treatments-xml/data/03/EA/87/03EA8795FFAF714466CAA87FFC1AF903.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.3790385" ID-GBIF-Dataset="b2ba6675-f6f1-4ace-a151-a385e3142ee0" ID-GBIF-Taxon="163755074" ID-ISSN="13132970" ID-Zenodo-Dep="3790385" ID-ZooBank="57917E4C-32AB-460F-A7A7-64BF79B87908" approvalRequired="14" approvalRequired_for_textStreams="14" checkinTime="1588000168194" checkinUser="plazi" docAuthor="Torres-Carvajal, Omar, de Queiroz, Kevin &amp; Etheridge, Richard" docDate="2009" docId="03EA8795FFAF714466CAA87FFC1AF903" docLanguage="en" docName="ZK_article_2102.pdf" docOrigin="ZooKeys 27 (27)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@56872fcb" docStyleName="zookeys.2008.journal_article" docTitle="Enyalioides rubrigularis Torres-Carvajal &amp; de Queiroz &amp; Etheridge 2009, sp. n." docType="treatment" docVersion="4" lastPageId="9" lastPageNumber="68" masterDocId="FFD3FFEDFFAD714D6646AA56FFF2FF8E" masterDocTitle="A new species of iguanid lizard (Hoplocercinae, Enyalioides) from southern Ecuador with a key to eastern Ecuadorian Enyalioides" masterLastPageNumber="71" masterPageNumber="59" pageId="2" pageNumber="61" updateTime="1643568103201" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of iguanid lizard (Hoplocercinae, Enyalioides) from southern Ecuador with a key to eastern Ecuadorian Enyalioides</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torres-Carvajal, Omar</mods:namePart>
<mods:affiliation>Pontificia Universidad Católica del Ecuador, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Queiroz, Kevin</mods:namePart>
<mods:affiliation>National Museum of Natural History, Smithsonian Institution MRC 162, 20560, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Etheridge, Richard</mods:namePart>
<mods:affiliation>San Diego State University, San Diego ,,</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-11-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>27</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>27</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>59</mods:start>
<mods:end>71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.27.273</mods:identifier>
<mods:identifier type="GBIF-Dataset">b2ba6675-f6f1-4ace-a151-a385e3142ee0</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576568</mods:identifier>
<mods:identifier type="ZooBank">57917E4C-32AB-460F-A7A7-64BF79B87908</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790385" ID-GBIF-Taxon="163755074" ID-Zenodo-Dep="3790385" LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903" httpUri="http://treatment.plazi.org/id/03EA8795FFAF714466CAA87FFC1AF903" lastPageId="9" lastPageNumber="68" pageId="2" pageNumber="61">
<subSubSection pageId="2" pageNumber="61" type="nomenclature">
<paragraph blockId="2.[140,976,553,616]" box="[140,501,553,580]" pageId="2" pageNumber="61">
<heading box="[140,501,553,580]" fontSize="11" level="2" pageId="2" pageNumber="61" reason="3">
<emphasis bold="true" box="[140,501,553,580]" pageId="2" pageNumber="61">
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[140,429,553,580]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis bold="true" box="[140,429,553,580]" italics="true" pageId="2" pageNumber="61">Enyalioides rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[436,501,554,580]" pageId="2" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[140,976,553,616]" box="[140,976,589,616]" pageId="2" pageNumber="61">
<uri box="[140,976,589,616]" pageId="2" pageNumber="61">
urn:lsid:zoobank.org:act:
<uuid box="[424,976,589,616]" pageId="2" pageNumber="61">1819C358-E8CC-4076-BB34-5DD0BF3F55F8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="61" type="materials_examined">
<paragraph blockId="2.[140,1108,659,1672]" pageId="2" pageNumber="61">
<materialsCitation ID-GBIF-Occurrence="2608994675" collectingDate="2008-06-23" collectionCode="QCAZ" collectorName="de Mesias San Martin &amp; O. Torres-Carvajal &amp; E. Arbelaez &amp; Campos &amp; D. Salazar." country="Ecuador" elevation="1154" latitude="-3.8563888" location="Piuntza" longLatPrecision="21" longitude="-78.86472" municipality="Provincia" pageId="2" pageNumber="61" specimenCode="QCAZ 8483" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
<emphasis box="[140,235,659,685]" italics="true" pageId="2" pageNumber="61">
<typeStatus box="[140,235,659,685]" pageId="2" pageNumber="61">Holotype</typeStatus>
</emphasis>
.
<specimenCode box="[255,408,660,686]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 8483</specimenCode>
(Fig. 1), an adult male from finca
<collectorName box="[844,1107,659,686]" pageId="2" pageNumber="61">de Mesías San Martín</collectorName>
(
<geoCoordinate box="[149,266,694,721]" degrees="3" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="23" value="-3.8563888">3°51'23&quot;S</geoCoordinate>
,
<geoCoordinate box="[282,424,694,721]" degrees="78" direction="west" minutes="51" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="53" value="-78.86472">78°51'53&quot;W</geoCoordinate>
,
<quantity box="[440,526,694,721]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="2" pageNumber="61" unit="m" value="1154.0">
<elevation box="[440,526,694,721]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="2" pageNumber="61" unit="m" value="1154.0">1154m</elevation>
</quantity>
), near
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F6424A8E1FD49FD5F" box="[610,699,695,721]" country="Ecuador" latitude="-3.8563888" longLatPrecision="21" longitude="-78.86472" municipality="Provincia" name="Piuntza" pageId="2" pageNumber="61" stateProvince="Zamora-Chinchipe">Piuntza</location>
,
<collectingMunicipality box="[714,819,695,721]" pageId="2" pageNumber="61">Provincia</collectingMunicipality>
<collectingRegion box="[831,1053,695,721]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="61">Zamora Chinchipe</collectingRegion>
,
<collectingCountry name="Ecuador" pageId="2" pageNumber="61">Ecuador</collectingCountry>
, collected on
<date box="[374,536,730,756]" pageId="2" pageNumber="61" value="2008-06-23">
<collectingDate box="[374,536,730,756]" pageId="2" pageNumber="61" value="2008-06-23">23 June 2008</collectingDate>
</date>
by
<collectorName box="[585,803,730,756]" pageId="2" pageNumber="61">O. Torres-Carvajal</collectorName>
,
<collectorName box="[818,953,730,756]" pageId="2" pageNumber="61">E. Arbeláez</collectorName>
, A. Carvajal-
<collectorName box="[140,237,765,791]" pageId="2" pageNumber="61">Campos</collectorName>
, and
<collectorName box="[298,420,765,792]" pageId="2" pageNumber="61">D. Salazar.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="2.[140,1108,659,1672]" pageId="2" pageNumber="61">
<materialsCitation ID-GBIF-Occurrence="2608994680" collectingDate="2008-06-23" collectionCode="QCAZ" collectorName="de Mesias San Martin &amp; O. Torres-Carvajal &amp; E. Arbelaez &amp; Campos &amp; D. Salazar." country="Ecuador" elevation="1154" latitude="-3.8563888" location="Provincia" longLatPrecision="21" longitude="-78.86472" municipality="Provincia" pageId="2" pageNumber="61" specimenCode="QCAZ 8484" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<emphasis box="[185,285,801,826]" italics="true" pageId="2" pageNumber="61">
<typeStatus box="[185,285,801,826]" pageId="2" pageNumber="61">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry box="[305,451,800,826]" name="Ecuador" pageId="2" pageNumber="61">ECUADOR</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F6795A976FDCEFCB4" box="[467,572,800,826]" country="Ecuador" name="Provincia" pageId="2" pageNumber="61" stateProvince="Zamora-Chinchipe">Provincia</location>
<collectingRegion box="[584,809,800,827]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="61">Zamora Chinchipe</collectingRegion>
:
<specimenCode box="[825,980,800,826]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 8484</specimenCode>
, same collection data as the holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994676" collectingDate="2008-06-20" collectingDateMax="2008-06-23" collectingDateMin="2008-06-20" collectionCode="QCAZ" country="Ecuador" elevation="1258" latitude="-3.8569443" location="Piuntza" longLatPrecision="21" longitude="-78.865555" pageId="2" pageNumber="61" specimenCode="QCAZ 8454, 8456-58, 8460, 8481-82" specimenCount="1" typeStatus="paratype">
<specimenCode box="[479,632,835,862]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 8454</specimenCode>
,
<specimenCode box="[646,752,835,862]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">845658</specimenCode>
,
<specimenCode box="[767,829,835,861]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">8460</specimenCode>
,
<specimenCode box="[843,948,836,862]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">848182</specimenCode>
, near
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F65B8A915FBA6FCD3" box="[1022,1108,835,861]" country="Ecuador" latitude="-3.8569443" longLatPrecision="21" longitude="-78.865555" name="Piuntza" pageId="2" pageNumber="61">Piuntza</location>
