treatments-xml/data/B1/5D/87/B15D87DEFFFCBE7B6EDEFF100CC49B05.xml

<document id="5B333762FFF76B7CC116DB505BBB8726" ID-CLB-Dataset="55729" ID-DOI="10.1080/00222933.2020.1763491" ID-GBIF-Dataset="fb05d004-0f2d-44da-b91e-b6e8304ab2a5" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1615924067624" checkinUser="carolina" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2020" docId="B15D87DEFFFCBE7B6EDEFF100CC49B05" docLanguage="en" docName="JNATHIST.54.7-8.445-532.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 54 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2020.1763491" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Shinobium Naruse &amp; Ng 2020, gen. nov." docType="treatment" docUuid="3C61559C-C727-4513-8F43-B624FC785407" docUuidSource="ZooBank" docVersion="8" lastPageNumber="469" masterDocId="4D64FFA6FFEABE626F33FF9E0F509F28" masterDocTitle="Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species" masterLastPageNumber="532" masterPageNumber="445" pageNumber="466" updateTime="1698933465109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2064A31F4A2098679CD0C86FA6BA62D7">Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species</mods:title>
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<treatment id="B15D87DEFFFCBE7B6EDEFF100CC49B05" ID-DOI="http://doi.org/10.5281/zenodo.6518951" ID-GBIF-Taxon="195191646" ID-Zenodo-Dep="6518951" LSID="urn:lsid:zoobank.org:act:3C61559C-C727-4513-8F43-B624FC785407" httpUri="http://treatment.plazi.org/id/B15D87DEFFFCBE7B6EDEFF100CC49B05" lastPageId="25" lastPageNumber="469" pageId="22" pageNumber="466">
<subSubSection id="71EE6543FFFCBE746EDEFF100C679F80" box="[493,823,142,168]" pageId="22" pageNumber="466" type="nomenclature">
<paragraph id="394B36C8FFFCBE746EDEFF100C679F80" blockId="22.[493,823,142,168]" box="[493,823,142,168]" pageId="22" pageNumber="466">
<heading id="620381A4FFFCBE746EDEFF100C679F80" box="[493,823,142,168]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="466" reason="2">
Genus 
<taxonomicName id="FEF44D4BFFFCBE746D0CFF100D929F80" ID-CoL="84GP8" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[575,706,142,168]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="466" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="0B80EADAFFFCBE746D0CFF100D929F80" bold="true" box="[575,706,142,168]" italics="true" pageId="22" pageNumber="466">Shinobium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10B357A1FFFCBE746DFAFF100C679F80" box="[713,823,142,168]" pageId="22" pageNumber="466" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFFCBE746F93FF6C0D849E8B" pageId="22" pageNumber="466" type="type_taxon">
<paragraph id="394B36C8FFFCBE746F93FF6C0E639E24" blockId="22.[160,1135,242,302]" box="[160,307,242,268]" pageId="22" pageNumber="466">
<emphasis id="0B80EADAFFFCBE746F93FF6C0E639E24" bold="true" box="[160,307,242,268]" italics="true" pageId="22" pageNumber="466">
<typeStatus id="E64F886AFFFCBE746F93FF6C0F889E24" box="[160,216,242,268]" pageId="22" pageNumber="466">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="394B36C8FFFCBE746F93FE8B0B3F9E06" blockId="22.[160,1135,242,302]" box="[160,1135,277,302]" pageId="22" pageNumber="466">
<taxonomicName id="FEF44D4BFFFCBE746F93FE8B0ECA9E06" ID-CoL="9DBPM" authorityName=": Rathbun" authorityYear="1907" box="[160,410,277,302]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="466" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="0B80EADAFFFCBE746F93FE8B0ECA9E06" box="[160,410,277,302]" italics="true" pageId="22" pageNumber="466">Sesarma trapezoideum</emphasis>
</taxonomicName>
H. Milne Edwards, 1837, by present designation. Gender neuter.
</paragraph>
<paragraph id="394B36C8FFFCBE746F93FEF90E0F9EA9" blockId="22.[160,724,359,419]" box="[160,351,359,385]" pageId="22" pageNumber="466">
<emphasis id="0B80EADAFFFCBE746F93FEF90E0F9EA9" bold="true" box="[160,351,359,385]" italics="true" pageId="22" pageNumber="466">Included species</emphasis>
</paragraph>
<paragraph id="394B36C8FFFCBE746F93FE140D849E8B" blockId="22.[160,724,359,419]" box="[160,724,394,419]" pageId="22" pageNumber="466">
<taxonomicName id="FEF44D4BFFFCBE746F93FE140EE29E8B" ID-CoL="84HC8" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1837" box="[160,434,394,419]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="466" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="0B80EADAFFFCBE746F93FE140EE29E8B" box="[160,434,394,419]" italics="true" pageId="22" pageNumber="466">Shinobium trapezoideum</emphasis>
</taxonomicName>
(H. Milne Edwards, 1837).
