treatments-xml/data/03/9D/41/039D41535461955636FE7C27FECAFED2.xml
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<document id="D7C81A50C258B4CA7F3DC7185B93DE7C" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625623959389" checkinUser="felipe" docAuthor="Published, First" docDate="2007" docId="039D41535461955636FE7C27FECAFED2" docLanguage="en" docName="zt01396p084.pdf" docOrigin="Zootaxa 1396" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Arrup kyushuensis Published 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FFA4392B5473954236697820FFDBFFA8" masterDocTitle="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)" masterLastPageNumber="84" masterPageNumber="1" pageNumber="19" updateTime="1698992576941" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4FBC9855A655A2304C2858A01BDECAA4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3D17ECD5F84F7189642D502D336D24B2">The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)</mods:title>
</mods:titleInfo>
<mods:name id="ADF635E5F3B28D64E7681130DFB90182" type="personal">
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<mods:namePart id="196BF94BF1AA1A281D91CBFC5E61CBE7">Published, First</mods:namePart>
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<mods:title id="4B271197395FD01076FFAF10BF4F4BE7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2C4A5A17559F262F82B4BB2D08C8353F">
<mods:date id="7C3CFC9A4E5C8CB9981C3B678804FAD0">2007</mods:date>
<mods:detail id="1367BFC2E24B9BC3AEFD040B69841FBA" type="pubDate">
<mods:number id="87B5C4CC51C9975E0812359C7C0611D9">2007-01-22</mods:number>
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<mods:number id="79C273DF383D2664A723DA512EB8945E">1396</mods:number>
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<mods:classification id="26000CFFBCE3467AD235FA7ECE27535B">journal article</mods:classification>
<mods:identifier id="2D72BA31AD863AA592620DE384556B45" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="039D41535461955636FE7C27FECAFED2" ID-DOI="http://doi.org/10.5281/zenodo.5076727" ID-Zenodo-Dep="5076727" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2" httpUri="http://treatment.plazi.org/id/039D41535461955636FE7C27FECAFED2" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C32EA3CE5461955036FE7C27FE1EFB89" box="[151,453,1031,1057]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B8BF0455461955036FE7C27FE1EFB89" blockId="18.[151,453,1031,1092]" box="[151,453,1031,1057]" pageId="18" pageNumber="19">
<heading id="D0C347295461955036FE7C27FE1EFB89" bold="true" box="[151,453,1031,1057]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B9402C575461955036FE7C27FE1EFB89" bold="true" box="[151,453,1031,1057]" pageId="18" pageNumber="19">
<taxonomicName id="4C348BC65461955036FE7C27FEA0FB89" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,379,1031,1057]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kyushuensis" status="sp. nov.">
<emphasis id="B9402C575461955036FE7C27FEA0FB89" bold="true" box="[151,379,1031,1057]" italics="true" pageId="18" pageNumber="19">Arrup kyushuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273912C5461955037EA7C27FE1EFB89" box="[387,453,1031,1057]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EA3CE5461955636FE7C0AFECAFED2" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="8B8BF0455461955036FE7C0AFEFAFBEC" blockId="18.[151,453,1031,1092]" box="[151,289,1066,1092]" pageId="18" pageNumber="19">
<figureCitation id="130FECC05461955036FE7C0AFEFAFBEC" box="[151,289,1066,1092]" captionStart="FIGURES 1821" captionStartId="20.[153,271,1785,1809]" captionTargetBox="[284,1300,438,1735]" captionTargetId="figure-151@20.[264,1323,400,1761]" captionTargetPageId="20" captionText="FIGURES 1821. Arrup kyushuensis n. sp., holotype: 18, head, dorsal; 19, anterior part of head, ventral (maxillary complex removed); 20, maxillary complex, ventral; 21, forcipular segment and left forcipule, dorsal (the poison calyx is indicated by solid lines, setae not drawn). Areolation is drawn only partially." pageId="18" pageNumber="19">Figs. 1821</figureCitation>
</paragraph>
<paragraph id="8B8BF0455461955036FE7C5BFB54FA9C" blockId="18.[151,1436,1146,2012]" pageId="18" pageNumber="19">
<emphasis id="B9402C575461955036FE7C5BFED5FB3C" box="[151,270,1147,1172]" italics="true" pageId="18" pageNumber="19">Diagnosis</emphasis>
. An
<taxonomicName id="4C348BC65461955037247C5BFE2AFB3C" box="[333,497,1146,1172]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9402C575461955037247C5BFE48FB3C" box="[333,403,1147,1172]" italics="true" pageId="18" pageNumber="19">Arrup</emphasis>
species
</taxonomicName>
with body length of adults ranging between about 1.5 and
<quantity id="4CCC5DA05461955032AB7C5AFADAFB3C" box="[1218,1281,1146,1172]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="cm" value="3.0">3 cm</quantity>
. Frontal line rounded. Distal article of the telopodite of the second maxillae without claw. Tooth of the forcipular article I triangular, large; tooth of the tarsungulum sharp and well developed. Poison calyx almost reaching the posterior end of the forcipular coxosternum in males (but see Remarks), only reaching the distal part of the forcipular article I in females. Sternum of the last leg­bearing segment as wide as long.
