Korn D." box="[1194,1268,1963,1990]" pageId="17" pageNumber="18" pagination="245 - 260" refId="ref114430" refString="Korn D. 1995 a. Impact of environmental perturbations on heterochronic development in Palaeozoic ammonoids. In: McNamara K. J. (ed.) Evolutionary Change and Heterochrony: 245 - 260. Wiley, Chichester." type="book chapter" year="1995">1995a</bibRefCitation>
,
<bibRefCitation id="EFD290F3CA18855EFACAF83BFAFD83AB" author="Korn D." box="[1284,1359,1963,1990]" pageId="17" pageNumber="18" pagination="155 - 165" refId="ref114471" refString="Korn D. 1995 b. Paedomorphosis of ammonoids as a result of sealevel fluctuations in the Late Devonian Wocklumeria Stufe. Lethaia 28: 155 - 165. https: // doi. org / 10.1111 / j. 1502 - 3931.1995. tb 01606. x" type="journal article" year="1995">1995b</bibRefCitation>
) is distinguished by divergent morphologies. Several side branches formed in the earliest Carboniferous. The most successful of these is the subfamily
<taxonomicName id="4C439681CA1B855DFD17FE99FC0F854E" authorityName="Korn" authorityYear="1994" box="[729,957,265,291]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitina" pageId="18" pageNumber="19" phylum="Mollusca" rank="subFamily" subFamily="Acutimitoceratinae">Acutimitoceratinae</taxonomicName>
, described in more detail below. Less successful side branches are the genera
<taxonomicName id="4C439681CA1B855DFD60FEBDFCD7852A" authorityName="Korn" authorityYear="1993" box="[686,869,301,327]" class="Cephalopoda" family="Prionoceratidae" genus="Globimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA1B855DFD60FEBDFCD7852A" box="[686,869,301,327]" italics="true" pageId="18" pageNumber="19">Globimitoceras</emphasis>
</taxonomicName>
,
<taxonomicName id="4C439681CA1B855DFCB6FEBDFC79852B" authorityName="Ebbighausen &amp; Bockwinkel" authorityYear="2007" box="[888,971,301,326]" class="Cephalopoda" family="Prionoceratidae" genus="Kornia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitina" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA1B855DFCB6FEBDFC79852B" box="[888,971,301,326]" italics="true" pageId="18" pageNumber="19">Kornia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C439681CA1B855DFBC1FEBDFB6B852A" authorityName="Schindewolf" authorityYear="1924" box="[1039,1241,301,327]" class="Cephalopoda" family="Prionoceratidae" genus="Paragattendorfia" kingdom="Animalia" order="Goniatitina" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA1B855DFBC1FEBDFB6B852A" box="[1039,1241,301,327]" italics="true" pageId="18" pageNumber="19">Paragattendorfia</emphasis>
</taxonomicName>
, all of them restricted to the Early and Middle Tournaisian, but apparently extinct without descendants.
