441 lines
53 KiB
XML
441 lines
53 KiB
XML
<document id="E43B8A1E14E7F452B4DCEB160D0970FF" ID-DOI="10.26028/cybium/2023-034" ID-ISSN="2101-0315" ID-Zenodo-Dep="12713757" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720651148950" checkinUser="felipe" docAuthor="Keith, Philippe & Mennesson, Marion I." docDate="2023" docId="03EA87D826070D0EFED0FD2DF1FFFA03" docLanguage="en" docName="Cybium.47.4.431-466.pdf" docOrigin="Cybium 47 (4)" docSource="http://dx.doi.org/10.26028/cybium/2023-034" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Butis gymnopomus" docType="treatment" docVersion="2" lastPageNumber="448" masterDocId="FFD3FFA026170D1FFFD5F857F53CFF10" masterDocTitle="Review of Butis (Teleostei: Butidae) from Indo-Pacific islands with description of three new species" masterLastPageNumber="466" masterPageNumber="431" pageNumber="447" updateTime="1720655022157" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="A16DE95B9DD12A68DCBD0C308BA86E24">
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<mods:title id="1B4B4123F9A930095B11E7800800868E">Review of Butis (Teleostei: Butidae) from Indo-Pacific islands with description of three new species</mods:title>
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<mods:namePart id="DF82192671FCCDF62EC08EB027FE4134">Keith, Philippe</mods:namePart>
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<mods:namePart id="3F04CA7C11E097DC5CCE9A755035EF1E">Mennesson, Marion I.</mods:namePart>
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<mods:typeOfResource id="A1F322B019B793891768F02F7871D01B">text</mods:typeOfResource>
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<mods:title id="3B704656731B5C8DC1066C96BEBB1636">Cybium</mods:title>
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<mods:part id="D64080F96EF8CF0F8E52D96596EBA4B9">
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<mods:date id="C35B9BC4E665AD0D5B5A6940658FBC67">2023</mods:date>
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<mods:number id="63B8649CD5B112D74F8B364C918A341E">47</mods:number>
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<mods:identifier id="537D26CBB45B3BD8B0BDEE4AEF1CE77F" type="DOI">10.26028/cybium/2023-034</mods:identifier>
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<mods:identifier id="6928DE6B3A650DBD9E1F088AD79EA262" type="ISSN">2101-0315</mods:identifier>
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<treatment id="03EA87D826070D0EFED0FD2DF1FFFA03" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03" httpUri="http://treatment.plazi.org/id/03EA87D826070D0EFED0FD2DF1FFFA03" lastPageId="17" lastPageNumber="448" pageId="16" pageNumber="447">
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<subSubSection id="C359654526070D0FFED0FD2DF7BBFA82" box="[261,647,1402,1426]" pageId="16" pageNumber="447" type="nomenclature">
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<paragraph id="8BFC36CE26070D0FFED0FD2DF7BBFA82" blockId="16.[261,647,1402,1426]" box="[261,647,1402,1426]" pageId="16" pageNumber="447">
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<heading id="D0B481A226070D0FFED0FD2DF7BBFA82" bold="true" box="[261,647,1402,1426]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="447" reason="2">
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<taxonomicName id="4C434D4D26070D0FFED0FD2DF7BBFA82" ID-CoL="NZ3Z" authority="(Bleeker, 1853)" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[261,647,1402,1426]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="gymnopomus">
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<emphasis id="B937EADC26070D0FFED0FD2DF7BBFA82" bold="true" box="[261,647,1402,1426]" pageId="16" pageNumber="447">
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<emphasis id="B937EADC26070D0FFED0FD2DF4E4FA82" bold="true" box="[261,472,1402,1426]" italics="true" pageId="16" pageNumber="447">Butis gymnopomus</emphasis>
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(
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<bibRefCitation id="EFD24B3F26070D0FFE30FD2DF743FA82" author="BLEEKER P." box="[485,639,1402,1426]" pageId="16" pageNumber="447" pagination="317 - 352" refId="ref25778" refString="BLEEKER P., 1853. - Vierde bijdrage tot de kennis der ichthyolo- gische fauna van Amboina. Natuurk. Tijdschr. Ned. - Indie, 5 (3): 317 - 352." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C359654526070D0FFEFAFDCDF6AFFEE1" pageId="16" pageNumber="447" type="description">
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<paragraph id="8BFC36CE26070D0FFEFAFDCDF760FAA3" blockId="16.[303,604,1434,1459]" box="[303,604,1434,1459]" pageId="16" pageNumber="447">
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(
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<figureCitation id="13782A4B26070D0FFEE2FDCDF49FFAA3" box="[311,419,1434,1459]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[1077,1141,1813,1835]" captionStartId-1="5.[126,189,1574,1596]" captionStartId-2="7.[132,195,1245,1267]" captionTargetBox-0="[138,1058,221,1894]" captionTargetBox-1="[132,1425,407,1517]" captionTargetBox-2="[133,1454,216,1229]" captionTargetId-0="figure-54@2.[132,1060,215,1900]" captionTargetId-1="figure-154@5.[132,1455,401,1558]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." captionText-1="Figure 2 (see also following page). – A: Bayesian tree of the cytochrome c oxidase subunit (COI; 452 bp) for sequenced specimens of Butis. Numbers at each node represent posterior probabilities. B: Barcoding NJ K2P tree of the COI gene (196 bp) for sequenced specimens of Butis including the holotype of Butis humeralis from the MNHN collection. Numbers at each node represent bootstrap values. C, D: Barcoding NJ K2P trees of the 12S and the 16S genes (129 and 270 bp respectively) for sequenced specimens of Butis collected during the 19th century from the MNHN collection including a paratype of B. audebertae (MNHN A-1503). Numbers at each node represent bootstrap values. [A high resolution version is available online.]" captionText-2="Figure 3. – A, B: Distribution of Butis species." figureDoi-0="http://doi.org/10.5281/zenodo.12713759" figureDoi-1="http://doi.org/10.5281/zenodo.12713761" figureDoi-2="http://doi.org/10.5281/zenodo.12713763" httpUri-0="https://zenodo.org/record/12713759/files/figure.png" httpUri-1="https://zenodo.org/record/12713761/files/figure.png" httpUri-2="https://zenodo.org/record/12713763/files/figure.png" pageId="16" pageNumber="447">Figs 1-3B</figureCitation>
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,
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<figureCitation id="13782A4B26070D0FFE7BFDCDF482FAA2" box="[430,446,1434,1458]" captionStart="Figure 6" captionStartId="17.[132,197,720,742]" captionTargetBox="[132,775,215,703]" captionTargetId="figure-730@17.[132,775,450,703]" captionTargetPageId="17" captionText="Figure 6. – Butis gymnopomus, uncatalogued, BIF 0311 (59 mm SL), Java (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12713769" httpUri="https://zenodo.org/record/12713769/files/figure.png" pageId="16" pageNumber="447">6</figureCitation>
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; Tables III-IV)
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</paragraph>
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<paragraph id="8BFC36CE26070D0FFF7FFD8DF6AFFEE1" blockId="16.[132,776,1498,1907]" lastBlockId="16.[813,1456,217,497]" pageId="16" pageNumber="447">
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According to
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<bibRefCitation id="EFD24B3F26070D0FFE9FFD8DF715FAE3" author="FRICKE R. & ESCHMEYER W. N. & VAN DER LAAN R. & EDS" box="[330,553,1498,1523]" pageId="16" pageNumber="447" refId="ref26140" refString="FRICKE R., ESCHMEYER W. N. & VAN DER LAAN R. (EDS), 2022. - Eschmeyer's catalog of fishes: genera, species, references. Electronic version accessed 5 March 2023. http: // researcharchive. calacademy. org / research / ichthyology / catalog / fish- catmain. asp" type="url" year="2022">