</paragraph>
<paragraph id="8B8BF0455461955036AF7D63FB21FAF4" blockId="18.[151,1436,1146,2012]" box="[198,1274,1346,1372]" pageId="18" pageNumber="19">
<emphasis id="B9402C575461955036AF7D63FEBCFAF4" box="[198,359,1347,1372]" italics="true" pageId="18" pageNumber="19">Type material</emphasis>
.
<typeStatus id="548F4EE754619550371F7D62FE3CFAF4" box="[374,487,1346,1372]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: male,
<quantity id="4CCC5DA05461955034537D62FD57FAF4" box="[570,652,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="mm" value="28.0">28 mm</quantity>
long, adult.
<typeStatus id="548F4EE75461955035767D62FC51FAF4" box="[799,906,1346,1372]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
: female,
<quantity id="4CCC5DA054619550359A7D62FB9EFAF4" box="[1011,1093,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="19" unit="mm" value="26.0">26 mm</quantity>
long, subadult.
</paragraph>
<paragraph id="8B8BF0455461955036AF7D4BFC30FA2C" blockId="18.[151,1436,1146,2012]" box="[198,1003,1386,1412]" pageId="18" pageNumber="19">
<materialsCitation id="3B5CFA185461955036AF7D4BFC3CFA2C" box="[198,999,1386,1412]" country="Japan" county="Kyushu" location="Unzen" municipality="Shimabara-shi" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Nagasaki" typeStatus="holotype">
<emphasis id="B9402C575461955036AF7D4BFE80FA2C" box="[198,347,1387,1412]" italics="true" pageId="18" pageNumber="19">
<typeStatus id="548F4EE75461955036AF7D4BFF26FA2C" box="[198,253,1387,1412]" pageId="18" pageNumber="19">Type</typeStatus>
locality
</emphasis>
.
<location id="8EEBA69E5461955037007D4AFE63FA2C" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955037007D4AFE63FA2C" box="[361,440,1386,1412]" country="Japan" county="Kyushu" municipality="Shimabara-shi" name="Unzen" pageId="18" pageNumber="19" stateProvince="Nagasaki">Unzen</location>
,
<collectingMunicipality id="6BEF6A3F5461955037AD7D4AFDABFA2C" box="[452,624,1386,1412]" pageId="18" pageNumber="19">Shimabara­shi</collectingMunicipality>
,
<collectingRegion id="49F03EA75461955034157D4AFCF7FA2C" box="[636,812,1386,1412]" country="Japan" name="Nagasaki" pageId="18" pageNumber="19">Nagasaki Pref.</collectingRegion>
,
<collectingCounty id="62EA88C954619550355E7D4AFC4EFA2C" box="[823,917,1386,1412]" pageId="18" pageNumber="19">Kyushu</collectingCounty>
,
<collectingCountry id="F323B0D55461955035C87D4AFC3CFA2C" box="[929,999,1386,1412]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF0455461955036AF7DB3FBECFA04" blockId="18.[151,1436,1146,2012]" box="[198,1079,1426,1452]" pageId="18" pageNumber="19">
<emphasis id="B9402C575461955036AF7DB3FDD2FA04" box="[198,521,1427,1452]" italics="true" pageId="18" pageNumber="19">Depository of type material</emphasis>
.
<materialsCitation id="3B5CFA185461955034717DB2FBE8FA04" box="[536,1075,1426,1452]" location="National Science Museum" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Tokyo" typeStatus="holotype">
<location id="8EEBA69E5461955034717DB2FC8AFA04" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955034717DB2FC8AFA04" box="[536,849,1426,1452]" name="National Science Museum" pageId="18" pageNumber="19" stateProvince="Tokyo">National Science Museum</location>
,
<collectingRegion id="49F03EA75461955035357DB2FC72FA04" box="[860,937,1426,1452]" country="Japan" name="Tokyo" pageId="18" pageNumber="19">Tokyo</collectingRegion>
, (
<typeStatus id="548F4EE75461955035D47DB2FBF1FA04" box="[957,1066,1426,1452]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF0455461955036AF7D9BFBABF894" blockId="18.[151,1436,1146,2012]" pageId="18" pageNumber="19">
<emphasis id="B9402C575461955036AF7D9BFE7EFA7C" box="[198,421,1467,1492]" italics="true" pageId="18" pageNumber="19">Material examined</emphasis>
.
