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<document id="7E08FA137EE532DDBD70E12A2D4B7370" ID-CLB-Dataset="22297" ID-DOI="10.5852/ejt.2018.455" ID-GBIF-Dataset="44a2fcf6-cc1b-4c07-9554-dd6dd279a76d" ID-Zenodo-Dep="3817707" ID-ZooBank="C445E95B-446A-4601-AAA3-C1CCBAB627F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1588963536872" checkinUser="valdenar" docAuthor="Páll-Gergely, Barna" docDate="2018" docId="03CC87A06C158A1BFD82F9964479F842" docLanguage="en" docName="ejt-455_pall-gergely.pdf.imf" docOrigin="European Journal of Taxonomy 455" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chersaecia scabra Páll-Gergely, 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="61" masterDocId="FFF5FFD86C2C8A27FFF8FFC04709FFA9" masterDocTitle="Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="58" updateTime="1698839078568" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CD2287F2CAFEE91DB0DCBBBE94C20232">Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
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<mods:name id="0D5BE03DC5415C392A87AB32FEC8470C" type="personal">
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<mods:namePart id="35940D006AF3175106F38E624C246335">Páll-Gergely, Barna</mods:namePart>
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<mods:affiliation id="5D0E8CB70067A92EB3F676E1C8B8069C">Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Ottó Street 15, Budapest, H- 1022, Hungary.</mods:affiliation>
<mods:nameIdentifier id="CECAD688D4E85E00EABED09D150FB90E" type="email">pall-gergely.barna@agrar.mta.hu</mods:nameIdentifier>
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<mods:title id="EFF18F571DE968BE9C1CD7C4CD9CF46C">European Journal of Taxonomy</mods:title>
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<mods:date id="A1A063D4D3DB470636D554687F3283E0">2018</mods:date>
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<mods:number id="2216E24F45BB60EECF92CAC47DADB312">2018-08-16</mods:number>
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<treatment id="03CC87A06C158A1BFD82F9964479F842" ID-DOI="http://doi.org/10.5281/zenodo.3818792" ID-GBIF-Taxon="163993812" ID-Zenodo-Dep="3818792" LSID="urn:lsid:plazi:treatment:03CC87A06C158A1BFD82F9964479F842" httpUri="http://treatment.plazi.org/id/03CC87A06C158A1BFD82F9964479F842" lastPageId="60" lastPageNumber="61" pageId="57" pageNumber="58">
<subSubSection id="C37F653D6C158A1EFD82F99643DCF93F" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BDA36B66C158A1EFD82F99644B3F9D8" blockId="57.[350,1237,1622,1723]" box="[634,954,1622,1649]" pageId="57" pageNumber="58">
<heading id="D09281DA6C158A1EFD82F99644B3F9D8" box="[634,954,1622,1649]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="58" reason="2">
<taxonomicName id="4C654D356C158A1EFD82F9964451F9D8" ID-CoL="TY3Y" authorityName="Pall-Gergely" authorityYear="2018" box="[634,856,1622,1649]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C158A1EFD82F9964451F9D8" bold="true" box="[634,856,1622,1649]" italics="true" pageId="57" pageNumber="58">Chersaecia scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C158A1EFCA7F99744B3F9D8" box="[863,954,1623,1649]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BDA36B66C158A1EFEA6F9BC43DCF93F" blockId="57.[350,1237,1622,1723]" box="[350,1237,1660,1686]" pageId="57" pageNumber="58">
<uri id="FFF43AB46C158A1EFEA6F9BC43DCF93F" box="[350,1237,1660,1686]" pageId="57" pageNumber="58">
urn:lsid:zoobank.org:act:
<uuid id="FFC30C636C158A1EFD78F9BC43DCF93F" box="[640,1237,1660,1686]" pageId="57" pageNumber="58">B6A4AF8F-9A14-4CA1-AE9D-BDF4F8A8F77A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C158A1EFD99F96044DBF912" box="[609,978,1696,1723]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BDA36B66C158A1EFD99F96044DBF912" blockId="57.[350,1237,1622,1723]" box="[609,978,1696,1723]" pageId="57" pageNumber="58">
<figureCitation id="135E2A336C158A1EFD99F96045BFF913" box="[609,694,1696,1722]" captionStart="Fig" captionStartId="31.[190,233,1849,1875]" captionTargetBox="[189,1400,265,1808]" captionTargetId="figure@31.[189,1400,265,1808]" captionTargetPageId="31" captionText="Fig. 16. Radula of species of Chersaecia Gude, 1899. A, C, E. Central and first lateral teeth. B, D, F. Marginal teeth. AB. C. auffenbergi sp. nov. (NHMW MO-112 221) CD. C. perrierae (Gude, 1898) (NHMUK 20170148). EF. C. scabra sp. nov. (NHMUK 20170149). All images: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.3817739" httpUri="https://zenodo.org/record/3817739/files/figure.png" pageId="57" pageNumber="58">Figs 16</figureCitation>