</paragraph>
<paragraph id="8BFCED02CA1B855DFF73FE02FE4485C0" blockId="18.[189,502,402,429]" box="[189,502,402,429]" pageId="18" pageNumber="19">
<heading id="D0B45A6ECA1B855DFF73FE02FE4485C0" bold="true" box="[189,502,402,429]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9373110CA1B855DFF73FE02FE4485C0" bold="true" box="[189,502,402,429]" pageId="18" pageNumber="19">Stratigraphic occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA1B855DFF73FE51FB598709" blockId="18.[188,1400,449,869]" pageId="18" pageNumber="19">
The subfamily
<taxonomicName id="4C439681CA1B855DFEA2FE51FD9085B6" authorityName="Hyatt" authorityYear="1884" box="[364,546,449,475]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="19" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
has its main distribution in the middle and late Famennian, where several species can be used as index species (
<bibRefCitation id="EFD290F3CA1B855DFDA5FE75FC9F8592" author="Korn D. &amp; Bockwinkel J. &amp; Ebbighausen V." box="[619,813,485,511]" pageId="18" pageNumber="19" pagination="167 - 204" refId="ref115642" refString="Korn D., Bockwinkel J. &amp; Ebbighausen V. 2014. Middle Famennian (Late Devonian) ammonoids from the Anti-Atlas of Morocco. 1. Prionoceras. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 272: 167 - 204. https: // doi. org / 10.1127 / 0077 - 7749 / 2014 / 0405" type="journal article" year="2014">
Korn
<emphasis id="B9373110CA1B855DFD7EFE76FD588592" box="[688,746,485,511]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2014
</bibRefCitation>
, 2015) and help to establish a stratigraphic scheme that complements the clymeniid-based zonation.According to current knowledge, the
<taxonomicName id="4C439681CA1B855DFB59FD99FAFF864E" authorityName="Hyatt" authorityYear="1884" box="[1175,1357,521,547]" class="Cephalopoda" family="Prionoceratinae" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="116" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
are one of the few ammonoid lineages that survived the Hangenberg Event at the DevonianCarboniferous boundary (
<bibRefCitation id="EFD290F3CA1B855DFEF5FDC1FE0F8607" author="Korn D." box="[315,445,592,619]" pageId="18" pageNumber="19" pagination="49 - 54" refId="ref114032" refString="Korn D. 1986. Ammonoid evolution in late Famennian and early Tournaisian. Annales de la Societe geologique de Belgique 109: 49 - 54." type="journal article" year="1986">Korn 1986</bibRefCitation>
, 1993, 2000; Becker 1993;
<bibRefCitation id="EFD290F3CA1B855DFCCCFDC0FC088606" author="Kullmann J." box="[770,954,592,619]" pageId="18" pageNumber="19" pagination="169 - 180" refId="ref115839" refString="Kullmann J. 2000. Ammonoid turnover at the Devonian - Carboniferous boundary. Revue de Paleobiologie, Volume special 8: 169 - 180." type="journal article" year="2000">Kullmann 2000</bibRefCitation>
). In contrast to the “failed survivors”, the cymaclymeniids, which survived the biocrisis with few forms but became extinct shortly afterwards (
<bibRefCitation id="EFD290F3CA1B855DFF0BFD08FEFA86DF" author="Korn D." box="[197,328,664,690]" pageId="18" pageNumber="19" pagination="371 - 372" refId="ref114164" refString="Korn D. 1990. Cymaclymenia aus der Acutimitoceras - Fauna (prorsum - Zone) vom Mussenberg (Devon / Karbon-Grenze; Rheinisches Schiefergebirge). Bulletin de la Societe belge de Geologie 98: 371 - 372." type="journal article" year="1990">Korn 1990</bibRefCitation>
; Korn
<emphasis id="B9373110CA1B855DFE59FD09FE6086DF" box="[407,466,664,690]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2004), the evolution of the prionoceratids is, in contrast to the cymaclymeniids, not a “dead clade walking” (
