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<document id="AC275214F75AEA9E7CD3FCE69474C7F2" ID-CLB-Dataset="58033" ID-DOI="10.5852/ejt.2023.882.2177" ID-GBIF-Dataset="57d1d191-2bf5-4229-a0f5-9ab472a114a6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177581" ID-ZooBank="67C909E4-C700-4F8D-B8CE-5FD9B2C5D549" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1690201644379" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Weyer, Dieter" docDate="2023" docId="03EA5C14CA658522FDB7FE98FC1B810A" docLanguage="en" docName="ejt-882_1-230.pdf" docOrigin="European Journal of Taxonomy 882" docSource="http://dx.doi.org/10.5852/ejt.2023.882.2177" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Imitoceras initium Korn &amp; Weyer 2023, sp. nov." docType="treatment" docUuid="F4100C55-64E5-48AF-8EDC-87CDA35580F1" docUuidSource="ZooBank" docVersion="16" lastPageNumber="110" masterDocId="FFD3246CCA09854FFFCEFF90FFB2846D" masterDocTitle="The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)" masterLastPageNumber="230" masterPageNumber="1" pageNumber="109" updateTime="1699545181523" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E3395A4A11D32647D03AB11DA96E3564">The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)</mods:title>
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<mods:namePart id="15116A6B2058FBA6F4475C742ADC544C">Korn, Dieter</mods:namePart>
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<mods:affiliation id="70612CB97C57C2EFBEA4D6D5AF79ECF0">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="3863861209C666C0A4105650266E0186">Weyer, Dieter</mods:namePart>
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<mods:affiliation id="8865B1E1808383C3F6EBC73340DEDFA6">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<subSubSection id="C359BE89CA658523FDB7FE98FB77852B" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="8BFCED02CA658523FDB7FE98FC09854E" blockId="108.[367,1221,264,362]" box="[633,955,264,291]" pageId="108" pageNumber="109">
<heading id="D0B45A6ECA658523FDB7FE98FC09854E" box="[633,955,264,291]" centered="true" fontSize="11" level="2" pageId="108" pageNumber="109" reason="2">
<taxonomicName id="4C439681CA658523FDB7FE98FCEB854F" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[633,857,264,290]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="108" pageNumber="109" phylum="Mollusca" rank="species" species="initium" status="sp. nov.">
<emphasis id="B9373110CA658523FDB7FE98FCEB854F" bold="true" box="[633,857,264,290]" italics="true" pageId="108" pageNumber="109">Imitoceras initium</emphasis>
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<taxonomicNameLabel id="A2048C6BCA658523FCAEFE99FC09854E" box="[864,955,265,291]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C359BE89CA658523FD78FEDFFCCF8504" box="[694,893,335,362]" pageId="108" pageNumber="109" type="description">
<paragraph id="8BFCED02CA658523FD78FEDFFCCF8504" blockId="108.[367,1221,264,362]" box="[694,893,335,362]" pageId="108" pageNumber="109">
<figureCitation id="1378F187CA658523FD78FEDFFCBE8507" box="[694,780,335,362]" captionStart="Fig" captionStartId="108.[189,232,1929,1955]" captionTargetBox="[244,1340,1488,1883]" captionTargetId="figure-296@108.[238,1350,1484,1889]" captionTargetPageId="108" captionText="Fig. 65. Imitoceras initium sp. nov., holotype MB.C.31144 (Korn 1977 Coll.) from the forestry road cutting west of the Oberrödinghausen railway cutting. A. Lateral and dorsal view. B. Suture line, at ww =14.0 mm, wh =15.4 mm. Scale bar units=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177713" httpUri="https://zenodo.org/record/8177713/files/figure.png" pageId="108" pageNumber="109">Fig. 65</figureCitation>
;
<tableCitation id="C6C1D8B9CA658523FCD6FEDFFCCF8504" box="[792,893,335,361]" captionStart="Table 63" captionStartId="109.[189,258,265,291]" captionTargetPageId="109" captionText="Table 63. Conch measurements, ratios and rates of Imitoceras initium sp. nov. from Oberrödinghausen and Oese." pageId="108" pageNumber="109">Table 63</tableCitation>
</paragraph>
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<subSubSection id="C359BE89CA658523FF73FE00FAC5860B" pageId="108" pageNumber="109" type="diagnosis">
<paragraph id="8BFCED02CA658523FF73FE00FE8485C7" blockId="108.[189,310,400,426]" box="[189,310,400,426]" pageId="108" pageNumber="109">
<heading id="D0B45A6ECA658523FF73FE00FE8485C7" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B9373110CA658523FF73FE00FE8485C7" bold="true" box="[189,310,400,426]" pageId="108" pageNumber="109">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA658523FF73FE2FFAC5860B" blockId="108.[189,1399,447,614]" pageId="108" pageNumber="109">