(
<geoCoordinate box="[149,266,870,897]" degrees="3" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="25" value="-3.8569443">3°51'25&quot;S</geoCoordinate>
,
<geoCoordinate box="[280,422,870,897]" degrees="78" direction="west" minutes="51" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="56" value="-78.865555">78°51'56&quot;W</geoCoordinate>
,
<quantity box="[435,521,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="2" pageNumber="61" unit="m" value="1258.0">
<elevation box="[435,521,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="2" pageNumber="61" unit="m" value="1258.0">1258m</elevation>
</quantity>
), collected between
<date box="[753,955,871,897]" pageId="2" pageNumber="61" value="2008-06-20" valueMax="2008-06-23" valueMin="2008-06-20">
<collectingDate box="[753,955,871,897]" pageId="2" pageNumber="61" value="2008-06-20" valueMax="2008-06-23" valueMin="2008-06-20">2023 June 2008</collectingDate>
</date>
by same collectors as for the
<typeStatus box="[329,431,906,932]" pageId="2" pageNumber="61">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994677" collectingDate="2008-06-22" collectingDateMax="2008-06-24" collectingDateMin="2008-06-22" collectionCode="QCAZ" country="Ecuador" elevation="1192" latitude="-3.857222" location="Piuntza" longLatPrecision="21" longitude="-78.86111" pageId="2" pageNumber="61" specimenCode="QCAZ 8459, 8486" specimenCount="1" typeStatus="paratype">
<specimenCode box="[443,591,906,933]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 8459</specimenCode>
,
<specimenCode box="[603,665,906,932]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">8486</specimenCode>
, near
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F649DA9DCFCC3FC2A" box="[731,817,906,932]" country="Ecuador" latitude="-3.857222" longLatPrecision="21" longitude="-78.86111" name="Piuntza" pageId="2" pageNumber="61">Piuntza</location>
(
<geoCoordinate box="[833,949,906,933]" degrees="3" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="26" value="-3.857222">3°51'26&quot;S</geoCoordinate>
,
<geoCoordinate box="[961,1103,906,933]" degrees="78" direction="west" minutes="51" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="40" value="-78.86111">78°51'40&quot;W</geoCoordinate>
,
<quantity box="[140,226,941,967]" metricMagnitude="3" metricUnit="m" metricValue="1.192" pageId="2" pageNumber="61" unit="m" value="1192.0">
<elevation box="[140,226,941,967]" metricMagnitude="3" metricUnit="m" metricValue="1.192" pageId="2" pageNumber="61" unit="m" value="1192.0">1192m</elevation>
</quantity>
), collected between
<date box="[458,659,941,968]" pageId="2" pageNumber="61" value="2008-06-22" valueMax="2008-06-24" valueMin="2008-06-22">
<collectingDate box="[458,659,941,968]" pageId="2" pageNumber="61" value="2008-06-22" valueMax="2008-06-24" valueMin="2008-06-22">2224 June 2008</collectingDate>
</date>
by same collectors as for the
<typeStatus box="[1001,1103,941,967]" pageId="2" pageNumber="61">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994681" collectingDate="2008-06-24" collectionCode="QCAZ" country="Ecuador" elevation="1170" latitude="-3.857222" location="Piuntza" longLatPrecision="21" longitude="-78.861946" pageId="2" pageNumber="61" specimenCode="QCAZ 8485" specimenCount="1" typeStatus="paratype">
<specimenCode box="[140,292,976,1003]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 8485</specimenCode>
, near
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F6728A986FE36FC64" box="[366,452,976,1002]" country="Ecuador" latitude="-3.857222" longLatPrecision="21" longitude="-78.861946" name="Piuntza" pageId="2" pageNumber="61">Piuntza</location>
(
<geoCoordinate box="[471,587,976,1003]" degrees="3" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="26" value="-3.857222">3°51'26&quot;S</geoCoordinate>
,
<geoCoordinate box="[602,744,976,1003]" degrees="78" direction="west" minutes="51" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="43" value="-78.861946">78°51'43&quot;W</geoCoordinate>
,
<quantity box="[759,845,976,1002]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="61" unit="m" value="1170.0">
<elevation box="[759,845,976,1002]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="61" unit="m" value="1170.0">1170m</elevation>
</quantity>
), collected on
<date pageId="2" pageNumber="61" value="2008-06-24">
<collectingDate pageId="2" pageNumber="61" value="2008-06-24">24 June 2008</collectingDate>
</date>
by same collectors as for the
<typeStatus box="[558,660,1011,1037]" pageId="2" pageNumber="61">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994679" collectingDate="2009-04-12" collectionCode="QCAZ" collectorName="Aldas &amp; J. M. Guayasamin &amp; Holger" country="Ecuador" county="Alto Miazi" elevation="1318" latitude="-4.246111" location="Cordillera del Condor" longLatPrecision="21" longitude="-78.61639" municipality="upper Rio Nangaritza" pageId="2" pageNumber="61" specimenCode="QCAZ 9089" specimenCount="1" typeStatus="paratype">
<specimenCode box="[676,830,1012,1038]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="2" pageNumber="61" type="Museum">QCAZ 9089</specimenCode>
,
<collectingCounty box="[846,974,1011,1037]" pageId="2" pageNumber="61">Alto Miazi</collectingCounty>
,
<collectingMunicipality pageId="2" pageNumber="61">upper Río Nangaritza</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F675FAE41FDE9FBBF" box="[281,539,1047,1073]" country="Ecuador" county="Alto Miazi" latitude="-4.246111" longLatPrecision="21" longitude="-78.61639" municipality="upper Rio Nangaritza" name="Cordillera del Condor" pageId="2" pageNumber="61">Cordillera del Cóndor</location>
(
<geoCoordinate box="[559,676,1047,1073]" degrees="4" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="46" value="-4.246111">4°14'46&quot;S</geoCoordinate>
,
<geoCoordinate box="[691,833,1046,1073]" degrees="78" direction="west" minutes="36" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="59" value="-78.61639">78°36'59&quot;W</geoCoordinate>
,
<quantity box="[848,943,1047,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.318" pageId="2" pageNumber="61" unit="m" value="1318.0">
<elevation box="[848,943,1047,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.318" pageId="2" pageNumber="61" unit="m" value="1318.0">1318 m</elevation>
</quantity>
), collected on
<date box="[140,303,1082,1108]" pageId="2" pageNumber="61" value="2009-04-12">
<collectingDate box="[140,303,1082,1108]" pageId="2" pageNumber="61" value="2009-04-12">12 April 2009</collectingDate>
</date>
by S.
<collectorName box="[378,443,1082,1108]" pageId="2" pageNumber="61">Aldás</collectorName>
,
<collectorName box="[456,661,1082,1108]" pageId="2" pageNumber="61">J.M. Guayasamin</collectorName>
,
<collectorName box="[673,755,1082,1108]" pageId="2" pageNumber="61">Holger</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994674" collectingDate="2008-11-07" collectionCode="EPN" collectorName="Almendariz &amp; Salazar &amp; Angamarca" country="Ecuador" county="Los Encuentros" elevation="1460" latitude="-3.8338888" location="Zarza" longLatPrecision="21" longitude="-78.523056" municipality="Bosque Protector" pageId="2" pageNumber="61" specimenCount="1" typeStatus="paratype">
<accessionNumber box="[768,909,1082,1109]" httpUri="http://www.ncbi.nlm.nih.gov/protein/EPN11356" pageId="2" pageNumber="61">
<collectionCode box="[768,824,1082,1108]" country="Ecuador" httpUri="http://grbio.org/cool/y3s2-2p0h" name="Escuela Polytecnica Nacional" pageId="2" pageNumber="61">EPN</collectionCode>
11356
</accessionNumber>
,
<collectingCounty box="[923,1104,1082,1108]" pageId="2" pageNumber="61">Los Encuentros</collectingCounty>
,
<collectingMunicipality box="[140,333,1118,1144]" pageId="2" pageNumber="61">Bosque Protector</collectingMunicipality>
el
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F6728AE08FE5FFBF6" box="[366,429,1118,1144]" country="Ecuador" county="Los Encuentros" latitude="-3.8338888" longLatPrecision="21" longitude="-78.523056" municipality="Bosque Protector" name="Zarza" pageId="2" pageNumber="61">Zarza</location>
(
<geoCoordinate box="[445,547,1117,1144]" degrees="3" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="2" value="-3.8338888">3°50'2&quot;S</geoCoordinate>
,
<geoCoordinate box="[559,701,1117,1143]" degrees="78" direction="west" minutes="31" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="23" value="-78.523056">78°31'23&quot;W</geoCoordinate>
,
<quantity box="[712,804,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="61" unit="m" value="1460.0">
<elevation box="[712,804,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="61" unit="m" value="1460.0">1460 m</elevation>
</quantity>
), collected on
<date pageId="2" pageNumber="61" value="2008-11-07">
<collectingDate pageId="2" pageNumber="61" value="2008-11-07">7 November 2008</collectingDate>
</date>
by A.
<collectorName box="[283,417,1152,1178]" pageId="2" pageNumber="61">Almendáriz</collectorName>
, M.