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Fricke
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<emphasis id="B937EADC26070D0FFE4EFD8CF4E9FAE2" box="[411,469,1498,1522]" italics="true" pageId="16" pageNumber="447">et al.</emphasis>
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(2022)
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</bibRefCitation>
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, the
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<typeStatus id="54F8886C26070D0FFDB1FD8DF7F4FAE2" box="[612,712,1498,1522]" pageId="16" pageNumber="447" type="holotype">holotype</typeStatus>
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from Western
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<collectingRegion id="4987F82C26070D0FFF37FDACF407F903" box="[226,315,1531,1555]" country="Indonesia" name="Sumatera Utara" pageId="16" pageNumber="447">Sumatra</collectingRegion>
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of this species is “whereabouts unknown”. In the species description
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<bibRefCitation id="EFD24B3F26070D0FFE62FE4CF759F923" author="BLEEKER P." box="[439,613,1562,1587]" pageId="16" pageNumber="447" pagination="317 - 352" refId="ref25778" refString="BLEEKER P., 1853. - Vierde bijdrage tot de kennis der ichthyolo- gische fauna van Amboina. Natuurk. Tijdschr. Ned. - Indie, 5 (3): 317 - 352." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
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wrote “
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<emphasis id="B937EADC26070D0FFD10FE4CF5C2F983" italics="true" pageId="16" pageNumber="447">
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Mijne verzameling bevat reeds meerdere soorten van
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<taxonomicName id="4C434D4D26070D0FFD79FE6CF43FF962" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="subSpecies" species="met" subSpecies="spitsen">Eleotris met spitsen</taxonomicName>
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platten kop, langen snuit en groote zijschubben, zooals
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</emphasis>
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<taxonomicName id="4C434D4D26070D0FFED3FE2CF70AF982" authority="Blkr." authorityName="Bleeker" authorityYear="1849" box="[262,566,1659,1683]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="melanostigma">
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Eleotris melanostigma
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<emphasis id="B937EADC26070D0FFDD4FE2CF70CF983" box="[513,560,1659,1683]" italics="true" pageId="16" pageNumber="447">Blkr</emphasis>
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.
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFD91FE2CF5CAF9A2" authority="Blkr." authorityName="Bleeker" authorityYear="1853" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="melanopterus">
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Eleotris melanopterus
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<emphasis id="B937EADC26070D0FFF14FECCF5CCF9A3" box="[193,240,1691,1715]" italics="true" pageId="16" pageNumber="447">Blkr</emphasis>
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.
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFED0FECCF4DBF9A2" authority="Blkr." authorityName="Bleeker" authorityYear="1851" box="[261,487,1691,1715]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="wolffii">
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Eleotris wolffii
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<emphasis id="B937EADC26070D0FFE67FECCF4DDF9A3" box="[434,481,1691,1715]" italics="true" pageId="16" pageNumber="447">Blkr</emphasis>
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.
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFE23FECCF63DF9A3" authority="Blkr." authorityName="Bleeker" authorityYear="1849" box="[502,769,1691,1715]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="prismatica">
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Eleotris prismatica
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<emphasis id="B937EADC26070D0FFD18FECCF63DF9A3" box="[717,769,1691,1715]" italics="true" pageId="16" pageNumber="447">Blkr.</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFF51FEECF448F9C2" authority="CV" authorityName="CV" box="[132,372,1723,1747]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="humeralis">
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Eleotris humeralis
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<emphasis id="B937EADC26070D0FFE87FEECF448F9C2" box="[338,372,1723,1746]" italics="true" pageId="16" pageNumber="447">CV</emphasis>
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</taxonomicName>
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<emphasis id="B937EADC26070D0FFEAEFEEBF5F7F823" italics="true" pageId="16" pageNumber="447">en de boven beschrevene, welke zich echter gemakklejik van alle boven genoemde laat onderkennen aan hare onbeschubte wangen en snuit en tusschenoogstreek.</emphasis>
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”
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<emphasis id="B937EADC26070D0FFF09FF4BF438F823" box="[220,260,1820,1843]" italics="true" pageId="16" pageNumber="447">i.e.,</emphasis>
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“My collection already contains several species of
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<taxonomicName id="4C434D4D26070D0FFF71FF6CF5C1F843" box="[164,253,1851,1875]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="genus">
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<emphasis id="B937EADC26070D0FFF71FF6CF5C1F843" box="[164,253,1851,1875]" italics="true" pageId="16" pageNumber="447">Eleotris</emphasis>
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</taxonomicName>
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with a flat, pointed head, long snout and large lateral scales, such as
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<taxonomicName id="4C434D4D26070D0FFEA6FF0CF79BF863" authority="Blkr." authorityName="Bleeker" authorityYear="1849" box="[371,679,1883,1907]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="melanostigma">
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<emphasis id="B937EADC26070D0FFEA6FF0CF75BF863" box="[371,615,1883,1907]" italics="true" pageId="16" pageNumber="447">Eleotris melanostigma</emphasis>
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Blkr.
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFD64FF0CF13CFFE1" authority="Blkr." authorityName="Bleeker" authorityYear="1853" box="[689,1024,217,1907]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="melanopterus">
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<emphasis id="B937EADC26070D0FFD64FF0CF6FDFFE1" box="[689,961,217,1907]" italics="true" pageId="16" pageNumber="447">Eleotris melanopterus</emphasis>
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Blkr.
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</taxonomicName>
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,
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<taxonomicName id="4C434D4D26070D0FFBDEF88EF1D7FFE1" authority="Blkr." authorityName="Bleeker" authorityYear="1851" box="[1035,1259,217,241]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="wolffii">
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<emphasis id="B937EADC26070D0FFBDEF88EF190FFE1" box="[1035,1196,217,241]" italics="true" pageId="16" pageNumber="447">Eleotris wolffii</emphasis>
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Blkr.