EF, 24OP, 26AC, 27
</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C158A1EFF45F9234330F8FA" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BDA36B66C158A1EFF45F923463FF954" blockId="57.[189,310,1763,1789]" box="[189,310,1763,1789]" pageId="57" pageNumber="58">
<heading id="D09281DA6C158A1EFF45F923463FF954" bold="true" box="[189,310,1763,1789]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B911EAA46C158A1EFF45F923463FF954" bold="true" box="[189,310,1763,1789]" pageId="57" pageNumber="58">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C158A1EFF45F8D44330F8FA" blockId="57.[189,1398,1812,1876]" pageId="57" pageNumber="58">A small, sinistral, nearly-flat species with a weak apertural fold; parietal wall with two lamellae, and two equal horizontal plicae (main and intermediate) attached to anterior lamella.</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C158A1EFF45F8BC4542F845" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BDA36B66C158A1EFF45F8BC464DF83F" blockId="57.[189,324,1916,1942]" box="[189,324,1916,1942]" pageId="57" pageNumber="58">
<heading id="D09281DA6C158A1EFF45F8BC464DF83F" bold="true" box="[189,324,1916,1942]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B911EAA46C158A1EFF45F8BC464DF83F" bold="true" box="[189,324,1916,1942]" pageId="57" pageNumber="58">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C158A1EFF45F86C4542F845" blockId="57.[189,1399,1964,2028]" pageId="57" pageNumber="58">
The specific epithet refers to the fact that even the live collected specimens had an eroded shell surface (
<emphasis id="B911EAA46C158A1EFF3FF811461CF842" box="[199,277,2001,2027]" italics="true" pageId="57" pageNumber="58">scaber</emphasis>
/
<taxonomicName id="4C654D356C158A1EFEE5F8114664F842" authorityName="Pall-Gergely" authorityYear="2018" box="[285,365,2001,2027]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="scabra">
<emphasis id="B911EAA46C158A1EFEE5F8114664F842" box="[285,365,2001,2027]" italics="true" pageId="57" pageNumber="58">scabra</emphasis>
</taxonomicName>
: erroded in Latin).
</paragraph>
</subSubSection>
<caption id="DF1A663E6C168A1DFF45F8D64345F817" ID-DOI="http://doi.org/10.5281/zenodo.3817755" ID-Zenodo-Dep="3817755" httpUri="https://zenodo.org/record/3817755/files/figure.png" pageId="58" pageNumber="59" startId="58.[189,232,1814,1840]" targetBox="[189,1399,265,1774]" targetPageId="58">
<paragraph id="8BDA36B66C168A1DFF45F8D64345F817" blockId="58.[189,1399,1814,1982]" pageId="58" pageNumber="59">
<emphasis id="B911EAA46C168A1DFF45F8D64614F899" bold="true" box="[189,285,1814,1840]" pageId="58" pageNumber="59">Fig. 26.</emphasis>
Shells of species of
<taxonomicName id="4C654D356C168A1DFDE6F8D64437F899" authority="Gude, 1899" authorityName="Gude" authorityYear="1899" box="[542,830,1814,1840]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="genus">
<emphasis id="B911EAA46C168A1DFDE6F8D645ADF899" box="[542,676,1814,1840]" italics="true" pageId="58" pageNumber="59">Chersaecia</emphasis>
Gude, 1899
</taxonomicName>
.
<emphasis id="B911EAA46C168A1DFCB4F8D64468F899" bold="true" box="[844,865,1814,1840]" pageId="58" pageNumber="59">A</emphasis>
.
<taxonomicName id="4C654D356C168A1DFC8BF8D744E0F899" authorityName="Pall-Gergely" authorityYear="2018" box="[883,1001,1814,1841]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C168A1DFC8BF8D744E0F899" box="[883,1001,1814,1841]" italics="true" pageId="58" pageNumber="59">C. scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C168A1DFC0BF8D7435AF898" box="[1011,1107,1815,1841]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (UF 380244).
<emphasis id="B911EAA46C168A1DFF45F8FA47D8F8FD" bold="true" box="[189,209,1850,1876]" pageId="58" pageNumber="59">B</emphasis>
. Aperture of
<taxonomicName id="4C654D356C168A1DFE88F8FA46EAF8FD" authorityName="Pall-Gergely" authorityYear="2018" box="[368,483,1850,1876]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C168A1DFE88F8FA46EAF8FD" box="[368,483,1850,1876]" italics="true" pageId="58" pageNumber="59">C. scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C168A1DFE13F8FA4540F8FD" box="[491,585,1850,1876]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (NHMUK 20170149).
