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<document id="10E59AE1570217E3B8EC02210F80AC2C" ID-CLB-Dataset="55729" ID-DOI="10.1080/00222933.2020.1763491" ID-GBIF-Dataset="fb05d004-0f2d-44da-b91e-b6e8304ab2a5" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1615924067624" checkinUser="carolina" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2020" docId="B15D87DEFFDCBE556EE6FCDE0DF19D60" docLanguage="en" docName="JNATHIST.54.7-8.445-532.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 54 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2020.1763491" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scandarma raymondi Ng 2013" docType="treatment" docVersion="8" lastPageNumber="499" masterDocId="4D64FFA6FFEABE626F33FF9E0F509F28" masterDocTitle="Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species" masterLastPageNumber="532" masterPageNumber="445" pageNumber="498" updateTime="1698933465109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="590D1879373CD6BC1EBA0D9D0DC88F1C">Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species</mods:title>
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<mods:namePart id="5980930B558083672BEC937367E414A1">Naruse, Tohru</mods:namePart>
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<mods:namePart id="EE52D1D44FA9D829443A5C640B9569FC">Ng, Peter K. L.</mods:namePart>
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<mods:date id="D69D0805053F52A6318BA030C0A2E441">2020</mods:date>
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<paragraph id="394B36C8FFDCBE546EE6FCDE0C1D9C72" blockId="54.[469,845,832,858]" box="[469,845,832,858]" pageId="54" pageNumber="498">
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<taxonomicName id="FEF44D4BFFDCBE546EE6FCDE0C1D9C72" ID-CoL="6Y3Y9" authority="Ng, 2013" authorityName="Ng" authorityYear="2013" box="[469,845,832,858]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFDCBE546EE6FCDE0D8D9C72" bold="true" box="[469,733,832,858]" italics="true" pageId="54" pageNumber="498">Scandarma raymondi</emphasis>
<bibRefCitation id="5D654B39FFDCBE546DD6FCDE0C1D9C72" author="Ng PKL" box="[741,845,832,858]" pageId="54" pageNumber="498" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng, 2013</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="394B36C8FFDCBE546F93FCEA0BD19CA5" blockId="54.[160,1153,884,909]" box="[160,1153,884,909]" pageId="54" pageNumber="498">
<treatmentCitationGroup id="19E411E6FFDCBE546F93FCEA0BD19CA5" box="[160,1153,884,909]" pageId="54" pageNumber="498">
<treatmentCitation id="B85510D9FFDCBE546F93FCEA0D1C9CA5" author="Ng PKL" box="[160,588,884,909]" page="289" pageId="54" pageNumber="498" year="2013">
<taxonomicName id="FEF44D4BFFDCBE546F93FCEA0D1C9CA5" ID-CoL="6Y3Y9" authority="Ng, 2013, p. 289" authorityName="Ng" authorityPageNumber="289" authorityYear="2013" box="[160,588,884,909]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="498" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFDCBE546F93FCEA0EDB9CA5" box="[160,395,884,909]" italics="true" pageId="54" pageNumber="498">Scandarma raymondi</emphasis>
<bibRefCitation id="5D654B39FFDCBE546EA0FCEA0EA79CA5" author="Ng PKL" box="[403,503,884,909]" pageId="54" pageNumber="498" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng, 2013</bibRefCitation>
, p. 289
</taxonomicName>
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, figs 13 [
<typeStatus id="E64F886AFFDCBE546D89FCEA0DBE9CA5" box="[698,750,884,909]" pageId="54" pageNumber="498">type</typeStatus>
locality:
<location id="3C2B6013FFDCBE546C60FCEA0CE79CA5" LSID="urn:lsid:plazi:treatment:B15D87DEFFDCBE556EE6FCDE0DF19D60:3C2B6013FFDCBE546C60FCEA0CE79CA5" box="[851,951,884,909]" country="Malaysia" name="Kinabalu" pageId="54" pageNumber="498" stateProvince="Sabah">Kinabalu</location>
,
<collectingRegion id="FB30F82AFFDCBE546CF2FCEA0B599CA5" box="[961,1033,884,909]" country="Malaysia" name="Sabah" pageId="54" pageNumber="498">Sabah</collectingRegion>
,
<collectingCountry id="41E37658FFDCBE546B20FCEA0B289CA5" box="[1043,1144,884,909]" name="Malaysia" pageId="54" pageNumber="498">Malaysia</collectingCountry>
].