</paragraph>
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<subSubSection id="71EE6543FFFCBE746F93FE420CBE992E" pageId="22" pageNumber="466" type="diagnosis">
<paragraph id="394B36C8FFFCBE746F93FE420E449EDE" blockId="22.[160,1156,476,1542]" box="[160,276,476,502]" pageId="22" pageNumber="466">
<emphasis id="0B80EADAFFFCBE746F93FE420E449EDE" bold="true" box="[160,276,476,502]" italics="true" pageId="22" pageNumber="466">Diagnosis</emphasis>
</paragraph>
<paragraph id="394B36C8FFFCBE746F93FE610CBE992E" blockId="22.[160,1156,476,1542]" pageId="22" pageNumber="466">Carapace trapezoidal, longer than wide; divergent posteriorly, widest at bases of P3, with 1 distinct epibranchial tooth behind external orbital angle, posterolateral regions setose; dorsal surface slightly convex, regions well defined. Front deflexed at anterior margin of postfrontal lobes at more than 90°, distally sloping ventroposteriorly; frontal margin bilobed with rounded median concavity, each lobe recurved, directed anteriorly, median concave part attached to narrow, long, clearly exposed antennular septum. Two pairs of postfrontal lobes present, lateral lobes slightly exceeding mesial lobes anteriorly, lateral lobes only slightly narrower than mesial lobes, all lobes projected anteriorly, overhanging onto front. Orbit, in dorsal view, L-shaped, median part of supraorbital margin almost horizontal; inner orbital tooth triangular, long, directed anteriorly. External orbital angle with 1 longitudinal ridge on ventral surface. Epistome posterior margin with 3 triangular lobes, all lobes directed anteriorly. Antenna entering orbit through wide gap between inner orbital tooth and front. Mxp3 exopod with distinct flagellum. Chela palm with many irregular rows of granules on upper surface both in male and female; outer surface granulated but without prominent protuberance; inner surface without transverse rows of granules; narrow rim extending along the occlusal margin of immovable finger to dactylar articulation on both outer and inner surfaces, these narrow rims of both outer and inner surfaces not interrupted near dactylar articulation. Upper-inner margin of movable finger with 1 dense, regular row of horizontal, oblong, tiny granules over most of length in male, row shorter with smaller granules in female. Ambulatory legs (P2–5) long, dorso-ventrally flattened; distal anterior corner of each merus forming falcate lamella, followed proximally by subdistal tooth; carpi and propodi distinctly narrower than respective meri; dactyli very short, shorter than half length of respective propodi. Male thoracic sternum transversely narrow; sternite 8 clearly exposed. Male pleon relatively narrow, bell shaped, lateral margin weakly concave, telson not reaching proximal half of bases of cheliped coxae. G1 narrow, very long, constricted at distal two-fifths, reaching distal fifth to distal end of thoracic sternite 5; beak-like and corneous process narrow. Vulvae opening on distal half of sternite 6, anterior margin adjacent to thoracic sternal suture 5/6, ellipsoidal; 3 rounded sternal vulval covers developed from posterolateral margin, middle largest, connected with lower cover by narrow rim.</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFFCBE746F93F9A00CC399E8" pageId="22" pageNumber="466" type="etymology">
<paragraph id="394B36C8FFFCBE746F93F9A00E4D9970" blockId="22.[160,1155,1598,1728]" box="[160,285,1598,1624]" pageId="22" pageNumber="466">
<emphasis id="0B80EADAFFFCBE746F93F9A00E4D9970" bold="true" box="[160,285,1598,1624]" italics="true" pageId="22" pageNumber="466">Etymology</emphasis>
</paragraph>
<paragraph id="394B36C8FFFCBE746F93F9FC0CC399E8" blockId="22.[160,1155,1598,1728]" pageId="22" pageNumber="466">
The generic name ‘ 
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<emphasis id="0B80EADAFFFCBE746E48F9FC0EA09953" box="[379,496,1634,1659]" italics="true" pageId="22" pageNumber="466">Shinobium</emphasis>
</taxonomicName>
’ is derived from an arbitrary combination of ‘ 
<emphasis id="0B80EADAFFFCBE746CCAF9FC0B1A9953" box="[1017,1098,1634,1659]" italics="true" pageId="22" pageNumber="466">Shinobi</emphasis>
’ and the suffix for the 
<taxonomicName id="FEF44D4BFFFCBE746E57F91B0EB099B6" authorityName=", Serene and Soh" authorityYear="1970" box="[356,480,1669,1694]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFCBE746E57F91B0EB099B6" box="[356,480,1669,1694]" italics="true" pageId="22" pageNumber="466">Labuanium</emphasis>
</taxonomicName>
‘ 
<emphasis id="0B80EADAFFFCBE746EDDF91B0D7099B6" box="[494,544,1669,1694]" italics="true" pageId="22" pageNumber="466">-ium</emphasis>
’. ‘ 
<emphasis id="0B80EADAFFFCBE746D0BF91B0DD999B6" box="[568,649,1669,1694]" italics="true" pageId="22" pageNumber="466">Shinobi</emphasis>
’ is an alias of ‘Ninja’ in Japanese, alluding to the swift and cryptic behaviour of the 
<typeStatus id="E64F886AFFFCBE746D62F9390DD399E8" box="[593,643,1703,1728]" pageId="22" pageNumber="466">type</typeStatus>
species. Gender neuter.
</paragraph>
</subSubSection>
<caption id="6D8B6640FFFDBE756F93FB670E759A63" ID-DOI="http://doi.org/10.5281/zenodo.4609170" ID-Zenodo-Dep="4609170" httpUri="https://zenodo.org/record/4609170/files/figure.png" pageId="23" pageNumber="467" startId="23.[160,225,1273,1296]" targetBox="[163,1153,142,1241]" targetPageId="23">
<paragraph id="394B36C8FFFDBE756F93FB670E759A63" blockId="23.[160,1157,1273,1355]" pageId="23" pageNumber="467">
<emphasis id="0B80EADAFFFDBE756F93FB670E599A38" bold="true" box="[160,265,1273,1296]" pageId="23" pageNumber="467">Figure 8.</emphasis>
<taxonomicName id="FEF44D4BFFFDBE756E2DFB670D759A38" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1837" box="[286,549,1273,1296]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="0B80EADAFFFDBE756E2DFB670D759A38" box="[286,549,1273,1296]" italics="true" pageId="23" pageNumber="467">Shinobium trapezoideum</emphasis>
</taxonomicName>
(H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view.