<specimenCount id="9D323BCC5461955037DF7D9AFD81FA7C" box="[438,602,1466,1492]" pageId="18" pageNumber="19" type="generic">23 specimens</specimenCount>
. Type series:
<specimenCount id="9D323BCC5461955034947D9AFC8BFA7C" box="[765,848,1466,1492]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
, adult (
<quantity id="4CCC5DA05461955035C07D9AFBD9FA7C" box="[937,1026,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="mm" value="28.0">28 mm</quantity>
), and
<materialsCitation id="3B5CFA185461955032237D9AFD23FA54" collectingDate="1985-09-21" collectionCode="T" county="Unzen" location="Kyushu" municipality="Shimabara-shi" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D323BCC5461955032237D9AFB6EFA7C" box="[1098,1205,1466,1492]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
, subadult (
<quantity id="4CCC5DA054619550335F7D9AFA54FA7C" box="[1334,1423,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="19" unit="mm" value="26.0">26 mm</quantity>
), from
<collectingCounty id="62EA88C95461955036BF7DC2FEFEFA54" box="[214,293,1506,1532]" pageId="18" pageNumber="19">Unzen</collectingCounty>
,
<collectingMunicipality id="6BEF6A3F5461955037587DC2FE06FA54" box="[305,477,1506,1532]" pageId="18" pageNumber="19">Shimabara­shi</collectingMunicipality>
,
<location id="8EEBA69E5461955037817DC2FD9DFA54" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955037817DC2FD9DFA54" box="[488,582,1506,1532]" county="Unzen" municipality="Shimabara-shi" name="Kyushu" pageId="18" pageNumber="19">Kyushu</location>
,
<date id="FF8AD6855461955034387DC2FD02FA54" box="[593,729,1506,1532]" pageId="18" pageNumber="19" value="1985-09-21">
<collectingDate id="EFCE2F6D5461955034387DC2FD02FA54" box="[593,729,1506,1532]" pageId="18" pageNumber="19" value="1985-09-21">21.IX.1985</collectingDate>
</date>
,
<collectionCode id="ED25688054619550348C7DC2FD23FA54" box="[741,760,1506,1532]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5CFA1854619550356B7DC2FEE9F98C" collectionCode="NSMT, AM" collectorName="Tanabe" pageId="18" pageNumber="19" specimenCount="2" typeStatus="holotype">
<collectorName id="26C1959354619550356B7DC2FC8DFA54" box="[770,854,1506,1532]" pageId="18" pageNumber="19">Tanabe</collectorName>
leg., coll.
<collectionCode id="ED2568805461955035B87DC2FBF9FA54" box="[977,1058,1506,1532]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="18" pageNumber="19">NSMT</collectionCode>
and
<collectionCode id="ED2568805461955032337DC2FB52FA54" box="[1114,1161,1506,1532]" country="Australia" name="Australian Museum" pageId="18" pageNumber="19" type="Museum">AM</collectionCode>
(
<typeStatus id="548F4EE75461955032FE7DC2FB24FA54" box="[1175,1279,1506,1532]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
and
<typeStatus id="548F4EE754619550335E7DC2FA40FA54" box="[1335,1435,1506,1532]" pageId="18" pageNumber="19" type="paratype">paratype</typeStatus>
respectively)
</materialsCitation>
. Other specimens:
<specimenCount id="9D323BCC54619550347A7E2AFDBDF98C" box="[531,614,1546,1572]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
, subadult (
<quantity id="4CCC5DA05461955034817E2AFC9BF98C" box="[744,832,1546,1572]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="19" unit="mm" value="17.0">17 mm</quantity>
),
<specimenCount id="9D323BCC54619550353C7E2AFC65F98C" box="[853,958,1546,1572]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
, adult (
<quantity id="4CCC5DA054619550327E7E2AFBABF98C" box="[1047,1136,1546,1572]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="mm" value="16.0">16 mm</quantity>
),
<specimenCount id="9D323BCC5461955032EC7E2AFB21F98C" box="[1157,1274,1546,1572]" pageId="18" pageNumber="19" type="female">2 females</specimenCount>
, subadult (12 and
<quantity id="4CCC5DA05461955036A17E12FEC4F9E4" box="[200,287,1586,1612]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="19" unit="mm" value="22.0">22 mm</quantity>
),
<materialsCitation id="3B5CFA1854619550375A7E12FA59F9E4" box="[307,1410,1586,1612]" collectionCode="AM, TT" collectorName="Tanabe" pageId="18" pageNumber="19" specimenCount="9" specimenCount-juvenile="9" typeStatus="holotype">
<specimenCount id="9D323BCC54619550375A7E12FE68F9E4" box="[307,435,1586,1612]" pageId="18" pageNumber="19" type="juvenile">9 juveniles</specimenCount>
(9, 11, 13, 13, 13, 14, 14, 15 and
<quantity id="4CCC5DA05461955035577E12FC4FF9E4" box="[830,916,1586,1612]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="15.0">15 mm</quantity>
), same data as holotype, coll.