<bibRefCitation id="EFD290F3CA1B855DFDCEFD2BFD0386BB" author="Jablonski D." box="[512,689,699,726]" pageId="18" pageNumber="19" pagination="8139 - 8144" refId="ref113361" refString="Jablonski D. 2002. Survival without recovery after mass extinctions. Proceedings of the National Academy of Sciences of the United States of America 99: 8139 - 8144. https: // doi. org / 10.1073 / pnas. 102163299" type="journal article" year="2002">Jablonski 2002</bibRefCitation>
), but the evolution of a group with a very successful radiation in the Early Carboniferous. However, this successful radiation was caused by flourishing of the subfamily
<taxonomicName id="4C439681CA1B855DFF73FC93FE2C8770" authorityName="Korn" authorityYear="1994" box="[189,414,771,797]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitina" pageId="18" pageNumber="19" phylum="Mollusca" rank="subFamily" subFamily="Acutimitoceratinae">Acutimitoceratinae</taxonomicName>
, while the genera of the
<taxonomicName id="4C439681CA1B855DFD2DFC93FC2B8770" authorityName="Hyatt" authorityYear="1884" box="[739,921,771,797]" class="Cephalopoda" family="Prionoceratinae" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="116" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
have only a short stratigraphic range, apparently restricted to the early and middle Tournaisian. The late Tournaisian record of the problematic species
<taxonomicName id="4C439681CA1B855DFED6FCDBFD658708" authority="Weyer, 1972" authorityName="Weyer" authorityYear="1972" box="[280,727,842,869]" class="Cephalopoda" family="Prionoceratidae" genus="Cunitoceras" higherTaxonomySource="CoL" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="schindewolfi">
<emphasis id="B9373110CA1B855DFED6FCDBFD8C8709" box="[280,574,842,869]" italics="true" pageId="18" pageNumber="19">Cunitoceras schindewolfi</emphasis>
<bibRefCitation id="EFD290F3CA1B855DFD8BFCDBFD658708" author="Weyer D." box="[581,727,843,869]" pageId="18" pageNumber="19" pagination="325 - 350" refId="ref118335" refString="Weyer D. 1972. Zum Alter der Ammonoideen-Faunen des Marshall-Sandsteins (Unterkarbon; Michigan, USA). Berichte der deutschen Gesellschaft der geologischen Wissenschaften, A, Geologie, Palaontologie 17: 325 - 350." type="journal article" year="1972">
Weyer,
<quantity id="4CBB40E7CA1B855DFD53FCDBFD658708" box="[669,727,843,869]" metricMagnitude="1" metricUnit="m" metricValue="5.00888" pageId="18" pageNumber="19" unit="in" value="1972.0">1972</quantity>
</bibRefCitation>
</taxonomicName>
in the Harz Mountains requires confirmation.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA1B855DFF73FC1CFAFD80BD" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BFCED02CA1B855DFF73FC1CFE5387CA" blockId="18.[189,481,908,935]" box="[189,481,908,935]" pageId="18" pageNumber="19">
<heading id="D0B45A6ECA1B855DFF73FC1CFE5387CA" bold="true" box="[189,481,908,935]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9373110CA1B855DFF73FC1CFE5387CA" bold="true" box="[189,481,908,935]" pageId="18" pageNumber="19">Geographic occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA1B855DFF73FC2BFAFD80BD" blockId="18.[186,1399,955,1232]" pageId="18" pageNumber="19">
The
<taxonomicName id="4C439681CA1B855DFF21FC2BFE1587B8" authorityName="Hyatt" authorityYear="1884" box="[239,423,955,981]" class="Cephalopoda" family="Prionoceratinae" kingdom="Animalia" order="Goniatitida" pageId="18" pageNumber="116" phylum="Mollusca" rank="subFamily" subFamily="Prionoceratinae">Prionoceratinae</taxonomicName>