</paragraph>
</caption>
<subSubSection id="71EE6543FFFDBE7B6F93FAE70CC49B05" lastPageId="25" lastPageNumber="469" pageId="23" pageNumber="467" type="discussion">
<paragraph id="394B36C8FFFDBE756F93FAE70E569ABB" blockId="23.[160,1156,1401,1705]" box="[160,262,1401,1427]" pageId="23" pageNumber="467">
<emphasis id="0B80EADAFFFDBE756F93FAE70E569ABB" bold="true" box="[160,262,1401,1427]" italics="true" pageId="23" pageNumber="467">Remarks</emphasis>
</paragraph>
<paragraph id="394B36C8FFFDBE7A6F93FA030CE59BF3" blockId="23.[160,1156,1401,1705]" lastBlockId="24.[160,1156,940,1243]" lastPageId="24" lastPageNumber="468" pageId="23" pageNumber="467">
<taxonomicName id="FEF44D4BFFFDBE756F93FA030E479A9E" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[160,279,1437,1462]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756F93FA030E479A9E" box="[160,279,1437,1462]" italics="true" pageId="23" pageNumber="467">Shinobium</emphasis>
</taxonomicName>
is morphologically most similar to 
<taxonomicName id="FEF44D4BFFFDBE756D8DFA030C6C9A9E" authorityName=", Serene and Soh" authorityYear="1970" box="[702,828,1437,1462]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756D8DFA030C6C9A9E" box="[702,828,1437,1462]" italics="true" pageId="23" pageNumber="467">Labuanium</emphasis>
</taxonomicName>
in its proportionally longer carapace, but the former can be distinguished from the latter by many characters of the carapace, male thoracic sternum, cheliped and ambulatory legs (
<tableCitation id="74760373FFFDBE756B24FA7C0B269AD3" box="[1047,1142,1506,1531]" captionStart="Table 1" captionStartId="5.[133,189,438,461]" captionTargetPageId="5" captionText="Table 1. Diagnostic characters of Labuanium Serène and Soh, 1970, Circulium gen. nov., Shinobium gen. nov. and Scandarma Schubart, Liu and Cuesta, 2003." httpUri="http://table.plazi.org/id/6D8B6640FFEFBE676FB6FE2809C99EE5" pageId="23" pageNumber="467" tableUuid="6D8B6640FFEFBE676FB6FE2809C99EE5">Table 1</tableCitation>
). 
<taxonomicName id="FEF44D4BFFFDBE756F93F99B0E479936" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[160,279,1541,1566]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756F93F99B0E479936" box="[160,279,1541,1566]" italics="true" pageId="23" pageNumber="467">Shinobium</emphasis>
</taxonomicName>
is characterised by a distinctly trapezoidal carapace (
<figureCitation id="A1CF2A4DFFFDBE756CBDF99B0CA09936" box="[910,1008,1541,1566]" captionStart="Figure 8" captionStartId="23.[160,225,1273,1296]" captionTargetBox="[163,1153,142,1241]" captionTargetId="figure-114@23.[163,1153,142,1242]" captionTargetPageId="23" captionText="Figure 8. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609170" httpUri="https://zenodo.org/record/4609170/files/figure.png" pageId="23" pageNumber="467">Figure 8</figureCitation>
(a)), whereas the carapace of 
<taxonomicName id="FEF44D4BFFFDBE756E53F9B60E8E9969" authorityName=", Serene and Soh" authorityYear="1970" box="[352,478,1576,1601]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756E53F9B60E8E9969" box="[352,478,1576,1601]" italics="true" pageId="23" pageNumber="467">Labuanium</emphasis>
</taxonomicName>
is longitudinally rectangular (
<figureCitation id="A1CF2A4DFFFDBE756C0CF9B60CF59969" box="[831,933,1576,1601]" captionStart="Figure 1" captionStartId="8.[160,225,1675,1698]" captionTargetBox="[163,1153,1314,1644]" captionTargetId="figure-202@8.[163,1153,1314,1644]" captionTargetPageId="8" captionText="Figure 1. Labuanium politum (De Man, 1887). Lectotype, NHM 1886.52, male, 18.7 × 17.1 mm (Mergui Archipelago). a, habitus, dorsal view; b, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4609152" httpUri="https://zenodo.org/record/4609152/files/figure.png" pageId="23" pageNumber="467">Figures 1</figureCitation>
(a), 2(a)). The male pleon of 
<taxonomicName id="FEF44D4BFFFDBE756E2BF9D40EDF994B" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[280,399,1610,1635]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756E2BF9D40EDF994B" box="[280,399,1610,1635]" italics="true" pageId="23" pageNumber="467">Shinobium</emphasis>
</taxonomicName>
is proportionally shorter and there is a substantial distance between the anterior end of the sternopleonal cavity and the border between thoracic sternites 3 and 4 (
<figureCitation id="A1CF2A4DFFFDBE756EC6F90E0D399981" box="[501,617,1680,1705]" captionStart="Figure 10" captionStartId="24.[160,225,1643,1666]" captionTargetBox="[163,1153,1287,1611]" captionTargetId="figure-182@24.[163,1153,1287,1611]" captionTargetPageId="24" captionText="Figure 10. Shinobium trapezoideum (H. Milne Edwards, 1837) (Holotype of Sesarma trapezoideum longitarsis var. longitarsis De Man, 1889). SMF 1980, male, 31.4 × 28.9 mm (Fiji). a, habitus, dorsal view; b, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4609174" httpUri="https://zenodo.org/record/4609174/files/figure.png" pageId="23" pageNumber="467">Figure 10</figureCitation>
(b)). In 
<taxonomicName id="FEF44D4BFFFDBE756DF4F90E0C159981" authorityName=", Serene and Soh" authorityYear="1970" box="[711,837,1680,1705]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFFDBE756DF4F90E0C159981" box="[711,837,1680,1705]" italics="true" pageId="23" pageNumber="467">Labuanium</emphasis>
</taxonomicName>
, the distance is distinctly shorter (
<figureCitation id="A1CF2A4DFFF2BE7A6E34FC320E369CED" box="[263,358,940,965]" captionStart="Figure 1" captionStartId="8.[160,225,1675,1698]" captionTargetBox="[163,1153,1314,1644]" captionTargetId="figure-202@8.[163,1153,1314,1644]" captionTargetPageId="8" captionText="Figure 1. Labuanium politum (De Man, 1887). Lectotype, NHM 1886.52, male, 18.7 × 17.1 mm (Mergui Archipelago). a, habitus, dorsal view; b, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4609152" httpUri="https://zenodo.org/record/4609152/files/figure.png" pageId="24" pageNumber="468">Figure 1</figureCitation>