Species of
<taxonomicName id="4C439681CA658523FE8FFE50FE0F85B4" authorityName="Wurmi Schindewolf" authorityYear="1926" box="[321,445,448,473]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="108" pageNumber="109" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA658523FE8FFE50FE0F85B4" box="[321,445,448,473]" italics="true" pageId="108" pageNumber="109">Imitoceras</emphasis>
</taxonomicName>
with a conch reaching
<quantity id="4CBB40E7CA658523FD1CFE2FFC9785B4" box="[722,805,447,473]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="108" pageNumber="109" unit="mm" value="60.0">60 mm</quantity>
diameter. Conch at
<quantity id="4CBB40E7CA658523FBDAFE2FFBD585B4" box="[1044,1127,447,474]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="108" pageNumber="109" unit="mm" value="50.0">50 mm</quantity>
dm discoidal, involute (ww/dm ~0.45; uw/dm ~0.05); whorl profile at
<quantity id="4CBB40E7CA658523FD23FE72FC8C8590" box="[749,830,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="108" pageNumber="109" unit="mm" value="30.0">30 mm</quantity>
dm weakly compressed (ww/wh ~0.75); coiling rate very high (WER ~2.30). Venter rounded, umbilical margin rounded, flanks strongly convergent. Growth lines very fine, narrow-standing, with slightly biconvex course. Weak constrictions on the shell surface of the inner flank. Suture line with a weakly pouched external lobe and a V-shaped adventive lobe.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA658523FF73FD1DFDA48694" pageId="108" pageNumber="109" type="etymology">
<paragraph id="8BFCED02CA658523FF73FD1DFEF686CA" blockId="108.[189,324,653,679]" box="[189,324,653,679]" pageId="108" pageNumber="109">
<heading id="D0B45A6ECA658523FF73FD1DFEF686CA" bold="true" box="[189,324,653,679]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B9373110CA658523FF73FD1DFEF686CA" bold="true" box="[189,324,653,679]" pageId="108" pageNumber="109">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA658523FF73FD2CFDA48694" blockId="108.[189,1399,700,761]" pageId="108" pageNumber="109">
After Latin
<taxonomicName id="4C439681CA658523FE99FD2CFE1886B8" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[343,426,700,725]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="108" pageNumber="109" phylum="Mollusca" rank="species" species="initium">
<emphasis id="B9373110CA658523FE99FD2CFE1886B8" box="[343,426,700,725]" italics="true" pageId="108" pageNumber="109">initium</emphasis>
</taxonomicName>
= the beginning; because of the position of the species at the base of a long-ranging evolutionary lineage.
</paragraph>
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<subSubSection id="C359BE89CA658523FF73FCB0FC6C87C2" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="8BFCED02CA658523FF73FCB0FE188757" blockId="108.[189,426,800,826]" box="[189,426,800,826]" pageId="108" pageNumber="109">
<heading id="D0B45A6ECA658523FF73FCB0FE188757" bold="true" box="[189,426,800,826]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B9373110CA658523FF73FCB0FE188757" bold="true" box="[189,426,800,826]" pageId="108" pageNumber="109">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA658523FF24FCDFFEEE8704" blockId="108.[189,1398,847,944]" box="[234,348,847,873]" pageId="108" pageNumber="109">
<emphasis id="B9373110CA658523FF24FCDFFEEE8704" bold="true" box="[234,348,847,873]" pageId="108" pageNumber="109">
<typeStatus id="54F853A0CA658523FF24FCDFFEEE8704" box="[234,348,847,873]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFCED02CA658523FF73FCE2FC6C87C2" blockId="108.[189,1398,847,944]" pageId="108" pageNumber="109">
<materialsCitation id="3B2BE75FCA658523FF73FCE2FC6C87C2" ID-GBIF-Occurrence="4165071649" country="Germany" location="Rhenish Mountains" pageId="108" pageNumber="109" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F354AD92CA658523FF73FCE2FEE387E1" box="[189,337,882,908]" name="Germany" pageId="108" pageNumber="109">GERMANY</collectingCountry>
<location id="8E9CBBD9CA658523FEA6FCE2FDE287E1" LSID="urn:lsid:plazi:treatment:03EA5C14CA658522FDB7FE98FC1B810A:8E9CBBD9CA658523FEA6FCE2FDE287E1" box="[360,592,882,908]" country="Germany" name="Rhenish Mountains" pageId="108" pageNumber="109">Rhenish Mountains</location>
, Oberrödinghausen, west of railway cutting; Hangenberg Limestone, loose material; Korn 1977 Coll.; illustrated in