are a subfamily with an almost global distribution in the Middle and Late Famennian; in the Early and Middle Tournaisian the distribution is significantly restricted. The most important occurrences in Europe are in the Rhenish Mountains (
<bibRefCitation id="EFD290F3CA1B855DFC98FB93FBA88070" author="Vohringer E." box="[854,1050,1027,1053]" pageId="18" pageNumber="19" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer 1960</bibRefCitation>
; Korn 1994;
<bibRefCitation id="EFD290F3CA1B855DFB0FFB94FF4B802C" author="Korn D. &amp; Weyer D." pageId="18" pageNumber="19" pagination="79 - 124" refId="ref115026" refString="Korn D. &amp; Weyer D. 2003. High resolution stratigraphy of the Devonian - Carboniferous transitional beds in the Rhenish Mountains. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Geowissenschaftliche Reihe 6: 79 - 124. https: // doi. org / 10.1002 / mmng. 4860060105" type="journal article" year="2003">Korn &amp; Weyer 2003</bibRefCitation>
), Franconia (
<bibRefCitation id="EFD290F3CA1B855DFE51FBB7FDCD802C" author="Schindewolf O. H." box="[415,639,1063,1089]" pageId="18" pageNumber="19" pagination="393 - 509" refId="ref117178" refString="Schindewolf O. H. 1923. Beitrage zur Kenntnis des Palaozoikums in Oberfranken, Ostthuringen und dem Sachsischen Vogtlande. I. Stratigraphie und Ammoneenfauna des Oberdevons von Hof a. S. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, Beilage-Band 49: 250 - 357, 393 - 509." type="journal article" year="1923">Schindewolf 1923</bibRefCitation>
; Korn 1994),
<collectingRegion id="498723E0CA1B855DFCFFFBB7FC17802C" box="[817,933,1063,1089]" country="Germany" name="Thueringen" pageId="18" pageNumber="19">Thuringia</collectingRegion>
(
<bibRefCitation id="EFD290F3CA1B855DFC76FBB7FBFD802C" author="Weyer D." box="[952,1103,1063,1089]" pageId="18" pageNumber="19" pagination="167 - 185" refId="ref118397" refString="Weyer D. 1977. Ammonoideen aus dem Untertournai von Schleiz (Ostthuringisches Schiefergebirge). Zeitschrift fur geologische Wissenschaften 5: 167 - 185." type="journal article" year="1977">Weyer 1977</bibRefCitation>
), questionably the Harz Mountains (
<bibRefCitation id="EFD290F3CA1B855DFE85FBDAFD948008" author="Schindewolf O. H." box="[331,550,1098,1125]" pageId="18" pageNumber="19" pagination="23 - 116" refId="ref117369" refString="Schindewolf O. H. 1951. Uber ein neues Vorkommen unterkarbonischer Pericyclus - Schichten im Oberharz. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 93: 23 - 116." type="journal article" year="1951">Schindewolf 1951</bibRefCitation>
),
<collectingRegion id="498723E0CA1B855DFDF0FBDAFD3E8009" box="[574,652,1098,1124]" country="Poland" name="Slaskie" pageId="18" pageNumber="19">Silesia</collectingRegion>
(
<bibRefCitation id="EFD290F3CA1B855DFD50FBDAFCAE8008" author="Dzik J." box="[670,796,1098,1125]" pageId="18" pageNumber="19" pagination="57 - 170" refId="ref112635" refString="Dzik J. 1997. Emergence and succession of Carboniferous conodont and ammonoid communities in the Polish part of the Variscan sea. Acta Palaeontologica Polonica 42: 57 - 170." type="journal article" year="1997">Dzik 1997</bibRefCitation>
), the Carnic Alps (
<bibRefCitation id="EFD290F3CA1B855DFC31FBDBFB238009" author="Korn D." box="[1023,1169,1098,1125]" pageId="18" pageNumber="19" pagination="7 - 19" refId="ref114290" refString="Korn D. 1992 b. Ammonoideen aus dem Devon / Karbon-Grenzprofil an der Grunen Schneid (Karnische Alpen, Osterreich). Jahrbuch der Geologischen Bundesanstalt 135: 7 - 19." type="journal article" year="1992">Korn 1992b</bibRefCitation>
) and the Montagne Noire (