<collectionCode id="ED25688054619550329A7E12FAF9F9E4" box="[1267,1314,1586,1612]" country="Australia" name="Australian Museum" pageId="18" pageNumber="19" type="Museum">AM</collectionCode>
and
<collectionCode id="ED2568805461955033357E12FA59F9E4" box="[1372,1410,1586,1612]" pageId="18" pageNumber="19">TT</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B5CFA185461955033E47E12FDCBF934" collectingDate="1989-04-24" collectionCode="AM, TT" collectorName="S. Nomura" county="Mt. Kyogatake" location="Kyushu" municipality="Kashima-shi" pageId="18" pageNumber="19" specimenCount="3" specimenCount-female="3" stateProvince="Saga" typeStatus="holotype">
<specimenCount id="9D323BCC5461955033E47E12FF2EF9DC" pageId="18" pageNumber="19" type="female">3 females</specimenCount>
, adult (22, 22 and
<quantity id="4CCC5DA05461955037B57E7AFDEEF9DC" box="[476,565,1626,1652]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="19" unit="mm" value="23.0">23 mm</quantity>
), from
<collectingCounty id="62EA88C95461955034E67E7AFC9EF9DC" box="[655,837,1626,1652]" pageId="18" pageNumber="19">Mt. Kyogatake</collectingCounty>
,
<collectingMunicipality id="6BEF6A3F54619550353A7E7AFC31F9DC" box="[851,1002,1626,1652]" pageId="18" pageNumber="19">Kashima­shi</collectingMunicipality>
,
<collectingRegion id="49F03EA75461955035917E7AFBADF9DC" box="[1016,1142,1626,1652]" country="Japan" name="Saga" pageId="18" pageNumber="19">Saga Pref.</collectingRegion>
,
<location id="8EEBA69E5461955032EC7E7AFB38F9DC" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955032EC7E7AFB38F9DC" box="[1157,1251,1626,1652]" county="Mt. Kyogatake" municipality="Kashima-shi" name="Kyushu" pageId="18" pageNumber="19" stateProvince="Saga">Kyushu</location>
,
<date id="FF8AD6855461955032987E7AFAADF9DC" box="[1265,1398,1626,1652]" pageId="18" pageNumber="19" value="1989-04-24">
<collectingDate id="EFCE2F6D5461955032987E7AFAADF9DC" box="[1265,1398,1626,1652]" pageId="18" pageNumber="19" value="1989-04-24">24.IV.1989</collectingDate>
</date>
,
<collectorName id="26C195935461955033ED7E7AFF2CF934" pageId="18" pageNumber="19">S. Nomura</collectorName>
leg., coll.
<collectionCode id="ED2568805461955037157EA2FE70F934" box="[380,427,1666,1692]" country="Australia" name="Australian Museum" pageId="18" pageNumber="19" type="Museum">AM</collectionCode>
and
<collectionCode id="ED2568805461955037837EA2FDCBF934" box="[490,528,1666,1692]" pageId="18" pageNumber="19">TT</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B5CFA185461955034767EA2FC82F96C" collectingDate="1986-08-12" collectionCode="AM" collectorName="A. Moroto" county="Nagata-hodo" location="Ryukyu Ids" municipality="Yakushima Id." pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount id="9D323BCC5461955034767EA2FDAFF934" box="[543,628,1666,1692]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
, subadult (
<quantity id="4CCC5DA05461955034937EA2FC8FF934" box="[762,852,1666,1692]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="mm" value="16.0">16 mm</quantity>
), from
<collectingCounty id="62EA88C95461955035C67EA2FB92F934" box="[943,1097,1666,1692]" pageId="18" pageNumber="19">Nagata­hodo</collectingCounty>
,
<collectingMunicipality id="6BEF6A3F5461955032317EA2FADCF934" box="[1112,1287,1666,1692]" pageId="18" pageNumber="19">Yakushima Id.</collectingMunicipality>
,
<collectingRegion id="49F03EA754619550337F7EA2FF09F96C" country="Japan" name="Kagoshima" pageId="18" pageNumber="19">Kagoshima Pref.</collectingRegion>
,
<location id="8EEBA69E5461955036897E8AFEB7F96C" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955036897E8AFEB7F96C" box="[224,364,1706,1732]" county="Nagata-hodo" municipality="Yakushima Id." name="Ryukyu Ids" pageId="18" pageNumber="19" stateProvince="Kagoshima">Ryukyu Ids</location>
,
<date id="FF8AD6855461955037137E8AFDCDF96C" box="[378,534,1706,1732]" pageId="18" pageNumber="19" value="1986-08-12">
<collectingDate id="EFCE2F6D5461955037137E8AFDCDF96C" box="[378,534,1706,1732]" pageId="18" pageNumber="19" value="1986-08-12">12.VIII.1986</collectingDate>
</date>
,
<collectorName id="26C1959354619550344D7E8AFD78F96C" box="[548,675,1706,1732]" pageId="18" pageNumber="19">A. Moroto</collectorName>
leg., coll.