<figureCitation id="1378F187CA658523FD12FC05FC9C87DD" box="[732,814,917,944]" captionStart="Fig" captionStartId="108.[189,232,1929,1955]" captionTargetBox="[244,1340,1488,1883]" captionTargetId="figure-296@108.[238,1350,1484,1889]" captionTargetPageId="108" captionText="Fig. 65. Imitoceras initium sp. nov., holotype MB.C.31144 (Korn 1977 Coll.) from the forestry road cutting west of the Oberrödinghausen railway cutting. A. Lateral and dorsal view. B. Suture line, at ww =14.0 mm, wh =15.4 mm. Scale bar units=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177713" httpUri="https://zenodo.org/record/8177713/files/figure.png" pageId="108" pageNumber="109">Fig. 65</figureCitation>
; MB.C.31144.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA658523FF73FC46FDD4809E" pageId="108" pageNumber="109" type="description">
<paragraph id="8BFCED02CA658523FF73FC46FEFC879D" blockId="108.[189,334,982,1008]" box="[189,334,982,1008]" pageId="108" pageNumber="109">
<heading id="D0B45A6ECA658523FF73FC46FEFC879D" bold="true" box="[189,334,982,1008]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B9373110CA658523FF73FC46FEFC879D" bold="true" box="[189,334,982,1008]" pageId="108" pageNumber="109">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA658523FF73FB95FDD4809E" blockId="108.[188,1400,1029,1267]" pageId="108" pageNumber="109">
<typeStatus id="54F853A0CA658523FF73FB95FE988072" box="[189,298,1029,1055]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
MB.C.31144 is well preserved and has a conch diameter of
<quantity id="4CBB40E7CA658523FBC4FB95FBD2804D" box="[1034,1120,1029,1056]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="108" pageNumber="109" unit="mm" value="52.0">52 mm</quantity>
(
<figureCitation id="1378F187CA658523FBBCFB95FB50804D" box="[1138,1250,1029,1056]" captionStart="Fig" captionStartId="108.[189,232,1929,1955]" captionTargetBox="[244,1340,1488,1883]" captionTargetId="figure-296@108.[238,1350,1484,1889]" captionTargetPageId="108" captionText="Fig. 65. Imitoceras initium sp. nov., holotype MB.C.31144 (Korn 1977 Coll.) from the forestry road cutting west of the Oberrödinghausen railway cutting. A. Lateral and dorsal view. B. Suture line, at ww =14.0 mm, wh =15.4 mm. Scale bar units=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177713" httpUri="https://zenodo.org/record/8177713/files/figure.png" pageId="108" pageNumber="109">Fig. 65A</figureCitation>
). It is thinly discoid with a slightly opened umbilicus (ww/dm = 0.43; uw/dm = 0.05) and has a compressed whorl profile that is widest at the rounded umbilical margin; it has fairly strongly converging flanks and a rounded venter. The coiling rate is very high (WER = 2.30). The shell bears barely visible, extremely faint growth lines with a biconvex course and also a shell constriction limited to the inner flank. The suture line has a weakly pouched external lobe that is about two-thirds the depth of the nearly symmetric V-shaped adventive lobe (
<figureCitation id="1378F187CA658523FE23FB48FDEB809E" box="[493,601,1240,1267]" captionStart="Fig" captionStartId="108.[189,232,1929,1955]" captionTargetBox="[244,1340,1488,1883]" captionTargetId="figure-296@108.[238,1350,1484,1889]" captionTargetPageId="108" captionText="Fig. 65. Imitoceras initium sp. nov., holotype MB.C.31144 (Korn 1977 Coll.) from the forestry road cutting west of the Oberrödinghausen railway cutting. A. Lateral and dorsal view. B. Suture line, at ww =14.0 mm, wh =15.4 mm. Scale bar units=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177713" httpUri="https://zenodo.org/record/8177713/files/figure.png" pageId="108" pageNumber="109">Fig. 65B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3CBD8ACA658523FF73F819FC8B8387" ID-DOI="http://doi.org/10.5281/zenodo.8177713" ID-Zenodo-Dep="8177713" httpUri="https://zenodo.org/record/8177713/files/figure.png" pageId="108" pageNumber="109" startId="108.[189,232,1929,1955]" targetBox="[244,1340,1488,1883]" targetPageId="108">
<paragraph id="8BFCED02CA658523FF73F819FC8B8387" blockId="108.[189,1398,1929,2026]" pageId="108" pageNumber="109">
<emphasis id="B9373110CA658523FF73F819FEAE83CE" bold="true" box="[189,284,1929,1955]" pageId="108" pageNumber="109">Fig. 65.</emphasis>
<taxonomicName id="4C439681CA658523FEE8F81AFDB283CE" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[294,512,1930,1955]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="108" pageNumber="109" phylum="Mollusca" rank="species" species="initium" status="sp. nov.">
<emphasis id="B9373110CA658523FEE8F81AFDB283CE" box="[294,512,1930,1955]" italics="true" pageId="108" pageNumber="109">Imitoceras initium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCA658523FDC4F81AFDD883C9" box="[522,618,1930,1956]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
, holotype MB.C.31144 (Korn 1977 Coll.) from the forestry road cutting west of the Oberrödinghausen railway cutting.
