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<document id="74CB0228682DAC706A33ACE28AB72C3F" ID-CLB-Dataset="55729" ID-DOI="10.1080/00222933.2020.1763491" ID-GBIF-Dataset="fb05d004-0f2d-44da-b91e-b6e8304ab2a5" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1615924067624" checkinUser="carolina" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2020" docId="B15D87DEFFDFBE546EB1FF100DEF9DDB" docLanguage="en" docName="JNATHIST.54.7-8.445-532.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 54 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2020.1763491" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scandarma splendidum Naruse and Ng 2007" docType="treatment" docVersion="9" lastPageNumber="498" masterDocId="4D64FFA6FFEABE626F33FF9E0F509F28" masterDocTitle="Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species" masterLastPageNumber="532" masterPageNumber="445" pageNumber="497" updateTime="1698933465109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="40D0E5081E5DA99BC022628699D37E83" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2C32CE5BFB3FCBC7C9B1AF1AF4B71A5B">Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species</mods:title>
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<mods:namePart id="12A8E1BA2E59B08DB046DED9CAB7211C">Naruse, Tohru</mods:namePart>
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<mods:namePart id="DF95C656F65FA008184BD1CD2ACF9894">Ng, Peter K. L.</mods:namePart>
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<mods:title id="D10BF17C92B2FD6F899E41729B2EC7E7">Journal of Natural History</mods:title>
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<mods:date id="FC073968BEF5DEAA5F8FA31082BF5F04">2020</mods:date>
<mods:detail id="6C2C02152D9DE62C68CE4416EC182B4B" type="series">
<mods:title id="29E7509BCBEA2AC06AE2B775107DCAFB">J. Nat. Hist.</mods:title>
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<mods:number id="B2AD77047512B89CD1BC6BBC17A36747">2020-08-12</mods:number>
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<mods:detail id="1C035882459E4A27F8E9B48D70CC97B7" type="volume">
<mods:number id="1E2460C2C22E411228635B5507DA3DED">54</mods:number>
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<mods:number id="783D379C129EBE8690C6F50F634CA7A3">7 - 8</mods:number>
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<mods:classification id="E2D3C299E55DE98A9A2880A0DA595099">journal article</mods:classification>
<mods:identifier id="182E8A4595501B148C6142A5A909DE7C" type="CLB-Dataset">55729</mods:identifier>
<mods:identifier id="0D2CC49944B0DAAA981438E837FA7A2F" type="DOI">10.1080/00222933.2020.1763491</mods:identifier>
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<treatment id="B15D87DEFFDFBE546EB1FF100DEF9DDB" ID-DOI="http://doi.org/10.5281/zenodo.4609131" ID-GBIF-Taxon="195191665" ID-Zenodo-Dep="4609131" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB" httpUri="http://treatment.plazi.org/id/B15D87DEFFDFBE546EB1FF100DEF9DDB" lastPageId="54" lastPageNumber="498" pageId="53" pageNumber="497">
<subSubSection id="71EE6543FFDFBE576EB1FF100CF19F80" box="[386,929,142,168]" pageId="53" pageNumber="497" type="nomenclature">
<paragraph id="394B36C8FFDFBE576EB1FF100CF19F80" blockId="53.[386,929,142,168]" box="[386,929,142,168]" pageId="53" pageNumber="497">
<heading id="620381A4FFDFBE576EB1FF100CF19F80" box="[386,929,142,168]" centered="true" fontSize="11" level="2" pageId="53" pageNumber="497" reason="2">