<collectionCode id="ED25688054619550354F7E8AFC82F96C" box="[806,857,1706,1732]" country="Australia" name="Australian Museum" pageId="18" pageNumber="19" type="Museum">AM</collectionCode>
</materialsCitation>
;
<specimenCount id="9D323BCC54619550350E7E8AFC1DF96C" box="[871,966,1706,1732]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
, subadult (both
<quantity id="4CCC5DA05461955032E07E8AFB39F96C" box="[1161,1250,1706,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="mm" value="13.0">13 mm</quantity>
), and
<materialsCitation id="3B5CFA185461955033447E8AFE98F8BC" collectingDate="1985-09-18" collectionCode="T" collectorName="Yakushima Id." county="Hanayama-hodo" location="Ryukyu Ids" municipality="Kumage-gun" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount id="9D323BCC5461955033447E8AFA43F96C" box="[1325,1432,1706,1732]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
, subadult (
<quantity id="4CCC5DA054619550377B7EF2FEAAF944" box="[274,369,1746,1772]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="mm" value="12.0">12 mm</quantity>
), from
<collectingCounty id="62EA88C95461955037BD7EF2FD42F944" box="[468,665,1746,1772]" pageId="18" pageNumber="19">Hanayama­hodo</collectingCounty>
,
<collectorName id="26C195935461955034C57EF2FCBBF944" box="[684,864,1746,1772]" pageId="18" pageNumber="19">Yakushima Id.</collectorName>
,
<collectingMunicipality id="6BEF6A3F54619550351D7EF2FBCBF944" box="[884,1040,1746,1772]" pageId="18" pageNumber="19">Kumage­gun</collectingMunicipality>
,
<collectingRegion id="49F03EA754619550324A7EF2FB28F944" box="[1059,1267,1746,1772]" country="Japan" name="Kagoshima" pageId="18" pageNumber="19">Kagoshima Pref.</collectingRegion>
,
<location id="8EEBA69E54619550336F7EF2FA4CF944" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E54619550336F7EF2FA4CF944" box="[1286,1431,1746,1772]" county="Hanayama-hodo" municipality="Kumage-gun" name="Ryukyu Ids" pageId="18" pageNumber="19" stateProvince="Kagoshima">Ryukyu Ids</location>
,
<date id="FF8AD6855461955036FE7EDAFEC4F8BC" box="[151,287,1786,1812]" pageId="18" pageNumber="19" value="1985-09-18">
<collectingDate id="EFCE2F6D5461955036FE7EDAFEC4F8BC" box="[151,287,1786,1812]" pageId="18" pageNumber="19" value="1985-09-18">18.IX.1985</collectingDate>
</date>
,
<collectionCode id="ED2568805461955037597EDAFE98F8BC" box="[304,323,1786,1812]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
</materialsCitation>
.
<collectorName id="26C1959354619550373A7EDAFE7CF8BC" box="[339,423,1786,1812]" pageId="18" pageNumber="19">Tanabe</collectorName>
leg., coll. TT;
<materialsCitation id="3B5CFA185461955034017EDAFC85F894" collectingDate="1985-11-11" collectionCode="T" county="Shikanosawa" location="Yakushima Id." municipality="Hanayama-hodo" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount id="9D323BCC5461955034017EDAFD65F8BC" box="[616,702,1786,1812]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
, adult (
<quantity id="4CCC5DA05461955035777EDAFCA1F8BC" box="[798,890,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="19" unit="mm" value="17.0">17 mm</quantity>
), from
<collectingCounty id="62EA88C95461955035B17EDAFBAEF8BC" box="[984,1141,1786,1812]" pageId="18" pageNumber="19">Shikanosawa</collectingCounty>
, along
<collectingMunicipality id="6BEF6A3F5461955032BA7EDAFA43F8BC" box="[1235,1432,1786,1812]" pageId="18" pageNumber="19">Hanayama­hodo</collectingMunicipality>
,
<location id="8EEBA69E5461955036FE7F02FE99F894" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5461955036FE7F02FE99F894" box="[151,322,1826,1852]" county="Shikanosawa" municipality="Hanayama-hodo" name="Yakushima Id." pageId="18" pageNumber="19" stateProvince="Kagoshima">Yakushima Id.</location>
,
<collectingRegion id="49F03EA75461955037267F02FDCCF894" box="[335,535,1826,1852]" country="Japan" name="Kagoshima" pageId="18" pageNumber="19">Kagoshima Pref.</collectingRegion>
,
<location id="8EEBA69E54619550344A7F02FD70F894" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E54619550344A7F02FD70F894" box="[547,683,1826,1852]" county="Shikanosawa" municipality="Hanayama-hodo" name="Ryukyu Ids" pageId="18" pageNumber="19" stateProvince="Kagoshima">Ryukyu Ids</location>
,
<date id="FF8AD6855461955034D17F02FCE4F894" box="[696,831,1826,1852]" pageId="18" pageNumber="19" value="1985-11-11">
<collectingDate id="EFCE2F6D5461955034D17F02FCE4F894" box="[696,831,1826,1852]" pageId="18" pageNumber="19" value="1985-11-11">11.XI.1985</collectingDate>
</date>
,
<collectionCode id="ED2568805461955035227F02FC85F894" box="[843,862,1826,1852]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
</materialsCitation>
.