<emphasis id="B9373110CA658523FC9DF83CFCDA83AB" bold="true" box="[851,872,1964,1990]" pageId="108" pageNumber="109">A</emphasis>
. Lateral and dorsal view.
<emphasis id="B9373110CA658523FB69F83CFB0983AB" bold="true" box="[1191,1211,1964,1990]" pageId="108" pageNumber="109">B</emphasis>
. Suture line, at ww =14.0 mm, wh =15.4 mm. Scale bar units=1 mm.
</paragraph>
</caption>
<caption id="DF3CBD8ACA648522FF73FE99FE83852A" pageId="109" pageNumber="110" startId="109.[189,258,265,291]" targetBox="[197,1379,380,481]" targetIsTable="true" targetPageId="109">
<paragraph id="8BFCED02CA648522FF73FE99FE83852A" blockId="109.[189,1399,265,327]" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FF73FE99FE9D854E" bold="true" box="[189,303,265,291]" pageId="109" pageNumber="110">Table 63.</emphasis>
Conch measurements, ratios and rates of
<taxonomicName id="4C439681CA648522FCD5FE9AFC41854E" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[795,1011,266,291]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium" status="sp. nov.">
<emphasis id="B9373110CA648522FCD5FE9AFC41854E" box="[795,1011,266,291]" italics="true" pageId="109" pageNumber="110">Imitoceras initium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCA648522FC35FE9AFBE48549" box="[1019,1110,266,292]" pageId="109" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
from Oberrödinghausen and Oese.
</paragraph>
</caption>
<paragraph id="8BFCED02CA648522FF2DFEECFAE0858C" pageId="109" pageNumber="110">
<table id="F9431FA2CA647AB0FF0BFEECFAD1858C" box="[197,1379,380,481]" colsContinueFrom="94.[197,1372,372,607]" colsContinueIn="121.[197,1377,344,546]" gridcols="11" gridrows="3" pageId="109" pageNumber="110">
<tr id="3573EF40CA647AB0FF0BFEECFAD185F9" box="[197,1379,380,404]" gridrow="0" pageId="109" pageNumber="110">
<th id="76A2863CCA647AB0FF0BFEECFEFD85F9" box="[197,335,380,404]" gridcol="0" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FF2DFEECFEFE85F9" bold="true" box="[227,332,380,404]" pageId="109" pageNumber="110">specimen</emphasis>
</th>
<th id="76A2863CCA647AB0FE5EFEECFE0D85F9" box="[400,447,380,404]" gridcol="1" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FE5BFEECFE0885F9" bold="true" box="[405,442,380,404]" pageId="109" pageNumber="110">dm</emphasis>
</th>
<th id="76A2863CCA647AB0FE37FEECFD9A85F9" box="[505,552,380,404]" gridcol="2" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FE33FEECFD9685F9" bold="true" box="[509,548,380,404]" pageId="109" pageNumber="110">ww</emphasis>
</th>
<th id="76A2863CCA647AB0FDADFEECFD2085F9" box="[611,658,380,404]" gridcol="3" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FDA7FEECFD3985F9" bold="true" box="[617,651,380,404]" pageId="109" pageNumber="110">wh</emphasis>
</th>
<th id="76A2863CCA647AB0FD06FEECFD5885F9" box="[712,746,380,404]" gridcol="4" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FD06FEECFD5885F9" bold="true" box="[712,746,380,404]" pageId="109" pageNumber="110">uw</emphasis>
</th>
<th id="76A2863CCA647AB0FCD7FEECFCFA85F9" box="[793,840,380,404]" gridcol="5" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FCECFEECFC8C85F9" bold="true" box="[802,830,380,404]" pageId="109" pageNumber="110">ah</emphasis>
</th>
<th id="76A2863CCA647AB0FCA1FEECFC7385F9" box="[879,961,380,404]" gridcol="6" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FCA1FEECFC7385F9" bold="true" box="[879,961,380,404]" pageId="109" pageNumber="110">ww/dm</emphasis>
</th>
<th id="76A2863CCA647AB0FC29FEECFB8485F9" box="[999,1078,380,404]" gridcol="7" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FC29FEECFB8485F9" bold="true" box="[999,1078,380,404]" pageId="109" pageNumber="110">ww/wh</emphasis>
</th>
<th id="76A2863CCA647AB0FB92FEECFB1885F9" box="[1116,1194,380,404]" gridcol="8" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FB92FEECFB1885F9" bold="true" box="[1116,1194,380,404]" pageId="109" pageNumber="110">uw/dm</emphasis>
</th>