<bibRefCitation id="EFD290F3CA1B855DFEC0FBFEFDA780E5" author="Becker R. T. &amp; Weyer D." box="[270,533,1134,1161]" pageId="18" pageNumber="19" pagination="11 - 36" refId="ref111430" refString="Becker R. T. &amp; Weyer D. 2004. Bartzschiceras n. gen. (Ammonoidea) from the Lower Tournaisian of Southern France. Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg 88: 11 - 36." type="journal article" year="2004">Becker &amp; Weyer 2004</bibRefCitation>
;
<bibRefCitation id="EFD290F3CA1B855DFDEEFBFFFD4C80E5" author="Korn D. &amp; Feist R." box="[544,766,1134,1161]" pageId="18" pageNumber="19" pagination="99 - 124" refId="ref114786" refString="Korn D. &amp; Feist R. 2007. Early Carboniferous ammonoid faunas and stratigraphy of the Montagne Noire (France). Fossil Record 10: 99 - 124. https: // doi. org / 10.5194 / fr- 10 - 99 - 2007" type="journal article" year="2007">Korn &amp; Feist 2007</bibRefCitation>
). Early Carboniferous species are also known from the Anti-Atlas (
<bibRefCitation id="EFD290F3CA1B855DFE86FB02FD5280C1" author="Bockwinkel J. &amp; Ebbighausen V." box="[328,736,1170,1196]" pageId="18" pageNumber="19" pagination="87 - 129" refId="ref112039" refString="Bockwinkel J. &amp; Ebbighausen V. 2006. A new ammonoid fauna from the Gattendorfia - Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco). Fossil Record 9: 87 - 129. https: // doi. org / 10.5194 / fr- 9 - 87 - 2006" type="journal article" year="2006">Bockwinkel &amp; Ebbighausen 2006</bibRefCitation>
), the South Urals (Popov 1975),
<collectingRegion id="498723E0CA1B855DFBA2FB02FB6380C1" box="[1132,1233,1170,1196]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou</collectingRegion>
(
<bibRefCitation id="EFD290F3CA1B855DFB2AFB02FAD480C1" author="Ruan Y." box="[1252,1382,1170,1196]" pageId="18" pageNumber="19" pagination="1 - 152" refId="ref116988" refString="Ruan Y. 1981. Devonian and earliest Carboniferous Ammonoids from Guangxi and Guizhou. Memoires of Nanjing Institute of Geology and Paleontology, Academia Sinica 15: 1 - 152." type="journal article" year="1981">Ruan 1981</bibRefCitation>
), Karaganda (
<bibRefCitation id="EFD290F3CA1B855DFE85FB26FDBE80BD" author="Librovitch L. S." box="[331,524,1206,1232]" pageId="18" pageNumber="19" pagination="1 - 395" refId="ref116035" refString="Librovitch L. S. 1940. Ammonoidea iz kamennougolnykh otlozheniy Severnogo Kazakhstana. Paleontologiya SSSR 4: 1 - 395." type="journal article" year="1940">Librovitch 1940</bibRefCitation>
) and (questionably)
<collectingRegion id="498723E0CA1B855DFD31FB26FCDD80BD" box="[767,879,1206,1232]" country="United States of America" name="Michigan" pageId="18" pageNumber="19">Michigan</collectingRegion>
(
<bibRefCitation id="EFD290F3CA1B855DFCB0FB26FB8280BD" author="Winchell A." box="[894,1072,1206,1232]" pageId="18" pageNumber="19" pagination="352 - 366" refId="ref118520" refString="Winchell A. 1862. Notice of the rocks lying between the Carboniferous limestone of the Lower Peninsula of Michigan and the limestones of the Hamilton group; with descriptions of some cephalopods supposed to be new to science. American Journal of Science and Arts 33: 352 - 366. https: // doi. org / 10.2475 / ajs. s 2 - 33.99.352" type="journal article" year="1862">Winchell 1862</bibRefCitation>
;
<bibRefCitation id="EFD290F3CA1B855DFBF2FB26FA8D80BD" author="Miller A. K. &amp; Garner H. F." box="[1084,1343,1205,1232]" pageId="18" pageNumber="19" pagination="113 - 173" refId="ref116190" refString="Miller A. K. &amp; Garner H. F. 1955. Lower Mississippian cephalopods if Michigan. Part III. Ammonoids and summary. Contributions of the Museum of Paleontology, University of Michigan 12: 113 - 173." type="journal article" year="1955">Miller &amp; Garner 1955</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>