<collectorName id="26C195935461955035007F02FC66F894" box="[873,957,1826,1852]" pageId="18" pageNumber="19">Tanabe</collectorName>
leg., coll. AM.
</paragraph>
<paragraph id="8B8BF0455461955136AF7F6BFE4BFCA2" blockId="18.[151,1436,1146,2012]" lastBlockId="19.[151,1440,152,2018]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B9402C575461955036AF7F6BFDD3F8CC" box="[198,520,1867,1892]" italics="true" pageId="18" pageNumber="19">
Description of the
<typeStatus id="548F4EE75461955037CD7F6BFDD3F8CC" box="[420,520,1867,1892]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
</emphasis>
. Male, adult, body length
<quantity id="4CCC5DA054619550352A7F6AFC41F8CC" box="[835,922,1866,1892]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="mm" value="28.0">28 mm</quantity>
. Body colour yellow, without dark patches. Head 1.4 times as long as wide; frontal line curved. Antennae 3.2 times longer than the head width. Apical sensilla about
<quantity id="4CCC5DA05461955037577FBAFEA0F81C" box="[318,379,1946,1972]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="mm" value="8.0">8 µm</quantity>
long. Clypeal setae: a median pair anterior to the plagulae and a group on each side of the clypeus, reaching the anterior corners. Clypeal ratio about 2.8. Labrum: internal margins of side­pieces convergent posteriorly, not touching each other; posterior margin of each side­piece slightly sinuous, convex at both ends and concave in the middle. First maxillae: anterior corners of the coxosternum not projecting; medial projection about 2.0 times as long as wide, with seven setae, internal margin with few spines and sinuate near the base, distal lobe about 2.0 times as long as wide, aligned to the external margin of the projection, with parallel margins and only slightly clavate at the tip, and a row of many hyaline scales on the dorsal side; telopodite about 3.1 times as long as wide, curved, with five long setae in the middle, its distal lobe attenuated. Second maxillae: article I of telopodite about 2.1 times as long as wide, the distal­external end bearing a small spine; article III about 2.9 times as long as wide, with about 10 long setae; apical claw absent replaced by 34 small tubercles. Forcipular segment: width to length ratio of exposed part of coxosternum about 1.2; cerrus absent; dorsal ridge of pleuron not evident; scapular point not reaching the anterior margin of coxosternum. Forcipules: article I about 1.7 times as long as wide, the distal tooth large, subtriangular, about as long as wide, and clearly projecting outwards; article II smooth; article III with a distinct tubercle; tarsungulum with a well developed basal tooth; poison calyx nearly reaching the posterior end of forcipular coxosternum. A total of 41 leg­bearing segments. Last leg­bearing segment: sternum subtriangular, as wide as long; telopodites with dense and short additional setae on the ventral side and a very small spine at the tip.
</paragraph>
<paragraph id="8B8BF0455460955136AF7B38FE46FB32" blockId="19.[151,1440,152,2018]" pageId="19" pageNumber="20">
<emphasis id="B9402C575460955136AF7B38FE33FC99" box="[198,488,792,817]" italics="true" pageId="19" pageNumber="20">Interindividual variation</emphasis>
(based on
<specimenCount id="9D323BCC5460955134017B38FCD7FC9A" box="[616,780,792,818]" pageId="19" pageNumber="20" type="generic">23 specimens</specimenCount>
). Total length
<quantity id="4CCC5DA05460955135DB7B38FBE8FC9A" box="[946,1075,792,818]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.8" metricValueMin="1.7" pageId="19" pageNumber="20" unit="mm" value="22.5" valueMax="28.0" valueMin="17.0">1728 mm</quantity>
. Head 1.41.5 times as long as wide. Antennae 2.73.2 times as long as the head width. Clypeal ratio 2.43.2. Medial projection of the first maxillae 1.72.2 times as long as wide, bearing 27 setae, basal sinus present or absent. Telopodite 2.63.2 times as long as wide, bearing 25 long setae in the middle. Second maxillae: article I of telopodite 1.52.5 times as long as wide, the distal­external end bearing a small spine or not; article III 1.72.7 times as long as wide. Forcipular article I 1.62.0 times as long as wide, distal tooth slightly variable in shape and size, but always large and triangular, always projecting. Poison calyx possibly polymorphic in full­grown males (see Remarks), invariably reaching the distal part of the forcipular article I in females and juveniles. Legs of the last pair with dense, short additional setae on ventral side in full­grown males, without additional setae in females and juveniles.