(b)). Rim-like structures of the cheliped palm are also present in 
<taxonomicName id="FEF44D4BFFF2BE7A6F93FC510E479CC0" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[160,279,975,1000]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6F93FC510E479CC0" box="[160,279,975,1000]" italics="true" pageId="24" pageNumber="468">Shinobium</emphasis>
</taxonomicName>
(
<figureCitation id="A1CF2A4DFFF2BE7A6E18FC510EDF9CC0" box="[299,399,975,1000]" captionStart="Figure 9" captionStartId="24.[160,225,810,833]" captionTargetBox="[163,1153,142,778]" captionTargetId="figure-142@24.[163,1153,142,778]" captionTargetPageId="24" captionText="Figure 9. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609172" httpUri="https://zenodo.org/record/4609172/files/figure.png" pageId="24" pageNumber="468">Figure 9</figureCitation>
(b)), but it is entire throughout its length without interruption, unlike 
<taxonomicName id="FEF44D4BFFF2BE7A6FDCFC6C0E3D9B23" authorityName=", Serene and Soh" authorityYear="1970" box="[239,365,1010,1035]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6FDCFC6C0E3D9B23" box="[239,365,1010,1035]" italics="true" pageId="24" pageNumber="468">Labuanium</emphasis>
</taxonomicName>
(
<figureCitation id="A1CF2A4DFFF2BE7A6E4FFC6C0E8E9B23" box="[380,478,1010,1035]" captionStart="Figure 3" captionStartId="10.[160,225,683,706]" captionTargetBox="[163,1153,142,651]" captionTargetId="figure-9@10.[163,1153,142,651]" captionTargetPageId="10" captionText="Figure 3. Labuanium politum (De Man, 1887). ZRC 2003.0381, male, 40.1 × 36.2 mm (Bohol, Philippines). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5. Arrow denotes rim." figureDoi="http://doi.org/10.5281/zenodo.4609158" httpUri="https://zenodo.org/record/4609158/files/figure.png" pageId="24" pageNumber="468">Figure 3</figureCitation>
(b)). In addition, the anterodistal corner of the ambulatory meri in 
<taxonomicName id="FEF44D4BFFF2BE7A6FCFFB8A0E239B05" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[252,371,1044,1069]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6FCFFB8A0E239B05" box="[252,371,1044,1069]" italics="true" pageId="24" pageNumber="468">Shinobium</emphasis>
</taxonomicName>
has a falcate lamella (
<figureCitation id="A1CF2A4DFFF2BE7A6D49FB8A0D8B9B05" box="[634,731,1044,1069]" captionStart="Figure 9" captionStartId="24.[160,225,810,833]" captionTargetBox="[163,1153,142,778]" captionTargetId="figure-142@24.[163,1153,142,778]" captionTargetPageId="24" captionText="Figure 9. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609172" httpUri="https://zenodo.org/record/4609172/files/figure.png" pageId="24" pageNumber="468">Figure 9</figureCitation>
(d)), but that of 
<taxonomicName id="FEF44D4BFFF2BE7A6CA4FB8A0B459B05" authorityName=", Serene and Soh" authorityYear="1970" box="[919,1045,1044,1069]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6CA4FB8A0B459B05" box="[919,1045,1044,1069]" italics="true" pageId="24" pageNumber="468">Labuanium</emphasis>
</taxonomicName>
is simply pointed (
<figureCitation id="A1CF2A4DFFF2BE7A6E3CFBA90E249B78" box="[271,372,1079,1104]" captionStart="Figure 3" captionStartId="10.[160,225,683,706]" captionTargetBox="[163,1153,142,651]" captionTargetId="figure-9@10.[163,1153,142,651]" captionTargetPageId="10" captionText="Figure 3. Labuanium politum (De Man, 1887). ZRC 2003.0381, male, 40.1 × 36.2 mm (Bohol, Philippines). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5. Arrow denotes rim." figureDoi="http://doi.org/10.5281/zenodo.4609158" httpUri="https://zenodo.org/record/4609158/files/figure.png" pageId="24" pageNumber="468">Figure 3</figureCitation>
(d)). The ambulatory dactyli of 
<taxonomicName id="FEF44D4BFFF2BE7A6DC2FBA90C389B78" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[753,872,1079,1104]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6DC2FBA90C389B78" box="[753,872,1079,1104]" italics="true" pageId="24" pageNumber="468">Shinobium</emphasis>
</taxonomicName>
are also proportionally longer than those of 
<taxonomicName id="FEF44D4BFFF2BE7A6EAAFBC40D479B5B" authorityName=", Serene and Soh" authorityYear="1970" box="[409,535,1114,1139]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6EAAFBC40D479B5B" box="[409,535,1114,1139]" italics="true" pageId="24" pageNumber="468">Labuanium</emphasis>
</taxonomicName>
(
<figureCitation id="A1CF2A4DFFF2BE7A6D1BFBC40DC29B5B" box="[552,658,1114,1139]" captionStart="Figure 8" captionStartId="23.[160,225,1273,1296]" captionTargetBox="[163,1153,142,1241]" captionTargetId="figure-114@23.[163,1153,142,1242]" captionTargetPageId="23" captionText="Figure 8. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609170" httpUri="https://zenodo.org/record/4609170/files/figure.png" pageId="24" pageNumber="468">Figures 8</figureCitation>
(a), 10 vs 
<figureCitation id="A1CF2A4DFFF2BE7A6C30FBC40C3E9B5B" box="[771,878,1114,1139]" captionStart="Figure 1" captionStartId="8.[160,225,1675,1698]" captionTargetBox="[163,1153,1314,1644]" captionTargetId="figure-202@8.[163,1153,1314,1644]" captionTargetPageId="8" captionText="Figure 1. Labuanium politum (De Man, 1887). Lectotype, NHM 1886.52, male, 18.7 × 17.1 mm (Mergui Archipelago). a, habitus, dorsal view; b, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4609152" httpUri="https://zenodo.org/record/4609152/files/figure.png" pageId="24" pageNumber="468">Figures 1</figureCitation>
, 