<th id="76A2863CCA647AB0FB00FEECFABC85F9" box="[1230,1294,380,404]" gridcol="9" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FB00FEECFABC85F9" bold="true" box="[1230,1294,380,404]" pageId="109" pageNumber="110">WER</emphasis>
</th>
<th id="76A2863CCA647AB0FAFDFEECFAD185F9" box="[1331,1379,380,404]" gridcol="10" gridrow="0" pageId="109" pageNumber="110">
<emphasis id="B9373110CA648522FAFDFEECFAD085F9" bold="true" box="[1331,1378,380,404]" pageId="109" pageNumber="110">IZR</emphasis>
</th>
</tr>
<tr id="3573EF40CA647AB0FF0BFE37FAD185AD" box="[197,1379,423,448]" gridrow="1" pageId="109" pageNumber="110">
<th id="76A2863CCA647AB0FF0BFE37FEFD85AD" box="[197,335,423,448]" gridcol="0" gridrow="1" pageId="109" pageNumber="110">MB.C.31144</th>
<td id="76A2863CCA647AB0FE5EFE37FE0D85AD" box="[400,447,423,448]" gridcol="1" gridrow="1" pageId="109" pageNumber="110">52.0</td>
<td id="76A2863CCA647AB0FE37FE37FD9A85AD" box="[505,552,423,448]" gridcol="2" gridrow="1" pageId="109" pageNumber="110">22.2</td>
<td id="76A2863CCA647AB0FDADFE37FD2085AD" box="[611,658,423,448]" gridcol="3" gridrow="1" pageId="109" pageNumber="110">29.6</td>
<td id="76A2863CCA647AB0FD06FE37FD5885AD" box="[712,746,423,448]" gridcol="4" gridrow="1" pageId="109" pageNumber="110">2.6</td>
<td id="76A2863CCA647AB0FCD7FE37FCFA85AD" box="[793,840,423,448]" gridcol="5" gridrow="1" pageId="109" pageNumber="110">17.7</td>
<td id="76A2863CCA647AB0FCA1FE37FC7385AD" box="[879,961,423,448]" gridcol="6" gridrow="1" pageId="109" pageNumber="110">0.43</td>
<td id="76A2863CCA647AB0FC29FE37FB8485AD" box="[999,1078,423,448]" gridcol="7" gridrow="1" pageId="109" pageNumber="110">0.75</td>
<td id="76A2863CCA647AB0FB92FE37FB1885AD" box="[1116,1194,423,448]" gridcol="8" gridrow="1" pageId="109" pageNumber="110">0.05</td>
<td id="76A2863CCA647AB0FB00FE37FABC85AD" box="[1230,1294,423,448]" gridcol="9" gridrow="1" pageId="109" pageNumber="110">2.30</td>
<td id="76A2863CCA647AB0FAFDFE37FAD185AD" box="[1331,1379,423,448]" gridcol="10" gridrow="1" pageId="109" pageNumber="110">0.40</td>
</tr>
<tr id="3573EF40CA647AB0FF0BFE59FAD1858C" box="[197,1379,457,481]" gridrow="2" pageId="109" pageNumber="110">
<th id="76A2863CCA647AB0FF0BFE59FEFD858C" box="[197,335,457,481]" gridcol="0" gridrow="2" pageId="109" pageNumber="110">MB.C.31144</th>
<td id="76A2863CCA647AB0FE5EFE59FE0D858C" box="[400,447,457,481]" gridcol="1" gridrow="2" pageId="109" pageNumber="110">34.2</td>
<td id="76A2863CCA647AB0FE37FE59FD9A858C" box="[505,552,457,481]" gridcol="2" gridrow="2" pageId="109" pageNumber="110">16.6</td>
<td id="76A2863CCA647AB0FDADFE59FD20858C" box="[611,658,457,481]" gridcol="3" gridrow="2" pageId="109" pageNumber="110">19.0</td>
<td id="76A2863CCA647AB0FD06FE59FD58858C" box="[712,746,457,481]" gridcol="4" gridrow="2" pageId="109" pageNumber="110">2.0</td>
<td id="76A2863CCA647AB0FCD7FE59FCFA858C" box="[793,840,457,481]" gridcol="5" gridrow="2" pageId="109" pageNumber="110"></td>
<td id="76A2863CCA647AB0FCA1FE59FC73858C" box="[879,961,457,481]" gridcol="6" gridrow="2" pageId="109" pageNumber="110">0.49</td>
<td id="76A2863CCA647AB0FC29FE59FB84858C" box="[999,1078,457,481]" gridcol="7" gridrow="2" pageId="109" pageNumber="110">0.87</td>
<td id="76A2863CCA647AB0FB92FE59FB18858C" box="[1116,1194,457,481]" gridcol="8" gridrow="2" pageId="109" pageNumber="110">0.06</td>
<td id="76A2863CCA647AB0FB00FE59FABC858C" box="[1230,1294,457,481]" gridcol="9" gridrow="2" pageId="109" pageNumber="110"></td>
<td id="76A2863CCA647AB0FAFDFE59FAD1858C" box="[1331,1379,457,481]" gridcol="10" gridrow="2" pageId="109" pageNumber="110"></td>
</tr>
</table>
</paragraph>
<subSubSection id="C359BE89CA648522FF73FD97FC1B810A" pageId="109" pageNumber="110" type="discussion">
<paragraph id="8BFCED02CA648522FF73FD97FE9D864C" blockId="109.[189,303,519,545]" box="[189,303,519,545]" pageId="109" pageNumber="110">
<heading id="D0B45A6ECA648522FF73FD97FE9D864C" bold="true" box="[189,303,519,545]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B9373110CA648522FF73FD97FE9D864C" bold="true" box="[189,303,519,545]" pageId="109" pageNumber="110">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA648522FF73FDA5FD5586B1" blockId="109.[189,1399,565,732]" pageId="109" pageNumber="110">