</paragraph>
<paragraph id="8B8BF0455460955136AF7C88FDB3FB6A" blockId="19.[151,1440,152,2018]" box="[198,616,1192,1218]" pageId="19" pageNumber="20">
<emphasis id="B9402C575460955136AF7C88FDB9FB69" box="[198,610,1192,1217]" italics="true" pageId="19" pageNumber="20">Distribution in the considered area</emphasis>
.
</paragraph>
<paragraph id="8B8BF0455460955136AF7CF0FB97FB42" blockId="19.[151,1440,152,2018]" box="[198,1100,1232,1258]" pageId="19" pageNumber="20">
<materialsCitation id="3B5CFA185460955136AF7CF0FB93FB42" box="[198,1096,1232,1258]" location="Mt. Kyogatake" municipality="Kashima-shi" pageId="19" pageNumber="20" specimenCount="1">
Kyushu: Unzen, Shimabara­shi (
<typeStatus id="548F4EE754609551342C7CF0FDA2FB42" box="[581,633,1232,1258]" pageId="19" pageNumber="20">type</typeStatus>
locality);
<location id="8EEBA69E5460955134987CF0FC7FFB42" LSID="urn:lsid:plazi:treatment:039D41535461955636FE7C27FECAFED2:8EEBA69E5460955134987CF0FC7FFB42" box="[753,932,1232,1258]" municipality="Kashima-shi" name="Mt. Kyogatake" pageId="19" pageNumber="20">Mt. Kyogatake</location>
,
<collectingMunicipality id="6BEF6A3F5460955135D97CF0FB93FB42" box="[944,1096,1232,1258]" pageId="19" pageNumber="20">Kashima­shi</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF0455460955136AF7CD8FB76FABA" blockId="19.[151,1440,152,2018]" box="[198,1197,1272,1298]" pageId="19" pageNumber="20">Ryukyu Islands: Nagata­hodo; Hanayama­hodo; Shikanosawa (all in Yakushima Id.).</paragraph>
<paragraph id="8B8BF0455460955136AF7D00FCD8FA92" blockId="19.[151,1440,152,2018]" box="[198,771,1312,1338]" pageId="19" pageNumber="20">
<emphasis id="B9402C575460955136AF7D00FE65FA91" box="[198,446,1312,1337]" italics="true" pageId="19" pageNumber="20">General distribution.</emphasis>
Kyushu and Yakushima Id.
</paragraph>
<paragraph id="8B8BF0455460955136AF7D68FDA8FACA" blockId="19.[151,1440,152,2018]" box="[198,627,1352,1378]" pageId="19" pageNumber="20">
<emphasis id="B9402C575460955136AF7D68FE40FAC9" box="[198,411,1352,1377]" italics="true" pageId="19" pageNumber="20">Derivatio nominis</emphasis>
. From Kyushu Id.
</paragraph>
<paragraph id="8B8BF0455460955136AF7D50FD73F90A" blockId="19.[151,1440,152,2018]" pageId="19" pageNumber="20">
<emphasis id="B9402C575460955136AF7D50FEF6FA21" box="[198,301,1392,1417]" italics="true" pageId="19" pageNumber="20">Remarks</emphasis>
. Males of
<taxonomicName id="4C348BC65460955137D37D50FDB7FA21" authorityName="Published" authorityYear="2007" box="[442,620,1392,1417]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kyushuensis">
<emphasis id="B9402C575460955137D37D50FDB7FA21" box="[442,620,1392,1417]" italics="true" pageId="19" pageNumber="20">A. kyushuensis</emphasis>
</taxonomicName>
are possibly polymorphic for the size of the poison calyx, as we examined four other males (not mentioned in the material listed above) which agree fully with the diagnosis of this species but for a shorter calyx. In three of these, all subadult (15 and
<quantity id="4CCC5DA054609551324A7DE0FBADFA72" box="[1059,1142,1472,1498]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="20" unit="mm" value="16.0">16 mm</quantity>
long from Nagata­hodo, Yakushima Id.,
<collectingRegion id="49F03EA754609551373A7DC8FDC6F9AA" box="[339,541,1512,1538]" country="Japan" name="Kagoshima" pageId="19" pageNumber="20">Kagoshima Pref.</collectingRegion>
,
<collectingCountry id="F323B0D55460955134437DC8FDABF9AA" box="[554,624,1512,1538]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
,
<date id="FF8AD6855460955134147DC8FCC2F9AA" box="[637,793,1512,1538]" pageId="19" pageNumber="20" value="1986-08-12">12.VIII.1986</date>
, A. Moroto leg., coll. AM and TT;
<quantity id="4CCC5DA05460955132AC7DC8FAC2F9AA" box="[1221,1305,1512,1538]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="20" unit="mm" value="21.0">21 mm</quantity>
long, from Mt. Shiratori­san, Izumi­mura, Yatsushiro­gun,
<collectingRegion id="49F03EA75460955134AE7E30FC55F982" box="[711,910,1552,1578]" country="Japan" name="Kumamoto" pageId="19" pageNumber="20">Kumamoto Pref.</collectingRegion>
,
<collectingCountry id="F323B0D55460955135F07E30FC05F982" box="[921,990,1552,1578]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
,
<date id="FF8AD6855460955135827E30FBBAF982" box="[1003,1121,1552,1578]" pageId="19" pageNumber="20" value="1987-04-05">5.IV.1987</date>
, S. Nomura leg., coll. AM) the calyx reaches the distal part of the first forcipular article (as in females), whereas in another apparently adult male (
<quantity id="4CCC5DA05460955137427E40FE59F9D2" box="[299,386,1632,1658]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="20" unit="mm" value="16.0">16 mm</quantity>
long, from Mt. Shiratori­san,
<date id="FF8AD6855460955134927E40FCABF9D2" box="[763,880,1632,1658]" pageId="19" pageNumber="20" value="1987-04-05">5.IV.1987</date>
, S. Nomura leg., coll. AM) the calyx almost reaches the base of the first forcipular article.
</paragraph>
<paragraph id="8B8BF0455460955136AF7E90FB82F8C2" blockId="19.[151,1440,152,2018]" pageId="19" pageNumber="20">
However we cannot rule out the possibility that these males actually belong to a different species, even though the detailed morphological and morphometrical analysis we carried out failed to produce evidence in support of this hypothesis. Indeed, we found preliminary evidence that the calyx length is associated with the presence of additional short setae on the last legs, as three of those
<specimenCount id="9D323BCC5460955135A47F08FB97F8EA" box="[973,1100,1832,1858]" pageId="19" pageNumber="20" type="male">four males</specimenCount>
with shorter calyx have no additional setae or only a few. Further material is needed to assess this hypothesis.
</paragraph>
<paragraph id="8B8BF0455460955136AF7F58FD31F812" blockId="19.[151,1440,152,2018]" pageId="19" pageNumber="20">
<typeStatus id="548F4EE75460955136AF7F58FEDBF83A" box="[198,256,1912,1938]" pageId="19" pageNumber="20">Type</typeStatus>
specimens are represented by a male with long calyx and a female from the same locality. No males with short calyx were collected at the
<typeStatus id="548F4EE754609551343A7F80FD5EF812" box="[595,645,1952,1978]" pageId="19" pageNumber="20">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B8BF0455460955636AF7FE8FB61FF1A" blockId="19.[151,1440,152,2018]" lastBlockId="20.[151,1438,152,378]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Worth notice is that, in
<taxonomicName id="4C348BC65460955137B17FE8FDA7F84A" box="[472,636,1992,2018]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9402C575460955137B17FE8FDC5F849" box="[472,542,1992,2017]" italics="true" pageId="19" pageNumber="20">Arrup</emphasis>
species
</taxonomicName>
, all juveniles examined have a short calyx, suggesting that the calyx elongates with body growth and that the difference between sexes emerges at a later stage.
</paragraph>
<paragraph id="8B8BF0455467955636AF78E0FECAFED2" blockId="20.[151,1438,152,378]" pageId="20" pageNumber="21">
Also remarkable in this species is the great variability in the relationship between body size and size of gonopods, as we found no trace of gonopods in specimens
<quantity id="4CCC5DA054679556352378C8FC62FEAA" box="[842,953,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="20" pageNumber="21" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">915 mm</quantity>
long, developing gonopods in specimens
<quantity id="4CCC5DA05467955636FE7930FECEFE82" box="[151,277,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="20" pageNumber="21" unit="mm" value="19.0" valueMax="26.0" valueMin="12.0">1226 mm</quantity>
long and fully developed gonopods in specimens
<quantity id="4CCC5DA05467955635037930FC32FE82" box="[874,1001,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.8" metricValueMin="1.6" pageId="20" pageNumber="21" unit="mm" value="22.0" valueMax="28.0" valueMin="16.0">1628 mm</quantity>
long. But we also found in the same population a
<quantity id="4CCC5DA054679556375D7918FE52FEFA" box="[308,393,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="21" unit="mm" value="16.0">16 mm</quantity>
female with fully developed gonopods and a
<quantity id="4CCC5DA05467955635DB7918FBDDFEFA" box="[946,1030,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="21" unit="mm" value="22.0">22 mm</quantity>
long female with still developing gonopods.
</paragraph>
</subSubSection>
</treatment>
</document>