<collectorName box="[476,555,1152,1178]" pageId="2" pageNumber="61">Salazar</collectorName>
, M.
<collectorName box="[615,745,1152,1178]" pageId="2" pageNumber="61">Angamarca</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994678" collectingDate="2008-03-27" collectionCode="EPN" collectorName="Almendariz &amp; Vivanco &amp; Sarango" country="Ecuador" elevation="1450" latitude="-3.8077776" location="Concesion Cuy" longLatPrecision="21" longitude="-78.605835" municipality="Los Encuentros" pageId="2" pageNumber="61" specimenCode="EPN 12432-33" specimenCount="1" typeStatus="paratype">
<specimenCode box="[759,946,1152,1179]" collectionCode="EPN" country="Ecuador" httpUri="http://grbio.org/cool/y3s2-2p0h" name="Escuela Polytecnica Nacional" pageId="2" pageNumber="61">
<accessionNumber box="[759,902,1152,1179]" httpUri="http://www.ncbi.nlm.nih.gov/protein/EPN12432" pageId="2" pageNumber="61">EPN 12432</accessionNumber>
33
</specimenCode>
,
<collectingMunicipality pageId="2" pageNumber="61">Los Encuentros</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03EA8795FFAF714466CAA87FFC1AF903:8E9C6058FFAF714F6683AEF5FE87FB30" box="[197,373,1187,1214]" country="Ecuador" latitude="-3.8077776" longLatPrecision="21" longitude="-78.605835" municipality="Los Encuentros" name="Concesion Cuy" pageId="2" pageNumber="61">Concesión Cuy</location>
(
<geoCoordinate box="[389,506,1188,1214]" degrees="3" direction="south" minutes="48" orientation="latitude" pageId="2" pageNumber="61" precision="15" seconds="28" value="-3.8077776">3°48'28&quot;S</geoCoordinate>
,
<geoCoordinate box="[518,660,1187,1213]" degrees="78" direction="west" minutes="36" orientation="longitude" pageId="2" pageNumber="61" precision="15" seconds="21" value="-78.605835">78°36'21&quot;W</geoCoordinate>
,
<quantity box="[672,764,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="61" unit="m" value="1450.0">
<elevation box="[672,764,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="61" unit="m" value="1450.0">1450 m</elevation>
</quantity>
), collected on
<date box="[930,1108,1187,1214]" pageId="2" pageNumber="61" value="2008-03-27">
<collectingDate box="[930,1108,1187,1214]" pageId="2" pageNumber="61" value="2008-03-27">27 March 2008</collectingDate>
</date>
by A.
<collectorName box="[207,341,1223,1249]" pageId="2" pageNumber="61">Almendáriz</collectorName>
, P.
<collectorName box="[378,474,1223,1249]" pageId="2" pageNumber="61">Vivanco</collectorName>
, C.
<collectorName box="[520,614,1223,1249]" pageId="2" pageNumber="61">Sarango</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="67" pageId="2" pageNumber="61" type="diagnosis">
<paragraph blockId="2.[140,1108,659,1672]" pageId="2" pageNumber="61">
<emphasis bold="true" box="[185,312,1257,1283]" pageId="2" pageNumber="61">
<emphasis box="[185,307,1257,1283]" inLineHeading="true" pageId="2" pageNumber="61" reason="1">Diagnosis</emphasis>
.
</emphasis>
The new species differs from all other species of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[896,1018,1258,1284]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis box="[896,1018,1258,1284]" italics="true" pageId="2" pageNumber="61">Enyalioides</emphasis>
</taxonomicName>
, except for
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[181,333,1293,1319]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[181,333,1293,1319]" italics="true" pageId="2" pageNumber="61">E. praestabilis</emphasis>
</taxonomicName>
, in having distinct caudal whorls, smooth or feebly keeled ventrals, fewer than 32 longitudinal rows of dorsals in a transverse line between dorsolateral crests at midbody, and in lacking projecting dorsal and limb scales. Males of the new species can be distinguished from
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[528,679,1399,1425]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[528,679,1399,1425]" italics="true" pageId="2" pageNumber="61">E. praestabilis</emphasis>
</taxonomicName>
by having larger scales on the ventral surface of the thighs (
<figureCitation box="[391,461,1434,1460]" captionStart="Figure 2" captionStartId="4.[140,212,1074,1096]" captionTargetBox="[140,1108,165,1054]" captionTargetId="figure@4.[140,1108,165,1055]" captionTargetPageId="4" captionText="Figure 2. Ventral view of thigh in two species of Enyalioides. Top: E. praestabilis (QCAZ 8821, adult male, SVL = 110 mm); bottom: E. rubrigularis sp. n. (QCAZ 8459, adult male, SVL = 122 mm). Photographs by O. Torres-Carvajal." figureDoi="http://doi.org/10.5281/zenodo.3769944" httpUri="https://zenodo.org/record/3769944/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
), and by having gulars with black margins (
<figureCitation box="[971,1040,1434,1460]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="2" pageNumber="61">Fig. 3</figureCitation>
). The skin between gulars is black in some male specimens of
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[775,926,1469,1495]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[775,926,1469,1495]" italics="true" pageId="2" pageNumber="61">E. praestabilis</emphasis>
</taxonomicName>
, but gulars lack black margins. In addition, gular scales in males of the new species vary between bright orange and red, and there is no black mark on the gular region posteromedially; males of
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[171,323,1575,1601]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[171,323,1575,1601]" italics="true" pageId="2" pageNumber="61">E. praestabilis</emphasis>
</taxonomicName>
have cream or yellow gular scales, and some specimens have a black patch covering the gular fold and posteromedial portion of the gular region (
<figureCitation box="[1023,1092,1610,1636]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="2" pageNumber="61">Fig. 3</figureCitation>
). The new species usually has two femoral pores, whereas
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[760,909,1645,1671]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[760,909,1645,1671]" italics="true" pageId="2" pageNumber="61">E. praestabilis</emphasis>
</taxonomicName>
has normally one
</paragraph>
<paragraph blockId="3.[140,1108,1622,1676]" pageId="3" pageNumber="62">
<emphasis bold="true" box="[140,237,1622,1644]" pageId="3" pageNumber="62">Figure |.</emphasis>
<typeStatus box="[241,329,1622,1644]" pageId="3" pageNumber="62">Holotype</typeStatus>
of
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[357,567,1622,1643]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis box="[357,567,1622,1643]" italics="true" pageId="3" pageNumber="62">Enyalioides rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[572,621,1623,1644]" pageId="3" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
(QCAZ 8483, adult male, SVL =
<quantity box="[933,1016,1623,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="62" unit="mm" value="123.0">123 mm</quantity>
). Top: lateral view; middle: close-up of head; bottom: ventral view. Photographs by O. Torres-Carvajal.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769944" ID-Zenodo-Dep="3769944" httpUri="https://zenodo.org/record/3769944/files/figure.png" pageId="4" pageNumber="63" startId="4.[140,212,1074,1096]" targetBox="[140,1108,165,1054]" targetPageId="4">
<paragraph blockId="4.[140,1108,1074,1160]" pageId="4" pageNumber="63">
<emphasis bold="true" box="[140,240,1074,1096]" pageId="4" pageNumber="63">Figure 2.</emphasis>
Ventral view of thigh in two species of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[621,721,1074,1095]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="genus">
<emphasis box="[621,721,1074,1095]" italics="true" pageId="4" pageNumber="63">Enyalioides</emphasis>
</taxonomicName>
. Top:
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[784,909,1074,1096]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[784,909,1074,1096]" italics="true" pageId="4" pageNumber="63">E. praestabilis</emphasis>
</taxonomicName>
(QCAZ 8821, adult male, SVL = 110 mm); bottom:
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[440,570,1106,1128]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis box="[440,570,1106,1128]" italics="true" pageId="4" pageNumber="63">E. rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[576,626,1107,1128]" pageId="4" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
(QCAZ 8459, adult male, SVL = 122 mm). Photographs by O. Torres-Carvajal.
</paragraph>
</caption>
<paragraph blockId="4.[140,1108,1223,1672]" pageId="4" pageNumber="63">femoral pore; otherwise, both species are very similar in scale counts and morphometric characters (Table 1).</paragraph>
<paragraph blockId="4.[140,1108,1223,1672]" lastBlockId="6.[140,1108,1469,1672]" lastPageId="6" lastPageNumber="65" pageId="4" pageNumber="63">
<emphasis box="[185,441,1293,1319]" italics="true" pageId="4" pageNumber="63">
Description of
<typeStatus box="[345,437,1293,1319]" pageId="4" pageNumber="63">holotype</typeStatus>
.