The occurrence of the genus
<taxonomicName id="4C439681CA648522FDEBFDA6FD108622" authorityName="Wurmi Schindewolf" authorityYear="1926" box="[549,674,566,591]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FDEBFDA6FD108622" box="[549,674,566,591]" italics="true" pageId="109" pageNumber="110">Imitoceras</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C439681CA648522FD31FDA5FBAE8622" authority="Limestone" authorityName="Limestone" box="[767,1052,565,591]" class="Cephalopoda" family="Gattendorfiidae" genus="Gattendorfia" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FD31FDA5FC278622" box="[767,917,565,591]" italics="true" pageId="109" pageNumber="110">Gattendorfia</emphasis>
Limestone
</taxonomicName>
near Oberrödinghausen was previously unknown, but is actually not a great surprise. Representatives of
<taxonomicName id="4C439681CA648522FB99FDC9FB66861F" authorityName="Wurmi Schindewolf" authorityYear="1926" box="[1111,1236,601,626]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FB99FDC9FB66861F" box="[1111,1236,601,626]" italics="true" pageId="109" pageNumber="110">Imitoceras</emphasis>
</taxonomicName>
have already been found together with early Tournaisian ammonoids in
<collectingRegion id="498723E0CA648522FC4EFDEBFC5786F8" box="[896,997,635,661]" country="China" name="Guizhou" pageId="109" pageNumber="110">Guizhou</collectingRegion>
(
<bibRefCitation id="EFD290F3CA648522FC36FDECFB6686FB" author="Sun Y. &amp; Shen Y." box="[1016,1236,635,662]" pageId="109" pageNumber="110" pagination="33 - 100" refId="ref117793" refString="Sun Y. &amp; Shen Y. 1965. On the late Upper Devonian ammonite fauna of the Wocklumeria beds of south Kweichow and its stratigraphical significance. Professional Papers of the Academy of Geological Science, Ministry of Geology, section B 1: 33 - 100." type="journal article" year="1965">Sun &amp; Shen 1965</bibRefCitation>
;
<bibRefCitation id="EFD290F3CA648522FB2DFDECFAD586FB" author="Ruan Y." box="[1251,1383,636,662]" pageId="109" pageNumber="110" pagination="1 - 152" refId="ref116988" refString="Ruan Y. 1981. Devonian and earliest Carboniferous Ammonoids from Guangxi and Guizhou. Memoires of Nanjing Institute of Geology and Paleontology, Academia Sinica 15: 1 - 152." type="journal article" year="1981">Ruan 1981</bibRefCitation>
), originally described as “
<taxonomicName id="4C439681CA648522FE27FD0FFCA486D4" box="[489,790,671,697]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="planolobatum">
<emphasis id="B9373110CA648522FE27FD0FFCA486D4" box="[489,790,671,697]" italics="true" pageId="109" pageNumber="110">Imitoceras planolobatum</emphasis>
</taxonomicName>
” and “
<taxonomicName id="4C439681CA648522FCBDFD0FFB4E86D5" authorityName="Ruan" authorityYear="1981" box="[883,1276,671,697]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="crassum" subGenus="Imitoceras">
<emphasis id="B9373110CA648522FCBDFD0FFC4286D5" box="[883,1008,671,696]" italics="true" pageId="109" pageNumber="110">Imitoceras</emphasis>
(
<emphasis id="B9373110CA648522FBC8FD0FFB3186D5" box="[1030,1155,671,696]" italics="true" pageId="109" pageNumber="110">Imitoceras</emphasis>
)
<emphasis id="B9373110CA648522FB54FD0FFB4E86D5" box="[1178,1276,671,696]" italics="true" pageId="109" pageNumber="110">crassum</emphasis>
</taxonomicName>
”, and the Anti-Atlas (
<bibRefCitation id="EFD290F3CA648522FE86FD52FD6A86B1" author="Bockwinkel J. &amp; Ebbighausen V." box="[328,728,706,732]" pageId="109" pageNumber="110" pagination="87 - 129" refId="ref112039" refString="Bockwinkel J. &amp; Ebbighausen V. 2006. A new ammonoid fauna from the Gattendorfia - Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco). Fossil Record 9: 87 - 129. https: // doi. org / 10.5194 / fr- 9 - 87 - 2006" type="journal article" year="2006">Bockwinkel &amp; Ebbighausen 2006</bibRefCitation>
).