<emphasis id="B911EAA46C168A1DFC2AF8F944EEF8FA" bold="true" box="[978,999,1849,1875]" pageId="58" pageNumber="59">C</emphasis>
. Protoconch of
<taxonomicName id="4C654D356C168A1DFB5EF8FA4210F8FD" authorityName="Pall-Gergely" authorityYear="2018" box="[1190,1305,1850,1876]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C168A1DFB5EF8FA4210F8FD" box="[1190,1305,1850,1876]" italics="true" pageId="58" pageNumber="59">C. scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C168A1DFAD9F8FA427EF8FD" box="[1313,1399,1850,1876]" pageId="58" pageNumber="59" rank="species">sp. nov</taxonomicNameLabel>
(UF 380244).
<emphasis id="B911EAA46C168A1DFE96F89D468AF8DE" bold="true" box="[366,387,1885,1911]" pageId="58" pageNumber="59">D</emphasis>
.
<taxonomicName id="4C654D356C168A1DFE6FF89D4405F8DE" authority="(Stoliczka, 1873)" baseAuthorityName="Stoliczka" baseAuthorityYear="1873" box="[407,780,1885,1911]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="shanensis">
<emphasis id="B911EAA46C168A1DFE6FF89D4538F8DE" box="[407,561,1885,1911]" italics="true" pageId="58" pageNumber="59">C. shanensis</emphasis>
(
<bibRefCitation id="EFF44B476C168A1DFDBDF89D440DF8DE" author="Stoliczka F." box="[581,772,1885,1911]" pageId="58" pageNumber="59" pagination="169 - 171" refId="ref56265" refString="Stoliczka F. 1873. Descriptions of two new species of Indian Landshells. Journal of the Asiatic Society of Bengal 42 (2): 169 - 171. Available from https: // biodiversitylibrary. org / page / 35546066 [accessed 30 Jul. 2018]." type="journal article" year="1873">Stoliczka, 1873</bibRefCitation>
)
</taxonomicName>
(NHMUK 20170293).
<emphasis id="B911EAA46C168A1DFBC9F89D434CF8DE" bold="true" box="[1073,1093,1885,1911]" pageId="58" pageNumber="59">E</emphasis>
.
<taxonomicName id="4C654D356C168A1DFBA0F89D43F8F8DE" baseAuthorityName="Stoliczka" baseAuthorityYear="1873" box="[1112,1265,1885,1911]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="shanensis">
<emphasis id="B911EAA46C168A1DFBA0F89D43F8F8DE" box="[1112,1265,1885,1911]" italics="true" pageId="58" pageNumber="59">C. shanensis</emphasis>
</taxonomicName>
(NHMUK 20170292).
<emphasis id="B911EAA46C168A1DFEB2F8404655F833" bold="true" box="[330,348,1920,1946]" pageId="58" pageNumber="59">F</emphasis>
. Protoconch of
<taxonomicName id="4C654D356C168A1DFDEDF84045A1F833" baseAuthorityName="Stoliczka" baseAuthorityYear="1873" box="[533,680,1920,1946]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="shanensis">
<emphasis id="B911EAA46C168A1DFDEDF84045A1F833" box="[533,680,1920,1946]" italics="true" pageId="58" pageNumber="59">C. shanensis</emphasis>
</taxonomicName>
, loc. 20091018B (coll. PGB). Scale bar (bottom left): A, DE = 10 mm; B not to scale. Images: T. Deli (A, DE), B. Páll-Gergely (BC, F).
</paragraph>
</caption>
<subSubSection id="C37F653D6C178A1BFF45FECB47F0FE3B" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="8BDA36B66C178A1CFF45FECB46A3FE8C" blockId="59.[189,426,267,293]" box="[189,426,267,293]" pageId="59" pageNumber="60">
<heading id="D09281DA6C178A1CFF45FECB46A3FE8C" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="B911EAA46C178A1CFF45FECB46A3FE8C" bold="true" box="[189,426,267,293]" pageId="59" pageNumber="60">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B0D3CEB6C178A1CFF12FE82462EFE77" ID-GBIF-Occurrence="2611489335" collectingDate="1988-05-09" collectionCode="UF" collectorName="F. G. Thompson" country="Thailand" elevation="550" latitude="20.353611" location="Chiang Rae Province" longLatPrecision="21" longitude="99.846115" pageId="59" pageNumber="60" specimenCount="1" typeStatus="holotype">
<paragraph id="8BDA36B66C178A1CFF12FE824655FEF5" blockId="59.[189,1399,322,478]" box="[234,348,322,348]" pageId="59" pageNumber="60">
<emphasis id="B911EAA46C178A1CFF12FE824655FEF5" bold="true" box="[234,348,322,348]" pageId="59" pageNumber="60">
<typeStatus id="54DE88146C178A1CFF12FE824655FEF5" box="[234,348,322,348]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDA36B66C178A1CFF45FEAD462EFE77" blockId="59.[189,1399,322,478]" pageId="59" pageNumber="60">
<collectingCountry id="F37276266C178A1CFF45FEAD4650FE2E" box="[189,345,365,391]" name="Thailand" pageId="59" pageNumber="60">THAILAND</collectingCountry>
: 1 shell (D =
<quantity id="4C9D9B536C178A1CFDFFFEAD4568FE21" box="[519,609,365,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="59" pageNumber="60" unit="mm" value="14.0">14 mm</quantity>