</emphasis>
Male (Fig. 1); SVL =
<quantity box="[696,798,1293,1320]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="4" pageNumber="63" unit="mm" value="123.0">123 mm</quantity>
; TL =
<quantity box="[877,978,1293,1320]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="63" unit="mm" value="180.0">180 mm</quantity>
; maximum head width =
<quantity box="[300,423,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="2.596" pageId="4" pageNumber="63" unit="mm" value="25.96">25.96 mm</quantity>
; head length =
<quantity box="[601,725,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="3.2369999999999997" pageId="4" pageNumber="63" unit="mm" value="32.37">32.37 mm</quantity>
; head height =
<quantity box="[903,1027,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="2.373" pageId="4" pageNumber="63" unit="mm" value="23.73">23.73 mm</quantity>
; dorsal head scales keeled or multicarinate, those on parietal region strongly projected dorsally; scales immediately posterior to supraciliares conical and dorsally projected, forming longitudinal row of seven (left) or six (right) scales that extends posteriorly over supratemporal region; temporal scales small, smooth or keeled, juxtaposed; two large, projected conical temporal scales aligned anterodorsally from anterodorsal aspect of tympanum; one enlarged pretympanic scale; supraciliares 16; canthals five; postrostrals four; left supralabials nine if counted to a point right below middle of eye, and 13 if counted to commisure of mouth (10 and 13 on right side, respectively); rostral (3.67 ×
<quantity box="[164,273,1645,1672]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="4" pageNumber="63" unit="mm" value="1.64">1.64 mm</quantity>
) about twice as wide as adjacent supralabials; single longitudinal row of lorilabials between suboculars and supralabials at level of middle of eye, longitudinal rows of lorilabials anterior to this point 23; loreal region broken into small, smooth, and juxtaposed scales; nasal at level of supralabial III; left infralabials eight if counted to a point right below middle of eye, and 13 if counted to commisure of mouth (eight and 11 on right side, respectively); mental (4.11 ×
<quantity box="[727,835,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="6" pageNumber="65" unit="mm" value="2.65">2.65 mm</quantity>
) wider and higher than adjacent infralabials; postmentals three; gulars ventrally projected; gular fold complete midventrally; neck with several longitudinal and oblique folds.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769946" ID-Zenodo-Dep="3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="5" pageNumber="64" startId="5.[140,212,1449,1471]" targetBox="[141,1108,165,1430]" targetPageId="5">
<paragraph blockId="5.[140,1108,1449,1536]" pageId="5" pageNumber="64">
<emphasis bold="true" box="[140,241,1449,1471]" pageId="5" pageNumber="64">Figure 3.</emphasis>
Two species of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[395,495,1450,1471]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis box="[395,495,1450,1471]" italics="true" pageId="5" pageNumber="64">Enyalioides</emphasis>
</taxonomicName>
. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[610,743,1482,1503]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis box="[610,743,1482,1503]" italics="true" pageId="5" pageNumber="64">E. rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[751,805,1482,1503]" pageId="5" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
; male of
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[898,1024,1482,1503]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[898,1024,1482,1503]" italics="true" pageId="5" pageNumber="64">E. praestabilis</emphasis>
</taxonomicName>
(H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H).
</paragraph>
</caption>
<paragraph blockId="6.[140,1108,166,284]" pageId="6" pageNumber="65">
<tableNote pageId="6" pageNumber="65" targetBox="[147,1097,306,1429]" targetPageId="6">
<emphasis bold="true" box="[140,227,166,188]" pageId="6" pageNumber="65">Table |.</emphasis>
Summary of morphological characters, measurements (mm), and color patterns of
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="praestabilis">
<emphasis italics="true" pageId="6" pageNumber="65">Enyalioides praestabilis</emphasis>
</taxonomicName>
and
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[288,421,198,220]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="rubrigularis" status="sp. nov.">
<emphasis box="[288,421,198,220]" italics="true" pageId="6" pageNumber="65">E. rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[430,503,198,219]" pageId="6" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
For each quantitative character, the
<emphasis box="[852,865,199,220]" italics="true" pageId="6" pageNumber="65">F</emphasis>
-value,
<emphasis box="[934,941,199,220]" italics="true" pageId="6" pageNumber="65">t</emphasis>
-value, and corresponding
<emphasis box="[232,245,231,252]" italics="true" pageId="6" pageNumber="65">P</emphasis>
-values are given. Range (first line), and mean ± standard deviation or mode (second line) are given for quantitative characters.
</tableNote>
</paragraph>
<paragraph pageId="6" pageNumber="65">
<table box="[147,1097,306,1429]" gridcols="7" gridrows="37" pageId="6" pageNumber="65">
<tr box="[147,1097,306,328]" gridrow="0" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[458,596,306,328]" gridcol="1" gridrow="0" pageId="6" pageNumber="65">
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[458,596,306,328]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="praestabilis">
<emphasis bold="true" box="[458,596,306,328]" italics="true" pageId="6" pageNumber="65">E. praestabilis</emphasis>
</taxonomicName>
</th>
<th box="[620,764,306,328]" gridcol="2" gridrow="0" pageId="6" pageNumber="65">
<taxonomicName authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[620,764,306,328]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="rubrigularis">
<emphasis bold="true" box="[620,764,306,328]" italics="true" pageId="6" pageNumber="65">E. rubrigularis</emphasis>
</taxonomicName>
</th>
</tr>
<tr box="[147,1097,335,357]" gridrow="1" pageId="6" pageNumber="65">
<th box="[147,404,335,357]" gridcol="0" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[147,244,335,357]" pageId="6" pageNumber="65">Character</emphasis>
</th>
<td box="[458,596,335,357]" gridcol="1" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[494,561,335,356]" pageId="6" pageNumber="65">N = 50</emphasis>
</td>
<td box="[620,764,335,357]" gridcol="2" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[659,725,335,356]" pageId="6" pageNumber="65">N = 16</emphasis>
</td>
<td box="[780,852,335,357]" gridcol="3" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[780,852,335,356]" pageId="6" pageNumber="65">F-value</emphasis>
</td>
<td box="[873,927,335,357]" gridcol="4" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[893,907,335,356]" pageId="6" pageNumber="65">P</emphasis>
</td>
<td box="[947,1014,335,357]" gridcol="5" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[947,1014,335,356]" pageId="6" pageNumber="65">t-value</emphasis>
</td>
<td box="[1032,1097,335,357]" gridcol="6" gridrow="1" pageId="6" pageNumber="65">
<emphasis bold="true" box="[1058,1072,335,356]" pageId="6" pageNumber="65">P</emphasis>
</td>
</tr>
<tr box="[147,1097,368,390]" gridrow="2" pageId="6" pageNumber="65">
<th box="[147,404,368,390]" gridcol="0" gridrow="2" pageId="6" pageNumber="65">Dorsals in transverse row</th>
<td box="[458,596,368,390]" gridcol="1" gridrow="2" pageId="6" pageNumber="65">2139</td>
<td box="[620,764,368,390]" gridcol="2" gridrow="2" pageId="6" pageNumber="65">2532</td>
<td box="[780,852,368,390]" gridcol="3" gridrow="2" pageId="6" pageNumber="65">2.249</td>
<td box="[873,927,368,390]" gridcol="4" gridrow="2" pageId="6" pageNumber="65">0.115</td>
<td box="[947,1014,368,390]" gridcol="5" gridrow="2" pageId="6" pageNumber="65">0.369</td>
<td box="[1032,1097,368,390]" gridcol="6" gridrow="2" pageId="6" pageNumber="65">0.714</td>
</tr>
<tr box="[147,1097,397,419]" gridrow="3" pageId="6" pageNumber="65" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[147,404,397,419]" gridcol="0" gridrow="3" pageId="6" pageNumber="65">between</th>
<td box="[458,596,397,419]" gridcol="1" gridrow="3" pageId="6" pageNumber="65">28.37 ± 3.91</td>
<td box="[620,764,397,419]" gridcol="2" gridrow="3" pageId="6" pageNumber="65">28.79 ± 2.61</td>
</tr>
<tr box="[147,1097,426,448]" gridrow="4" pageId="6" pageNumber="65">
<th box="[147,1097,426,448]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="6" pageNumber="65">dorsolateral crests at midbody</th>
</tr>
<tr box="[147,1097,458,480]" gridrow="5" pageId="6" pageNumber="65">
<th box="[147,404,458,480]" gridcol="0" gridrow="5" pageId="6" pageNumber="65">Ventrals in transverse row</th>
<td box="[458,596,458,480]" gridcol="1" gridrow="5" pageId="6" pageNumber="65">2139</td>
<td box="[620,764,458,480]" gridcol="2" gridrow="5" pageId="6" pageNumber="65">2231</td>
<td box="[780,852,458,480]" gridcol="3" gridrow="5" pageId="6" pageNumber="65">2.665</td>
<td box="[873,927,458,480]" gridcol="4" gridrow="5" pageId="6" pageNumber="65">0.059</td>
<td box="[947,1014,458,480]" gridcol="5" gridrow="5" pageId="6" pageNumber="65">2.381</td>
<td box="[1032,1097,458,480]" gridcol="6" gridrow="5" pageId="6" pageNumber="65">0.021</td>
</tr>
<tr box="[147,1097,487,509]" gridrow="6" pageId="6" pageNumber="65" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[147,404,487,509]" gridcol="0" gridrow="6" pageId="6" pageNumber="65">at midbody</th>
<td box="[458,596,487,509]" gridcol="1" gridrow="6" pageId="6" pageNumber="65">28.69 ± 3.87</td>
<td box="[620,764,487,509]" gridcol="2" gridrow="6" pageId="6" pageNumber="65">26.07 ± 2.37</td>
</tr>
<tr box="[147,1097,519,542]" gridrow="7" pageId="6" pageNumber="65">
<th box="[147,404,519,542]" gridcol="0" gridrow="7" pageId="6" pageNumber="65">Vertebrals from occiput to</th>
<td box="[458,596,519,542]" gridcol="1" gridrow="7" pageId="6" pageNumber="65">4466</td>
<td box="[620,764,519,542]" gridcol="2" gridrow="7" pageId="6" pageNumber="65">4958</td>
<td box="[780,852,519,542]" gridcol="3" gridrow="7" pageId="6" pageNumber="65">4.243</td>
<td box="[873,927,519,542]" gridcol="4" gridrow="7" pageId="6" pageNumber="65">0.007</td>
<td box="[947,1014,519,542]" gridcol="5" gridrow="7" pageId="6" pageNumber="65">1.224</td>
<td box="[1032,1097,519,542]" gridcol="6" gridrow="7" pageId="6" pageNumber="65">0.226</td>
</tr>
<tr box="[147,1097,548,570]" gridrow="8" pageId="6" pageNumber="65" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[147,404,548,570]" gridcol="0" gridrow="8" pageId="6" pageNumber="65">base of tail</th>
<td box="[458,596,548,570]" gridcol="1" gridrow="8" pageId="6" pageNumber="65">54.64 ± 5.43</td>
<td box="[620,764,548,570]" gridcol="2" gridrow="8" pageId="6" pageNumber="65">52.79 ± 2.64</td>
</tr>
<tr box="[147,1097,581,603]" gridrow="9" pageId="6" pageNumber="65">
<th box="[147,404,581,603]" gridcol="0" gridrow="9" pageId="6" pageNumber="65">Gulars</th>
<td box="[458,596,581,603]" gridcol="1" gridrow="9" pageId="6" pageNumber="65">2234</td>
<td box="[620,764,581,603]" gridcol="2" gridrow="9" pageId="6" pageNumber="65">2634</td>
<td box="[780,852,581,603]" gridcol="3" gridrow="9" pageId="6" pageNumber="65">1.274</td>
<td box="[873,927,581,603]" gridcol="4" gridrow="9" pageId="6" pageNumber="65">0.657</td>
<td box="[947,1014,581,603]" gridcol="5" gridrow="9" pageId="6" pageNumber="65">1.776</td>
<td box="[1032,1097,581,603]" gridcol="6" gridrow="9" pageId="6" pageNumber="65">0.081</td>
</tr>
<tr box="[147,1097,610,632]" gridrow="10" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,610,632]" gridcol="1" gridrow="10" pageId="6" pageNumber="65">27.91 ± 2.99</td>
<td box="[620,764,610,632]" gridcol="2" gridrow="10" pageId="6" pageNumber="65">29.50 ± 2.65</td>
</tr>
<tr box="[147,1097,642,664]" gridrow="11" pageId="6" pageNumber="65">
<th box="[147,404,642,664]" gridcol="0" gridrow="11" pageId="6" pageNumber="65">Infralabials</th>
<td box="[458,596,642,664]" gridcol="1" gridrow="11" pageId="6" pageNumber="65">711</td>
<td box="[620,764,642,664]" gridcol="2" gridrow="11" pageId="6" pageNumber="65">710</td>
<td box="[780,852,642,664]" gridcol="3" gridrow="11" pageId="6" pageNumber="65">1.755</td>
<td box="[873,927,642,664]" gridcol="4" gridrow="11" pageId="6" pageNumber="65">0.271</td>
<td box="[947,1014,642,664]" gridcol="5" gridrow="11" pageId="6" pageNumber="65">0.025</td>
<td box="[1032,1097,642,664]" gridcol="6" gridrow="11" pageId="6" pageNumber="65">0.979</td>
</tr>
<tr box="[147,1097,671,693]" gridrow="12" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,671,693]" gridcol="1" gridrow="12" pageId="6" pageNumber="65">8.86 ± 0.88</td>
<td box="[620,764,671,693]" gridcol="2" gridrow="12" pageId="6" pageNumber="65">8.86 ± 0.66</td>
</tr>
<tr box="[147,1097,703,726]" gridrow="13" pageId="6" pageNumber="65">
<th box="[147,404,703,726]" gridcol="0" gridrow="13" pageId="6" pageNumber="65">Supralabials</th>
<td box="[458,596,703,726]" gridcol="1" gridrow="13" pageId="6" pageNumber="65">812</td>
<td box="[620,764,703,726]" gridcol="2" gridrow="13" pageId="6" pageNumber="65">911</td>
<td box="[780,852,703,726]" gridcol="3" gridrow="13" pageId="6" pageNumber="65">4.060</td>
<td box="[873,927,703,726]" gridcol="4" gridrow="13" pageId="6" pageNumber="65">0.008</td>
<td box="[947,1014,703,726]" gridcol="5" gridrow="13" pageId="6" pageNumber="65">0.659</td>
<td box="[1032,1097,703,726]" gridcol="6" gridrow="13" pageId="6" pageNumber="65">0.512</td>
</tr>
<tr box="[147,1097,732,754]" gridrow="14" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,732,754]" gridcol="1" gridrow="14" pageId="6" pageNumber="65">10.34 ± 1.08</td>
<td box="[620,764,732,754]" gridcol="2" gridrow="14" pageId="6" pageNumber="65">10.14 ± 0.53</td>
</tr>
<tr box="[147,1097,765,787]" gridrow="15" pageId="6" pageNumber="65">
<th box="[147,404,765,787]" gridcol="0" gridrow="15" pageId="6" pageNumber="65">Canthals</th>
<td box="[458,596,765,787]" gridcol="1" gridrow="15" pageId="6" pageNumber="65">46</td>
<td box="[620,764,765,787]" gridcol="2" gridrow="15" pageId="6" pageNumber="65">46</td>
<td box="[780,852,765,787]" gridcol="3" gridrow="15" pageId="6" pageNumber="65">1.097</td>
<td box="[873,927,765,787]" gridcol="4" gridrow="15" pageId="6" pageNumber="65">0.895</td>
<td box="[947,1014,765,787]" gridcol="5" gridrow="15" pageId="6" pageNumber="65">1.531</td>
<td box="[1032,1097,765,787]" gridcol="6" gridrow="15" pageId="6" pageNumber="65">0.133</td>
</tr>
<tr box="[147,1097,794,816]" gridrow="16" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,794,816]" gridcol="1" gridrow="16" pageId="6" pageNumber="65">5</td>
<td box="[620,764,794,816]" gridcol="2" gridrow="16" pageId="6" pageNumber="65">5</td>
</tr>
<tr box="[147,1097,826,848]" gridrow="17" pageId="6" pageNumber="65">
<th box="[147,404,826,848]" gridcol="0" gridrow="17" pageId="6" pageNumber="65">Superciliaries</th>
<td box="[458,596,826,848]" gridcol="1" gridrow="17" pageId="6" pageNumber="65">1220</td>
<td box="[620,764,826,848]" gridcol="2" gridrow="17" pageId="6" pageNumber="65">1319</td>
<td box="[780,852,826,848]" gridcol="3" gridrow="17" pageId="6" pageNumber="65">1.794</td>
<td box="[873,927,826,848]" gridcol="4" gridrow="17" pageId="6" pageNumber="65">0.263</td>
<td box="[947,1014,826,848]" gridcol="5" gridrow="17" pageId="6" pageNumber="65">0.144</td>
<td box="[1032,1097,826,848]" gridcol="6" gridrow="17" pageId="6" pageNumber="65">0.886</td>
</tr>
<tr box="[147,1097,855,877]" gridrow="18" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,855,877]" gridcol="1" gridrow="18" pageId="6" pageNumber="65">15.56 ± 1.87</td>
<td box="[620,764,855,877]" gridcol="2" gridrow="18" pageId="6" pageNumber="65">15.64 ± 1.39</td>
</tr>
<tr box="[147,1097,887,910]" gridrow="19" pageId="6" pageNumber="65">
<th box="[147,404,887,910]" gridcol="0" gridrow="19" pageId="6" pageNumber="65">Transverse rows of ventrals</th>
<td box="[458,596,887,910]" gridcol="1" gridrow="19" pageId="6" pageNumber="65">3148</td>
<td box="[620,764,887,910]" gridcol="2" gridrow="19" pageId="6" pageNumber="65">3140</td>
<td box="[780,852,887,910]" gridcol="3" gridrow="19" pageId="6" pageNumber="65">1.604</td>
<td box="[873,927,887,910]" gridcol="4" gridrow="19" pageId="6" pageNumber="65">0.359</td>
<td box="[947,1014,887,910]" gridcol="5" gridrow="19" pageId="6" pageNumber="65">1.213</td>
<td box="[1032,1097,887,910]" gridcol="6" gridrow="19" pageId="6" pageNumber="65">0.230</td>
</tr>
<tr box="[147,1097,916,938]" gridrow="20" pageId="6" pageNumber="65" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[147,404,916,938]" gridcol="0" gridrow="20" pageId="6" pageNumber="65">between fore and hind limb</th>
<td box="[458,596,916,938]" gridcol="1" gridrow="20" pageId="6" pageNumber="65">37.77 ± 3.36</td>
<td box="[620,764,916,938]" gridcol="2" gridrow="20" pageId="6" pageNumber="65">36.57 ± 2.65</td>
</tr>
<tr box="[147,1097,949,971]" gridrow="21" pageId="6" pageNumber="65">
<th box="[147,404,949,971]" gridcol="0" gridrow="21" pageId="6" pageNumber="65">Subdigitals Finger IV</th>
<td box="[458,596,949,971]" gridcol="1" gridrow="21" pageId="6" pageNumber="65">1724</td>
<td box="[620,764,949,971]" gridcol="2" gridrow="21" pageId="6" pageNumber="65">1722</td>
<td box="[780,852,949,971]" gridcol="3" gridrow="21" pageId="6" pageNumber="65">1.036</td>
<td box="[873,927,949,971]" gridcol="4" gridrow="21" pageId="6" pageNumber="65">0.998</td>
<td box="[947,1014,949,971]" gridcol="5" gridrow="21" pageId="6" pageNumber="65">0.761</td>
<td box="[1032,1097,949,971]" gridcol="6" gridrow="21" pageId="6" pageNumber="65">0.450</td>
</tr>
<tr box="[147,1097,978,1000]" gridrow="22" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,978,1000]" gridcol="1" gridrow="22" pageId="6" pageNumber="65">19.82 ± 1.32</td>
<td box="[620,764,978,1000]" gridcol="2" gridrow="22" pageId="6" pageNumber="65">19.50 ± 1.34</td>
</tr>
<tr box="[147,1097,1010,1032]" gridrow="23" pageId="6" pageNumber="65">
<th box="[147,404,1010,1032]" gridcol="0" gridrow="23" pageId="6" pageNumber="65">Subdigitals Toe IV</th>
<td box="[458,596,1010,1032]" gridcol="1" gridrow="23" pageId="6" pageNumber="65">2228</td>
<td box="[620,764,1010,1032]" gridcol="2" gridrow="23" pageId="6" pageNumber="65">2329</td>
<td box="[780,852,1010,1032]" gridcol="3" gridrow="23" pageId="6" pageNumber="65">1.586</td>
<td box="[873,927,1010,1032]" gridcol="4" gridrow="23" pageId="6" pageNumber="65">0.254</td>
<td box="[947,1014,1010,1032]" gridcol="5" gridrow="23" pageId="6" pageNumber="65">0.252</td>
<td box="[1032,1097,1010,1032]" gridcol="6" gridrow="23" pageId="6" pageNumber="65">0.802</td>
</tr>
<tr box="[147,1097,1039,1061]" gridrow="24" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1039,1061]" gridcol="1" gridrow="24" pageId="6" pageNumber="65">25.84 ± 1.54</td>
<td box="[620,764,1039,1061]" gridcol="2" gridrow="24" pageId="6" pageNumber="65">25.71 ± 1.94</td>
</tr>
<tr box="[147,1097,1071,1093]" gridrow="25" pageId="6" pageNumber="65">
<th box="[147,404,1071,1093]" gridcol="0" gridrow="25" pageId="6" pageNumber="65">Femoral pores</th>
<td box="[458,596,1071,1093]" gridcol="1" gridrow="25" pageId="6" pageNumber="65">12</td>
<td box="[620,764,1071,1093]" gridcol="2" gridrow="25" pageId="6" pageNumber="65">12</td>
<td box="[780,852,1071,1093]" gridcol="3" gridrow="25" pageId="6" pageNumber="65">1.262</td>
<td box="[873,927,1071,1093]" gridcol="4" gridrow="25" pageId="6" pageNumber="65">0.420</td>
<td box="[947,1014,1071,1093]" gridcol="5" gridrow="25" pageId="6" pageNumber="65">2.947</td>
<td box="[1032,1097,1071,1093]" gridcol="6" gridrow="25" pageId="6" pageNumber="65">&lt;0.005</td>
</tr>
<tr box="[147,1097,1100,1121]" gridrow="26" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1100,1121]" gridcol="1" gridrow="26" pageId="6" pageNumber="65">1</td>
<td box="[620,764,1100,1121]" gridcol="2" gridrow="26" pageId="6" pageNumber="65">2</td>
</tr>
<tr box="[147,1097,1133,1155]" gridrow="27" pageId="6" pageNumber="65">
<th box="[147,404,1133,1155]" gridcol="0" gridrow="27" pageId="6" pageNumber="65">Head length/Head width</th>
<td box="[458,596,1133,1155]" gridcol="1" gridrow="27" pageId="6" pageNumber="65">1.201.45</td>
<td box="[620,764,1133,1155]" gridcol="2" gridrow="27" pageId="6" pageNumber="65">1.201.39</td>
<td box="[780,852,1133,1155]" gridcol="3" gridrow="27" pageId="6" pageNumber="65">1.146</td>
<td box="[873,927,1133,1155]" gridcol="4" gridrow="27" pageId="6" pageNumber="65">0.827</td>
<td box="[947,1014,1133,1155]" gridcol="5" gridrow="27" pageId="6" pageNumber="65">1.819</td>
<td box="[1032,1097,1133,1155]" gridcol="6" gridrow="27" pageId="6" pageNumber="65">0.074</td>
</tr>
<tr box="[147,1097,1161,1183]" gridrow="28" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1161,1183]" gridcol="1" gridrow="28" pageId="6" pageNumber="65">1.32 ± 0.07</td>
<td box="[620,764,1161,1183]" gridcol="2" gridrow="28" pageId="6" pageNumber="65">1.28 ± 0.05</td>
</tr>
<tr box="[147,1097,1194,1216]" gridrow="29" pageId="6" pageNumber="65">
<th box="[147,404,1194,1216]" gridcol="0" gridrow="29" pageId="6" pageNumber="65">Fore limb length/SVL</th>
<td box="[458,596,1194,1216]" gridcol="1" gridrow="29" pageId="6" pageNumber="65">0.470.61</td>
<td box="[620,764,1194,1216]" gridcol="2" gridrow="29" pageId="6" pageNumber="65">0.380.54</td>
<td box="[780,852,1194,1216]" gridcol="3" gridrow="29" pageId="6" pageNumber="65">2.897</td>
<td box="[873,927,1194,1216]" gridcol="4" gridrow="29" pageId="6" pageNumber="65">0.010</td>
<td box="[947,1014,1194,1216]" gridcol="5" gridrow="29" pageId="6" pageNumber="65">2.931</td>
<td box="[1032,1097,1194,1216]" gridcol="6" gridrow="29" pageId="6" pageNumber="65">0.005</td>
</tr>
<tr box="[147,1097,1223,1245]" gridrow="30" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1223,1245]" gridcol="1" gridrow="30" pageId="6" pageNumber="65">0.54 ± 0.03</td>
<td box="[620,764,1223,1245]" gridcol="2" gridrow="30" pageId="6" pageNumber="65">0.49 ± 0.04</td>
</tr>
<tr box="[147,1097,1255,1277]" gridrow="31" pageId="6" pageNumber="65">
<th box="[147,404,1255,1277]" gridcol="0" gridrow="31" pageId="6" pageNumber="65">Hind limb length/SVL</th>
<td box="[458,596,1255,1277]" gridcol="1" gridrow="31" pageId="6" pageNumber="65">0.720.95</td>
<td box="[620,764,1255,1277]" gridcol="2" gridrow="31" pageId="6" pageNumber="65">0.720.86</td>
<td box="[780,852,1255,1277]" gridcol="3" gridrow="31" pageId="6" pageNumber="65">1.888</td>
<td box="[873,927,1255,1277]" gridcol="4" gridrow="31" pageId="6" pageNumber="65">0.217</td>
<td box="[947,1014,1255,1277]" gridcol="5" gridrow="31" pageId="6" pageNumber="65">0.951</td>
<td box="[1032,1097,1255,1277]" gridcol="6" gridrow="31" pageId="6" pageNumber="65">0.346</td>
</tr>
<tr box="[147,1097,1284,1306]" gridrow="32" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1284,1306]" gridcol="1" gridrow="32" pageId="6" pageNumber="65">0.82 ± 0.05</td>
<td box="[620,764,1284,1306]" gridcol="2" gridrow="32" pageId="6" pageNumber="65">0.79 ± 0.05</td>
</tr>
<tr box="[147,1097,1317,1339]" gridrow="33" pageId="6" pageNumber="65">
<th box="[147,404,1317,1339]" gridcol="0" gridrow="33" pageId="6" pageNumber="65">Tail length/Total length</th>
<td box="[458,596,1317,1339]" gridcol="1" gridrow="33" pageId="6" pageNumber="65">0.590.66</td>
<td box="[620,764,1317,1339]" gridcol="2" gridrow="33" pageId="6" pageNumber="65">0.590.62</td>
<td box="[780,852,1317,1339]" gridcol="3" gridrow="33" pageId="6" pageNumber="65">2.812</td>
<td box="[873,927,1317,1339]" gridcol="4" gridrow="33" pageId="6" pageNumber="65">0.086</td>
<td box="[947,1014,1317,1339]" gridcol="5" gridrow="33" pageId="6" pageNumber="65">2.806</td>
<td box="[1032,1097,1317,1339]" gridcol="6" gridrow="33" pageId="6" pageNumber="65">0.007</td>
</tr>
<tr box="[147,1097,1346,1368]" gridrow="34" pageId="6" pageNumber="65" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td box="[458,596,1346,1368]" gridcol="1" gridrow="34" pageId="6" pageNumber="65">0.62 ± 0.02</td>
<td box="[620,764,1346,1368]" gridcol="2" gridrow="34" pageId="6" pageNumber="65">0.60 ± 0.01</td>
</tr>
<tr box="[147,1097,1378,1400]" gridrow="35" pageId="6" pageNumber="65" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[147,404,1378,1400]" gridcol="0" gridrow="35" pageId="6" pageNumber="65">Dark gular patch</th>
<td box="[458,596,1378,1400]" gridcol="1" gridrow="35" pageId="6" pageNumber="65">Present</td>
<td box="[620,764,1378,1400]" gridcol="2" gridrow="35" pageId="6" pageNumber="65">absent</td>
</tr>
<tr box="[147,1097,1407,1429]" gridrow="36" pageId="6" pageNumber="65">
<th box="[147,1097,1407,1429]" colspan="7" colspanRight="6" gridcol="0" gridrow="36" pageId="6" pageNumber="65">(polymorphic)</th>
</tr>
</table>
</paragraph>
<paragraph blockId="6.[140,1108,1469,1672]" lastBlockId="7.[140,1108,165,1636]" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">Vertebral crest strongly projected and decreasing in size posteriorly, with vertebrals on neck at least four times higher than vertebrals between hind limbs; crest bifurcates posteriorly and extends onto tail about ¼ its length; body flanks between fore and hind limbs with dorsolateral and ventrolateral folds, as well as several oblique folds; scales on dorsolateral folds slightly larger than adjacent scales giving the fold the appearance of a crest; dorsal scales between dorsolateral folds and vertebral crest small, prominently keeled, and imbricate; scales on flanks (i.e., ventral to dorsolateral folds) similar in size to dorsal scales, with a few scattered enlarged scales 23 times larger than adjacent scales; ventral scales imbricate, smooth, rectangular, with a posterolateral mucron; ventrals more than twice the size of dorsals.</paragraph>
<paragraph blockId="7.[140,1108,165,1636]" pageId="7" pageNumber="66">Limb scales keeled dorsally and smooth or slightly keeled ventrally; scales on dorsal and posterior aspects of thighs heterogeneous in size, with most scales less than half the size of those scales on anterior and ventral aspects; subdigitals on Finger IV 21; subdigitals on Toe IV 26; two femoral pores on each side; tail laterally compressed and gradually decreasing in height towards tip; caudal scales strongly keeled and imbricate, increasing in size posteriorly on lateral and dorsal aspects of each autotomic segment; ventral larger than dorsal caudals, with individual autotomic segments being three scales long ventrally and six scales long dorsally.</paragraph>
<paragraph blockId="7.[140,1108,165,1636]" pageId="7" pageNumber="66">
<emphasis bold="true" box="[185,548,729,756]" pageId="7" pageNumber="66">
<emphasis box="[185,317,729,755]" inLineHeading="true" pageId="7" pageNumber="66" reason="1">Coloration</emphasis>
in life of
<typeStatus box="[442,548,730,756]" pageId="7" pageNumber="66">holotype</typeStatus>
</emphasis>
(Fig. 1). Scales on dorsal and lateral surfaces of head mostly green or yellow, with black margins causing a reticulate pattern; some head scales entirely black; labials and mental yellowish orange with black margins; rostral green medially and black laterally; gulars orange with black margins, a few lateral gulars green or yellow; skin between gulars black; black gular patch absent; six enlarged greenish-cream scales form a distinct spot posterior to tympanum; paraverterbrals and caudals green or dark brown; flanks mottled with lemon-green scales; dorsal limb scales light green, some with dark brown margins; ventral surface of body, limbs, and tail white medially and light green laterally, with irregular light green stripes projecting medially; lining of mouth whitish cream; iris light brown peripherially with dark brown projections originating from the dark brown center.
</paragraph>
<paragraph blockId="7.[140,1108,165,1636]" pageId="7" pageNumber="66">
<emphasis bold="true" box="[185,379,1117,1143]" pageId="7" pageNumber="66">
<emphasis box="[185,255,1117,1143]" inLineHeading="true" pageId="7" pageNumber="66" reason="1">Color</emphasis>
variation.
</emphasis>
Adult males QCAZ 8456, 8460 differed from the
<typeStatus box="[976,1075,1117,1143]" pageId="7" pageNumber="66">holotype</typeStatus>
in having a dark brown to black reticulate pattern on dorsal and lateral aspects of body. In addition, male QCAZ 8460 had the dorsal surface of head black with green and yellow dots; loreal and subocular regions with black and red scales; labials red with black margins; gulars red with black margins (
<figureCitation box="[541,608,1258,1284]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="7" pageNumber="66">Fig. 3</figureCitation>
). Metachromatism was observed in some individuals, in which the green tones were replaced with yellow or brown tones (
<figureCitation box="[1025,1093,1293,1319]" captionStart="Figure 4" captionStartId="8.[140,212,758,780]" captionTargetBox="[140,1108,165,738]" captionTargetId="figure@8.[140,1108,165,739]" captionTargetPageId="8" captionText="Figure 4. Metachromatism in Enyalioides rubrigularis sp. n. Top left and right: adult male (QCAZ 8460, SVL = 102); bottom left and right: adult male (QCAZ 8456, SVL = 89). Photographs by O. Torres- Carvajal (left) and L. Coloma (right)." figureDoi="http://doi.org/10.5281/zenodo.3769948" httpUri="https://zenodo.org/record/3769948/files/figure.png" pageId="7" pageNumber="66">Fig. 4</figureCitation>
).
</paragraph>
<paragraph blockId="7.[140,1108,165,1636]" lastBlockId="8.[140,1108,906,1636]" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
Adult female QCAZ 8457 (
<figureCitation box="[513,584,1328,1354]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="7" pageNumber="66">Fig. 3</figureCitation>
): dorsal background of body, limbs, and tail light brown spotted with yellow scales; dorsal background of head dark brown with scattered black scales; labials yellowish green with grey margins; enlarged pretympanic scale yellow; faint yellowish strip extends longitudinally from tympanum to scapular region; each vertebral yellow anteriorly and grey posteriorly; gular region brown with a few scattered orange scales; ventral surface of body, limbs, and tail brownish cream; iris copper with dark brown reticulations. Adult female QCAZ 8458 (
<figureCitation box="[937,1004,1539,1566]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="7" pageNumber="66">Fig. 3</figureCitation>
) differed from the latter in having a dark olive green dorsal background, a black stripe extending longitudinally from posterior margin of eye to dorsal margin of tympanum, a black stripe extending dorsolaterally from comisure of mouth to eye, and a yellowish cream gular region with scattered black dots.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769948" ID-Zenodo-Dep="3769948" httpUri="https://zenodo.org/record/3769948/files/figure.png" pageId="8" pageNumber="67" startId="8.[140,212,758,780]" targetBox="[140,1108,165,738]" targetPageId="8">
<paragraph blockId="8.[140,1108,758,844]" pageId="8" pageNumber="67">
<emphasis bold="true" box="[140,237,758,780]" pageId="8" pageNumber="67">Figure 4.</emphasis>
Metachromatism in
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[430,640,758,779]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis box="[430,640,758,779]" italics="true" pageId="8" pageNumber="67">Enyalioides rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[645,695,758,779]" pageId="8" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
Top left and right: adult male (QCAZ 8460, SVL = 102); bottom left and right: adult male (QCAZ 8456, SVL = 89). Photographs by O. Torres- Carvajal (left) and L. Coloma (right).
</paragraph>
</caption>
<paragraph blockId="8.[140,1108,906,1636]" pageId="8" pageNumber="67">
Juvenile QCAZ 8454 (
<figureCitation box="[451,521,976,1002]" captionStart="Figure 3" captionStartId="5.[140,212,1449,1471]" captionTargetBox="[141,1108,165,1430]" captionTargetId="figure@5.[140,1108,165,1430]" captionTargetPageId="5" captionText="Figure 3. Two species of Enyalioides. Juvenile (A, B, QCAZ 8454), females (C, D, QCAZ 8457; E, F, QCAZ 8458), and male (G, QCAZ 8460) of E. rubrigularis sp. n.; male of E. praestabilis (H, KU 169854). Photographs by O. Torres-Carvajal (A-G) and W. E. Duellman (H)." figureDoi="http://doi.org/10.5281/zenodo.3769946" httpUri="https://zenodo.org/record/3769946/files/figure.png" pageId="8" pageNumber="67">Fig. 3</figureCitation>
): dorsal background yellowish green with irregular dark brown marks on head, body, flanks, tail, and black transverse lines on limbs; black stripe extends from posterior margin of eye to dorsal margin of tympanum; another black stripe extends from subocular region to anterior margin of tympanum, from where it extends ventrally into gular region; dorsolateral light green band with brown and black margins extends from temporal to scapular region; vertebrals black and yellowish green; gular region yellowish cream with brown, yellow, and orange scales laterally; ventral surface of body and limbs whitish cream, with faint grey dots; ventral surface of tail similar in color but fainter than dorsal surface; tongue cream with anterior tip grey; lining of mouth whitish cream; iris copper. Juvenile QCAZ 8485 differs from the latter in having a yellowish-brown background, with dark brown transverse bands arranged longitudinally from the scapular region to the tip of the tail.
</paragraph>
<paragraph blockId="8.[140,1108,906,1636]" pageId="8" pageNumber="67">
<emphasis bold="true" box="[185,376,1398,1425]" pageId="8" pageNumber="67">
<emphasis box="[185,277,1399,1425]" inLineHeading="true" pageId="8" pageNumber="67" reason="1">Natural</emphasis>
history.
</emphasis>
Juvenile QCAZ 8454 was encountered at 5 pm basking in the sun on the ground; when approached it ran quickly into a hole
<quantity box="[874,933,1434,1461]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="67" unit="cm" value="5.0">5 cm</quantity>
wide located
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="67" unit="m" value="1.0">1 m</quantity>
higher in the ground. All other specimens were found sleeping at night (7:00 pm 12:00 am) between
<quantity box="[363,435,1504,1531]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="67" unit="cm" value="30.0">30 cm</quantity>
and
<quantity box="[490,556,1504,1531]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="67" unit="m" value="2.5">2.5 m</quantity>
above ground. These specimens were found with their heads facing up on vertical stems with diameters varying between
<quantity box="[964,1070,1540,1566]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="8" pageNumber="67" unit="cm" value="6.0" valueMax="10.0" valueMin="2.0">210 cm</quantity>
, or lying horizontally on ferns with stems
<quantity box="[571,657,1575,1601]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="67" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">23 cm</quantity>
wide. Most specimens were collected in secondary forest close to pasture.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="68" type="distribution">
<paragraph blockId="9.[140,1108,166,475]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[185,344,166,192]" pageId="9" pageNumber="68">
<emphasis box="[185,339,166,192]" inLineHeading="true" pageId="9" pageNumber="68" reason="1">Distribution</emphasis>
.
</emphasis>
<taxonomicName authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[352,610,167,193]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="rubrigularis">
<emphasis box="[352,610,167,193]" italics="true" pageId="9" pageNumber="68">Enyalioides rubrigularis</emphasis>
</taxonomicName>
inhabits rainforests on the eastern slopes of the Andes and western slopes of Cordillera del Cóndor in southern
<collectingCountry box="[892,986,202,228]" name="Ecuador" pageId="9" pageNumber="68">Ecuador</collectingCountry>
(Fig. 5). It occurs at elevations of
<quantity box="[389,549,237,264]" metricMagnitude="3" metricUnit="m" metricValue="1.28" metricValueMax="1.46" metricValueMin="1.1" pageId="9" pageNumber="68" unit="m" value="1280.0" valueMax="1460.0" valueMin="1100.0">11001460 m</quantity>
in the upper basins of the Zamora and Nangaritza rivers, Provincia
<collectingRegion box="[328,548,272,299]" country="Ecuador" name="Zamora-Chinchipe" pageId="9" pageNumber="68">Zamora Chinchipe</collectingRegion>
. Th e
<typeStatus box="[610,658,273,299]" pageId="9" pageNumber="68">type</typeStatus>
locality of
<taxonomicName authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[786,945,272,298]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="rubrigularis">
<emphasis box="[786,945,272,298]" italics="true" pageId="9" pageNumber="68">E. rubrigularis</emphasis>
</taxonomicName>
is surrounded by secondary forest, pasture,
<taxonomicName authorityName="Smith in Smith" authorityYear="1840" box="[466,546,307,333]" class="Actinopterygii" family="Cichlidae" genus="Tilapia" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis box="[466,546,307,333]" italics="true" pageId="9" pageNumber="68">Tilapia</emphasis>
</taxonomicName>
ponds, and bullfrog (
<taxonomicName authorityName="Shaw" authorityYear="1802" box="[792,978,307,333]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="catesbeiana">
<emphasis box="[792,978,307,333]" italics="true" pageId="9" pageNumber="68">Rana catesbeiana</emphasis>
</taxonomicName>
) farms.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="68" type="etymology">
<paragraph blockId="9.[140,1108,166,475]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[185,320,343,369]" pageId="9" pageNumber="68">
<emphasis box="[185,315,343,369]" inLineHeading="true" pageId="9" pageNumber="68" reason="1">Etymology</emphasis>
.
</emphasis>
The name
<taxonomicName authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[450,578,343,369]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="rubrigularis">
<emphasis box="[450,578,343,369]" italics="true" pageId="9" pageNumber="68">rubrigularis</emphasis>
</taxonomicName>
is an adjective in the nominative singular and derives from the Latin words
<emphasis box="[481,540,378,404]" italics="true" pageId="9" pageNumber="68">ruber</emphasis>
(=red) and
<emphasis box="[678,724,378,404]" italics="true" pageId="9" pageNumber="68">gula</emphasis>
(=throat). It refers to the characteristic orange or reddish throat and chin of adult males, which distinguishes the new species from other species of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[467,589,448,474]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis box="[467,589,448,474]" italics="true" pageId="9" pageNumber="68">Enyalioides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="9.[230,962,516,536]" box="[230,962,516,536]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[230,962,516,536]" pageId="9" pageNumber="68">8ı° 80° 79° 78° 77° 76° 75°</emphasis>
</paragraph>
<paragraph blockId="9.[165,197,588,605]" box="[165,197,588,605]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[165,197,588,605]" pageId="9" pageNumber="68">ı°N</emphasis>
</paragraph>
<paragraph blockId="9.[180,196,705,722]" box="[180,196,705,722]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[180,196,705,722]" pageId="9" pageNumber="68"></emphasis>
</paragraph>
<paragraph blockId="9.[171,197,822,839]" box="[171,197,822,839]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,822,839]" pageId="9" pageNumber="68">ı°S</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,941,958]" box="[171,197,941,958]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,941,958]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,941,958]" degrees="2" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-2.0">2°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,1058,1075]" box="[171,197,1058,1075]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,1058,1075]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,1058,1075]" degrees="3" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-3.0">3°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,1176,1193]" box="[171,197,1176,1193]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,1176,1193]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,1176,1193]" degrees="4" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-4.0">4°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,1280,1297]" box="[171,197,1280,1297]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,1280,1297]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,1280,1297]" degrees="5" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-5.0">5°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,1405,1422]" box="[171,197,1405,1422]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,1405,1422]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,1405,1422]" degrees="6" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-6.0">6°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[171,197,1517,1534]" box="[171,197,1517,1534]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[171,197,1517,1534]" pageId="9" pageNumber="68">
<geoCoordinate box="[171,197,1517,1534]" degrees="7" direction="south" orientation="latitude" pageId="9" pageNumber="68" precision="55555" value="-7.0">7°S</geoCoordinate>
</emphasis>
</paragraph>
<paragraph blockId="9.[140,1000,1612,1677]" box="[230,962,1612,1629]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[230,962,1612,1629]" pageId="9" pageNumber="68">8ı° 80° 79° 78° 77° 76° 75°</emphasis>
</paragraph>
<paragraph blockId="9.[140,1000,1612,1677]" box="[140,1000,1654,1677]" pageId="9" pageNumber="68">
<emphasis bold="true" box="[140,238,1654,1676]" pageId="9" pageNumber="68">Figure 5.</emphasis>
Distribution of
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1881" box="[390,594,1654,1675]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="praestabilis">
<emphasis box="[390,594,1654,1675]" italics="true" pageId="9" pageNumber="68">Enyalioides praestabilis</emphasis>
</taxonomicName>
(circles) and
<taxonomicName authority="Torres-Carvajal &amp; de Queiroz &amp; Etheridge, 2009" authorityName="Torres-Carvajal &amp; de Queiroz &amp; Etheridge" authorityYear="2009" box="[718,848,1654,1676]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="rubrigularis" status="sp. n.">
<emphasis box="[718,848,1654,1676]" italics="true" pageId="9" pageNumber="68">E. rubrigularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[855,905,1655,1676]" pageId="9" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
(squares).
</paragraph>
</subSubSection>
</treatment>
</document>