</paragraph>
<paragraph id="8BFCED02CA648522FF73FC93FC5487EA" blockId="109.[189,1400,771,903]" pageId="109" pageNumber="110">
The only specimen available is unique in the ammonoid assemblage from Oberrödinghausen because of its high coiling rate of 2.30. Such a value is rarely reached by other members of the prionoceratid ammonoids. In the subfamily
<taxonomicName id="4C439681CA648522FDE8FCD9FCB8870E" authorityName="Korn" authorityYear="1994" box="[550,778,841,867]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitina" pageId="109" pageNumber="110" phylum="Mollusca" rank="subFamily" subFamily="Acutimitoceratinae">Acutimitoceratinae</taxonomicName>
, the whorl expansion rate is normally between 1.70 and
<quantity id="4CBB40E7CA648522FF3FFCFCFEF187EB" box="[241,323,876,902]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="109" pageNumber="110" unit="in" value="1.9">1.90 in</quantity>
<taxonomicName id="4C439681CA648522FE89FCFCFE6187EB" baseAuthorityName="Korn &amp; Weyer" baseAuthorityYear="2023" box="[327,467,876,902]" class="Cephalopoda" family="Prionoceratidae" genus="Stockumites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FE89FCFCFE6187EB" box="[327,467,876,902]" italics="true" pageId="109" pageNumber="110">Stockumites</emphasis>
</taxonomicName>
and between 1.90 and
<quantity id="4CBB40E7CA648522FD2CFCFCFC8687EB" box="[738,820,876,903]" metricMagnitude="-2" metricUnit="m" metricValue="5.207" pageId="109" pageNumber="110" unit="in" value="2.05">2.05 in</quantity>
<taxonomicName id="4C439681CA648522FCF5FCFDFC5287EB" authorityName="Korn" authorityYear="1993" box="[827,992,877,902]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FCF5FCFDFC5287EB" box="[827,992,877,902]" italics="true" pageId="109" pageNumber="110">Nicimitoceras</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCED02CA648522FF73FC3DFC048039" blockId="109.[189,1399,941,1108]" pageId="109" pageNumber="110">
Of these genera, only
<taxonomicName id="4C439681CA648522FE16FC3EFDCF87AA" authorityName="Korn" authorityYear="1993" box="[472,637,942,967]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FE16FC3EFDCF87AA" box="[472,637,942,967]" italics="true" pageId="109" pageNumber="110">Nicimitoceras</emphasis>
</taxonomicName>
has a short external lobe like in
<taxonomicName id="4C439681CA648522FBE5FC3EFABB87AA" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[1067,1289,942,967]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium" status="sp. nov.">
<emphasis id="B9373110CA648522FBE5FC3EFABB87AA" box="[1067,1289,942,967]" italics="true" pageId="109" pageNumber="110">Imitoceras initium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCA648522FAD8FC3EFAC587A5" box="[1302,1399,942,968]" pageId="109" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
However,
<taxonomicName id="4C439681CA648522FEF6FC41FE148787" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[312,422,977,1002]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium">
<emphasis id="B9373110CA648522FEF6FC41FE148787" box="[312,422,977,1002]" italics="true" pageId="109" pageNumber="110">I. initium</emphasis>
</taxonomicName>
is stouter than the size-equivalent
<typeStatus id="54F853A0CA648522FC82FC40FC0C8787" box="[844,958,976,1002]" pageId="109" pageNumber="110" type="holotype">holotypes</typeStatus>
of other
<taxonomicName id="4C439681CA648522FBFFFC41FA868787" box="[1073,1332,976,1002]" class="Cephalopoda" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca">
<emphasis id="B9373110CA648522FBFFFC41FB648787" box="[1073,1238,977,1002]" italics="true" pageId="109" pageNumber="110">Nicimitoceras</emphasis>
species
</taxonomicName>
from Oberrödinghausen: its ww/dm ratio is 0.43 at
<quantity id="4CBB40E7CA648522FD08FC63FCA48063" box="[710,790,1011,1038]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="109" pageNumber="110" unit="mm" value="52.0">52 mm</quantity>
dm in
<taxonomicName id="4C439681CA648522FC90FC64FC758060" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[862,967,1012,1037]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium">
<emphasis id="B9373110CA648522FC90FC64FC758060" box="[862,967,1012,1037]" italics="true" pageId="109" pageNumber="110">I. initium</emphasis>
</taxonomicName>
, while the value in
<taxonomicName id="4C439681CA648522FB6DFC64FADF8060" baseAuthorityName="Vohringer" baseAuthorityYear="1960" box="[1187,1389,1011,1037]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="heterolobatum">