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||
</taxonomicName>
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||
,
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<taxonomicName id="4C434D4D26070D0FFB23F88EF6B5FE01" authority="Blkr" authorityName="Bleeker" authorityYear="1849" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="prismatica">
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<emphasis id="B937EADC26070D0FFB23F88EF672FE01" italics="true" pageId="16" pageNumber="447">Eleotris prismatica</emphasis>
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Blkr
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</taxonomicName>
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,
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||
<taxonomicName id="4C434D4D26070D0FFC40F8AEF1B3FE01" authority="CV" authorityName="CV" box="[917,1167,249,273]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="humeralis">
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||
<emphasis id="B937EADC26070D0FFC40F8AEF15EFE01" box="[917,1122,249,273]" italics="true" pageId="16" pageNumber="447">Eleotris humeralis</emphasis>
|
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CV
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</taxonomicName>
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and the species described above, which can easily be distinguished from all of these species by its cheeks, snout and inter-orbital region without scales”. This description is sufficiently clear and no
|
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<typeStatus id="54F8886C26070D0FFA8FF90DF093FE62" box="[1370,1455,346,370]" pageId="16" pageNumber="447" type="neotype">neotype</typeStatus>
|
||
designation is needed.
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<taxonomicName id="4C434D4D26070D0FFBFDF92DF1EBFE81" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[1064,1239,378,401]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="447" phylum="Chordata" rank="species" species="gymnopomus">
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||
<emphasis id="B937EADC26070D0FFBFDF92DF1EBFE81" box="[1064,1239,378,401]" italics="true" pageId="16" pageNumber="447">B. gymnopomus</emphasis>
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</taxonomicName>
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is, indeed, the only species with an interorbital space without scales (with IOS 0/0/0) and with a flat pointed head and a long snout (see species key).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C359654526070D0FFCF8FA4EF126FC82" pageId="16" pageNumber="447" type="materials_examined">
|
||
<paragraph id="8BFC36CE26070D0FFCF8FA4EF138FD21" blockId="16.[813,1456,537,914]" box="[813,1028,537,561]" pageId="16" pageNumber="447">
|
||
<emphasis id="B937EADC26070D0FFCF8FA4EF138FD21" bold="true" box="[813,1028,537,561]" pageId="16" pageNumber="447">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BFC36CE26070D0FFC87FA6DF126FC82" blockId="16.[813,1456,537,914]" pageId="16" pageNumber="447">
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<materialsCitation id="3B2B3C9326070D0FFC87FA6DF1FFFD62" collectingDate="1908-10" collectionCode="SMF" collectorName="Sunda" country="Indonesia" location="Kabaena" municipality="Sunda" pageId="16" pageNumber="447" specimenCode="SMF 9964" specimenCount="1">
|
||
<specimenCode id="DBE59EB526070D0FFC87FA6DF6EFFD42" box="[850,979,570,594]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="447">SMF 9964</specimenCode>
|
||
, female (
|
||
<quantity id="4CBB9B2B26070D0FFB94FA6DF1AAFD41" box="[1089,1174,570,594]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="447" unit="mm" value="72.0">72 mm</quantity>
|
||
SL),
|
||
<collectingCountry id="F354765E26070D0FFB0EFA6DF06EFD42" box="[1243,1362,570,594]" name="Indonesia" pageId="16" pageNumber="447">Indonesia</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6B98ACB426070D0FFA8AFA6DF096FD42" box="[1375,1450,570,594]" pageId="16" pageNumber="447">Sunda</collectingMunicipality>
|
||
,
|
||
<location id="8E9C601526070D0FFCF8FA0DF6ACFD62" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03:8E9C601526070D0FFCF8FA0DF6ACFD62" box="[813,912,602,626]" country="Indonesia" municipality="Sunda" name="Kabaena" pageId="16" pageNumber="447">Kabaena</location>
|
||
,
|
||
<date id="FFFD100E26070D0FFC4EFA0DF136FD62" box="[923,1034,602,626]" pageId="16" pageNumber="447" value="1908-10">
|
||
<collectingDate id="EFB9E9E626070D0FFC4EFA0DF136FD62" box="[923,1034,602,626]" pageId="16" pageNumber="447" value="1908-10">Oct. 1908</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B6531826070D0FFBC0FA0DF166FD62" box="[1045,1114,602,626]" pageId="16" pageNumber="447">Sunda</collectorName>
|
||
exp. coll
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C9326070D0FFB1BFA0DF179FDA2" collectingDate="1909-10" collectionCode="SMF" collectorName="Sunda" country="Indonesia" location="Rumbia" pageId="16" pageNumber="447" specimenCode="SMF 9965" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sulawesi Utara">
|
||
<specimenCode id="DBE59EB526070D0FFB1BFA0DF079FD61" box="[1230,1349,601,626]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="447">SMF 9965</specimenCode>
|
||
,
|
||
<specimenCount id="9D45FD4726070D0FFA85FA0DF08CFD62" box="[1360,1456,602,626]" count="1" pageId="16" pageNumber="447" type="female">1 female</specimenCount>
|
||
and
|
||
<specimenCount id="9D45FD4726070D0FFC8EFA2DF68CFD82" box="[859,944,634,658]" count="2" pageId="16" pageNumber="447" type="male">2 males</specimenCount>
|
||
(
|
||
<quantity id="4CBB9B2B26070D0FFC15FA2DF10FFD81" box="[960,1075,634,658]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" metricValueMax="7.3" metricValueMin="4.9" pageId="16" pageNumber="447" unit="mm" value="61.0" valueMax="73.0" valueMin="49.0">49-73 mm</quantity>
|
||
SL),
|
||
<collectingCountry id="F354765E26070D0FFBA7FA2DF1DDFD82" box="[1138,1249,634,658]" name="Indonesia" pageId="16" pageNumber="447">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4987F82C26070D0FFB3EFA2DF07AFD82" box="[1259,1350,634,658]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="447">Celebes</collectingRegion>
|
||
,
|
||