),
<location id="8EBA606D6C178A1CFD80FEAD4472FE2E" LSID="urn:lsid:plazi:treatment:03CC87A06C158A1BFD82F9964479F842:8EBA606D6C178A1CFD80FEAD4472FE2E" box="[632,891,365,392]" country="Thailand" latitude="20.353611" longLatPrecision="21" longitude="99.846115" name="Chiang Rae Province" pageId="59" pageNumber="60">Chiang Rae Province</location>
,
<quantity id="4C9D9B536C178A1CFC72FEAD44DEFE2E" box="[906,983,365,391]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="59" pageNumber="60" unit="km" value="12.0">12 km</quantity>
S,
<locationDeviation id="9A682F946C178A1CFBFCFEAD43F5FE2E" box="[1028,1276,365,392]" location="Mae Sai" pageId="59" pageNumber="60">
<quantity id="4C9D9B536C178A1CFBFCFEAD434BFE2E" box="[1028,1090,365,392]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="59" pageNumber="60" unit="km" value="5.0">5 km</quantity>
W of Mae Sai
</locationDeviation>
, ravine in limestone mountains,
<geoCoordinate id="EE5150716C178A1CFE3CFE574555FE1A" box="[452,604,407,435]" degrees="20" direction="north" minutes="21" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="13" value="20.353611">20°2113″ N</geoCoordinate>
,
<geoCoordinate id="EE5150716C178A1CFD93FE5745F6FE1A" box="[619,767,407,435]" degrees="99" direction="east" minutes="50" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="46" value="99.846115">99°5046″ E</geoCoordinate>
,
<elevation id="0048D1856C178A1CFCF6FE584495FE1B" box="[782,924,408,435]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="59" pageNumber="60" unit="m" value="550.0">
<quantity id="4C9D9B536C178A1CFCF6FE584452FE1A" box="[782,859,408,435]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="59" pageNumber="60" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDB10766C178A1CFC54FE584348FE1B" box="[940,1089,408,435]" pageId="59" pageNumber="60" value="1988-05-09">
<collectingDate id="EF9FE99E6C178A1CFC54FE584348FE1B" box="[940,1089,408,435]" pageId="59" pageNumber="60" value="1988-05-09">9 May 1988</collectingDate>
</date>
,
<collectorName id="269053606C178A1CFBA8FE584201FE1B" box="[1104,1288,408,434]" pageId="59" pageNumber="60">F.G. Thompson</collectorName>
leg. (
<collectionCode id="ED74AE736C178A1CFAB7FE58427FFE1B" box="[1359,1398,408,434]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="59" pageNumber="60" type="Museum">UF</collectionCode>
380244).
</paragraph>
</materialsCitation>
<paragraph id="8BDA36B66C178A1CFF12FDCE466EFD81" blockId="59.[189,1399,526,942]" box="[234,359,526,552]" pageId="59" pageNumber="60">
<emphasis id="B911EAA46C178A1CFF12FDCE466EFD81" bold="true" box="[234,359,526,552]" pageId="59" pageNumber="60">
<typeStatus id="54DE88146C178A1CFF12FDCE466EFD81" box="[234,359,526,552]" pageId="59" pageNumber="60" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDA36B66C178A1CFF45FDF94255FC07" blockId="59.[189,1399,526,942]" pageId="59" pageNumber="60">
<collectingCountry id="F37276266C178A1CFF45FDF94650FDFA" box="[189,345,569,595]" name="Thailand" pageId="59" pageNumber="60">THAILAND</collectingCountry>
: 3 shells, same data as for
<typeStatus id="54DE88146C178A1CFD7AFDF945E0FDFA" box="[642,745,569,595]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED74AE736C178A1CFD0EFDF94414FDFA" box="[758,797,569,595]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="59" pageNumber="60" type="Museum">UF</collectionCode>
448576); 1 shell, same data as for
<typeStatus id="54DE88146C178A1CFB5FFDF94207FDFA" box="[1191,1294,569,595]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED74AE736C178A1CFAE3FDF9427EFDFA" box="[1307,1399,569,595]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="59" pageNumber="60">HNHM</collectionCode>
97460); 1 shell,
<collectingRegion id="49A1F8546C178A1CFE8FFDA44579FDD7" box="[375,624,612,639]" inferredBySuffix="true" pageId="59" pageNumber="60">Chiang Rae Province</collectingRegion>
, Tham Tumluang (cave),
<quantity id="4C9D9B536C178A1CFC6EFDA544D9FDD7" box="[918,976,612,639]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="59" pageNumber="60" unit="km" value="7.0">7 km</quantity>
S,
<quantity id="4C9D9B536C178A1CFC0BFDA44324FDD7" box="[1011,1069,612,638]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="km" value="2.0">2 km</quantity>
W of Mae Sai,
<geoCoordinate id="EE5150716C178A1CFB26FDA2427BFDD6" box="[1246,1394,610,639]" degrees="20" direction="north" minutes="22" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="10" value="20.369444">20°2210″ N</geoCoordinate>