<emphasis id="B9373110CA648522FB6DFC64FADF8060" box="[1187,1389,1011,1037]" italics="true" pageId="109" pageNumber="110">N. heterolobatum</emphasis>
</taxonomicName>
, for example, is 0.36 at
<quantity id="4CBB40E7CA648522FE1EFB86FD91805C" box="[464,547,1046,1073]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="109" pageNumber="110" unit="mm" value="54.0">54 mm</quantity>
dm. In addition, the whorl profile shows rapidly converging flanks in
<taxonomicName id="4C439681CA648522FAABFB87FEA2803E" authorityName="Korn &amp; Weyer" authorityYear="2023" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium">
<emphasis id="B9373110CA648522FAABFB87FEA2803E" italics="true" pageId="109" pageNumber="110">I. initium</emphasis>
</taxonomicName>
, which tend to be subparallel in species of
<taxonomicName id="4C439681CA648522FCC5FBAAFC02803E" authorityName="Korn" authorityYear="1993" box="[779,944,1082,1107]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA648522FCC5FBAAFC02803E" box="[779,944,1082,1107]" italics="true" pageId="109" pageNumber="110">Nicimitoceras</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCED02CA648522FF73FBEBFC1B810A" blockId="109.[187,1399,1147,1384]" pageId="109" pageNumber="110">
A superficially similar species to
<taxonomicName id="4C439681CA648522FD94FBECFC8580F8" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[602,823,1148,1173]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium" status="sp. nov.">
<emphasis id="B9373110CA648522FD94FBECFC8580F8" box="[602,823,1148,1173]" italics="true" pageId="109" pageNumber="110">Imitoceras initium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCA648522FC8DFBEBFC1080F8" box="[835,930,1147,1173]" pageId="109" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
is
<taxonomicName id="4C439681CA648522FC03FBEBFAB380F8" authorityName="Korn" authorityYear="1984" box="[973,1281,1147,1173]" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="procedens">
<emphasis id="B9373110CA648522FC03FBEBFAB380F8" box="[973,1281,1147,1173]" italics="true" pageId="109" pageNumber="110">Acutimitoceras procedens</emphasis>
</taxonomicName>
from the Stockum limestone equivalent of the Müssenberg (
<bibRefCitation id="EFD290F3CA648522FCEBFB0EFC1880D5" author="Korn D." box="[805,938,1182,1208]" pageId="109" pageNumber="110" pagination="513 - 526" refId="ref113936" refString="Korn D. 1981. Ein neues, Ammonoiden-fuhrendes Profil an der Devon-Karbon-Grenze im Sauerland (Rhein. Schiefergebirge). Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte 1981: 513 - 526. https: // doi. org / 10.1127 / njgpm / 1981 / 1981 / 513" type="journal article" year="1981">Korn 1981</bibRefCitation>
). This also shows a high coiling rate, rapidly convergent flanks, and a shell ornament with weak growth lines and constrictions. The
<typeStatus id="54F853A0CA648522FADEFB51FAC580B6" box="[1296,1399,1217,1243]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
of this species is much stouter (ww/dm = 0.54 at
<quantity id="4CBB40E7CA648522FCCBFB74FCEB8093" box="[773,857,1252,1278]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="109" pageNumber="110" unit="mm" value="33.0">33 mm</quantity>
dm) than
<typeStatus id="54F853A0CA648522FC1CFB74FB8B8093" box="[978,1081,1252,1278]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
MB.C.31144 of
<taxonomicName id="4C439681CA648522FACDFB74FAC58093" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[1283,1399,1252,1278]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="109" pageNumber="110" phylum="Mollusca" rank="species" species="initium">
<emphasis id="B9373110CA648522FACDFB74FAC58093" box="[1283,1399,1252,1278]" italics="true" pageId="109" pageNumber="110">A. initium</emphasis>
</taxonomicName>
(ww/dm = 0.43 at
<quantity id="4CBB40E7CA648522FE43FA97FE6F814C" box="[397,477,1287,1314]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="109" pageNumber="110" unit="mm" value="52.0">52 mm</quantity>
dm). This difference cannot be explained by the advanced ontogenetic evolution towards a more slender conch in specimen MB.C.31144, as this has half a whorl in front of the largest diameter, a ww/dm ratio of only 0.47 at
<quantity id="4CBB40E7CA648522FD5BFADDFD5A8105" box="[661,744,1357,1384]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="109" pageNumber="110" unit="mm" value="36.0">36 mm</quantity>
conch diameter.
</paragraph>
</subSubSection>
</treatment>
</document>