<location id="8E9C601526070D0FFA84FA2DF097FD82" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03:8E9C601526070D0FFA84FA2DF097FD82" box="[1361,1451,634,658]" country="Indonesia" name="Rumbia" pageId="16" pageNumber="447" stateProvince="Sulawesi Utara">Rumbia</location>
|
||
,
|
||
<date id="FFFD100E26070D0FFCF8FACDF6ABFDA2" box="[813,919,666,690]" pageId="16" pageNumber="447" value="1909-10">
|
||
<collectingDate id="EFB9E9E626070D0FFCF8FACDF6ABFDA2" box="[813,919,666,690]" pageId="16" pageNumber="447" value="1909-10">Oct. 1909</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B6531826070D0FFC75FACDF6DFFDA2" box="[928,995,666,690]" pageId="16" pageNumber="447">Sunda</collectorName>
|
||
exp. coll
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C9326070D0FFB9BFACDF68DFDE2" collectionCode="MNHN" collectorName="d'Aubenton & Fily" country="Cambodia" location="Sihanoukville" pageId="16" pageNumber="447" specimenCode="MNHN 1963-0596" specimenCount="1" stateProvince="Sihanoukville">
|
||
<specimenCode id="DBE59EB526070D0FFB9BFACDF026FDA2" box="[1102,1306,666,690]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="447">MNHN 1963-0596</specimenCode>
|
||
, male (
|
||
<quantity id="4CBB9B2B26070D0FFAB1FACDF093FDA1" box="[1380,1455,666,690]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="447" unit="mm" value="50.0">50 mm</quantity>
|
||
SL),
|
||
<collectingCountry id="F354765E26070D0FFCB2FAEDF6DCFDC2" box="[871,992,698,722]" name="Cambodia" pageId="16" pageNumber="447">Cambodia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4987F82C26070D0FFC39FAEDF1ACFDC2" box="[1004,1168,698,722]" country="Cambodia" name="Sihanoukville" pageId="16" pageNumber="447">Sihanoukville</collectingRegion>
|
||
, 1961-1962,
|
||
<collectorName id="26B6531826070D0FFAF2FAEDF093FDC2" box="[1319,1455,698,722]" pageId="16" pageNumber="447">d’Aubenton</collectorName>
|
||
&
|
||
<collectorName id="26B6531826070D0FFC9FFA8DF64BFDE2" box="[842,887,730,754]" pageId="16" pageNumber="447">Fily</collectorName>
|
||
coll.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2B3C9326070D0FFC6CFA8DF1C9FC02" collectionCode="MNHN" collectorName="d'Aubenton & Fily" country="Cambodia" location="Sihanoukville" pageId="16" pageNumber="447" specimenCode="MNHN 1985-0991, 1" specimenCount="1" stateProvince="Sihanoukville">
|
||
<specimenCode id="DBE59EB526070D0FFC6CFA8DF1ADFDE2" box="[953,1169,730,754]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="447">MNHN 1985-0991</specimenCode>
|
||
,
|
||
<specimenCode id="DBE59EB526070D0FFB4BFA8DF197FDE2" box="[1182,1195,730,754]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="447">1</specimenCode>
|
||
spm,
|
||
<collectingCountry id="F354765E26070D0FFB24FA8DF05BFDE2" box="[1265,1383,730,754]" name="Cambodia" pageId="16" pageNumber="447">Cambodia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4987F82C26070D0FFAA1FA8DF6A5FC02" country="Cambodia" name="Sihanoukville" pageId="16" pageNumber="447">Sihanoukville</collectingRegion>
|
||
, 1962,
|
||
<collectorName id="26B6531826070D0FFC3CFAADF151FC02" box="[1001,1133,762,786]" pageId="16" pageNumber="447">d’Aubenton</collectorName>
|
||
&
|
||
<collectorName id="26B6531826070D0FFB45FAADF181FC02" box="[1168,1213,762,786]" pageId="16" pageNumber="447">Fily</collectorName>
|
||
coll.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2B3C9326070D0FFB29FAADF08CFC22" collectingDate="2001-12" collectorName="Onishi" country="Myanmar" location="Gwa" municipality="Rakhaine" pageId="16" pageNumber="447" specimenCode="NSMT-P 71103, 6" specimenCount="1">
|
||
<specimenCode id="DBE59EB526070D0FFB29FAADF097FC02" box="[1276,1451,762,786]" collectionCode="NSMT-P" pageId="16" pageNumber="447">NSMT-P 71103</specimenCode>
|
||
,
|
||
<specimenCode id="DBE59EB526070D0FFCF8FB4DF607FC22" box="[813,827,794,818]" collectionCode="NSMT-P" pageId="16" pageNumber="447">6</specimenCode>
|
||
spm,
|
||
<collectingCountry id="F354765E26070D0FFCA8FB4DF6D6FC22" box="[893,1002,794,818]" name="Myanmar" pageId="16" pageNumber="447">Myanmar</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6B98ACB426070D0FFC20FB4DF15CFC22" box="[1013,1120,794,818]" pageId="16" pageNumber="447">Rakhaine</collectingMunicipality>
|
||
,
|
||
<location id="8E9C601526070D0FFBB9FB4DF19FFC22" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03:8E9C601526070D0FFBB9FB4DF19FFC22" box="[1132,1187,794,818]" country="Myanmar" municipality="Rakhaine" name="Gwa" pageId="16" pageNumber="447">Gwa</location>
|
||
,
|
||
<date id="FFFD100E26070D0FFB7BFB4DF01EFC22" box="[1198,1314,794,818]" pageId="16" pageNumber="447" value="2001-12">
|
||
<collectingDate id="EFB9E9E626070D0FFB7BFB4DF01EFC22" box="[1198,1314,794,818]" pageId="16" pageNumber="447" value="2001-12">Dec. 2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B6531826070D0FFAFBFB4DF044FC22" box="[1326,1400,794,818]" pageId="16" pageNumber="447">Onishi</collectorName>
|
||
coll.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2B3C9326070D0FFCF8FB6DF6D7FC62" collectingDate="2001-12" collectorName="Onishi" country="Myanmar" location="Gwa" municipality="Rakhaine" pageId="16" pageNumber="447" specimenCode="NSMT-P 71104, 3" specimenCount="1">
|
||
<specimenCode id="DBE59EB526070D0FFCF8FB6DF6E2FC42" box="[813,990,826,850]" collectionCode="NSMT-P" pageId="16" pageNumber="447">NSMT-P 71104</specimenCode>
|
||
,
|
||
<specimenCode id="DBE59EB526070D0FFC3FFB6DF6C4FC42" box="[1002,1016,826,850]" collectionCode="NSMT-P" pageId="16" pageNumber="447">3</specimenCode>
|
||
spms,
|
||
<collectingCountry id="F354765E26070D0FFB92FB6DF189FC42" box="[1095,1205,826,850]" name="Myanmar" pageId="16" pageNumber="447">Myanmar</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6B98ACB426070D0FFB15FB6DF011FC42" box="[1216,1325,826,850]" pageId="16" pageNumber="447">Rakhaine</collectingMunicipality>
|
||
,
|
||
<location id="8E9C601526070D0FFAECFB6DF04CFC42" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03:8E9C601526070D0FFAECFB6DF04CFC42" box="[1337,1392,826,850]" country="Myanmar" municipality="Rakhaine" name="Gwa" pageId="16" pageNumber="447">Gwa</location>
|
||
,
|
||
<date id="FFFD100E26070D0FFAA9FB6DF659FC62" pageId="16" pageNumber="447" value="2001-12">
|
||