,
<geoCoordinate id="EE5150716C178A1CFF45FD4E4644FD03" box="[189,333,654,682]" degrees="99" direction="east" minutes="51" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="28" value="99.85777">99°5128″ E</geoCoordinate>
,
<quantity id="4C9D9B536C178A1CFEA0FD5046A8FD03" box="[344,417,656,682]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="59" pageNumber="60" unit="m" value="470.0">470 m</quantity>
a.s.l.,
<date id="FFDB10766C178A1CFE13FD504571FD03" box="[491,632,656,682]" pageId="59" pageNumber="60" value="1988-05-08">8 May 1988</date>
, F.G. Thompson leg. (
<collectionCode id="ED74AE736C178A1CFC8FFD504497FD03" box="[887,926,656,682]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="59" pageNumber="60" type="Museum">UF</collectionCode>
347154); 2 shells,
<collectingRegion id="49A1F8546C178A1CFB81FD4F427BFD00" box="[1145,1394,655,682]" inferredBySuffix="true" pageId="59" pageNumber="60">Chiang Rae Province</collectingRegion>
, Doi Thung Mountain,
<geoCoordinate id="EE5150716C178A1CFE30FD794557FD7F" box="[456,606,697,726]" degrees="20" direction="north" minutes="20" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="32" value="20.342222">20°2032″ N</geoCoordinate>
,
<geoCoordinate id="EE5150716C178A1CFD93FD7945F7FD7F" box="[619,766,697,726]" degrees="99" direction="east" minutes="50" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="21" value="99.83917">99°5021″ E</geoCoordinate>
,
<quantity id="4C9D9B536C178A1CFCF4FD7B446FFD7F" box="[780,870,699,726]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="59" pageNumber="60" unit="m" value="1320.0">1320 m</quantity>
a.s.l.,
<date id="FFDB10766C178A1CFC4CFD7B434EFD7C" box="[948,1095,699,726]" pageId="59" pageNumber="60" value="1988-05-08">8 May 1988</date>
, F.G. Thompson leg. (
<collectionCode id="ED74AE736C178A1CFAB7FD7B427FFD7C" box="[1359,1398,699,725]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="59" pageNumber="60" type="Museum">UF</collectionCode>
347122);
<specimenCount id="9D63FD3F6C178A1CFECAFD2646DDFCA9" box="[306,468,742,769]" pageId="59" pageNumber="60" type="generic">50 specimens</specimenCount>
(preserved in ethanol: PL053PL054 used for molecular study),
<collectingRegion id="49A1F8546C178A1CFB16FD264621FC85" country="Thailand" name="Chiang Rai" pageId="59" pageNumber="60">Chiang Rai Province</collectingRegion>
, Doi Tung,
<quantity id="4C9D9B536C178A1CFE4AFCD246E4FC85" box="[434,493,786,813]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="59" pageNumber="60" unit="m" value="50.0">50 m</quantity>
from Wat Phra That Doi Tung,
<geoCoordinate id="EE5150716C178A1CFC99FCD04308FC85" box="[865,1025,784,812]" degrees="20" direction="north" minutes="19.540" orientation="latitude" pageId="59" pageNumber="60" precision="1" value="20.325666">20°19.540 N</geoCoordinate>
,
<geoCoordinate id="EE5150716C178A1CFBF4FCD043A1FC85" box="[1036,1192,784,812]" degrees="99" direction="east" minutes="49.987" orientation="longitude" pageId="59" pageNumber="60" precision="1" value="99.833115">99°49.987 E</geoCoordinate>
,
<quantity id="4C9D9B536C178A1CFB4CFCD24204FC85" box="[1204,1293,786,813]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="59" pageNumber="60" unit="m" value="1350.0">1350 m</quantity>
a.s.l.,
<date id="FFDB10766C178A1CFAA1FCD24633FCFE" pageId="59" pageNumber="60" value="2016-03-30">30 Mar. 2016</date>
, A. Hunyadi leg. (coll.
<collectionCode id="ED74AE736C178A1CFDB7FCFD4577FCFE" box="[591,638,829,855]" pageId="59" pageNumber="60">HA</collectionCode>
);
<specimenCount id="9D63FD3F6C178A1CFD6BFCFD4406FCF1" box="[659,783,829,856]" pageId="59" pageNumber="60" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(anatomically examined: PL052 used for molecular study), same data as for preceding (
<collectionCode id="ED74AE736C178A1CFD9AFCA945DAFC2A" box="[610,723,873,899]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="59" pageNumber="60" type="Museum">NHMUK</collectionCode>
20170149); 1 shell,
<collectingRegion id="49A1F8546C178A1CFC30FCA843C9FC2B" box="[968,1216,872,898]" country="Thailand" name="Chiang Rai" pageId="59" pageNumber="60">Chiang Rai Province</collectingRegion>
, Wat Phra That Doi Tung,
<date id="FFDB10766C178A1CFEC3FC5446DEFC07" box="[315,471,916,942]" pageId="59" pageNumber="60" value="2016-10-22">22 Oct. 2016</date>
, K. Okubo leg (coll.