<figureCitation id="A1CF2A4DFFF2BE7A6C4EFBC40CDB9B5B" box="[893,907,1114,1139]" captionStart="Figure 2" captionStartId="9.[160,225,1370,1393]" captionTargetBox="[163,1153,142,1338]" captionTargetId="figure-84@9.[163,1153,142,1338]" captionTargetPageId="9" captionText="Figure 2. Labuanium politum (De Man, 1887). ZRC 2003.0381, male, 40.1 × 36.2 mm (Bohol, Philippines). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609154" httpUri="https://zenodo.org/record/4609154/files/figure.png" pageId="24" pageNumber="468">2</figureCitation>
(a)). The shape of the G1 also differs significantly between the two genera; 
<taxonomicName id="FEF44D4BFFF2BE7A6C7BFBE20CEF9BBD" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[840,959,1148,1173]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6C7BFBE20CEF9BBD" box="[840,959,1148,1173]" italics="true" pageId="24" pageNumber="468">Shinobium</emphasis>
</taxonomicName>
has a medially narrowed shaft of the G1 with a bulb-like subdistal part (
<figureCitation id="A1CF2A4DFFF2BE7A6C6CFB010C9C9B90" box="[863,972,1183,1208]" captionStart="Figure 11" captionStartId="26.[160,225,778,801]" captionTargetBox="[173,1150,144,728]" captionTargetId="figure-302@26.[163,1153,142,746]" captionTargetPageId="26" captionText="Figure 11. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, pleon; b–e, left G1; b, ventral view; c, dorsal view; d, distal part, ventral view, denuded; e, distal part, dorsal view, denuded. Scale bars: a= 5 mm; b–e = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4609176" httpUri="https://zenodo.org/record/4609176/files/figure.png" pageId="24" pageNumber="468">Figure 11</figureCitation>
(b–e)), whereas that of 
<taxonomicName id="FEF44D4BFFF2BE7A6FC9FB5C0E289BF3" authorityName=", Serene and Soh" authorityYear="1970" box="[250,376,1218,1243]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF2BE7A6FC9FB5C0E289BF3" box="[250,376,1218,1243]" italics="true" pageId="24" pageNumber="468">Labuanium</emphasis>
</taxonomicName>
has a more or less straight shaft (
<figureCitation id="A1CF2A4DFFF2BE7A6C26FB5C0C289BF3" box="[789,888,1218,1243]" captionStart="Figure 4" captionStartId="10.[160,225,1649,1672]" captionTargetBox="[166,1150,835,1615]" captionTargetId="figure-52@10.[163,1153,832,1618]" captionTargetPageId="10" captionText="Figure 4. Labuanium politum (De Man, 1887). ZRC 1965.7.29.58, male 40.8 × 36.2 mm (Borneo, Malaysia). a, pleon and telson; b, left G1, ventral view; c, distal part of left G1, dorsal view. Scale bars: a= 5 mm; b, c= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4609160" httpUri="https://zenodo.org/record/4609160/files/figure.png" pageId="24" pageNumber="468">Figure 4</figureCitation>
(b,c)).
</paragraph>
<caption id="6D8B6640FFF2BE7A6F93FCB40E3A9C54" ID-DOI="http://doi.org/10.5281/zenodo.4609172" ID-Zenodo-Dep="4609172" httpUri="https://zenodo.org/record/4609172/files/figure.png" pageId="24" pageNumber="468" startId="24.[160,225,810,833]" targetBox="[163,1153,142,778]" targetPageId="24">
<paragraph id="394B36C8FFF2BE7A6F93FCB40E3A9C54" blockId="24.[160,1157,810,892]" pageId="24" pageNumber="468">
<emphasis id="0B80EADAFFF2BE7A6F93FCB40E599C69" bold="true" box="[160,265,810,833]" pageId="24" pageNumber="468">Figure 9.</emphasis>
<taxonomicName id="FEF44D4BFFF2BE7A6E2DFCB40D759C69" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1837" box="[286,549,810,833]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="0B80EADAFFF2BE7A6E2DFCB40D759C69" box="[286,549,810,833]" italics="true" pageId="24" pageNumber="468">Shinobium trapezoideum</emphasis>
</taxonomicName>
(H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5.
</paragraph>
</caption>
<caption id="6D8B6640FFF2BE7A6F93F9F50EDE9995" ID-DOI="http://doi.org/10.5281/zenodo.4609174" ID-Zenodo-Dep="4609174" httpUri="https://zenodo.org/record/4609174/files/figure.png" pageId="24" pageNumber="468" startId="24.[160,225,1643,1666]" targetBox="[163,1153,1287,1611]" targetPageId="24">
<paragraph id="394B36C8FFF2BE7A6F93F9F50EDE9995" blockId="24.[160,1156,1643,1725]" pageId="24" pageNumber="468">
<emphasis id="0B80EADAFFF2BE7A6F93F9F50E5A99AA" bold="true" box="[160,266,1643,1666]" pageId="24" pageNumber="468">Figure 10.</emphasis>
<taxonomicName id="FEF44D4BFFF2BE7A6E26F9F50D5899AA" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1837" box="[277,520,1643,1666]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="0B80EADAFFF2BE7A6E26F9F50D5899AA" box="[277,520,1643,1666]" italics="true" pageId="24" pageNumber="468">Shinobium trapezoideum</emphasis>
</taxonomicName>
(H. Milne Edwards, 1837) (Holotype of 
<taxonomicName id="FEF44D4BFFF2BE7A6C96F9F50D4D99B7" authority="var. longitarsis De Man, 1889" authorityName="var. longitarsis De Man" authorityYear="1889" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="468" phylum="Arthropoda" rank="variety" species="trapezoideum" subSpecies="longitarsis" variety="longitarsis">
<emphasis id="0B80EADAFFF2BE7A6C96F9F50E5299B7" italics="true" pageId="24" pageNumber="468">Sesarma trapezoideum longitarsis</emphasis>
var. 
<emphasis id="0B80EADAFFF2BE7A6E03F9160EC299B7" box="[304,402,1672,1695]" italics="true" pageId="24" pageNumber="468">longitarsis</emphasis>
<bibRefCitation id="5D654B39FFF2BE7A6EA4F9160D4D99B7" author="De Man JG" box="[407,541,1672,1695]" pageId="24" pageNumber="468" pagination="409 - 452" refId="ref41120" refString="De Man JG. 1889. Uber einige neue oder seltene indopacifische Brachyuren. Zool Jahrb Abt Syst Geogr Biol Thiere. 4: 409 - 452, pls. 9 - 10." type="journal article" year="1889">De Man, 1889</bibRefCitation>
</taxonomicName>
). SMF 1980, male, 31.4 × 28.9 mm (Fiji). a, habitus, dorsal view; b, habitus, ventral view.