<collectingDate id="EFB9E9E626070D0FFAA9FB6DF659FC62" pageId="16" pageNumber="447" value="2001-12">Dec. 2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B6531826070D0FFCBAFB0DF68AFC62" box="[879,950,858,882]" pageId="16" pageNumber="447">Onishi</collectorName>
|
||
coll.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2B3C9326070D0FFC24FB0DF126FC82" collectingDate="2003-03-09" collectorName="Shibukawa" country="Thailand" location="Libong" pageId="16" pageNumber="447" specimenCode="NSMT-P 66100" specimenCount="1">
|
||
<specimenCode id="DBE59EB526070D0FFC24FB0DF1A6FC62" box="[1009,1178,858,882]" collectionCode="NSMT-P" pageId="16" pageNumber="447">NSMT-P 66100</specimenCode>
|
||
,
|
||
<collectingCountry id="F354765E26070D0FFB70FB0DF03BFC62" box="[1189,1287,858,882]" name="Thailand" pageId="16" pageNumber="447">Thailand</collectingCountry>
|
||
,
|
||
<location id="8E9C601526070D0FFAC5FB0DF05CFC62" LSID="urn:lsid:plazi:treatment:03EA87D826070D0EFED0FD2DF1FFFA03:8E9C601526070D0FFAC5FB0DF05CFC62" box="[1296,1376,858,882]" country="Thailand" name="Libong" pageId="16" pageNumber="447">Libong</location>
|
||
,
|
||
<date id="FFFD100E26070D0FFABFFB0DF659FC82" pageId="16" pageNumber="447" value="2003-03-09">
|
||
<collectingDate id="EFB9E9E626070D0FFABFFB0DF659FC82" pageId="16" pageNumber="447" value="2003-03-09">9 Mar. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B6531826070D0FFCBAFB2DF6DAFC82" box="[879,998,890,914]" pageId="16" pageNumber="447">Shibukawa</collectorName>
|
||
coll.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359654526070D0FFCF8FBEDF0A9FB42" pageId="16" pageNumber="447" type="diagnosis">
|
||
<paragraph id="8BFC36CE26070D0FFCF8FBEDF6A7FCC2" blockId="16.[813,1455,954,1106]" box="[813,923,954,978]" pageId="16" pageNumber="447">
|
||
<emphasis id="B937EADC26070D0FFCF8FBEDF6A7FCC2" bold="true" box="[813,923,954,978]" pageId="16" pageNumber="447">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26070D0FFC87FB8DF0A9FB42" blockId="16.[813,1455,954,1106]" pageId="16" pageNumber="447">
|
||
Interorbital space without scales with IOS 0/0/0 (
|
||
<figureCitation id="13782A4B26070D0FFCE0FBADF6B3FB05" box="[821,911,1018,1045]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="16" pageNumber="447">
|
||
Fig. 1a
|
||
<subScript id="17C7348B26070D0FFC53FC53F6B3FB05" attach="left" box="[902,911,1028,1045]" fontSize="7" pageId="16" pageNumber="447">1</subScript>
|
||
</figureCitation>
|
||
); jaws reaching back to below front part of eye (
|
||
<figureCitation id="13782A4B26070D0FFCE0FC4DF6BBFB22" box="[821,903,1050,1074]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="16" pageNumber="447">Fig. 1b</figureCitation>
|
||
<figureCitation id="13782A4B26070D0FFC52FC73F6ACFB25" box="[903,912,1060,1077]" captionStart="Figure 2" captionStartId="5.[126,189,1574,1596]" captionTargetBox="[132,1425,407,1517]" captionTargetId="figure-154@5.[132,1455,401,1558]" captionTargetPageId="5" captionText="Figure 2 (see also following page). – A: Bayesian tree of the cytochrome c oxidase subunit (COI; 452 bp) for sequenced specimens of Butis. Numbers at each node represent posterior probabilities. B: Barcoding NJ K2P tree of the COI gene (196 bp) for sequenced specimens of Butis including the holotype of Butis humeralis from the MNHN collection. Numbers at each node represent bootstrap values. C, D: Barcoding NJ K2P trees of the 12S and the 16S genes (129 and 270 bp respectively) for sequenced specimens of Butis collected during the 19th century from the MNHN collection including a paratype of B. audebertae (MNHN A-1503). Numbers at each node represent bootstrap values. [A high resolution version is available online.]" figureDoi="http://doi.org/10.5281/zenodo.12713761" httpUri="https://zenodo.org/record/12713761/files/figure.png" pageId="16" pageNumber="447">
|
||
<subScript id="17C7348B26070D0FFC52FC73F6ACFB25" attach="left" box="[903,912,1060,1077]" fontSize="7" pageId="16" pageNumber="447">2</subScript>
|
||
</figureCitation>
|
||
); no auxiliary scales on the body (
|
||
<figureCitation id="13782A4B26070D0FFAF5FC4DF04CFB22" box="[1312,1392,1050,1074]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="16" pageNumber="447">Fig. 1c</figureCitation>
|
||
<figureCitation id="13782A4B26070D0FFAA5FC73F045FB25" box="[1392,1401,1060,1077]" captionStart="Figure 4" captionStartId="13.[132,197,1575,1597]" captionTargetBox="[132,775,215,1558]" captionTargetPageId="13" captionText="Figure 4. – A: Butis amboinensis, neotype, RMNH.PISC.85037 (77.5 mm SL), Indonesia (Photo F. Loggen, RMNH); B: Butis amboinensis, in MNHN 2005-1897 (74,4 mm SL), Vanuatu (Photo P. Keith); C: Butis amboinensis, MNHN 2022-0237 (67 mm SL), New Caledonia, tag RTNC_053 (Photo N. Charpin); D: Butis amboinensis, Kolobangara, Solomon (Photo C. Lord)." figureDoi="http://doi.org/10.5281/zenodo.12713765" httpUri="https://zenodo.org/record/12713765/files/figure.png" pageId="16" pageNumber="447">
|
||
<subScript id="17C7348B26070D0FFAA5FC73F045FB25" attach="left" box="[1392,1401,1060,1077]" fontSize="7" pageId="16" pageNumber="447">4</subScript>
|
||
</figureCitation>
|
||
); no black spot on the pectoral fin base; predorsal scales 12-15.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359654526070D0EFCF8FC2DF6E8FD02" lastPageId="17" lastPageNumber="448" pageId="16" pageNumber="447" type="description">
|
||
<paragraph id="8BFC36CE26070D0FFCF8FC2DF68DFB82" blockId="16.[813,1456,1146,1907]" box="[813,945,1146,1170]" pageId="16" pageNumber="447">
|
||
<emphasis id="B937EADC26070D0FFCF8FC2DF68DFB82" bold="true" box="[813,945,1146,1170]" pageId="16" pageNumber="447">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26070D0FFC87FCCDF020FA82" blockId="16.[813,1456,1146,1907]" pageId="16" pageNumber="447">
|
||
The scale counts are given in Table III and selected morphometrics in Table IV. The body is elongated, the body depth at anal fin origin is 14-21% SL, at first dorsal fin 16-21% SL, and the caudal peduncle depth is 8-10% SL. Predorsal length 40-44% SL and preanal length 61-64% SL. Bony ridges on head barely serrated to smooth (12-24 medium spines), usually smooth on snout (4-8 smooth spines). Size: up to
|
||
<quantity id="4CBB9B2B26070D0FFC76FD2DF6C5FA82" box="[931,1017,1402,1426]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="447" unit="mm" value="115.0">115 mm</quantity>
|
||
SL. Twenty-five vertebrae.