<collectionCode id="ED74AE736C178A1CFD35FC544402FC07" box="[717,779,916,942]" pageId="59" pageNumber="60">PGB</collectionCode>
); 3 shells, same data as for preceding (coll.
<collectionCode id="ED74AE736C178A1CFAF3FC544259FC07" box="[1291,1360,916,942]" pageId="59" pageNumber="60">OKA</collectionCode>
).
</paragraph>
<caption id="DF1A663E6C178A1CFF45F8A74420F842" ID-DOI="http://doi.org/10.5281/zenodo.3817757" ID-Zenodo-Dep="3817757" httpUri="https://zenodo.org/record/3817757/files/figure.png" pageId="59" pageNumber="60" startId="59.[189,232,1895,1921]" targetBox="[202,1399,1040,1853]" targetPageId="59">
<paragraph id="8BDA36B66C178A1CFF45F8A74420F842" blockId="59.[189,1399,1895,2027]" pageId="59" pageNumber="60">
<emphasis id="B911EAA46C178A1CFF45F8A74615F828" bold="true" box="[189,284,1895,1921]" pageId="59" pageNumber="60">Fig. 27.</emphasis>
Genitalia of
<taxonomicName id="4C654D356C178A1CFE39F8A745A8F828" authorityName="Pall-Gergely" authorityYear="2018" box="[449,673,1895,1921]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C178A1CFE39F8A745A8F828" box="[449,673,1895,1921]" italics="true" pageId="59" pageNumber="60">Chersaecia scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C178A1CFD54F8A84403F82B" box="[684,778,1896,1922]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
(NHMUK 20170149).
<emphasis id="B911EAA46C178A1CFBD1F8A74337F828" bold="true" box="[1065,1086,1895,1921]" pageId="59" pageNumber="60">A</emphasis>
. Male genitalia.
<emphasis id="B911EAA46C178A1CFAF6F8A7422BF828" bold="true" box="[1294,1314,1895,1921]" pageId="59" pageNumber="60">B</emphasis>
. Cross sectional diagram of the penis.
<emphasis id="B911EAA46C178A1CFDC8F84A454CF80D" bold="true" box="[560,581,1930,1956]" pageId="59" pageNumber="60">C</emphasis>
. Distal genitalia.
<emphasis id="B911EAA46C178A1CFCE0F84A4447F80D" bold="true" box="[792,846,1930,1956]" pageId="59" pageNumber="60">DF</emphasis>
. Inner structure of genital organs (D = vagina, E = vas deferens, F = diverticulum). Scale bars: A, DF = 0.5 mm; C = 2 mm; B not to scale. For abbreviations see Fig. 2. All photos: B. Páll-Gergely.
</paragraph>
</caption>
<paragraph id="8BDA36B66C108A1BFF12FECB46E8FE8C" blockId="60.[189,1399,267,402]" box="[234,481,267,293]" pageId="60" pageNumber="61">
<emphasis id="B911EAA46C108A1BFF12FECB46E8FE8C" bold="true" box="[234,481,267,293]" pageId="60" pageNumber="61">Additional material</emphasis>
</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FEEF47F0FE3B" blockId="60.[189,1399,267,402]" pageId="60" pageNumber="61">
<collectingCountry id="F37276266C108A1BFF45FEEF4650FEE0" box="[189,345,303,329]" name="Thailand" pageId="60" pageNumber="61">THAILAND</collectingCountry>
:
<specimenCount id="9D63FD3F6C108A1BFE9CFEEF46AFFEE0" box="[356,422,303,329]" pageId="60" pageNumber="61" type="generic">2 juv.</specimenCount>
specimens (preserved in ethanol),
<collectingRegion id="49A1F8546C108A1BFCB9FEEF4330FEE0" box="[833,1081,303,329]" country="Thailand" name="Chiang Rai" pageId="60" pageNumber="61">Chiang Rai Province</collectingRegion>
, Doi Tung,
<quantity id="4C9D9B536C108A1BFB3CFEEF4209FEE3" box="[1220,1280,303,330]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="60" pageNumber="61" unit="m" value="50.0">50 m</quantity>
from Wat Phra That Doi Tung,
<geoCoordinate id="EE5150716C108A1BFE44FE924554FEC7" box="[444,605,338,366]" degrees="20" direction="north" minutes="19.540" orientation="latitude" pageId="60" pageNumber="61" precision="1" value="20.325666">20°19.540 N</geoCoordinate>
,
<geoCoordinate id="EE5150716C108A1BFD93FE924401FEC7" box="[619,776,338,366]" degrees="99" direction="east" minutes="49.987" orientation="longitude" pageId="60" pageNumber="61" precision="1" value="99.833115">99°49.987 E</geoCoordinate>
,
<quantity id="4C9D9B536C108A1BFCEFFE94447BFEC7" box="[791,882,340,367]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="60" pageNumber="61" unit="m" value="1350.0">1350 m</quantity>
a.s.l.,
<date id="FFDB10766C108A1BFC3AFE94436FFEC7" box="[962,1126,340,366]" pageId="60" pageNumber="61" value="2016-03-30">30 Mar. 2016</date>
, A. Hunyadi leg. (coll.