</paragraph>
</caption>
<paragraph id="394B36C8FFF3BE7B6FF3FF100CC49B05" blockId="25.[160,1156,142,1069]" pageId="25" pageNumber="469">
When 
<bibRefCitation id="5D654B39FFF3BE7B6E22FF100E959F8F" author="De Man JG" box="[273,453,142,167]" pageId="25" pageNumber="469" pagination="265 - 527" refId="ref41187" refString="De Man JG. 1892. Decapoden des indischen Archipels. In: Weber M, editor. Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Vol. 2, Leiden: E. J. Brill; p. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man (1892)</bibRefCitation>
described 
<taxonomicName id="FEF44D4BFFF3BE7B6D62FF100CEF9F8F" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[593,959,142,167]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6D62FF100C569F8F" box="[593,774,142,167]" italics="true" pageId="25" pageNumber="469">Sesarma weberi</emphasis>
<bibRefCitation id="5D654B39FFF3BE7B6C27FF100CEF9F8F" author="De Man JG" box="[788,959,142,167]" pageId="25" pageNumber="469" pagination="265 - 527" refId="ref41187" refString="De Man JG. 1892. Decapoden des indischen Archipels. In: Weber M, editor. Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Vol. 2, Leiden: E. J. Brill; p. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
(now placed in 
<taxonomicName id="FEF44D4BFFF3BE7B6F93FF2F0D259FE2" authority=", sensu Serene and Soh 1970" authorityName=", sensu Serene and Soh" authorityYear="1970" box="[160,629,177,202]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6F93FF2F0E469FE2" box="[160,278,177,202]" italics="true" pageId="25" pageNumber="469">Sesarmops</emphasis>
, sensu 
<bibRefCitation id="5D654B39FFF3BE7B6E46FF2F0D259FE2" author="Serene R &amp; Soh CL" box="[373,629,177,202]" pageId="25" pageNumber="469" pagination="387 - 416" refId="ref46714" refString="Serene R, Soh CL. 1970. New Indo-Pacific genera allied to Sesarma Say, 1817 (Brachyura, Decapoda, Crustacea). Treubia. 27 (4): 387 - 416." type="journal article" year="1970">Serène and Soh 1970</bibRefCitation>
</taxonomicName>
), he compared it with 
<emphasis id="0B80EADAFFF3BE7B6CBDFF2F0B029FE2" box="[910,1106,177,202]" italics="true" pageId="25" pageNumber="469">
Se. 
<taxonomicName id="FEF44D4BFFF3BE7B6C8AFF2F0B029FE2" authorityName=": Rathbun" authorityYear="1907" box="[953,1106,177,202]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="trapezoideum">trapezoideum</taxonomicName>
</emphasis>
. 
<bibRefCitation id="5D654B39FFF3BE7B6B50FF2F0E639FC5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" pageId="25" pageNumber="469" pagination="1 - 286" refId="ref44591" refString="Ng PKL, Guinot D, Davie PJF. 2008 a. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl. 17: 1 - 286." type="journal article" year="2008">Ng et al. (2008a</bibRefCitation>
, p. 224) commented that ‘ 
<taxonomicName id="FEF44D4BFFF3BE7B6D45FF4A0C329FC5" authorityName=": Nobili" authorityYear="1900" box="[630,866,212,237]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="trapezoidea">
<emphasis id="0B80EADAFFF3BE7B6D45FF4A0C329FC5" box="[630,866,212,237]" italics="true" pageId="25" pageNumber="469">Sesarma trapezoidea</emphasis>
</taxonomicName>
H. Milne Edwards, 1837 and 
<taxonomicName id="FEF44D4BFFF3BE7B6FE5FF680D679E27" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[214,567,246,271]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6FE5FF680ED89E27" box="[214,392,246,271]" italics="true" pageId="25" pageNumber="469">Sesarma weberi</emphasis>
<bibRefCitation id="5D654B39FFF3BE7B6EA0FF680D679E27" author="De Man JG" box="[403,567,246,271]" pageId="25" pageNumber="469" pagination="265 - 527" refId="ref41187" refString="De Man JG. 1892. Decapoden des indischen Archipels. In: Weber M, editor. Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Vol. 2, Leiden: E. J. Brill; p. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
should be placed in a same genus as they share a suite of cheliped, gonopodal and larval features’. These characters are the presence of a dense row of tiny tubercles on the upper margin of chelal movable fingers (
<figureCitation id="A1CF2A4DFFF3BE7B6F9BFEC00E459E5F" box="[168,277,350,375]" captionStart="Figure 9" captionStartId="24.[160,225,810,833]" captionTargetBox="[163,1153,142,778]" captionTargetId="figure-142@24.[163,1153,142,778]" captionTargetPageId="24" captionText="Figure 9. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609172" httpUri="https://zenodo.org/record/4609172/files/figure.png" pageId="25" pageNumber="469">Figures 9</figureCitation>
(c), 13(b)), trapezoidal carapace shape as well as the general form of the antennule and antenna (
<figureCitation id="A1CF2A4DFFF3BE7B6EF5FE1F0D629EB2" box="[454,562,385,410]" captionStart="Figure 8" captionStartId="23.[160,225,1273,1296]" captionTargetBox="[163,1153,142,1241]" captionTargetId="figure-114@23.[163,1153,142,1242]" captionTargetPageId="23" captionText="Figure 8. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609170" httpUri="https://zenodo.org/record/4609170/files/figure.png" pageId="25" pageNumber="469">Figures 8</figureCitation>
(a), 12, 13(a, b, d), 14(b–d)) (
<bibRefCitation id="5D654B39FFF3BE7B6CB5FE1F0B6D9EB2" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[902,1085,385,410]" pageId="25" pageNumber="469" pagination="1 - 286" refId="ref44591" refString="Ng PKL, Guinot D, Davie PJF. 2008 a. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl. 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008a</bibRefCitation>
; 
<bibRefCitation id="5D654B39FFF3BE7B6B7DFE1F0E499E95" author="Jeng MS &amp; Liu HC &amp; Tzeng CS &amp; Ng PKL" pageId="25" pageNumber="469" pagination="227 - 240" refId="ref42333" refString="Jeng MS, Liu HC, Tzeng CS, Ng PKL. 2003. On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana. 76: 227 - 240. doi: 10.1163 / 156854003321824576" type="journal article" year="2003">Jeng et al. 2003</bibRefCitation>
. 