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26070D0FFC87FDCDF062F943" blockId="16.[813,1456,1146,1907]" pageId="16" pageNumber="447">
|
||
The head 33-36% SL is depressed as is the snout; snout length 12-14% SL. Mouth terminal, lower jaw prominent, jaws (14-16% SL) reaching back to below front part of eye (
|
||
<figureCitation id="13782A4B26070D0FFCE0FDACF6BEF903" box="[821,898,1531,1555]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="16" pageNumber="447">Fig. 1b</figureCitation>
|
||
<figureCitation id="13782A4B26070D0FFC57FE53F6B7F905" box="[898,907,1540,1557]" captionStart="Figure 2" captionStartId="5.[126,189,1574,1596]" captionTargetBox="[132,1425,407,1517]" captionTargetId="figure-154@5.[132,1455,401,1558]" captionTargetPageId="5" captionText="Figure 2 (see also following page). – A: Bayesian tree of the cytochrome c oxidase subunit (COI; 452 bp) for sequenced specimens of Butis. Numbers at each node represent posterior probabilities. B: Barcoding NJ K2P tree of the COI gene (196 bp) for sequenced specimens of Butis including the holotype of Butis humeralis from the MNHN collection. Numbers at each node represent bootstrap values. C, D: Barcoding NJ K2P trees of the 12S and the 16S genes (129 and 270 bp respectively) for sequenced specimens of Butis collected during the 19th century from the MNHN collection including a paratype of B. audebertae (MNHN A-1503). Numbers at each node represent bootstrap values. [A high resolution version is available online.]" figureDoi="http://doi.org/10.5281/zenodo.12713761" httpUri="https://zenodo.org/record/12713761/files/figure.png" pageId="16" pageNumber="447">
|
||
<subScript id="17C7348B26070D0FFC57FE53F6B7F905" attach="left" box="[898,907,1540,1557]" fontSize="7" pageId="16" pageNumber="447">2</subScript>
|
||
</figureCitation>
|
||
). Outermost row of teeth in each jaw enlarged and widely spaced with strong caniniform teeth. Interorbital width 6-8% SL. Gill opening below rear edge of eye.
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26070D0FFC87FE0CF00AF9E3" blockId="16.[813,1456,1146,1907]" pageId="16" pageNumber="447">Dorsal fins VI-I,8 with no filamentous rays; first dorsal fin triangular. Anal fin I,7-8 directly opposite to the second dorsal fin. Pectoral fins 18 with the posterior margin slightly rounded. The caudal fin is with 12 branched- rays and its posterior margin is rounded. Pelvic fins separate, I,5.</paragraph>
|
||
<paragraph id="8BFC36CE26070D0EFC87FEACF7F7FB2F" blockId="16.[813,1456,1146,1907]" lastBlockId="17.[126,775,800,1218]" lastPageId="17" lastPageNumber="448" pageId="16" pageNumber="447">
|
||
Lateral scales 25-28, with ctenoid scales on flanks, opercle and caudal peduncle. Cheek usually naked. Scales in transverse back series 8-9, in transverse forward 11-14, in predorsal 12-15 and in zigzag usually 6. Interopercle naked or with few scattered scales; no scales between orbital ridge and eyes, IOS 0/0/0 (
|
||
<figureCitation id="13782A4B26060D0EFEBFFB16F4FCFC4B" box="[362,448,833,859]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="17" pageNumber="448">
|
||
Fig. 1a
|
||
<subScript id="17C7348B26060D0EFE62FB1DF4FCFC4B" attach="left" box="[439,448,842,859]" fontSize="7" pageId="17" pageNumber="448">1</subScript>
|
||
</figureCitation>
|
||
). No scales on cheeks. Scales absent from snout. Pectoral base, breast and belly scales cycloid. No auxiliary scales on body (
|
||
<figureCitation id="13782A4B26060D0EFDF5FBD5F750FC8A" box="[544,620,898,922]" captionStart="Figure 1" captionStartId="2.[1077,1141,1813,1835]" captionTargetBox="[138,1058,221,1894]" captionTargetId="figure-54@2.[132,1060,215,1900]" captionTargetPageId="2" captionText="Figure 1. – A: The different patterns of interorbital scales (IOS); head in dorsal view. B: Size of the jaw; head in lateral view. C: Auxiliary scales on flanks." figureDoi="http://doi.org/10.5281/zenodo.12713759" httpUri="https://zenodo.org/record/12713759/files/figure.png" pageId="17" pageNumber="448">Fig. 1c</figureCitation>
|
||
<figureCitation id="13782A4B26060D0EFDB9FBDBF749FC8D" box="[620,629,908,925]" captionStart="Figure 4" captionStartId="13.[132,197,1575,1597]" captionTargetBox="[132,775,215,1558]" captionTargetPageId="13" captionText="Figure 4. – A: Butis amboinensis, neotype, RMNH.PISC.85037 (77.5 mm SL), Indonesia (Photo F. Loggen, RMNH); B: Butis amboinensis, in MNHN 2005-1897 (74,4 mm SL), Vanuatu (Photo P. Keith); C: Butis amboinensis, MNHN 2022-0237 (67 mm SL), New Caledonia, tag RTNC_053 (Photo N. Charpin); D: Butis amboinensis, Kolobangara, Solomon (Photo C. Lord)." figureDoi="http://doi.org/10.5281/zenodo.12713765" httpUri="https://zenodo.org/record/12713765/files/figure.png" pageId="17" pageNumber="448">
|
||
<subScript id="17C7348B26060D0EFDB9FBDBF749FC8D" attach="left" box="[620,629,908,925]" fontSize="7" pageId="17" pageNumber="448">4</subScript>
|
||
</figureCitation>
|
||
). Head pores present, with usually incomplete sensory canal and pores (A’, B, C, (D), (E), F, (G), H, I, J, K, L’); 5 preopercular pores (M’, N, O, P, Q’). The pore numbers are asymmetrical in some individuals, with a pore on one side of the head, which is absent on the other side. D, E, G sometimes missing.