<taxonomicName id="FEF44D4BFFDFBE576EB1FF100CF19F80" ID-CoL="6Y49X" authority="Naruse and Ng, 2007" authorityName="Naruse and Ng" authorityYear="2007" box="[386,929,142,168]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="497" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDFBE576EB1FF100DF29F80" bold="true" box="[386,674,142,168]" italics="true" pageId="53" pageNumber="497">Scandarma splendidum</emphasis>
<bibRefCitation id="5D654B39FFDFBE576D99FF100CF19F80" author="Naruse T &amp; Ng PKL" box="[682,929,142,168]" pageId="53" pageNumber="497" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng, 2007</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDFBE576F93FF5D0B439FD7" pageId="53" pageNumber="497" type="reference_group">
<paragraph id="394B36C8FFDFBE576F93FF5D0B439FD7" blockId="53.[160,1155,195,255]" pageId="53" pageNumber="497">
<treatmentCitationGroup id="19E411E6FFDFBE576F93FF5D0B439FD7" pageId="53" pageNumber="497">
<treatmentCitation id="B85510D9FFDFBE576F93FF5D0DB59FF4" LSID="urn:lsid:zoobank.org:act:0B850C3C-8D05-4DD0-BE77-E43503317325" author="Naruse T &amp; Ng PKL" box="[160,741,195,220]" httpUri="http://treatment.plazi.org/id/AA1E8792955EFFE6FDE6FAB211082F43" page="337" pageId="53" pageNumber="497" year="2007">
<taxonomicName id="FEF44D4BFFDFBE576F93FF5D0DB59FF4" ID-CoL="6Y49X" authority="Naruse and Ng, 2007, p. 337" authorityName="Naruse and Ng" authorityPageNumber="337" authorityYear="2007" box="[160,741,195,220]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="497" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDFBE576F93FF5D0EF19FF4" box="[160,417,195,220]" italics="true" pageId="53" pageNumber="497">Scandarma splendidum</emphasis>
<bibRefCitation id="5D654B39FFDFBE576E94FF5D0DC19FF4" author="Naruse T &amp; Ng PKL" box="[423,657,195,220]" pageId="53" pageNumber="497" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng, 2007</bibRefCitation>
, p. 337
</taxonomicName>
</treatmentCitation>
, figs 1–3 [
<typeStatus id="E64F886AFFDFBE576C62FF5D0CD59FF4" box="[849,901,195,220]" pageId="53" pageNumber="497">type</typeStatus>
locality:
<location id="3C2B6013FFDFBE576CDBFF5D0FA39FD7" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB:3C2B6013FFDFBE576CDBFF5D0FA39FD7" country="Malaysia" name="Bako National Park" pageId="53" pageNumber="497">Bako National Park</location>
,
<collectingCountry id="41E37658FFDFBE576FCEFF780E0C9FD7" box="[253,348,230,255]" name="Malaysia" pageId="53" pageNumber="497">Sarawak</collectingCountry>
,
<collectingCountry id="41E37658FFDFBE576E54FF780E9C9FD7" box="[359,460,230,255]" name="Malaysia" pageId="53" pageNumber="497">Malaysia</collectingCountry>
];
<treatmentCitation id="B85510D9FFDFBE576EE8FF780D889FD7" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[475,728,230,255]" page="223" pageId="53" pageNumber="497" year="2008">
<bibRefCitation id="5D654B39FFDFBE576EE8FF780DD59FD7" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[475,645,230,255]" pageId="53" pageNumber="497" pagination="1 - 286" refId="ref44591" refString="Ng PKL, Guinot D, Davie PJF. 2008 a. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl. 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008a</bibRefCitation>
, p. 223
</treatmentCitation>
;
<treatmentCitation id="B85510D9FFDFBE576DD0FF780B5F9FD7" author="Lai F &amp; Olesen B" box="[739,1039,230,255]" page="72" pageId="53" pageNumber="497" year="2016">
<bibRefCitation id="5D654B39FFDFBE576DD0FF780C989FD7" author="Lai F &amp; Olesen B" box="[739,968,230,255]" pageId="53" pageNumber="497" refId="ref42514" refString="Lai F, Olesen B. 2016. A visual celebration of Borneo's wildlife. Singapore: Bjorn Olesen Wildlife Photography." type="book" year="2016">Lai and Olesen 2016</bibRefCitation>
, p. 72
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDFBE576F93FEAB0CC79DB3" pageId="53" pageNumber="497" type="materials_examined">
<paragraph id="394B36C8FFDFBE576F93FEAB0E2F9E67" blockId="53.[160,1156,309,404]" box="[160,383,309,335]" pageId="53" pageNumber="497">
<emphasis id="0B80EADAFFDFBE576F93FEAB0E2F9E67" bold="true" box="[160,383,309,335]" italics="true" pageId="53" pageNumber="497">Material examined</emphasis>
</paragraph>
<paragraph id="394B36C8FFDFBE576F93FEC90E1E9EBC" blockId="53.[160,1156,309,404]" pageId="53" pageNumber="497">
<materialsCitation id="899C3C95FFDFBE576F93FEC90E1E9EBC" ID-GBIF-Occurrence="3762866377" collectingDate="2002-03-15" collectionCode="ZRC" country="Malaysia" location="Bako National Park" pageId="53" pageNumber="497" specimenCode="ZRC 2002.0265" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="0B80EADAFFDFBE576F93FEC90E429E59" bold="true" box="[160,274,343,369]" italics="true" pageId="53" pageNumber="497">
<typeStatus id="E64F886AFFDFBE576F93FEC90E5D9E59" box="[160,269,343,369]" pageId="53" pageNumber="497" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="69529EB3FFDFBE576E12FEC60E9A9E59" box="[289,458,344,369]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="497">ZRC 2002.0265</specimenCode>
,
<specimenCount id="2FF2FD41FFDFBE576EE2FEC60D4E9E59" box="[465,542,344,369]" pageId="53" pageNumber="497" type="male">1 male</specimenCount>
, 19.1 ×
<quantity id="FE0C9B2DFFDFBE576D43FEC60D839E59" box="[624,723,344,369]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="53" pageNumber="497" unit="mm" value="18.5">18.5 mm</quantity>
,
<location id="3C2B6013FFDFBE576DEFFEC60CFE9E59" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB:3C2B6013FFDFBE576DEFFEC60CFE9E59" box="[732,942,344,369]" country="Malaysia" name="Bako National Park" pageId="53" pageNumber="497">Bako National Park</location>
,
<collectingCountry id="41E37658FFDFBE576C85FEC60B459E59" box="[950,1045,344,369]" name="Malaysia" pageId="53" pageNumber="497">Sarawak</collectingCountry>
,
<collectingCountry id="41E37658FFDFBE576B2DFEC60BD09E59" box="[1054,1152,344,369]" name="Malaysia" pageId="53" pageNumber="497">Malaysia</collectingCountry>
,
<date id="4D4A1008FFDFBE576F93FEE50E199EBC" box="[160,329,379,404]" pageId="53" pageNumber="497" value="2002-03-15">
<collectingDate id="5D0EE9E0FFDFBE576F93FEE50E199EBC" box="[160,329,379,404]" pageId="53" pageNumber="497" value="2002-03-15">15 March 2002</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="394B36C8FFDFBE576F93FE540D099D2E" blockId="53.[160,1156,458,518]" pageId="53" pageNumber="497">
<materialsCitation id="899C3C95FFDFBE576F93FE540D059D2E" ID-GBIF-Occurrence="3762866326" collectingDate="2001-09-13" collectionCode="ZRC" country="Malaysia" location="Bako National Park" pageId="53" pageNumber="497" specimenCode="ZRC 2007.0102" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="0B80EADAFFDFBE576F93FE540E4C9ECC" bold="true" box="[160,284,458,484]" italics="true" pageId="53" pageNumber="497">
<typeStatus id="E64F886AFFDFBE576F93FE540E479ECC" box="[160,279,458,484]" pageId="53" pageNumber="497" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="69529EB3FFDFBE576E1FFE550E899ECC" box="[300,473,459,484]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="497">ZRC 2007.0102</specimenCode>
,
<specimenCount id="2FF2FD41FFDFBE576ED6FE550D119ECC" box="[485,577,459,484]" pageId="53" pageNumber="497" type="male">2 males</specimenCount>
, 18.0 × 20.0, 18.1 ×
<quantity id="FE0C9B2DFFDFBE576C03FE550CC89ECC" box="[816,920,459,484]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="53" pageNumber="497" unit="mm" value="19.1">19.1 mm</quantity>
,
<location id="3C2B6013FFDFBE576C96FE550BD09ECC" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB:3C2B6013FFDFBE576C96FE550BD09ECC" box="[933,1152,459,484]" country="Malaysia" name="Bako National Park" pageId="53" pageNumber="497">Bako National Park</location>
,
<collectingCountry id="41E37658FFDFBE576F93FE730FAF9D2E" box="[160,255,493,518]" name="Malaysia" pageId="53" pageNumber="497">Sarawak</collectingCountry>
,
<collectingCountry id="41E37658FFDFBE576E3AFE730E3B9D2E" box="[265,363,493,518]" name="Malaysia" pageId="53" pageNumber="497">Malaysia</collectingCountry>
,
<date id="4D4A1008FFDFBE576E45FE730D059D2E" box="[374,597,493,518]" pageId="53" pageNumber="497" value="2001-09-13">
<collectingDate id="5D0EE9E0FFDFBE576E45FE730D059D2E" box="[374,597,493,518]" pageId="53" pageNumber="497" value="2001-09-13">13 September 2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="394B36C8FFDFBE576F93FDA20CC79DB3" blockId="53.[160,1156,572,667]" pageId="53" pageNumber="497">
<emphasis id="0B80EADAFFDFBE576F93FDA20FA59D7E" bold="true" box="[160,245,572,598]" italics="true" pageId="53" pageNumber="497">Others.</emphasis>
<materialsCitation id="899C3C95FFDFBE576E36FDA30C0D9D51" ID-GBIF-Occurrence="3762866324" collectingDate="2002-05" collectionCode="ZRC" collectorName="Borneo" country="Malaysia" location="Bako" pageId="53" pageNumber="497" specimenCode="ZRC 2011.1013" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCode id="69529EB3FFDFBE576E36FDA30EE29D7E" box="[261,434,573,598]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="497">ZRC 2011.1013</specimenCode>
,
<specimenCount id="2FF2FD41FFDFBE576E8DFDA30D4A9D7E" box="[446,538,573,598]" pageId="53" pageNumber="497" type="male">2 males</specimenCount>
, 10.2 × 11.3, 15.4 ×
<quantity id="FE0C9B2DFFDFBE576C39FDA30C229D7E" box="[778,882,573,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="53" pageNumber="497" unit="mm" value="16.9">16.9 mm</quantity>
,
<specimenCount id="2FF2FD41FFDFBE576C4CFDA30CBF9D7E" box="[895,1007,573,598]" pageId="53" pageNumber="497" type="female">2 females</specimenCount>
, 11.8 × 12.8, 12.2 ×
<quantity id="FE0C9B2DFFDFBE576FCCFDFE0E3C9D51" box="[255,364,608,633]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="53" pageNumber="497" unit="mm" value="13.1">13.1 mm</quantity>
,
<location id="3C2B6013FFDFBE576E4CFDFE0EE89D51" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB:3C2B6013FFDFBE576E4CFDFE0EE89D51" box="[383,440,608,633]" country="Malaysia" name="Bako" pageId="53" pageNumber="497">Bako</location>
,
<collectingCountry id="41E37658FFDFBE576EFFFDFE0D7B9D51" box="[460,555,608,633]" name="Malaysia" pageId="53" pageNumber="497">Sarawak</collectingCountry>
,
<collectorName id="9401531EFFDFBE576D0DFDFE0DC19D51" box="[574,657,608,633]" pageId="53" pageNumber="497">Borneo</collectorName>
, coll.
<date id="4D4A1008FFDFBE576DD2FDFE0C099D51" box="[737,857,608,633]" pageId="53" pageNumber="497" value="2002-05">
<collectingDate id="5D0EE9E0FFDFBE576DD2FDFE0C099D51" box="[737,857,608,633]" pageId="53" pageNumber="497" value="2002-05">May 2002</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="899C3C95FFDFBE576C5FFDFE0CC39DB3" ID-GBIF-Occurrence="3762866366" collectingDate="2002-05" collectionCode="ZRC" country="Malaysia" pageId="53" pageNumber="497" specimenCode="ZRC 2011.1013" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
RUMF-ZC-4985,
<specimenCount id="2FF2FD41FFDFBE576B1AFDFE0BD09D51" box="[1065,1152,608,633]" pageId="53" pageNumber="497" type="male">1 male</specimenCount>
, 16.6 ×
<quantity id="FE0C9B2DFFDFBE576FDEFD1C0E029DB3" box="[237,338,642,667]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="53" pageNumber="497" unit="mm" value="18.3">18.3 mm</quantity>
,
<specimenCount id="2FF2FD41FFDFBE576E6EFD1C0E919DB3" box="[349,449,642,667]" pageId="53" pageNumber="497" type="female">1 female</specimenCount>
, 11.8 ×
<quantity id="FE0C9B2DFFDFBE576D2BFD1C0D2D9DB3" box="[536,637,642,667]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="53" pageNumber="497" unit="mm" value="12.9">12.9 mm</quantity>
, same as
<specimenCode id="69529EB3FFDFBE576DDAFD1C0CC39DB3" box="[745,915,642,667]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="497">ZRC 2011.1013</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDFBE576F93FD4F0B0499E8" pageId="53" pageNumber="497" type="diagnosis">
<paragraph id="394B36C8FFDFBE576F93FD4F0E449DC3" blockId="53.[160,1156,721,1475]" box="[160,276,721,747]" pageId="53" pageNumber="497">
<emphasis id="0B80EADAFFDFBE576F93FD4F0E449DC3" bold="true" box="[160,276,721,747]" italics="true" pageId="53" pageNumber="497">Diagnosis</emphasis>
</paragraph>
<paragraph id="394B36C8FFDFBE576F93FD6B0DF99AEB" blockId="53.[160,1156,721,1475]" pageId="53" pageNumber="497">Carapace squarish, external orbital angle triangular, lateral margins slightly convex; 2 pairs of postfrontal lobes present, lateral lobes slightly exceeding mesial lobes anteriorly; anterior margins of all lobes far from frontal margin in dorsal view. Palm of male chela swollen; outer surface granulated, granules smaller on lower part, with small median protuberance on proximal part of smooth sloping area around bases of fingers; upper surface with 1 straight, longitudinally traversing row of densely packed small granules, and with several short, oblique rows on inner side. Immovable finger almost straight, gradually tapering towards tip, occlusal margin lacking marked proximal elevation, lined with small teeth on proximal two-fifths, 1 large tooth submedially, followed distally by 2 or 3 smaller teeth, subdistally with 1 large tooth and 1 low tooth; lower margin with irregular row of small teeth. Movable finger curved downwards; occlusal margin lined with 2 proximal teeth, followed distally by small teeth, and 1 subdistal tooth; inner side of upper margin with regularly lined rounded granules, these granules indistinct distally. Subdistal tooth of movable finger fitting between 2 subdistal teeth of immovable finger when closed. Male pleonal somite 3 widest, somite 4 abruptly narrowed distally, lateral margins of somites 3–5 clearly concave. G1 short, almost straight, stout, narrowed medially, distal end with anterolaterally directed narrow corneous process. Vulvae located on distal two-fifths of sternite 6, anterior margin adjacent to thoracic sternal suture 5/6, ellipsoidal, margin rimmed posteriorly, sternal vulval cover developed from posterolateral corner, covering almost entire vulva, except for anteromesial hole, cover produced ventroanteriorly as trigonal pyramid structure.</paragraph>
<paragraph id="394B36C8FFDFBE576F93FA670E7C993B" blockId="53.[160,1156,1529,1728]" box="[160,300,1529,1555]" pageId="53" pageNumber="497">
<emphasis id="0B80EADAFFDFBE576F93FA670E7C993B" bold="true" box="[160,300,1529,1555]" italics="true" pageId="53" pageNumber="497">Colouration</emphasis>
</paragraph>
<paragraph id="394B36C8FFDFBE576F93F9830B0499E8" blockId="53.[160,1156,1529,1728]" pageId="53" pageNumber="497">
In life, anterior half of carapace (suborbital to pterygostomial regions and frontal to epibranchial regions), chelipeds and third maxillipeds reddish, gradually changing from darker anterior inner parts (chelae and third maxillipeds) to orange posterior outer parts. Posterior half of dorsal surface of carapace varying from cream to dark khaki mixed with dark purple patterns. Ambulatory legs dark purple. See
<bibRefCitation id="5D654B39FFDFBE576DFFF9390CE599E8" author="Naruse T &amp; Ng PKL" box="[716,949,1703,1728]" pageId="53" pageNumber="497" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng (2007</bibRefCitation>
, p. 338, fig. 1).
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDCBE546F93FF100FB49FC5" pageId="54" pageNumber="498" type="distribution">
<paragraph id="394B36C8FFDCBE546F93FF100E7C9F80" blockId="54.[160,1155,142,237]" box="[160,300,142,168]" pageId="54" pageNumber="498">
<emphasis id="0B80EADAFFDCBE546F93FF100E7C9F80" bold="true" box="[160,300,142,168]" italics="true" pageId="54" pageNumber="498">Distribution</emphasis>
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FF2C0FB49FC5" blockId="54.[160,1155,142,237]" pageId="54" pageNumber="498">
<location id="3C2B6013FFDCBE546F93FF2C0FBE9FE3" LSID="urn:lsid:plazi:treatment:B15D87DEFFDFBE546EB1FF100DEF9DDB:3C2B6013FFDCBE546F93FF2C0FBE9FE3" box="[160,238,178,203]" country="Malaysia" name="Known" pageId="54" pageNumber="498">Known</location>
only from
<collectingCountry id="41E37658FFDCBE546E43FF2C0E9F9FE3" box="[368,463,178,203]" name="Malaysia" pageId="54" pageNumber="498">Sarawak</collectingCountry>
,
<collectingCountry id="41E37658FFDCBE546EEFFF2C0D6C9FE3" box="[476,572,178,203]" name="Malaysia" pageId="54" pageNumber="498">Malaysia</collectingCountry>
[
<typeStatus id="E64F886AFFDCBE546D78FF2C0D2F9FE3" box="[587,639,178,203]" pageId="54" pageNumber="498">type</typeStatus>
locality: Bako National Park] (
<bibRefCitation id="5D654B39FFDCBE546CE6FF2C0F879FC5" author="Naruse T &amp; Ng PKL" pageId="54" pageNumber="498" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDCBE546F93FEB70DEF9DDB" pageId="54" pageNumber="498" type="discussion">
<paragraph id="394B36C8FFDCBE546F93FEB70E569E6B" blockId="54.[160,1156,297,531]" box="[160,262,297,323]" pageId="54" pageNumber="498">
<emphasis id="0B80EADAFFDCBE546F93FEB70E569E6B" bold="true" box="[160,262,297,323]" italics="true" pageId="54" pageNumber="498">Remarks</emphasis>
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FED20C119D3B" blockId="54.[160,1156,297,531]" pageId="54" pageNumber="498">
<taxonomicName id="FEF44D4BFFDCBE546F93FED20EF29E4D" authorityName="Naruse and Ng" authorityYear="2007" box="[160,418,332,357]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDCBE546F93FED20EF29E4D" box="[160,418,332,357]" italics="true" pageId="54" pageNumber="498">Scandarma splendidum</emphasis>
</taxonomicName>
is morphologically closest to
<taxonomicName id="FEF44D4BFFDCBE546DC5FED20CD49E4D" authorityName="Ng" authorityYear="2013" box="[758,900,332,357]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFDCBE546DC5FED20CD49E4D" box="[758,900,332,357]" italics="true" pageId="54" pageNumber="498">Sc. raymondi</emphasis>
</taxonomicName>
. Among the five diagnostic characters listed by
<bibRefCitation id="5D654B39FFDCBE546EE8FEF10D169EA0" author="Ng PKL" box="[475,582,367,392]" pageId="54" pageNumber="498" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng (2013</bibRefCitation>
, p. 293), the distal corneous part of the G1 was described as being relatively wider in
<taxonomicName id="FEF44D4BFFDCBE546D48FE0C0C5F9E83" authorityName="Ng" authorityYear="2013" box="[635,783,402,427]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFDCBE546D48FE0C0C5F9E83" box="[635,783,402,427]" italics="true" pageId="54" pageNumber="498">Sc. raymondi</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEF44D4BFFDCBE546CB5FE0C0B609E83" authorityName="Naruse and Ng" authorityYear="2007" box="[902,1072,402,427]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDCBE546CB5FE0C0B609E83" box="[902,1072,402,427]" italics="true" pageId="54" pageNumber="498">Sc. splendidum</emphasis>
</taxonomicName>
. A reexamination of the present material, however, shows that the comparative characters were inadvertently transposed by
<bibRefCitation id="5D654B39FFDCBE546D15FE490DC59ED8" author="Ng PKL" box="[550,661,471,496]" pageId="54" pageNumber="498" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng (2013)</bibRefCitation>
and that
<taxonomicName id="FEF44D4BFFDCBE546C35FE490CFA9ED8" authorityName="Naruse and Ng" authorityYear="2007" box="[774,938,471,496]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDCBE546C35FE490CFA9ED8" box="[774,938,471,496]" italics="true" pageId="54" pageNumber="498">Sc. splendidum</emphasis>
</taxonomicName>
is actually the one with a proportionately wider distal corneous part of the G1.
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FDD00E049D40" blockId="54.[160,1155,590,755]" box="[160,340,590,616]" pageId="54" pageNumber="498">
<emphasis id="0B80EADAFFDCBE546F93FDD00E049D40" bold="true" box="[160,340,590,616]" italics="true" pageId="54" pageNumber="498">Ecological note</emphasis>
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FDEC0DEF9DDB" blockId="54.[160,1155,590,755]" pageId="54" pageNumber="498">
<bibRefCitation id="5D654B39FFDCBE546F93FDEC0ED99DA3" author="Naruse T &amp; Ng PKL" box="[160,393,626,651]" pageId="54" pageNumber="498" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng (2007</bibRefCitation>
, p. 338) reported that
<taxonomicName id="FEF44D4BFFDCBE546D4CFDEC0C709DA3" authorityName="Naruse and Ng" authorityYear="2007" box="[639,800,626,651]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDCBE546D4CFDEC0C709DA3" box="[639,800,626,651]" italics="true" pageId="54" pageNumber="498">Sc. splendidum</emphasis>
</taxonomicName>
was nocturnal and observed on plants to a height of
<quantity id="FE0C9B2DFFDCBE546EBBFD0A0EE29D85" box="[392,434,660,685]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="498" unit="m" value="2.0">2 m</quantity>
near streams or puddles, and sometimes found up to
<quantity id="FE0C9B2DFFDCBE546B3CFD0A0B179D85" box="[1039,1095,660,685]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="498" unit="m" value="10.0">10 m</quantity>
from any water source. They also commented that the ecology and behaviour of
<taxonomicName id="FEF44D4BFFDCBE546CD1FD290BD39DF8" authorityName="Naruse and Ng" authorityYear="2007" box="[994,1155,695,720]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDCBE546CD1FD290BD39DF8" box="[994,1155,695,720]" italics="true" pageId="54" pageNumber="498">Sc. splendidum</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="FEF44D4BFFDCBE546EAEFD440D519DDB" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[413,513,730,755]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="lintou">
<emphasis id="0B80EADAFFDCBE546EAEFD440D519DDB" box="[413,513,730,755]" italics="true" pageId="54" pageNumber="498">Sc. lintou</emphasis>
</taxonomicName>
in most aspects.
</paragraph>
</subSubSection>
</treatment>
</document>
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