<taxonomicName id="FEF44D4BFFF3BE7B6E14FE3A0EA09E95" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[295,496,420,445]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6E14FE3A0EA09E95" box="[295,496,420,445]" italics="true" pageId="25" pageNumber="469">Sesarmops weberi</emphasis>
</taxonomicName>
also differs from all known congeners in having a dense row of tiny tubercles on the upper margin of the movable finger of the cheliped in both the male and the female (
<figureCitation id="A1CF2A4DFFF3BE7B6D3EFE770D2C9D2A" box="[525,636,489,514]" captionStart="Figure 13" captionStartId="28.[160,225,840,863]" captionTargetBox="[163,1153,142,809]" captionTargetId="figure-9@28.[163,1153,142,809]" captionTargetPageId="28" captionText="Figure 13. Sesarmops weberi (De Man, 1892). a–c, lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia); d, ZRC 2018.0771, male, 16.2 × 16.3 (Cebu, Philippines). a, major chela, outer view; b, major chela, upper view; c, cephalothorax, ventral view; d, major chela, inner view." figureDoi="http://doi.org/10.5281/zenodo.4609180" httpUri="https://zenodo.org/record/4609180/files/figure.png" pageId="25" pageNumber="469">Figure 13</figureCitation>
(b)). 
<taxonomicName id="FEF44D4BFFF3BE7B6D9CFE770C279D2A" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[687,887,489,514]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6D9CFE770C279D2A" box="[687,887,489,514]" italics="true" pageId="25" pageNumber="469">Sesarmops weberi</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="FEF44D4BFFF3BE7B6B3EFE770BD49D2A" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[1037,1156,489,514]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6B3EFE770BD49D2A" box="[1037,1156,489,514]" italics="true" pageId="25" pageNumber="469">Shinobium</emphasis>
</taxonomicName>
in its relatively trapezoidal carapace (
<figureCitation id="A1CF2A4DFFF3BE7B6D50FD920D859D0D" box="[611,725,524,549]" captionStart="Figure 12" captionStartId="27.[160,225,1246,1269]" captionTargetBox="[163,1153,142,1214]" captionTargetId="figure-110@27.[163,1153,142,1215]" captionTargetPageId="27" captionText="Figure 12. Sesarmops weberi (De Man, 1892). Lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia). a, habitus, dorsal view; b, cephalothorax, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4609178" httpUri="https://zenodo.org/record/4609178/files/figure.png" pageId="25" pageNumber="469">Figure 12</figureCitation>
(a)), setose posterolateral regions of the carapace, and weakly lamellar anterodistal end of ambulatory meri (
<figureCitation id="A1CF2A4DFFF3BE7B6CDFFDB00B0A9D6F" box="[1004,1114,558,583]" captionStart="Figure 12" captionStartId="27.[160,225,1246,1269]" captionTargetBox="[163,1153,142,1214]" captionTargetId="figure-110@27.[163,1153,142,1215]" captionTargetPageId="27" captionText="Figure 12. Sesarmops weberi (De Man, 1892). Lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia). a, habitus, dorsal view; b, cephalothorax, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4609178" httpUri="https://zenodo.org/record/4609178/files/figure.png" pageId="25" pageNumber="469">Figure 12</figureCitation>
(a)). Nevertheless, 
<taxonomicName id="FEF44D4BFFF3BE7B6E71FDCF0D5A9D42" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[322,522,593,618]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6E71FDCF0D5A9D42" box="[322,522,593,618]" italics="true" pageId="25" pageNumber="469">Sesarmops weberi</emphasis>
</taxonomicName>
is not assigned to 
<taxonomicName id="FEF44D4BFFF3BE7B6DD8FDCF0C329D42" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[747,866,593,618]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6DD8FDCF0C329D42" box="[747,866,593,618]" italics="true" pageId="25" pageNumber="469">Shinobium</emphasis>
</taxonomicName>
because of its stout and almost straight G1 with distal corneous beak-like process that is directed laterally (
<figureCitation id="A1CF2A4DFFF3BE7B6F9BFD080E4A9D87" box="[168,282,662,687]" captionStart="Figure 14" captionStartId="28.[160,225,1572,1595]" captionTargetBox="[165,1151,1023,1538]" captionTargetId="figure-60@28.[163,1153,1021,1540]" captionTargetPageId="28" captionText="Figure 14. Sesarmops weberi (De Man, 1892). Lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia). a, pleonal somites 3–6 and telson; b–d, left G1; b, ventral view; c, distal part, ventral view; d, distal part, dorsal view. Scale bars: a= 5 mm; b–d = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4609182" httpUri="https://zenodo.org/record/4609182/files/figure.png" pageId="25" pageNumber="469">Figure 14</figureCitation>
(b–d)) (vs medially narrowed shaft of G1 with bulb-like subdistal part in 
<taxonomicName id="FEF44D4BFFF3BE7B6F93FD270E479DFA" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[160,279,697,722]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6F93FD270E479DFA" box="[160,279,697,722]" italics="true" pageId="25" pageNumber="469">Shinobium</emphasis>
</taxonomicName>
, 
<figureCitation id="A1CF2A4DFFF3BE7B6E15FD270EC09DFA" box="[294,400,697,722]" captionStart="Figure 11" captionStartId="26.[160,225,778,801]" captionTargetBox="[173,1150,144,728]" captionTargetId="figure-302@26.[163,1153,142,746]" captionTargetPageId="26" captionText="Figure 11. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, pleon; b–e, left G1; b, ventral view; c, dorsal view; d, distal part, ventral view, denuded; e, distal part, dorsal view, denuded. Scale bars: a= 5 mm; b–e = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4609176" httpUri="https://zenodo.org/record/4609176/files/figure.png" pageId="25" pageNumber="469">Figure 11</figureCitation>
(b–e)), lateral and median lobes of epistome directed anteroventrally and ventrally, respectively (
<figureCitation id="A1CF2A4DFFF3BE7B6D04FD420DFB9DDD" box="[567,683,732,757]" captionStart="Figure 12" captionStartId="27.[160,225,1246,1269]" captionTargetBox="[163,1153,142,1214]" captionTargetId="figure-110@27.[163,1153,142,1215]" captionTargetPageId="27" captionText="Figure 12. Sesarmops weberi (De Man, 1892). Lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia). a, habitus, dorsal view; b, cephalothorax, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4609178" httpUri="https://zenodo.org/record/4609178/files/figure.png" pageId="25" pageNumber="469">Figure 12</figureCitation>
(b)) (vs both lateral and median lobes directed anteriorly, 
<figureCitation id="A1CF2A4DFFF3BE7B6EBDFD600EA39C3F" box="[398,499,766,791]" captionStart="Figure 8" captionStartId="23.[160,225,1273,1296]" captionTargetBox="[163,1153,142,1241]" captionTargetId="figure-114@23.[163,1153,142,1242]" captionTargetPageId="23" captionText="Figure 8. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609170" httpUri="https://zenodo.org/record/4609170/files/figure.png" pageId="25" pageNumber="469">Figure 8</figureCitation>
(b)), the rim along the occlusal margin of immovable finger to dactylar articulation is disconnected only at the inner surface of male major chela (
<figureCitation id="A1CF2A4DFFF3BE7B6FC2FCDA0E339C75" box="[241,355,836,861]" captionStart="Figure 13" captionStartId="28.[160,225,840,863]" captionTargetBox="[163,1153,142,809]" captionTargetId="figure-9@28.[163,1153,142,809]" captionTargetPageId="28" captionText="Figure 13. Sesarmops weberi (De Man, 1892). a–c, lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia); d, ZRC 2018.0771, male, 16.2 × 16.3 (Cebu, Philippines). a, major chela, outer view; b, major chela, upper view; c, cephalothorax, ventral view; d, major chela, inner view." figureDoi="http://doi.org/10.5281/zenodo.4609180" httpUri="https://zenodo.org/record/4609180/files/figure.png" pageId="25" pageNumber="469">Figure 13</figureCitation>
(d)) (vs rim not disconnected on both inner and outer surfaces of both male and female chelae, 
<figureCitation id="A1CF2A4DFFF3BE7B6D37FCF80D349C57" box="[516,612,870,895]" captionStart="Figure 9" captionStartId="24.[160,225,810,833]" captionTargetBox="[163,1153,142,778]" captionTargetId="figure-142@24.[163,1153,142,778]" captionTargetPageId="24" captionText="Figure 9. Shinobium trapezoideum (H. Milne Edwards, 1837). ZRC 2018.0051, male, 29.5 × 26.8 mm (Vanuatu). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609172" httpUri="https://zenodo.org/record/4609172/files/figure.png" pageId="25" pageNumber="469">Figure 9</figureCitation>
(b)), and the male pleonal somite 6 is rounded laterally and has subequal proportions (
<figureCitation id="A1CF2A4DFFF3BE7B6DA2FC170C559C8A" box="[657,773,905,930]" captionStart="Figure 14" captionStartId="28.[160,225,1572,1595]" captionTargetBox="[165,1151,1023,1538]" captionTargetId="figure-60@28.[163,1153,1021,1540]" captionTargetPageId="28" captionText="Figure 14. Sesarmops weberi (De Man, 1892). Lectotype, RMNH D2615, male, 14.5 × 15.0 mm (Flores, Indonesia). a, pleonal somites 3–6 and telson; b–d, left G1; b, ventral view; c, distal part, ventral view; d, distal part, dorsal view. Scale bars: a= 5 mm; b–d = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4609182" httpUri="https://zenodo.org/record/4609182/files/figure.png" pageId="25" pageNumber="469">Figure 14</figureCitation>
(a)) (vs distolaterally angulated, proportionally longer in 
<taxonomicName id="FEF44D4BFFF3BE7B6EE2FC320D189CED" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[465,584,940,965]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6EE2FC320D189CED" box="[465,584,940,965]" italics="true" pageId="25" pageNumber="469">Shinobium</emphasis>
</taxonomicName>
, 
<figureCitation id="A1CF2A4DFFF3BE7B6D6EFC320D829CED" box="[605,722,940,965]" captionStart="Figure 10" captionStartId="24.[160,225,1643,1666]" captionTargetBox="[163,1153,1287,1611]" captionTargetId="figure-182@24.[163,1153,1287,1611]" captionTargetPageId="24" captionText="Figure 10. Shinobium trapezoideum (H. Milne Edwards, 1837) (Holotype of Sesarma trapezoideum longitarsis var. longitarsis De Man, 1889). SMF 1980, male, 31.4 × 28.9 mm (Fiji). a, habitus, dorsal view; b, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4609174" httpUri="https://zenodo.org/record/4609174/files/figure.png" pageId="25" pageNumber="469">Figure 10</figureCitation>
(b)). These differences indicate that 
<taxonomicName id="FEF44D4BFFF3BE7B6F93FC500E3F9CCF" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[160,367,974,999]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6F93FC500E3F9CCF" box="[160,367,974,999]" italics="true" pageId="25" pageNumber="469">Sesarmops weberi</emphasis>
</taxonomicName>
is not congeneric with 
<taxonomicName id="FEF44D4BFFF3BE7B6D96FC500C4C9CCF" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[677,796,974,999]" class="Malacostraca" family="Sesarmidae" genus="Shinobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6D96FC500C4C9CCF" box="[677,796,974,999]" italics="true" pageId="25" pageNumber="469">Shinobium</emphasis>
</taxonomicName>
. An ongoing revision of the genus 
<taxonomicName id="FEF44D4BFFF3BE7B6FC0FC6F0E399B22" authorityName=", sensu Serene and Soh" authorityYear="1970" box="[243,361,1009,1034]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6FC0FC6F0E399B22" box="[243,361,1009,1034]" italics="true" pageId="25" pageNumber="469">Sesarmops</emphasis>
</taxonomicName>
by the second author and C.D. Schubart will clarify the generic position of ‘ 
<taxonomicName id="FEF44D4BFFF3BE7B6E1CFB8A0EF59B05" authorityName=", sensu Serene and Soh" authorityYear="1970" box="[303,421,1044,1069]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF3BE7B6E1CFB8A0EF59B05" box="[303,421,1044,1069]" italics="true" pageId="25" pageNumber="469">Sesarmops</emphasis>
</taxonomicName>
’ 
<taxonomicName id="FEF44D4BFFF3BE7B6E85FB8A0EAF9B05" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[438,511,1044,1069]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="469" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="0B80EADAFFF3BE7B6E85FB8A0EAF9B05" box="[438,511,1044,1069]" italics="true" pageId="25" pageNumber="469">weberi</emphasis>
</taxonomicName>
and allied species at a later date.
</paragraph>
</subSubSection>
</treatment>
</document>