<collectionCode id="ED74AE736C108A1BFF45FEB847E5FE3B" box="[189,236,376,402]" pageId="60" pageNumber="61">HA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C108A1BFF45FE7B4392FA41" pageId="60" pageNumber="61" type="description">
<paragraph id="8BDA36B66C108A1BFF45FE7B4647FE7C" blockId="60.[189,334,443,469]" box="[189,334,443,469]" pageId="60" pageNumber="61">
<heading id="D09281DA6C108A1BFF45FE7B4647FE7C" bold="true" box="[189,334,443,469]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B911EAA46C108A1BFF45FE7B4647FE7C" bold="true" box="[189,334,443,469]" pageId="60" pageNumber="61">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FE2B4661FD12" blockId="60.[189,1398,491,699]" pageId="60" pageNumber="61">SHELL. Sinistral, light brown (although all shells, including live collected ones, were weathered), almost flat, only protoconch protrudes above dorsal surface; protoconch consists of 2.52.75 whorls, with fine granular structure and inconspicuous wrinkles; teleoconch with dense ribs and spiral sculpture, resulting in a reticulated structure; whorls 66.5, separated by a rather shallow suture; aperture rounded; peristome only slightly thickened and reflected; parietal callus low, apertural fold very weak, free from parietal callus.</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FD2346DCFC71" blockId="60.[189,1399,739,984]" pageId="60" pageNumber="61">Three shells were opened. Parietal wall with two vertical lamellae, posterior one longer and C-shaped, with the arms of the C pointing posteriorly; anterior lamella short and connected at both ends to horizontal plicae; main and intermediate plicae of comparable length; ends of plicae visible through oblique view into aperture; main plica and apertural fold situated far from each other. Palatal wall with six plicae; first straight and parallel with upper suture; second short and curved; third longest; fourth and fifth vertical and fused to each other, forming an inverted structure similar to the letter Y; last plica short and curved downwards.</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FBC14466FBB5" blockId="60.[189,879,1025,1052]" box="[189,879,1025,1052]" pageId="60" pageNumber="61">MEASUREMENTS (in mm). D = 11.213, H = 3.74.5 (n = 5).</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FB84423CFAF4" blockId="60.[189,1399,1092,1373]" pageId="60" pageNumber="61">CHARACTERS OF THE GENITAL SYSTEM. A single specimen was dissected (NHMUK 20170149). Right ommatophoral retractor runs between penis and vagina-vas-deferens; penis reduced, very short, blunt, ovoid, inner wall with a strongly thickened fold; retractor muscle short, inserts on proximal end of penis; vas deferens extremely long, thick proximally, slowly reduces in diameter towards penis, internally with oblique, converging longitudinal folds; vagina slender near atrium, slowly increases in width towards spermoviduct; interior of vagina with reticulated sculpture proximally, distal part of vagina with parallel longitudinal folds; diverticulum slender and relatively short, bursa copulatrix long and slender, both originate near base of spermoviduct; diverticulum internally with elevated, parallel, regular folds.</paragraph>
<paragraph id="8BDA36B66C108A1BFF45FA464392FA41" blockId="60.[189,1398,1413,1513]" pageId="60" pageNumber="61">RADULA. Centrals with small, triangular cusps with pointed tips; endocones of laterals subrhomboid with pointed tip; ectocones of first laterals larger than centrals; ectocones of marginals undivided or divided to two; endocones of marginals triangular, undivided or divided by shallow incision.</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C108A1BFF45F9D1453DF8FD" pageId="60" pageNumber="61" type="diagnosis">
<paragraph id="8BDA36B66C108A1BFF45F9D146C0F982" blockId="60.[189,457,1553,1579]" box="[189,457,1553,1579]" pageId="60" pageNumber="61">
<heading id="D09281DA6C108A1BFF45F9D146C0F982" bold="true" box="[189,457,1553,1579]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B911EAA46C108A1BFF45F9D146C0F982" bold="true" box="[189,457,1553,1579]" pageId="60" pageNumber="61">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C108A1BFF45F9814535F944" blockId="60.[189,1399,1601,1773]" pageId="60" pageNumber="61">
<taxonomicName id="4C654D356C108A1BFF45F9814697F9F2" authorityName="Pall-Gergely" authorityYear="2018" box="[189,414,1601,1627]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C108A1BFF45F9814697F9F2" box="[189,414,1601,1627]" italics="true" pageId="60" pageNumber="61">Chersaecia scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C108A1BFE50F981450EF9F2" box="[424,519,1601,1627]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