|
||
</paragraph>
|
||
<caption id="DF3C664626060D0EFF51FA87F4A4FDEE" ID-DOI="http://doi.org/10.5281/zenodo.12713769" ID-Zenodo-Dep="12713769" httpUri="https://zenodo.org/record/12713769/files/figure.png" pageId="17" pageNumber="448" startId="17.[132,197,720,742]" targetBox="[132,775,215,703]" targetPageId="17" targetType="figure">
|
||
<paragraph id="8BFC36CE26060D0EFF51FA87F4A4FDEE" blockId="17.[132,775,720,766]" pageId="17" pageNumber="448">
|
||
Figure 6. –
|
||
<taxonomicName id="4C434D4D26060D0EFF2CFA87F48AFDF5" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[249,438,720,741]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="gymnopomus">
|
||
<emphasis id="B937EADC26060D0EFF2CFA87F48AFDF5" box="[249,438,720,741]" italics="true" pageId="17" pageNumber="448">Butis gymnopomus</emphasis>
|
||
</taxonomicName>
|
||
, uncatalogued, BIF 0311 (59 mm SL), Java (Photo N. Hubert).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFC36CE26060D0EFF7FFC1FF44EFBD2" blockId="17.[126,775,800,1218]" pageId="17" pageNumber="448">Males with an elongated urogenital papilla with distal tip rounded. Females have an oval or chalice-shaped bulbous urogenital papilla with slightly crenulated outer edges around distal opening.</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFF51FCBCF4B9FA13" blockId="17.[132,776,1259,1546]" box="[132,389,1259,1283]" pageId="17" pageNumber="448">
|
||
<emphasis id="B937EADC26060D0EFF51FCBCF4B9FA13" bold="true" box="[132,389,1259,1283]" pageId="17" pageNumber="448">Colour in preservation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFF7FFD5BF592F91A" blockId="17.[132,776,1259,1546]" pageId="17" pageNumber="448">Male and female similar. Head, body and belly grey-beige to brown. Head with a distinct double broad dark brown to blackish band on snout. Seven to eight oblique dark bands from head to caudal fin. Pectoral and pelvic fins beige. Abdomen and gular region beige. First dorsal fin brown to blackish. Anal and second dorsal fins dark with spots aligned on rays. Caudal fin brownish to blackish with greyish margin.</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFF51FE64F426F95B" blockId="17.[132,776,1587,1907]" box="[132,282,1587,1611]" pageId="17" pageNumber="448">
|
||
<emphasis id="B937EADC26060D0EFF51FE64F426F95B" bold="true" box="[132,282,1587,1611]" pageId="17" pageNumber="448">Colour in life</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFF7FFE03F6E8FD02" blockId="17.[132,776,1587,1907]" lastBlockId="17.[813,1455,217,530]" pageId="17" pageNumber="448">
|
||
Male and female similar (
|
||
<figureCitation id="13782A4B26060D0EFE11FE03F734F97C" box="[452,520,1620,1644]" captionStart="Figure 6" captionStartId="17.[132,197,720,742]" captionTargetBox="[132,775,215,703]" captionTargetId="figure-730@17.[132,775,450,703]" captionTargetPageId="17" captionText="Figure 6. – Butis gymnopomus, uncatalogued, BIF 0311 (59 mm SL), Java (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12713769" httpUri="https://zenodo.org/record/12713769/files/figure.png" pageId="17" pageNumber="448">Fig. 6</figureCitation>
|
||
). Head, body and belly grey-beige to brown or red, with several lines of white spots framing scale rows or constituting several longitudinal white to pink lines. Head with a double broad dark brown to blackish band from snout tip running through eye, and scattered with numerous white spots on snout, preopercle and opercle. Opercle pinkish. Several specimens with seven to eight light- er oblique bands from head to caudal fin. Sometimes a dark band runs along the body and ends at caudal fin base. Lips plain brown; eyes red. First dorsal fin brown to blackish with a beige blotch at posterior part. Second dorsal fin blackish with whitish spots along rays and sometimes with a red basal part. Anal and pelvic fins mostly red with a black and white margin or yellowish to brown with some black and yellow to whitish spotting posteriorly. No black spots nor red spots on pectoral fin base; pectoral fin translucent with a base mottled with brown and blackish. Caudal fin translucent to blackish with fine whitish spots; lower and upper base reddish, lower margin whitish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359654526060D0EFCF8FA6DF685FDE2" pageId="17" pageNumber="448" type="biology_ecology">
|
||
<paragraph id="8BFC36CE26060D0EFCF8FA6DF6BBFD42" blockId="17.[813,1456,570,754]" box="[813,903,570,594]" pageId="17" pageNumber="448">
|
||
<emphasis id="B937EADC26060D0EFCF8FA6DF6BBFD42" bold="true" box="[813,903,570,594]" pageId="17" pageNumber="448">Ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFC87FA0DF685FDE2" blockId="17.[813,1456,570,754]" pageId="17" pageNumber="448">
|
||
This species occurs in many coastal streams in brackish to freshwaters over substrates of mud, gravels, sandy-mud or rocky bottom. It is an ambush hunter, feeding mainly on small fish and crustaceans (shrimps (
|
||
<taxonomicName id="4C434D4D26060D0EFB6CFAEDF028FDC2" box="[1209,1300,698,722]" class="Malacostraca" family="Atyidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="448" phylum="Arthropoda" rank="family">Atyidae</taxonomicName>
|
||
), prawns (
|
||
<taxonomicName id="4C434D4D26060D0EFA57FAEDF699FDE2" authorityName="Rafinesque" authorityYear="1815" class="Malacostraca" family="Palaemonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="448" phylum="Arthropoda" rank="family">Palaemonidae</taxonomicName>
|
||
)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359654526060D0EFCF8FB4DF1FFFA03" pageId="17" pageNumber="448" type="distribution">
|
||
<paragraph id="8BFC36CE26060D0EFCF8FB4DF684FC22" blockId="17.[813,1455,794,882]" box="[813,952,794,818]" pageId="17" pageNumber="448">
|
||
<emphasis id="B937EADC26060D0EFCF8FB4DF684FC22" bold="true" box="[813,952,794,818]" pageId="17" pageNumber="448">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFC87FB6DF1BAFC62" blockId="17.[813,1455,794,882]" pageId="17" pageNumber="448">
|
||
Known from
|
||