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</paragraph>
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<subSubSection id="71EE6543FFDCBE546F93FC570E979B00" pageId="54" pageNumber="498" type="materials_examined">
<paragraph id="394B36C8FFDCBE546F93FC570E2F9CCB" blockId="54.[160,1156,969,1064]" box="[160,383,969,995]" pageId="54" pageNumber="498">
<emphasis id="0B80EADAFFDCBE546F93FC570E2F9CCB" bold="true" box="[160,383,969,995]" italics="true" pageId="54" pageNumber="498">Material examined</emphasis>
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FC720E979B00" blockId="54.[160,1156,969,1064]" pageId="54" pageNumber="498">
<materialsCitation id="899C3C95FFDCBE546F93FC720E979B00" ID-GBIF-Occurrence="3762866381" collectingDate="1990-11-01" collectionCode="R" collectorName="R. Goh" country="Malaysia" location="Kota Kinabalu" municipality="Goh" pageId="54" pageNumber="498" specimenCode="ZRC 2013.0614" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="0B80EADAFFDCBE546F93FC720E429B2E" bold="true" box="[160,274,1004,1030]" italics="true" pageId="54" pageNumber="498">
<typeStatus id="E64F886AFFDCBE546F93FC720E5D9B2E" box="[160,269,1004,1030]" pageId="54" pageNumber="498" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="69529EB3FFDCBE546E12FC720E9E9B2D" box="[289,462,1004,1029]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="54" pageNumber="498">ZRC 2013.0614</specimenCode>
,
<specimenCount id="2FF2FD41FFDCBE546EE8FC720D429B2D" box="[475,530,1004,1029]" pageId="54" pageNumber="498" type="male">male</specimenCount>
11.9 ×
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,
<location id="3C2B6013FFDCBE546DD7FC720CD59B2D" LSID="urn:lsid:plazi:treatment:B15D87DEFFDCBE556EE6FCDE0DF19D60:3C2B6013FFDCBE546DD7FC720CD59B2D" box="[740,901,1004,1029]" country="Malaysia" municipality="Goh" name="Kota Kinabalu" pageId="54" pageNumber="498" stateProvince="Sabah">Kota Kinabalu</location>
,
<collectingRegion id="FB30F82AFFDCBE546CA1FC720C8A9B2D" box="[914,986,1004,1029]" country="Malaysia" name="Sabah" pageId="54" pageNumber="498">Sabah</collectingRegion>
,
<collectingCountry id="41E37658FFDCBE546CD4FC720B199B2D" box="[999,1097,1004,1029]" name="Malaysia" pageId="54" pageNumber="498">Malaysia</collectingCountry>
, coll.
<collectorName id="9401531EFFDCBE546F93FB910FBC9B00" box="[160,236,1039,1064]" pageId="54" pageNumber="498">R. Goh</collectorName>
,
<date id="4D4A1008FFDCBE546FC4FB910E939B00" box="[247,451,1039,1064]" pageId="54" pageNumber="498" value="1990-11-01">
<collectingDate id="5D0EE9E0FFDCBE546FC4FB910E939B00" box="[247,451,1039,1064]" pageId="54" pageNumber="498" value="1990-11-01">1 November 1990</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
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<subSubSection id="71EE6543FFDCBE556F93FBFD0EEF9FC5" lastPageId="55" lastPageNumber="499" pageId="54" pageNumber="498" type="diagnosis">
<paragraph id="394B36C8FFDCBE546F93FBFD0E449B55" blockId="54.[159,1156,1123,1704]" box="[160,276,1123,1149]" pageId="54" pageNumber="498">
<emphasis id="0B80EADAFFDCBE546F93FBFD0E449B55" bold="true" box="[160,276,1123,1149]" italics="true" pageId="54" pageNumber="498">Diagnosis</emphasis>
</paragraph>
<paragraph id="394B36C8FFDCBE546F93FB190DEA9980" blockId="54.[159,1156,1123,1704]" pageId="54" pageNumber="498">Carapace squarish, external orbital angle triangular, lateral margins with 2 small crenulations; 2 pairs of postfrontal lobes present, lateral lobes slightly exceeding mesial lobes anteriorly; anterior margins of all lobes far from frontal margin in dorsal view. Palm of male chela swollen; outer surface granulated, granules smaller on lower part, with small median protuberance on proximal part of sloping smooth area around bases of fingers; upper surface with 1 straight, longitudinally traversing, not densely packed row of relatively small granules, and with a few short, oblique rows on inner side. Immovable finger almost straight, gradually tapering towards tip, occlusal margin lacking marked proximal elevation, lined with small teeth and 1 large tooth over proximal third, followed distally by similar combination of small teeth and 1 large tooth, 1 large subdistal tooth. Movable finger curved, occlusal margin lined with 2 proximal large teeth, followed distally by small teeth, and 1 subdistal tooth; inner side of upper margin sparsely lined with acute granules, granules indistinct distally; subdistal tooth of movable finger fitting between 2 subdistal teeth of immovable finger when closed. Male pleonal somite 3 widest, somite 4 abruptly narrowed distally, lateral margins of somites 35 clearly concave. G1 short, almost straight, stout, narrowed medially, distal end with relatively narrow, anterolaterally directed corneous process.</paragraph>
<paragraph id="394B36C8FFDDBE556F93FF100E7C9F80" blockId="55.[160,1155,142,237]" box="[160,300,142,168]" pageId="55" pageNumber="499">
<emphasis id="0B80EADAFFDDBE556F93FF100E7C9F80" bold="true" box="[160,300,142,168]" italics="true" pageId="55" pageNumber="499">Colouration</emphasis>
</paragraph>
<paragraph id="394B36C8FFDDBE556F93FF2C0EEF9FC5" blockId="55.[160,1155,142,237]" pageId="55" pageNumber="499">
In preservative: carapace yellowish-brown, chelipeds yellow just after preservation in alcohol (
<bibRefCitation id="5D654B39FFDDBE556FCCFF4A0E0F9FC5" author="Ng PKL" box="[255,351,212,237]" pageId="55" pageNumber="499" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng 2013</bibRefCitation>
, p. 293).
</paragraph>
</subSubSection>
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<emphasis id="0B80EADAFFDDBE556F93FE870E7C9E1B" bold="true" box="[160,300,281,307]" italics="true" pageId="55" pageNumber="499">Distribution</emphasis>
</paragraph>
<paragraph id="394B36C8FFDDBE556F93FEA20C979E7D" blockId="55.[160,967,281,341]" box="[160,967,316,341]" pageId="55" pageNumber="499">
Known only from the
<typeStatus id="E64F886AFFDDBE556EA4FEA20E999E7D" box="[407,457,316,341]" pageId="55" pageNumber="499">type</typeStatus>
locality,
<location id="3C2B6013FFDDBE556D1EFEA20DC19E7D" LSID="urn:lsid:plazi:treatment:B15D87DEFFDCBE556EE6FCDE0DF19D60:3C2B6013FFDDBE556D1EFEA20DC19E7D" box="[557,657,316,341]" country="Malaysia" name="Kinabalu" pageId="55" pageNumber="499" stateProvince="Sabah">Kinabalu</location>
,
<collectingRegion id="FB30F82AFFDDBE556DA8FEA20DB39E7D" box="[667,739,316,341]" country="Malaysia" name="Sabah" pageId="55" pageNumber="499">Sabah</collectingRegion>
,
<collectingCountry id="41E37658FFDDBE556DDEFEA20C1D9E7D" box="[749,845,316,341]" name="Malaysia" pageId="55" pageNumber="499">Malaysia</collectingCountry>
(
<bibRefCitation id="5D654B39FFDDBE556C68FEA20CEB9E7D" author="Ng PKL" box="[859,955,316,341]" pageId="55" pageNumber="499" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFDDBE556F93FE1F0DF19D60" pageId="55" pageNumber="499" type="discussion">
<paragraph id="394B36C8FFDDBE556F93FE1F0E569EB3" blockId="55.[160,872,385,445]" box="[160,262,385,411]" pageId="55" pageNumber="499">
<emphasis id="0B80EADAFFDDBE556F93FE1F0E569EB3" bold="true" box="[160,262,385,411]" italics="true" pageId="55" pageNumber="499">Remarks</emphasis>
</paragraph>
<paragraph id="394B36C8FFDDBE556F93FE3A0C389E95" blockId="55.[160,872,385,445]" box="[160,872,420,445]" pageId="55" pageNumber="499">
See
<bibRefCitation id="5D654B39FFDDBE556FFCFE3A0E669E95" author="Ng PKL" box="[207,310,420,445]" pageId="55" pageNumber="499" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng (2013</bibRefCitation>
, p. 293) and Remarks for
<taxonomicName id="FEF44D4BFFDDBE556D52FE3A0C329E95" authorityName="Naruse and Ng" authorityYear="2007" box="[609,866,420,445]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="499" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFDDBE556D52FE3A0C329E95" box="[609,866,420,445]" italics="true" pageId="55" pageNumber="499">Scandarma splendidum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="394B36C8FFDDBE556F93FE770E049D2B" blockId="55.[160,1155,489,584]" box="[160,340,489,515]" pageId="55" pageNumber="499">
<emphasis id="0B80EADAFFDDBE556F93FE770E049D2B" bold="true" box="[160,340,489,515]" italics="true" pageId="55" pageNumber="499">Ecological note</emphasis>
</paragraph>
<paragraph id="394B36C8FFDDBE556F93FD920DF19D60" blockId="55.[160,1155,489,584]" pageId="55" pageNumber="499">
The only known specimen was collected from inside a mixed forest in a town centre near a river less than
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from the sea (
<bibRefCitation id="5D654B39FFDDBE556D07FDB10DC49D60" author="Ng PKL" box="[564,660,559,584]" pageId="55" pageNumber="499" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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