largely resembles
<taxonomicName id="4C654D356C108A1BFD15F98144AEF9F2" baseAuthorityName="Gude" baseAuthorityYear="1899" box="[749,935,1601,1627]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="woodthorpei">
<emphasis id="B911EAA46C108A1BFD15F98144AEF9F2" box="[749,935,1601,1627]" italics="true" pageId="60" pageNumber="61">C. woodthorpei</emphasis>
</taxonomicName>
in the arrangement of the palatal and parietal plicae and lamellae, but the main plica of the latter species reaches the parietal callus, the intermediate plica is shorter, and a very long lower plica reaches the parietal callus. Moreover,
<taxonomicName id="4C654D356C108A1BFAFDF94A427EF90D" authorityName="Pall-Gergely" authorityYear="2018" box="[1285,1399,1674,1700]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="scabra" status="sp. nov.">
<emphasis id="B911EAA46C108A1BFAFDF94A427EF90D" box="[1285,1399,1674,1700]" italics="true" pageId="60" pageNumber="61">C. scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C108A1BFF45F96F4611F960" box="[189,280,1711,1737]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
is larger, has less shouldered whorls and lacks the series of horizontal palatal plicae, which are characteristic in
<taxonomicName id="4C654D356C108A1BFE87F913453CF944" baseAuthorityName="Gude" baseAuthorityYear="1899" box="[383,565,1747,1773]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="woodthorpei">
<emphasis id="B911EAA46C108A1BFE87F913453CF944" box="[383,565,1747,1773]" italics="true" pageId="60" pageNumber="61">C. woodthorpei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA36B66C108A1BFF45F8D5453DF8FD" blockId="60.[189,1398,1813,1876]" pageId="60" pageNumber="61">
<taxonomicName id="4C654D356C108A1BFF45F8D546D8F886" authorityName="Pall-Gergely" authorityYear="2018" box="[189,465,1813,1839]" class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="auffenbergi" status="sp. nov.">
<emphasis id="B911EAA46C108A1BFF45F8D546D8F886" box="[189,465,1813,1839]" italics="true" pageId="60" pageNumber="61">Chersaecia auffenbergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257DF6C108A1BFE22F8D6453FF899" box="[474,566,1814,1840]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
also has a similar parietal lamellation, but that species is much flatter and has Y-shaped palatal plicae.
</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C108A1BFF45F8BC4479F842" pageId="60" pageNumber="61" type="distribution">
<paragraph id="8BDA36B66C108A1BFF45F8BC465FF83F" blockId="60.[189,342,1916,1942]" box="[189,342,1916,1942]" pageId="60" pageNumber="61">
<heading id="D09281DA6C108A1BFF45F8BC465FF83F" bold="true" box="[189,342,1916,1942]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B911EAA46C108A1BFF45F8BC465FF83F" bold="true" box="[189,342,1916,1942]" pageId="60" pageNumber="61">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C108A1BFF45F86C4479F842" blockId="60.[189,1398,1964,2027]" pageId="60" pageNumber="61">
The new species is known from four geographically-proximate sites situated within
<quantity id="4C9D9B536C108A1BFB62F86C43DCF86F" box="[1178,1237,1964,1990]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="60" pageNumber="61" unit="km" value="6.0">6 km</quantity>
of each other in northern Chiang Rae Province, north
<collectingCountry id="F37276266C108A1BFD6AF81145F3F842" box="[658,762,2001,2027]" name="Thailand" pageId="60" pageNumber="61">Thailand</collectingCountry>
(
<figureCitation id="135E2A336C108A1BFCF3F811446BF842" box="[779,866,2001,2027]" captionStart="Fig" captionStartId="34.[189,232,1824,1850]" captionTargetBox="[189,1399,940,1784]" captionTargetId="figure@34.[186,1400,938,1787]" captionTargetPageId="34" captionText="Fig. 17. Distribution of plectopylid species in Northern Thailand. Legends: circle = Chersaecia auffenbergi sp. nov.; triangle = Chersaecia leiophis (Benson, 1860) (empty triangle shows the type locality of Plectopylis degerbolae Solem, 1966); empty square = Chersaecia perarcta simplex (Solem, 1966); cross = Chersaecia perarcta perarcta (Blanford, 1865); filled square = Chersaecia scabra sp. nov.; rhomb = Chersaecia densegyrata sp. nov.; the upper filled triangle also shows the locality of Naggsia oligogyra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3817741" httpUri="https://zenodo.org/record/3817741/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>