<collectingCountry id="F354765E26060D0EFC3BFB6DF111FC42" box="[1006,1069,826,850]" name="India" pageId="17" pageNumber="448">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F354765E26060D0EFBECFB6DF193FC42" box="[1081,1199,826,850]" name="Sri Lanka" pageId="17" pageNumber="448">Sri Lanka</collectingCountry>
|
||
,
|
||
<collectingCountry id="F354765E26060D0EFB6EFB6DF010FC42" box="[1211,1324,826,850]" name="Myanmar" pageId="17" pageNumber="448">Myanmar</collectingCountry>
|
||
,
|
||
<collectingCountry id="F354765E26060D0EFAEDFB6DF097FC42" box="[1336,1451,826,850]" name="Indonesia" pageId="17" pageNumber="448">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F354765E26060D0EFCF8FB0DF6B0FC62" box="[813,908,858,882]" name="Thailand" pageId="17" pageNumber="448">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="F354765E26060D0EFC68FB0DF12BFC62" box="[957,1047,858,882]" name="Vietnam" pageId="17" pageNumber="448">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="13782A4B26060D0EFBF3FB0DF144FC62" box="[1062,1144,858,882]" captionStart="Figure 3" captionStartId="7.[132,195,1245,1267]" captionTargetBox="[133,1454,216,1229]" captionTargetPageId="7" captionText="Figure 3. – A, B: Distribution of Butis species." figureDoi="http://doi.org/10.5281/zenodo.12713763" httpUri="https://zenodo.org/record/12713763/files/figure.png" pageId="17" pageNumber="448">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFCF8FBCDF686FCA2" blockId="17.[813,1457,922,1299]" box="[813,954,922,946]" pageId="17" pageNumber="448">
|
||
<emphasis id="B937EADC26060D0EFCF8FBCDF686FCA2" bold="true" box="[813,954,922,946]" pageId="17" pageNumber="448">Comparison</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36CE26060D0EFC87FBECF1FFFA03" blockId="17.[813,1457,922,1299]" pageId="17" pageNumber="448">
|
||
<taxonomicName id="4C434D4D26060D0EFC87FBECF10AFCC2" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[850,1078,955,978]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="gymnopomus">
|
||
<emphasis id="B937EADC26060D0EFC87FBECF10AFCC2" box="[850,1078,955,978]" italics="true" pageId="17" pageNumber="448">Butis gymnopomus</emphasis>
|
||
</taxonomicName>
|
||
differs from the other species sequenced in having a significant percentage of divergence in
|
||
<emphasis id="B937EADC26060D0EFC9EFBACF646FB02" box="[843,890,1019,1042]" italics="true" pageId="17" pageNumber="448">COI</emphasis>
|
||
gene (15.7-17.6%) (Table II). Moreover, it differs from
|
||
<emphasis id="B937EADC26060D0EFCB3FC4CF100FB42" italics="true" pageId="17" pageNumber="448">
|
||
<taxonomicName id="4C434D4D26060D0EFCB3FC4CF6C1FB22" authorityName="Keith & Mennesson" authorityYear="2023" box="[870,1021,1050,1074]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="audebertae">B. audebertae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D4D26060D0EFBD2FC4CF1ACFB22" box="[1031,1168,1050,1074]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="humeralis">B. humeralis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D4D26060D0EFB4FFC4CF03AFB22" authorityName="Keith & Mennesson" authorityYear="2023" box="[1178,1286,1050,1074]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="huberti">B. huberti</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D4D26060D0EFAC5FC4CF097FB22" baseAuthorityName="Bleeker" baseAuthorityYear="1849" box="[1296,1451,1051,1074]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="prismaticus">B. prismaticus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D4D26060D0EFCF8FC6CF6B9FB42" authorityName=", MNHN" authorityYear="2022" box="[813,901,1082,1106]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="butis">B. butis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D4D26060D0EFC44FC6CF100FB42" box="[913,1084,1082,1106]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="amboinensis">B. amboinensis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C434D4D26060D0EFBA6FC6CF1E3FB42" authorityName="Keith & Mennesson" authorityYear="2023" box="[1139,1247,1082,1106]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="abdoui">
|
||
<emphasis id="B937EADC26060D0EFBA6FC6CF1E3FB42" box="[1139,1247,1082,1106]" italics="true" pageId="17" pageNumber="448">B. abdoui</emphasis>
|
||
</taxonomicName>
|
||
in having 0 scales
|
||
<emphasis id="B937EADC26060D0EFCF8FC0CF649FB62" box="[813,885,1115,1138]" italics="true" pageId="17" pageNumber="448">versus</emphasis>
|
||
2-11 between the two orbital ridges and predorsal scales 12-15
|
||
<emphasis id="B937EADC26060D0EFC68FC2CF13FFB82" box="[957,1027,1147,1170]" italics="true" pageId="17" pageNumber="448">versus</emphasis>
|
||
25-46. It differs from
|
||
<taxonomicName id="4C434D4D26060D0EFB2FFC2CF093FB82" baseAuthorityName="Bleeker" baseAuthorityYear="1849" box="[1274,1455,1146,1170]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="koilomatodon">
|
||
<emphasis id="B937EADC26060D0EFB2FFC2CF093FB82" box="[1274,1455,1146,1170]" italics="true" pageId="17" pageNumber="448">B. koilomatodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C434D4D26060D0EFC89FCCCF6C0FBA2" baseAuthorityName="Barnard" baseAuthorityYear="1927" box="[860,1020,1178,1202]" family="Eleotridae" genus="Butis" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="448" phylum="Chordata" rank="species" species="delagoensis">
|
||
<emphasis id="B937EADC26060D0EFC89FCCCF6C0FBA2" box="[860,1020,1178,1202]" italics="true" pageId="17" pageNumber="448">B. delagoensis</emphasis>
|
||
</taxonomicName>
|
||
in having jaws reaching back to below front part of eye
|
||
<emphasis id="B937EADC26060D0EFC3CFCECF10CFBC2" box="[1001,1072,1211,1234]" italics="true" pageId="17" pageNumber="448">versus</emphasis>
|
||
jaws reaching back to one third to middle of eye, pectoral fin rays 18
|
||
<emphasis id="B937EADC26060D0EFB7EFC8CF1CDFBE2" box="[1195,1265,1243,1266]" italics="true" pageId="17" pageNumber="448">versus</emphasis>
|
||
20-22, and snout length 12-14% SL (
|
||
<emphasis id="B937EADC26060D0EFC28FCACF17DFA02" box="[1021,1089,1275,1298]" italics="true" pageId="17" pageNumber="448">versus</emphasis>
|
||
7-10% SL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |