553 lines
79 KiB
XML
553 lines
79 KiB
XML
<document id="9008446A9A800E15F5D78C6595CC5692" ID-DOI="10.5281/zenodo.184516" ID-GBIF-Dataset="62a45f81-bcff-449c-82d8-fef278006ea8" ID-ISSN="1175-5326" ID-Zenodo-Dep="184516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459985733509" checkinUser="plazi" docAuthor="Artal, Pedro, Guinot, Danièle, Bakel, Barry Van & Castillo, Juan" docDate="2008" docId="03E5D65BEF323F78079CFE65FDBAD37F" docLanguage="en" docName="zt01907p027.pdf" docOrigin="Zootaxa 1907" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ibericancer sanchoi Artal, Guinot, Bakel & Castillo, 2008, sp. nov." docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FFDCAE23EF3A3F6A070BFF97FFB5D115" masterDocTitle="Ibericancridae, a new dakoticancroid family (Decapoda, Brachyura, Podotremata) from the upper Campanian (Upper Cretaceous) of Spain" masterLastPageNumber="27" masterPageNumber="1" pageNumber="9" updateTime="1698232278591" updateUser="plazi">
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<mods:titleInfo id="C3594D8D38A1582C4E213EA12131510B">
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<mods:title id="793FA8CC7F9503F1DB8871D44ECB756B">Ibericancridae, a new dakoticancroid family (Decapoda, Brachyura, Podotremata) from the upper Campanian (Upper Cretaceous) of Spain</mods:title>
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</mods:titleInfo>
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<mods:name id="AF23D555446CEC142545DC3DCECD8D99" type="personal">
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<mods:roleTerm id="B6ACE35D223332205C86D42141E82AA3">Author</mods:roleTerm>
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<mods:namePart id="51FE342C67E008420C165560B5CA4F50">Artal, Pedro</mods:namePart>
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<mods:roleTerm id="31C0F6A6CF33D32F07393A949AFBF2F6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BC3ACB0214FF2885CDE6EEA34F28EB50">Guinot, Danièle</mods:namePart>
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</mods:name>
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<mods:name id="53A5BC2C0BB28F19432E68FFD19B9C8B" type="personal">
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<mods:roleTerm id="402769F5480CF4E7D766DD5220E515DE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ADDC0972038472FEB54EDCD045BCED53">Bakel, Barry Van</mods:namePart>
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</mods:name>
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<mods:name id="72D70F16512BA0F5BA8CFE44B5D138E6" type="personal">
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<mods:roleTerm id="C156B1EBE67C88687BCA280A5E051B43">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F9A2323EFC271823958E40461FBB397D">Castillo, Juan</mods:namePart>
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<mods:typeOfResource id="0B8FF0044E25B9FE89E5165B9EBEAD5E">text</mods:typeOfResource>
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<mods:title id="2FF5C79DD1304CBF5548CE85A8C3966B">Zootaxa</mods:title>
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<mods:part id="82EC4F17F27CB847546B0A373EE1AF48">
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<mods:date id="9FBD583FB790501B0E7CAD493CCD7500">2008</mods:date>
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<mods:number id="F4C7F50A18E4E51487F8C71FA8E29878">1907</mods:number>
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<mods:classification id="4A664A1964EA28FB4C7249A6E8A455FC">journal article</mods:classification>
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<mods:identifier id="25791C31DCE90E0C248F0A87143C6211" type="DOI">10.5281/zenodo.184516</mods:identifier>
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<mods:identifier id="A502F1AC9EB92D3DC1AEE38B9049DA81" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="515F464576B86109F0578CA402196492" type="Zenodo-Dep">184516</mods:identifier>
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<treatment id="03E5D65BEF323F78079CFE65FDBAD37F" ID-DOI="http://doi.org/10.5281/zenodo.6233908" ID-GBIF-Taxon="119379218" ID-Zenodo-Dep="6233908" LSID="urn:lsid:plazi:treatment:03E5D65BEF323F78079CFE65FDBAD37F" httpUri="http://treatment.plazi.org/id/03E5D65BEF323F78079CFE65FDBAD37F" lastPageId="18" lastPageNumber="19" pageId="8" pageNumber="9">
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<subSubSection id="C35634C6EF323F62079CFE65FE93D33A" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BF3674DEF323F62079CFE65FE58D319" blockId="8.[151,493,498,559]" box="[151,493,498,524]" pageId="8" pageNumber="9">
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<heading id="D0BBD021EF323F62079CFE65FE58D319" bold="true" box="[151,493,498,524]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B938BB5FEF323F62079CFE65FE58D319" bold="true" box="[151,493,498,524]" pageId="8" pageNumber="9">
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<taxonomicName id="4C4C1CCEEF323F62079CFE65FE3CD319" ID-CoL="643ac661-6d3f-4be0-a28e-a0d2b5e4d4a2" box="[151,393,498,524]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanchoi" status="sp. nov.">
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<emphasis id="B938BB5FEF323F62079CFE65FE3CD319" bold="true" box="[151,393,498,524]" italics="true" pageId="8" pageNumber="9">Ibericancer sanchoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B0624EF323F620684FE65FE58D319" box="[399,493,498,524]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF3674DEF323F62079CFD82FE93D33A" blockId="8.[151,493,498,559]" box="[151,294,533,559]" pageId="8" pageNumber="9">
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(
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<figureCitation id="13777BC8EF323F620794FD82FEA8D33A" box="[159,285,533,559]" captionStart-0="FIGURE 4 A – E" captionStart-1="FIGURE 5 A – D" captionStart-2="FIGURE 6 A, B" captionStart-3="FIGURE 7 A – C" captionStart-4="FIGURE 8 A – D" captionStart-5="FIGURE 9 A – C" captionStart-6="FIGURE 10 A – D" captionStartId-0="9.[151,255,1902,1926]" captionStartId-1="10.[151,255,1897,1921]" captionStartId-2="11.[151,255,1919,1943]" captionStartId-3="12.[151,255,1919,1943]" captionStartId-4="13.[151,255,1903,1927]" captionStartId-5="14.[151,255,1919,1943]" captionStartId-6="15.[151,255,1191,1215]" captionTargetBox-0="[226,1360,181,1874]" captionTargetBox-1="[226,1360,178,1871]" captionTargetBox-2="[226,1360,194,1894]" captionTargetBox-3="[226,1360,194,1894]" captionTargetBox-4="[226,1360,178,1878]" captionTargetBox-5="[226,1360,194,1894]" captionTargetBox-6="[151,1434,194,1166]" captionTargetId-0="figure@9.[226,1360,175,1876]" captionTargetId-1="figure@10.[226,1360,172,1873]" captionTargetId-2="figure@11.[226,1360,194,1895]" captionTargetId-3="figure@12.[226,1360,194,1895]" captionTargetId-4="figure@13.[226,1360,178,1879]" captionTargetId-5="figure@14.[226,1360,194,1895]" captionTargetId-6="figure@15.[151,1436,194,1167]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="FIGURE 4 A – E. Ibericancer sanchoi n. gen., n. sp.; A. thoracic sternum, paratype MGSB 68573 a, the arrows indicating the abdominal holding mechanism on sternite 5 and G 1; B, C. enlargement of G 1; D. female thoracic sternum, paratype MGSB 68573 b, the arrow indicating the press button type protuberance on sternite 5; E. paratype MGSB 68573 c, outer surface of left chela." captionText-1="FIGURE 5 A – D. Ibericancer sanchoi n. gen., n. sp.; paratypes: A. MGSB 68573 d, oblique posterior view showing reduced P 4 and P 5; B. MGSB 68573 e, posterior view showing a 2 covering the male gonopore; C. MGSB 68574 f, partial male abdomen and thoracic sternum; D. MGSB 68574 g, complete female abdomen; the arrow indicating gonopore on P 3 coxa." captionText-2="FIGURE 6 A, B. Ibericancer sanchoi n. gen., n. sp.; holotype MGSB 68572, male: A. left lateral view showing lateral side and a portion of the pterygostomial region slightly displaced; B. ventral view." captionText-3="FIGURE 7 A – C. Ibericancer sanchoi n. gen., n. sp.; paratypes: A. MGSB 68573 h, oblique lateral view showing the reduced last pair of periopods; the arrows indicating P 4 and P 5; B-C. MGSB 68574 b, ventral and lateral views, respectively, displaying the pterygostomial region." captionText-4="FIGURE 8 A – D. Ibericancer sanchoi n. gen., n. sp.; paratypes: A. partial ventral view of a male, MGSB 68574 c; the arrow indicating the male gonopore on P 5 coxa; B. partial carapace in dorsal view, MGSB 68574 d, showing the complete front and orbit; C. oblique lateral view of the same specimen; the arrow indicating the infraorbital protuberance; D. oblique partial view, MGSB 68574 e, with detail of reduced P 4 and P 5." captionText-5="FIGURE 9 A – C. Ibericancer sanchoi n. gen., n. sp.; A. paratype MGSB 68574 a, dorsal view; B. holotype MGSB 68572, posterior view showing the concavity of the posterior margin; C. detail of P 1 - P 5 in the same specimen of A." captionText-6="FIGURE 10 A – D. Ibericancer sanchoi n. gen., n. sp., paratype MGSB 68574 f (specimen whitened with ammonium chloride prior to photography); A. frontal view; B. dorsal view; C. reconstruction of the complete frontal view; D. reconstruction of the complete carapace in dorsal view." httpUri-0="https://zenodo.org/record/184519/files/figure.png" httpUri-1="https://zenodo.org/record/184520/files/figure.png" httpUri-2="https://zenodo.org/record/184521/files/figure.png" httpUri-3="https://zenodo.org/record/184522/files/figure.png" httpUri-4="https://zenodo.org/record/184523/files/figure.png" httpUri-5="https://zenodo.org/record/184524/files/figure.png" httpUri-6="https://zenodo.org/record/184525/files/figure.png" pageId="8" pageNumber="9">Figs. 4–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF323F62079CFDF2FD94D36A" box="[151,545,613,639]" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BF3674DEF323F62079CFDF2FD94D36A" blockId="8.[151,1437,613,1959]" box="[151,545,613,639]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F62079CFDF2FEA2D36A" bold="true" box="[151,279,613,639]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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Same as in the family.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF323F6207CDFD1AFAC9D3B2" box="[198,1404,653,679]" pageId="8" pageNumber="9" type="etymology">
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<paragraph id="8BF3674DEF323F6207CDFD1AFAC9D3B2" blockId="8.[151,1437,613,1959]" box="[198,1404,653,679]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F6207CDFD1AFEE6D3B2" bold="true" box="[198,339,653,679]" pageId="8" pageNumber="9">Etymology.</emphasis>
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In honour of the late Mr. Fausto Sancho (Gandía,
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<collectingCountry id="F35B27DDEF323F6204A9FD1AFC5CD3B2" box="[930,1001,653,679]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
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), who discovered the new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF323F6207CDFD22FBD1D252" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BF3674DEF323F6207CDFD22FB4FD3E2" blockId="8.[151,1437,613,1959]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F6207CDFD22FEC9D3DA" bold="true" box="[198,380,693,719]" pageId="8" pageNumber="9">
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<typeStatus id="54F7D9EFEF323F6207CDFD22FEB6D3DA" box="[198,259,693,719]" pageId="8" pageNumber="9">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54F7D9EFEF323F620688FD22FE46D3DA" box="[387,499,693,719]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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, male, (
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<collectionCode id="ED5DFF88EF323F620546FD22FD16D3DA" box="[589,675,693,719]" pageId="8" pageNumber="9">MGSB</collectionCode>
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68572);
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<typeStatus id="54F7D9EFEF323F62040FFD22FCC2D3DA" box="[772,887,693,719]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
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, male and female specimens (
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<collectionCode id="ED5DFF88EF323F6203DFFD22FA9DD3DA" box="[1236,1320,693,719]" pageId="8" pageNumber="9">MGSB</collectionCode>
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68573a–h and
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<collectionCode id="ED5DFF88EF323F6207C3FD4AFEA9D3E2" box="[200,284,733,759]" pageId="8" pageNumber="9">MGSB</collectionCode>
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68574a–f). Maximum size (in millimeters): W 49, L 59. Minimum size: W 37, L 36.
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</paragraph>
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<paragraph id="8BF3674DEF323F6207CDFC92FB3BD20A" blockId="8.[151,1437,613,1959]" box="[198,1166,773,799]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F6207CDFC92FED3D20A" bold="true" box="[198,358,773,799]" pageId="8" pageNumber="9">
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<typeStatus id="54F7D9EFEF323F6207CDFC92FEB1D20A" box="[198,260,773,799]" pageId="8" pageNumber="9">Type</typeStatus>
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locality
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</emphasis>
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: Pla de la Basa, Valencia,
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<collectingCountry id="F35B27DDEF323F620596FC92FD51D20A" box="[669,740,773,799]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
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. Upper Cretaceous, late Campanian.
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</paragraph>
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<paragraph id="8BF3674DEF323F6207CDFCBAFBD1D252" blockId="8.[151,1437,613,1959]" box="[198,1124,813,839]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F6207CDFCBAFDBCD252" bold="true" box="[198,521,813,839]" pageId="8" pageNumber="9">Other material examined.</emphasis>
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Pla de la Basa, Valencia,
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<collectingCountry id="F35B27DDEF323F620433FCBAFCCED252" box="[824,891,813,839]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
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(
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<collectionCode id="ED5DFF88EF323F620481FCBAFC55D252" box="[906,992,813,839]" pageId="8" pageNumber="9">MGSB</collectionCode>
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68575a–h).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF323F7A07CDFCC2FAF9D0E7" lastPageId="16" lastPageNumber="17" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BF3674DEF323F6207CDFCC2FBE1D5C2" blockId="8.[151,1437,613,1959]" pageId="8" pageNumber="9">
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<emphasis id="B938BB5FEF323F6207CDFCC2FEEBD27A" bold="true" box="[198,350,853,879]" pageId="8" pageNumber="9">Description.</emphasis>
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Carapace of large size, nearly as broad as long; suboval shape with smoothly arched margins posterior to pronounced branchiocardiac furrow; dorsal surface strongly convex longitudinally, nearly flat transversely. Dorsal surface finely and densely granular. Fronto-orbital margin about 53% of maximum carapace width, rather advanced. Front occupying 46% of anterior margin, delimited by two protuberant nodes at internal orbital sides, slightly concave in dorsal view, clearly notched axially, bearing 2 blunt nodes at base of rostrum; rostrum narrow, strongly deflected downwards, about 90 degrees, deeply grooved medially, terminating in a bifid end with rounded lobes. Proepistome as a very thin plate, epistome well defined. Orbits small, directed forwards, bounded by frontal lobe and outer orbital spine; supraorbital margin strongly modified by a salient, triangular, robust spine, deep fissure between medial, extraorbital spines, short infraorbital margin with small terminal spine. Ocular peduncles probably well sheltered inside orbits.
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</paragraph>
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<paragraph id="8BF3674DEF323F6207CEFB72FB02D4D2" blockId="8.[151,1437,613,1959]" pageId="8" pageNumber="9">
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Lateral margins arched; anterolateral margin slightly shorter than posterior, ornamented by a thin spine close to outer-orbital, followed by a concavity and a short, salient ridge; anterolateral margin joining the posterolateral posteriorly to a rather large and swollen epibranchial node. Posterolateral margin gently arched, continuous, not interrupted by nodes or notches. Posterior margin concave
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<emphasis id="B938BB5FEF323F62032FFACAFBDBD463" box="[1060,1134,1373,1398]" italics="true" pageId="8" pageNumber="9">per se</emphasis>
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, gently convex in dorsal view, somewhat broader than fronto-orbital width; notably rimmed, affected axially by a strong conical protuberance. In posterior view, posterior border robust, V-shaped, with apex directed upwards.
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</paragraph>
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<paragraph id="8BF3674DEF323F6207CEFA42FD46D7CA" blockId="8.[151,1437,613,1959]" pageId="8" pageNumber="9">Dorsal regions relatively inflated, not strongly protuberant or grooved except for branchiocardiac groove, some depressions, rare swellings. Gastric regions broad, swollen, undivided, occupying major surface of anterior half of carapace. Hepatic region not well defined, marked by a slight depression. Epibranchial region long, transversely inflated, curving abruptly at level of cardiac region, forming small terminal protuberance. Branchiocardiac groove smooth, moderately deep, more marked axially. Cardiac region large, being individualized by a slightly swollen, nearly circular lobe. Postbranchial regions obliquely ridged, diverging backwards, bounded by broad, depressed intestinal area.</paragraph>
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<paragraph id="8BF3674DEF323F6207C6F97AFD73D6B2" blockId="8.[151,1437,613,1959]" pageId="8" pageNumber="9">Pterygostomial region inflated, ornamented with a row of three large tubercles. Pleural suture located on carapace sides. Third maxillipeds broad, not divergent, separated by a straight gap; noticeably long, exceeding front; ischium, merus in same plane; endopod much longer than wide, with deep longitudinal groove on ischium; coxa very broad, trapezoidal, with lower margin bearing two small protuberances at level of sternal articulation; exopod thin, long, sharp at termination.</paragraph>
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<caption id="DF3337C5EF333F63079CF8F9FEC4D6F8" httpUri="https://zenodo.org/record/184519/files/figure.png" pageId="9" pageNumber="10" targetBox="[226,1360,181,1874]" targetPageId="9">
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<paragraph id="8BF3674DEF333F63079CF8F9FEC4D6F8" blockId="9.[151,1436,1902,2029]" pageId="9" pageNumber="10">
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<emphasis id="B938BB5FEF333F63079CF8F9FEFED693" bold="true" box="[151,331,1902,1926]" pageId="9" pageNumber="10">FIGURE 4A–E.</emphasis>
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<taxonomicName id="4C4C1CCEEF333F63065AF8F8FD92D693" box="[337,551,1903,1926]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
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<emphasis id="B938BB5FEF333F63065AF8F8FD92D693" box="[337,551,1903,1926]" italics="true" pageId="9" pageNumber="10">Ibericancer sanchoi</emphasis>
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</taxonomicName>
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<emphasis id="B938BB5FEF333F630526F8F9FD0AD693" bold="true" box="[557,703,1902,1926]" pageId="9" pageNumber="10">
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<taxonomicNameLabel id="A20B0624EF333F630526F8F9FDCDD693" box="[557,632,1902,1926]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
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, n. sp.
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</emphasis>
|
||
; A. thoracic sternum, paratype MGSB 68573a, the arrows indicating the abdominal holding mechanism on sternite 5 and G1; B, C. enlargement of G1; D. female thoracic sternum, paratype MGSB 68573b, the arrow indicating the press button type protuberance on sternite 5; E. paratype MGSB 68573c, outer surface of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF303F60079CF8FEFF65D6FD" httpUri="https://zenodo.org/record/184520/files/figure.png" pageId="10" pageNumber="11" targetBox="[226,1360,178,1871]" targetPageId="10">
|
||
<paragraph id="8BF3674DEF303F60079CF8FEFF65D6FD" blockId="10.[151,1436,1897,2024]" pageId="10" pageNumber="11">
|
||
<emphasis id="B938BB5FEF303F60079CF8FEFEE5D694" bold="true" box="[151,336,1897,1921]" pageId="10" pageNumber="11">FIGURE 5A–D.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF303F600650F8FDFD81D694" box="[347,564,1898,1921]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF303F600650F8FDFD81D694" box="[347,564,1898,1921]" italics="true" pageId="10" pageNumber="11">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF303F600535F8FEFD68D694" bold="true" box="[574,733,1897,1921]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A20B0624EF303F600535F8FEFD3BD694" box="[574,654,1897,1921]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
; paratypes: A. MGSB 68573d, oblique posterior view showing reduced P4 and P5; B. MGSB 68573e, posterior view showing a2 covering the male gonopore; C. MGSB 68574f, partial male abdomen and thoracic sternum; D. MGSB 68574g, complete female abdomen; the arrow indicating gonopore on P3 coxa.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF313F61079CF8E8FBB1D6AC" httpUri="https://zenodo.org/record/184521/files/figure.png" pageId="11" pageNumber="12" targetBox="[226,1360,194,1894]" targetPageId="11">
|
||
<paragraph id="8BF3674DEF313F61079CF8E8FBB1D6AC" blockId="11.[151,1436,1919,1977]" pageId="11" pageNumber="12">
|
||
<emphasis id="B938BB5FEF313F61079CF8E8FEF8D682" bold="true" box="[151,333,1919,1943]" pageId="11" pageNumber="12">FIGURE 6A, B.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF313F61065EF8E8FD99D683" box="[341,556,1919,1942]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF313F61065EF8E8FD99D683" box="[341,556,1919,1942]" italics="true" pageId="11" pageNumber="12">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF313F610538F8E8FD7FD682" bold="true" box="[563,714,1919,1943]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A20B0624EF313F610538F8E8FD35D682" box="[563,640,1919,1943]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
; holotype MGSB 68572, male: A. left lateral view showing lateral side and a portion of the pterygostomial region slightly displaced; B. ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF363F66079CF8E8FDDBD6CE" httpUri="https://zenodo.org/record/184522/files/figure.png" pageId="12" pageNumber="13" targetBox="[226,1360,194,1894]" targetPageId="12">
|
||
<paragraph id="8BF3674DEF363F66079CF8E8FDDBD6CE" blockId="12.[151,1436,1919,2011]" pageId="12" pageNumber="13">
|
||
<emphasis id="B938BB5FEF363F66079CF8E8FEFAD682" bold="true" box="[151,335,1919,1943]" pageId="12" pageNumber="13">FIGURE 7A–C.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF363F660652F8E8FD84D683" box="[345,561,1919,1942]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF363F660652F8E8FD84D683" box="[345,561,1919,1942]" italics="true" pageId="12" pageNumber="13">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF363F660530F8E8FD63D682" bold="true" box="[571,726,1919,1943]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A20B0624EF363F660530F8E8FD3CD682" box="[571,649,1919,1943]" pageId="12" pageNumber="13" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
; paratypes: A. MGSB 68573h, oblique lateral view showing the reduced last pair of periopods; the arrows indicating P4 and P5; B-C. MGSB 68574b, ventral and lateral views, respectively, displaying the pterygostomial region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF373F67079CF8F8FC3CD6FB" httpUri="https://zenodo.org/record/184523/files/figure.png" pageId="13" pageNumber="14" targetBox="[226,1360,178,1878]" targetPageId="13">
|
||
<paragraph id="8BF3674DEF373F67079CF8F8FC3CD6FB" blockId="13.[151,1436,1903,2030]" pageId="13" pageNumber="14">
|
||
<emphasis id="B938BB5FEF373F67079CF8F8FEFAD692" bold="true" box="[151,335,1903,1927]" pageId="13" pageNumber="14">FIGURE 8A–D.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF373F67065CF8F8FD9AD693" box="[343,559,1903,1926]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF373F67065CF8F8FD9AD693" box="[343,559,1903,1926]" italics="true" pageId="13" pageNumber="14">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF373F67053CF8F8FD65D692" bold="true" box="[567,720,1903,1927]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="A20B0624EF373F67053CF8F8FD30D692" box="[567,645,1903,1927]" pageId="13" pageNumber="14" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
; paratypes: A. partial ventral view of a male, MGSB 68574c; the arrow indicating the male gonopore on P5 coxa; B. partial carapace in dorsal view, MGSB 68574d, showing the complete front and orbit; C. oblique lateral view of the same specimen; the arrow indicating the infraorbital protuberance; D. oblique partial view, MGSB 68574e, with detail of reduced P4 and P5.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF343F64079CF8E8FADDD6AC" httpUri="https://zenodo.org/record/184524/files/figure.png" pageId="14" pageNumber="15" targetBox="[226,1360,194,1894]" targetPageId="14">
|
||
<paragraph id="8BF3674DEF343F64079CF8E8FADDD6AC" blockId="14.[151,1436,1919,1977]" pageId="14" pageNumber="15">
|
||
<emphasis id="B938BB5FEF343F64079CF8E8FEE4D682" bold="true" box="[151,337,1919,1943]" pageId="14" pageNumber="15">FIGURE 9A–C.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF343F640657F8E8FD83D683" box="[348,566,1919,1942]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF343F640657F8E8FD83D683" box="[348,566,1919,1942]" italics="true" pageId="14" pageNumber="15">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF343F64054BF8E8FD55D682" bold="true" box="[576,736,1919,1943]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="A20B0624EF343F64054BF8E8FD25D682" box="[576,656,1919,1943]" pageId="14" pageNumber="15" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
; A. paratype MGSB 68574a, dorsal view; B. holotype MGSB 68572, posterior view showing the concavity of the posterior margin; C. detail of P1-P5 in the same specimen of A.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3337C5EF353F65079CFB30FD1DD416" httpUri="https://zenodo.org/record/184525/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1434,194,1166]" targetPageId="15">
|
||
<paragraph id="8BF3674DEF353F65079CFB30FD1DD416" blockId="15.[151,1436,1191,1283]" pageId="15" pageNumber="16">
|
||
<emphasis id="B938BB5FEF353F65079CFB30FEE8D5AA" bold="true" box="[151,349,1191,1215]" pageId="15" pageNumber="16">FIGURE 10A–D.</emphasis>
|
||
<taxonomicName id="4C4C1CCEEF353F65066CFB30FDF5D5AB" box="[359,576,1191,1214]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF353F65066CFB30FDF5D5AB" box="[359,576,1191,1214]" italics="true" pageId="15" pageNumber="16">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF353F650541FB30FD52D5AA" bold="true" box="[586,743,1191,1215]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A20B0624EF353F650541FB30FD2CD5AA" box="[586,665,1191,1215]" pageId="15" pageNumber="16" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
, paratype MGSB 68574f (specimen whitened with ammonium chloride prior to photography); A. frontal view; B. dorsal view; C. reconstruction of the complete frontal view; D. reconstruction of the complete carapace in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF3674DEF353F6507CEFAA0FBE6D61C" blockId="15.[151,1436,1335,2001]" pageId="15" pageNumber="16">Thoracic sternum relatively wide, stout, strongly elevated laterally; a rather large portion of sternum visible between P1, P2 coxae and male abdomen. Sternites 1–4 fused, forming a broad triangle visible nearly entirely in ventral view; anterior portion (1–3) narrow, pointed forwards, inserted between bases of mxp3; episternite 3 very small, only a reduced salient portion where coxa of mxp3 is articulated; sternite 4 well developed, with lateral margins strongly elevated, forming an acute ridge, episternite 4 small, extended laterally, slightly curved; sternite 5 as wide as sternite 4, subrectangular, nearly horizontal; sternite 6 narrower, slightly oblique; sternite 7 very reduced, oblique, mostly covered by third pleomere; sternite 8 reduced, oblique, completely covered by second pleomere. Thoracic sutures 4/5 to 7/8 not reaching axial area, preceding sutures 1/2, 2/3 absent. Presence of a small tubercle on sternite 5, at level of pleomere 6, which is supposed to be a press button for the abdominal holding mechanism. Female gonopore on P3 coxae very small, nearly elliptical, gently rimmed; male gonopore on P5 coxae, somewhat larger, fairly elliptical, not rimmed; spermatheca at end of sternal suture 7/8, aperture large, ovate, somewhat oblique.</paragraph>
|
||
<paragraph id="8BF3674DEF353F7A07CEF880FCC9D017" blockId="15.[151,1436,1335,2001]" lastBlockId="16.[151,1436,152,498]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
Abdomen rather wide in males, not much widened in females; nearly rectangular over its entire length in males, triangular in females; strongly sculptured, presenting two longitudinal grooves and transverse crests on its first pleomeres. All pleomeres free, subrectangular, with salient small lobes at upper margin of each segment. A1 very narrow, convex in male; a2 slightly wider, notably sculptured, medially keeled, with elevated lateral extensions covering male gonopore on P5 coxa; a1 to a6 with straight margins, increasing in length from a3 to a6; telson triangular. Female abdomen roughly similar in shape, only slightly broader; pleomeres a1–a6 and telson with more clearly arched lateral margins conferring a more suboval-triangular outline. G1 thick, long, very stout, probably rolled (see
|
||
<figureCitation id="13777BC8EF2A3F7A05FAFF7FFCDBD017" box="[753,878,232,258]" captionStart="FIGURE 4 A – E" captionStartId="9.[151,255,1902,1926]" captionTargetBox="[226,1360,181,1874]" captionTargetId="figure@9.[226,1360,175,1876]" captionTargetPageId="9" captionText="FIGURE 4 A – E. Ibericancer sanchoi n. gen., n. sp.; A. thoracic sternum, paratype MGSB 68573 a, the arrows indicating the abdominal holding mechanism on sternite 5 and G 1; B, C. enlargement of G 1; D. female thoracic sternum, paratype MGSB 68573 b, the arrow indicating the press button type protuberance on sternite 5; E. paratype MGSB 68573 c, outer surface of left chela." httpUri="https://zenodo.org/record/184519/files/figure.png" pageId="16" pageNumber="17">Fig. 4B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2A3F7A07CEFE87FAF9D0E7" blockId="16.[151,1436,152,498]" pageId="16" pageNumber="17">Chelipeds of equal size, finely granulated; merus long, robust, subtriangular in section; propodus slender, longer than wide, subelliptical in section; fingers very long, thin. P2, P3 very long, robust, coxa subtriangular, stout; basis narrow; ischium robust, curved; merus very long, circular in section, curved at first proximal portion. P4, P5 subdorsal, conspicuously reduced, subequal, rather long, with smaller coxae. P4 coxa subtriangular, basis narrow, ischium straight, long, merus very long. P5 coxa robust despite its reduced size; rather complex in shape, with acute, thin ridge at upper margin; merus long; dactylus small, probably curved.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35634C6EF2A3F7A07CEFDB0FF4BD227" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8BF3674DEF2A3F7A07CEFDB0FF4BD227" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
<emphasis id="B938BB5FEF2A3F7A07CEFDB0FD52D354" bold="true" box="[197,743,551,577]" pageId="16" pageNumber="17">Remarks on dakoticancroid spermathecae.</emphasis>
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A05F6FDBFFC58D357" author="Bishop" box="[765,1005,552,578]" pageId="16" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2A3F7A0450FDBFFC27D354" box="[859,914,552,577]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. (1998
|
||
</bibRefCitation>
|
||
: 239) stated in the diagnosis of the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A079CFDC7FEE2D37F" box="[151,343,592,618]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
that the female sternum lacked “longitudinal grooves”. The term “longitudinal grooves” was used by these authors as well as by many neontologists (e.g.,
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A046AFDEFFC4BD387" author="Gordon" box="[865,1022,632,658]" pageId="16" pageNumber="26" refString="Gordon, I. (1950) Crustacea Dromiacea. Part I: Systematic account of the Dromiacea collected by the " John Murray " Expedition; Part II. The morphology of the spermatheca in certain Dromiacea. The John Murray Expedition 1933 - 34, Scientific Reports, 9 (3), 201 - 253." type="journal article" year="1950">Gordon 1950</bibRefCitation>
|
||
) to designate the condition of these sutures found in the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A068FFD37FDB0D3AF" box="[388,517,672,698]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dromiidae</taxonomicName>
|
||
. The term “groove” is not appropriate to describe the suture 7/8, which is short (
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0794FD5FFEFDD3F7" box="[159,328,712,738]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dynomenidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A065DFD5FFD85D3F7" box="[342,560,712,738]" class="Malacostraca" family="Homolodromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Homolodromiidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0534FD5FFCADD3F7" box="[575,792,712,738]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subFamily" subFamily="Sphaerodromiinae">Sphaerodromiinae</taxonomicName>
|
||
) or developed as an elongated and salient tube in the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A079CFD67FEA0D21F" box="[151,277,752,778]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A065BFD67FDD0D21F" author="Guinot" box="[336,613,752,778]" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae de Haan, 1833, with diagnoses and description of new genera and species. Zoosystema, 25, 43 - 129." type="journal article" year="2003">Guinot & Tavares 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A057BFD67FC23D21F" author="Guinot" box="[624,918,752,778]" pageId="16" pageNumber="26" refString="Guinot, D. & Quenette, G. (2005) The spermatheca in podotreme crabs (Crustacea, Decapoda, Brachyura, Podotremata) and its phylogenetic implications. Zoosystema, 27, 267 - 342." type="journal article" year="2005">Guinot & Quenette 2005</bibRefCitation>
|
||
). We propose to use ‘suture’ as the appropriate term.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35634C6EF2A3F7A07CEFCD7FB45D407" pageId="16" pageNumber="17" type="reference_group">
|
||
<paragraph id="8BF3674DEF2A3F7A07CEFCD7FAFAD58F" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
In
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A07EEFCD7FE7AD24C" ID-CoL="643ac661-6d3f-4be0-a28e-a0d2b5e4d4a2" box="[229,463,832,857]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF2A3F7A07EEFCD7FE7AD24C" box="[229,463,832,857]" italics="true" pageId="16" pageNumber="17">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B0624EF2A3F7A06DDFCA8FD9DD24C" box="[470,552,831,857]" pageId="16" pageNumber="17" rank="genus">
|
||
<emphasis id="B938BB5FEF2A3F7A06DDFCA8FD9DD24C" bold="true" box="[470,552,831,857]" pageId="16" pageNumber="17">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B938BB5FEF2A3F7A053DFCA8FDCDD24C" bold="true" box="[566,632,831,857]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
(
|
||
<figureCitation id="13777BC8EF2A3F7A058CFCD7FD50D24F" box="[647,741,832,858]" captionStart="FIGURE 4 A – E" captionStartId="9.[151,255,1902,1926]" captionTargetBox="[226,1360,181,1874]" captionTargetId="figure@9.[226,1360,175,1876]" captionTargetPageId="9" captionText="FIGURE 4 A – E. Ibericancer sanchoi n. gen., n. sp.; A. thoracic sternum, paratype MGSB 68573 a, the arrows indicating the abdominal holding mechanism on sternite 5 and G 1; B, C. enlargement of G 1; D. female thoracic sternum, paratype MGSB 68573 b, the arrow indicating the press button type protuberance on sternite 5; E. paratype MGSB 68573 c, outer surface of left chela." httpUri="https://zenodo.org/record/184519/files/figure.png" pageId="16" pageNumber="17">Fig. 4D</figureCitation>
|
||
), as well as in
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0484FCD7FB20D24C" ID-CoL="cdc0c084-90c4-482f-85b7-7807435b8e40" box="[911,1173,832,857]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF2A3F7A0484FCD7FB20D24C" box="[911,1173,832,857]" italics="true" pageId="16" pageNumber="17">Dakoticancer overana</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A03D9FCD7FADFD24F" author="Guinot" box="[1234,1386,832,858]" pageId="16" pageNumber="26" refString="Guinot, D. (1993) Donnees nouvelles sur les crabes primitifs (Crustacea Decapoda Brachyura Podotremata). Comptes Rendus de l'Academie des Sciences, ser. 3, 316, 1225 - 1232." type="journal article" year="1993">Guinot 1993</bibRefCitation>
|
||
: fig. 7;
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A07BFFCFFFE7FD297" author="Guinot" box="[180,458,872,898]" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares 2001</bibRefCitation>
|
||
: fig. 7H), and
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0564FCFFFC03D294" ID-CoL="f4e46612-3a7b-4136-9f3e-4be11fb02f70" box="[623,950,872,897]" class="Malacostraca" family="Dakoticancridae" genus="Tetracarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="subquadratus">
|
||
<emphasis id="B938BB5FEF2A3F7A0564FCFFFC03D294" box="[623,950,872,897]" italics="true" pageId="16" pageNumber="17">Tetracarcinus subquadratus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A04F9FCFFFB26D297" author="Roberts" box="[1010,1171,872,898]" pageId="16" pageNumber="27" refString="Roberts, H. B. (1962) The Upper Cretaceous decapod crustaceans of New Jersey and Delaware, In: Richards, H. G., Ramsdell, R. G., Miller, A. K., Garner, H. F. & Reeside, J. B. (Eds.), The Cretaceous fossils of New Jersey. Paleontology Series, New Jersey Geological Survey, 61, 163 - 191." type="journal article" year="1962">Roberts 1962</bibRefCitation>
|
||
: pl. 87, fig. 3), the thoracic sternal suture 7/8 is rather short and ends in a large orifice. Referring the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A033CFC07FB42D2BF" ID-CoL="377b5adf-9ed1-498f-8aef-7a7c4133e597" box="[1079,1271,912,938]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
as not having “longitudinal grooves” might give the wrong impression that sutures 7/8 are not present and thus do not play a reproductive role. Such a role is performed by dakoticancroid sutures 7/8 since their distal portions bear a large orifice as in all podotreme crabs (
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A056FFB9FFBA5D537" author="Guinot" box="[612,1040,1032,1058]" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares 2001: 522, fig.10</bibRefCitation>
|
||
). Only the raninoid spermathecae present a special pattern (see
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A0506FBA7FD0ED55F" author="Hartnoll" box="[525,699,1072,1098]" pageId="16" pageNumber="26" refString="Hartnoll, R. G. (1979) The phyletic implications of spermathecal structure in the Raninidae (Decapoda: Brachyura). Journal of Zoology, London, 187, 75 - 83." type="journal article" year="1979">Hartnoll 1979</bibRefCitation>
|
||
). In the Dakoticancroidea, although dependent of the same phragma 7/8, the spermatheca with its large aperture at the extremity of suture 7/8 does not appear similar to that of the Dromiacea and Homoloidea. It can only be inferred that its internal organisation is different.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2A3F7A07CEFB3FFB45D407" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
The diagnosis of the family
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0505FB3FFD7BD5D7" ID-CoL="377b5adf-9ed1-498f-8aef-7a7c4133e597" box="[526,718,1192,1218]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A05F1FB3FFC50D5D7" box="[762,997,1192,1218]" pageId="16" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2A3F7A045FFB3FFC3DD5D4" box="[852,904,1192,1217]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
is proposed herein to be modified as: “sternum of females with oblique sutures 7/8 and presence of large rounded orifices at their distal portions, which are homologous to the spermathecal apertures” (see also
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A048BFB6FFB1DD407" author="Guinot" box="[896,1192,1272,1298]" pageId="16" pageNumber="26" refString="Guinot, D. & Quenette, G. (2005) The spermatheca in podotreme crabs (Crustacea, Decapoda, Brachyura, Podotremata) and its phylogenetic implications. Zoosystema, 27, 267 - 342." type="journal article" year="2005">Guinot & Quenette 2005</bibRefCitation>
|
||
: 317).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35634C6EF2A3F7A07CEFAB7FE10D67F" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BF3674DEF2A3F7A07CEFAB7FE78D717" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
The thoracic sternum is wider in the Dakoticancroidea than in the Dromiacea. Sutures 4/5, 5/6 and 6/7 are developed and, instead of being short and confined laterally as in the Dromiacea, the sutures reach the sternoabdominal cavity (
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0679FAE7FD83D49F" box="[370,566,1392,1418]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
) or the median depression (
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A0476FAE7FB92D49F" box="[893,1063,1392,1418]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
|
||
<emphasis id="B938BB5FEF2A3F7A0325FAF8FBCAD49C" bold="true" box="[1070,1151,1391,1417]" pageId="16" pageNumber="17">n. fam</emphasis>
|
||
.) (only a sternoabdominal depression in the Dromiacea; see
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A057CFA0FFC09D4A7" author="Guinot" box="[631,956,1432,1458]" pageId="16" pageNumber="26" refString="Guinot, D. & Bouchard, J. M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20, 613 - 694." type="journal article" year="1998">Guinot & Bouchard 1998</bibRefCitation>
|
||
). Sutures 7/8 are much shorter in the Dakoticancroidea than in the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A06E6FA57FDDBD4CF" box="[493,622,1472,1498]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
, and the distal aperture is large and rounded instead of a usually small aperture in the
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A064CFA7FFE7DD717" box="[327,456,1512,1538]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2A3F7A07CEF987FD26D7B7" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
No affinities are suggested with the thoracic sternum and sutures 7/8 of the Homoloidea, in which, additionally, suture 6/7 is complete. There is a resemblance with the Cyclodorippoidea Ortmann, 1892, which have a broad sternum, with (typically) slit-like distal spermathecal apertures (
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A0349F9F7FAD3D76F" author="Guinot" box="[1090,1382,1632,1658]" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares 2001</bibRefCitation>
|
||
: fig. 10F, I, J;
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A060FF91FFD9ED7B7" author="Guinot" box="[260,555,1672,1698]" pageId="16" pageNumber="26" refString="Guinot, D. & Quenette, G. (2005) The spermatheca in podotreme crabs (Crustacea, Decapoda, Brachyura, Podotremata) and its phylogenetic implications. Zoosystema, 27, 267 - 342." type="journal article" year="2005">Guinot & Quenette 2005</bibRefCitation>
|
||
: fig. 23).
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2A3F7A07CEF927FE10D67F" blockId="16.[151,1437,551,1898]" pageId="16" pageNumber="17">
|
||
The Etyidae
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A0657F927FDCFD7DF" author="Guinot" box="[348,634,1712,1738]" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares, 2001</bibRefCitation>
|
||
, is a fossil podotreme family in which the spermathecae are known. Two oblique and large slits at the extremities of the short sutures 7/8 have been described for
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A03F7F94FFE8AD60F" authority="Mantell, 1844" authorityName="Mantell" authorityYear="1844" class="Malacostraca" family="Etyiidae" genus="Etyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martini">
|
||
<emphasis id="B938BB5FEF2A3F7A03F7F94FFA29D7E4" box="[1276,1436,1752,1777]" italics="true" pageId="16" pageNumber="17">Etyus martini</emphasis>
|
||
Mantell, 1844
|
||
</taxonomicName>
|
||
, from the European mid-Cretaceous (
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A05F7F897FBA1D60F" author="Wright" box="[764,1044,1792,1818]" pageId="16" pageNumber="27" refString="Wright, C. W. & Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of Palaeontological Society London, 126 (533), 126, 1 - 114." type="journal article" year="1972">Wright & Collins 1972</bibRefCitation>
|
||
: 102, pl. 21, fig. 6d, e;
|
||
<bibRefCitation id="EFDD1ABCEF2A3F7A0220F897FE82D657" author="Guinot" pageId="16" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares 2001</bibRefCitation>
|
||
: figs 2, 3, 10J). The spermathecae of
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A05E2F8BFFC66D654" box="[745,979,1832,1857]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF2A3F7A05E2F8BFFC66D654" box="[745,979,1832,1857]" italics="true" pageId="16" pageNumber="17">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B0624EF2A3F7A04D1F8B0FB99D654" box="[986,1068,1831,1857]" pageId="16" pageNumber="17" rank="genus">
|
||
<emphasis id="B938BB5FEF2A3F7A04D1F8B0FB99D654" bold="true" box="[986,1068,1831,1857]" pageId="16" pageNumber="17">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B938BB5FEF2A3F7A0331F8B0FBC9D654" bold="true" box="[1082,1148,1831,1857]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
are large, ovate in shape and obliquely directed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35634C6EF2A3F7B07CEF808FB65D7B7" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8BF3674DEF2A3F7B07CEF808FD05D30F" blockId="16.[151,1437,1951,2018]" lastBlockId="17.[151,1436,152,538]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
<emphasis id="B938BB5FEF2A3F7A07CEF808FDECD6AC" bold="true" box="[197,601,1951,1977]" pageId="16" pageNumber="17">Remarks on abdominal holding.</emphasis>
|
||
Casts of
|
||
<taxonomicName id="4C4C1CCEEF2A3F7A05C2F837FC64D6AC" box="[713,977,1952,1977]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF2A3F7A05C2F837FC64D6AC" box="[713,977,1952,1977]" italics="true" pageId="16" pageNumber="17">Dakoticancer overana</emphasis>
|
||
</taxonomicName>
|
||
examined and illustrated show a small protuberance on sternite 5 at the level of the socket on the ventral surface of pleomere 6 (
|
||
<figureCitation id="13777BC8EF2A3F7A03B1F85FFAA2D6F7" box="[1210,1303,1992,2018]" captionStart="FIGURE 3 A – E" captionStartId="5.[151,255,1871,1895]" captionTargetBox="[243,1344,221,1860]" captionTargetId="figure@5.[243,1345,211,1863]" captionTargetPageId="5" captionText="FIGURE 3 A – E. Casts of Dakoticancer overana; A. MNHN B 22902 - 1 (cast of USNM 173529 - 1), ventral view of a male; B. posterior view of the same specimen; C. details of the abdominal holding mechanism, the arrow indicating the press button protuberance on sternite 5; D. MNHM B 22902 - 2 (cast of USNM 173529 - 2), ventral view of a female, without the abdomen; the arrow indicating the aperture of the spermatheca; E. MNHN B 22903 (cast of USNM 173544), ventral view of a female; the arrow indicating the female gonopore on P 3 coxa." httpUri="https://zenodo.org/record/184518/files/figure.png" pageId="16" pageNumber="17">Fig. 3C</figureCitation>
|
||
). A similar condition can be deduced from the abdomen of
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B05C4FF0FFCEED1A4" box="[719,859,152,177]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF2B3F7B05C4FF0FFCEED1A4" box="[719,859,152,177]" italics="true" pageId="17" pageNumber="18">D. australis</emphasis>
|
||
</taxonomicName>
|
||
illustrated by
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B030DFF0FFB5BD1A7" author="Bishop" box="[1030,1262,152,178]" pageId="17" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2B3F7B036AFF0FFB23D1A4" box="[1121,1174,152,177]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. (1998
|
||
</bibRefCitation>
|
||
: 243, fig. 1.8), where the presence of a socket is suspected below the produced latero-posterior angles of pleomere 6. An abdominal holding system, with a press button located on sternite 5 and an abdominal socket, is thus recorded for the first time in a podotreme crab. It is of the press button
|
||
<typeStatus id="54F7D9EFEF2B3F7B046CFE87FC29D03F" box="[871,924,272,298]" pageId="17" pageNumber="18">type</typeStatus>
|
||
, as in nearly all Eubrachyura Saint Laurent, 1980. A salient button for the same function is present in the
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B047FFEAFFBA9D047" box="[884,1052,312,338]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
|
||
<emphasis id="B938BB5FEF2B3F7B032FFEA0FBC8D044" bold="true" box="[1060,1149,311,337]" pageId="17" pageNumber="18">n. fam.</emphasis>
|
||
(
|
||
<figureCitation id="13777BC8EF2B3F7B0385FEAFFAA1D047" box="[1166,1300,312,338]" captionStart="FIGURE 4 A – E" captionStartId="9.[151,255,1902,1926]" captionTargetBox="[226,1360,181,1874]" captionTargetId="figure@9.[226,1360,175,1876]" captionTargetPageId="9" captionText="FIGURE 4 A – E. Ibericancer sanchoi n. gen., n. sp.; A. thoracic sternum, paratype MGSB 68573 a, the arrows indicating the abdominal holding mechanism on sternite 5 and G 1; B, C. enlargement of G 1; D. female thoracic sternum, paratype MGSB 68573 b, the arrow indicating the press button type protuberance on sternite 5; E. paratype MGSB 68573 c, outer surface of left chela." httpUri="https://zenodo.org/record/184519/files/figure.png" pageId="17" pageNumber="18">Fig. 4A, D</figureCitation>
|
||
). The press button of the Dakoticancroidea constitutes a robust synapomorphy of the superfamily among podotremes. The press button is not exclusive to the Eubrachyura since a socket (although not coupled with a typical “button”) also occurs in another podotreme group, the
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B05AFFE27FC73D0DF" authority="Guinot, 1993" authorityName="Guinot" authorityYear="1993" box="[676,966,432,458]" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Lyreidinae">
|
||
Lyreidinae
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0422FE27FC73D0DF" author="Guinot" box="[809,966,432,458]" pageId="17" pageNumber="26" refString="Guinot, D. (1993) Donnees nouvelles sur les crabes primitifs (Crustacea Decapoda Brachyura Podotremata). Comptes Rendus de l'Academie des Sciences, ser. 3, 316, 1225 - 1232." type="journal article" year="1993">Guinot, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
, among the Raninoidea
|
||
<collectingCountry id="F35B27DDEF2B3F7B03E8FE27FAB0D0DF" box="[1251,1285,432,458]" name="South Georgia and the South Sandwich Islands" pageId="17" pageNumber="18">De</collectingCountry>
|
||
Haan, 1839. The “homolid press button” of the Homoloidea also consists of a socket on pleomere 6, but the corresponding prominence is located anteriorly on sternite 4.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2B3F7B07CEFDD8FADFD477" blockId="17.[151,1437,591,2018]" pageId="17" pageNumber="18">
|
||
<emphasis id="B938BB5FEF2B3F7B07CEFDD8FC8CD37C" bold="true" box="[197,825,591,617]" pageId="17" pageNumber="18">Hypotheses on burrowing and carrying behaviour.</emphasis>
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B044AFDC7FB9ED37F" box="[833,1067,592,618]" pageId="17" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2B3F7B0490FDC7FC7BD37C" box="[923,974,592,617]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
stated that evidence of burrowing by dakoticancrid crabs was equivocal. Although
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B0409FDEFFBBCD384" box="[770,1033,632,657]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF2B3F7B0409FDEFFBBCD384" box="[770,1033,632,657]" italics="true" pageId="17" pageNumber="18">Dakoticancer overana</emphasis>
|
||
</taxonomicName>
|
||
has frequently been found associated with burrows, these were neither abundant nor large enough to have been their homes, so that the species was suggested not to have been an obligate burrower (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B042DFD5FFC76D3F7" author="Bishop" box="[806,963,712,738]" pageId="17" pageNumber="25" refString="Bishop, G. A. (1981) Occurrence and fossilization of the Dakoticancer Assemblage, Upper Cretaceous Pierre Shale, South Dakota. In: Gray, J., Boucot, A. J. & Berry, W. B. N. (Eds.), Communities of the past, 13, 383 - 413. Hutchinson Ross Publishing Company, Stroudsburg, PA." type="journal article" year="1981">Bishop 1981</bibRefCitation>
|
||
). The presence of moulds of more than twenty specimens of
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B0686FD67FDA2D21C" box="[397,535,752,778]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF2B3F7B0686FD67FE17D21C" box="[397,418,752,777]" italics="true" pageId="17" pageNumber="18">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF2B3F7B06BBFD67FDA2D21C" box="[432,535,752,777]" italics="true" pageId="17" pageNumber="18">australis</emphasis>
|
||
</taxonomicName>
|
||
(with articulated appendages and dislocated plastrons) preserved within boxworks in the Potrerillos Formation of
|
||
<collectingCountry id="F35B27DDEF2B3F7B055BFC8FFD19D227" box="[592,684,792,818]" name="Mexico" pageId="17" pageNumber="18">Mexico</collectingCountry>
|
||
, inferred to be exuviae (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B04CDFC8FFB57D227" author="Vega" box="[966,1250,792,818]" pageId="17" pageNumber="27" refString="Vega, F. J. & Feldmann, R. M. (1991) Fossil crabs (Crustacea, Decapoda) from the Maastrichtian Difunta Group, northeastern Mexico. Annals of Carnegie Museum, 60, 163 - 177." type="journal article" year="1991">Vega & Feldmann 1991</bibRefCitation>
|
||
), however, suggests that moulting within burrows did take place. Such a behaviour has also been hypothesised by
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0226FCD7FEB0D297" author="Fraaije" pageId="17" pageNumber="25" refString="Fraaije, R. H. B., van Bakel, B. W. M. & Jagt, J. W. M. (2008) Albunea turritellacola, a new sand crab (Anomura, Albuneidae) from the lower Miocene of southwest France. Bulletin of the Mizunami Fossil Museum, 34, 17 - 22." type="journal article" year="2008">
|
||
Fraaije
|
||
<emphasis id="B938BB5FEF2B3F7B028CFCD7FF1BD294" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. (2008
|
||
</bibRefCitation>
|
||
: 20) for a new albuneid (sand crab) from the Miocene of
|
||
<collectingCountry id="F35B27DDEF2B3F7B0494FCFFFC47D297" box="[927,1010,872,898]" name="France" pageId="17" pageNumber="18">France</collectingCountry>
|
||
, which was found within a large turritellid gastropod. Conversely,
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B050AFC07FD39D2BC" box="[513,652,912,937]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF2B3F7B050AFC07FD39D2BC" box="[513,652,912,937]" italics="true" pageId="17" pageNumber="18">D. australis</emphasis>
|
||
</taxonomicName>
|
||
, from the Cárdenas Formation in east-central
|
||
<collectingCountry id="F35B27DDEF2B3F7B03A3FC07FAB1D2BF" box="[1192,1284,912,938]" name="Mexico" pageId="17" pageNumber="18">Mexico</collectingCountry>
|
||
, is not found associated with burrows, and most specimens preserve articulated appendages.
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B034EFC2FFAAED2C7" box="[1093,1307,952,978]" pageId="17" pageNumber="27" refString="Vega, F. J., Feldmann, R. M. & Sour-Tovar, F. (1995) Fossil crabs (Crustacea: Decapoda) from the Late Cretaceous Cardenas Formation, east-central Mexico. Journal of Paleontology, 69, 340 - 350." type="journal article">
|
||
Vega
|
||
<emphasis id="B938BB5FEF2B3F7B0383FC2FFB08D2C4" box="[1160,1213,952,977]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. (1995)
|
||
</bibRefCitation>
|
||
concluded that either these specimens were corpses or that the observed differences in preservation between the two Mexican assemblages resulted from different taphonomic histories. All Spanish specimens studied here are interpreted as moults. The pterygostomian plates are separated and are preserved slightly laterally displaced from the carapace. The proximal end of the first pleomere is disconnected from the thick posterior margin; this segment has fallen inside the carapace, and is preserved under an angle relative to the second pleomere. Another fact that does not fit a burrowing life is the supposed carrying behaviour inferred from the dorsal location and reduction of P5 (
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B06F9FB47FD03D5FF" box="[498,694,1232,1258]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
) or both P4 and P5 (
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B04ACFB47FBE4D5FF" box="[935,1105,1232,1258]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
|
||
<emphasis id="B938BB5FEF2B3F7B0353FB58FB1DD5FC" bold="true" box="[1112,1192,1231,1257]" pageId="17" pageNumber="18">n. fam</emphasis>
|
||
.) (
|
||
<figureCitation id="13777BC8EF2B3F7B03C2FB47FA87D5FF" box="[1225,1330,1232,1258]" captionStart="FIGURE 7 A – C" captionStartId="12.[151,255,1919,1943]" captionTargetBox="[226,1360,194,1894]" captionTargetId="figure@12.[226,1360,194,1895]" captionTargetPageId="12" captionText="FIGURE 7 A – C. Ibericancer sanchoi n. gen., n. sp.; paratypes: A. MGSB 68573 h, oblique lateral view showing the reduced last pair of periopods; the arrows indicating P 4 and P 5; B-C. MGSB 68574 b, ventral and lateral views, respectively, displaying the pterygostomial region." httpUri="https://zenodo.org/record/184522/files/figure.png" pageId="17" pageNumber="18">Figs. 7A</figureCitation>
|
||
,
|
||
<figureCitation id="13777BC8EF2B3F7B0235FB47FAD1D5FF" box="[1342,1380,1232,1258]" captionStart="FIGURE 8 A – D" captionStartId="13.[151,255,1903,1927]" captionTargetBox="[226,1360,178,1878]" captionTargetId="figure@13.[226,1360,178,1879]" captionTargetPageId="13" captionText="FIGURE 8 A – D. Ibericancer sanchoi n. gen., n. sp.; paratypes: A. partial ventral view of a male, MGSB 68574 c; the arrow indicating the male gonopore on P 5 coxa; B. partial carapace in dorsal view, MGSB 68574 d, showing the complete front and orbit; C. oblique lateral view of the same specimen; the arrow indicating the infraorbital protuberance; D. oblique partial view, MGSB 68574 e, with detail of reduced P 4 and P 5." httpUri="https://zenodo.org/record/184523/files/figure.png" pageId="17" pageNumber="18">8D</figureCitation>
|
||
,
|
||
<figureCitation id="13777BC8EF2B3F7B027AFB47FF1BD407" captionStart="FIGURE 9 A – C" captionStartId="14.[151,255,1919,1943]" captionTargetBox="[226,1360,194,1894]" captionTargetId="figure@14.[226,1360,194,1895]" captionTargetPageId="14" captionText="FIGURE 9 A – C. Ibericancer sanchoi n. gen., n. sp.; A. paratype MGSB 68574 a, dorsal view; B. holotype MGSB 68572, posterior view showing the concavity of the posterior margin; C. detail of P 1 - P 5 in the same specimen of A." httpUri="https://zenodo.org/record/184524/files/figure.png" pageId="17" pageNumber="18">9A, C</figureCitation>
|
||
). If the crab conceals itself by holding something (shell, sponge, algae) over its carapace, a burrow is unnecessary.
|
||
<collectingCountry id="F35B27DDEF2B3F7B07E6FAB7FEBBD42F" box="[237,270,1312,1338]" name="American Samoa" pageId="17" pageNumber="18">As</collectingCountry>
|
||
the dactyl of the leg used for carrying, which would be hooked or form a subchelate end to hold the object, is not preserved in the Dakoticancroidea, there is no proof of such a grasping by these pereiopods.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF2B3F7B07CEFAE7FB65D7B7" blockId="17.[151,1437,591,2018]" pageId="17" pageNumber="18">
|
||
Our interpretation of functional morphology nevertheless provides a clear sign for camouflage by the hind leg(s) in dakoticancroid crabs, as in most other podotremes (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B045CFA0FFB84D4A7" author="Guinot" box="[855,1073,1432,1458]" pageId="17" pageNumber="26" refString="Guinot, D., Doumenc, D. & Chintiroglou, C. C. (1995) A review of the carrying behaviour in brachyuran crabs, with additional information on the symbioses with sea anemones. The Raffles Bulletin of Zoology, 43, 377 - 416." type="journal article" year="1995">
|
||
Guinot
|
||
<emphasis id="B938BB5FEF2B3F7B04BAFA0FFC51D4A4" box="[945,996,1432,1457]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. 1995
|
||
</bibRefCitation>
|
||
). It is a conservative behaviour actually found in all families with the exception of the extant
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B047BFA57FBADD4CF" box="[880,1048,1472,1498]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Dynomenidae</taxonomicName>
|
||
, which live in corals or on rocky bottoms (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B060FFA7FFE15D717" author="McLay" box="[260,416,1512,1538]" pageId="17" pageNumber="26" refString="McLay, C. L. (1999) Crustacea Decapoda: revision of the family Dynomenidae. In: Crosnier (Ed.), Resultats des Campagnes MUSORSTOM Volume 20. Memoires du Museum national d'Histoire naturelle, 180, 427 - 569" type="journal article" year="1999">McLay 1999</bibRefCitation>
|
||
), and of the
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B053BFA7FFD1FD717" box="[560,682,1512,1538]" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Raninidae</taxonomicName>
|
||
, which bury themselves in sand. Among the Cyclodorippoidea a carrying behaviour using sea urchin spines, pieces of shell and bits of seaweed has been noted for the
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B079CF9AFFEEFD747" box="[151,346,1592,1618]" class="Malacostraca" family="Cyclodorippidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Cyclodorippidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B069CF9AFFD43D747" authority="Bouvier, 1897" authorityName="Bouvier" authorityYear="1897" box="[407,758,1592,1618]" class="Malacostraca" family="Cymonomidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">
|
||
Cymonomidae
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0546F9AFFD43D747" author="Bouvier" box="[589,758,1592,1618]" pageId="17" pageNumber="25" refString="Bouvier, E. - L. (1897) Observations sur les crabes de la famille des Dorippides. Comptes Rendus de l'Academie des Sciences, Paris, 125 (20), 784 - 787." type="journal article" year="1897">Bouvier, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B040DF9AFFC27D747" author="Garth" box="[774,914,1592,1618]" pageId="17" pageNumber="26" refString="Garth, J. S. (1946) Littoral brachyuran fauna of the Galapagos Archipelago. Allan Hancock Pacific Expeditions, 5 (10), i - iv + 341 - 600." type="journal article" year="1946">Garth 1946</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B04AAF9AFFBE0D747" author="Wicksten" box="[929,1109,1592,1618]" pageId="17" pageNumber="27" refString="Wicksten, M. K. (1982) Behavior in Clythrocerus planus (Rathbun, 1900) (Brachyura, Dorippidae). Crustaceana, 43, 306 - 308." type="journal article" year="1982">Wicksten 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0368F9AFFAB3D747" author="Tavares" box="[1123,1286,1592,1618]" pageId="17" pageNumber="27" refString="Tavares, M. (1994) Brachyoures bathyaux recoltes par le " Marion Dufresne " au large du Bresil. Systematique et phylogenie des Cyclodorippoidea mondiaux (Crustacea, Decapoda, Brachyura), 342 pp. Unpublished Doctoral Thesis, Universite Paris 6, Oceanologie Biologique." type="book" year="1994">Tavares 1994</bibRefCitation>
|
||
). A carrying behaviour is probable, perhaps combined with a burying behaviour (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B04BFF9F7FAB0D76F" author="Guinot" box="[948,1285,1632,1658]" pageId="17" pageNumber="26" refString="Guinot, D. & Tavares, M. (2001) Une nouvelle famille de crabes du Cretace, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema, 23, 507 - 546." type="journal article" year="2001">Guinot & Tavares 2001: 529</bibRefCitation>
|
||
) in the
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B0256F9F7FE52D7B7" authority="Tavares, 1998" authorityName="Tavares" authorityYear="1998" class="Malacostraca" family="Phyllotymolinidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Phyllotymolinidae Tavares, 1998</taxonomicName>
|
||
, which similarly has reduced subchelate and mobile P4 and P5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35634C6EF2B3F7807CEF927FDBAD37F" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="reference_group">
|
||
<paragraph id="8BF3674DEF2B3F7807CEF927FB45D017" blockId="17.[151,1437,591,2018]" lastBlockId="18.[151,1437,152,618]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B07CEF927FE47D7DC" ID-CoL="4f9190af-7f4e-40d0-8bd4-4a306cf8ad91" box="[197,498,1712,1737]" class="Malacostraca" family="Dakoticancridae" genus="Avitelmessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="grapsoideus">
|
||
<emphasis id="B938BB5FEF2B3F7B07CEF927FE47D7DC" box="[197,498,1712,1737]" italics="true" pageId="17" pageNumber="18">Avitelmessus grapsoideus</emphasis>
|
||
</taxonomicName>
|
||
, a dakoticancrid with P5 probably quite contrasting to the thick and long P4, is an unusual crab on account of its large size (a giant among North American Cretaceous crabs) and its flattened and ornamented carapace. It was supposed to live “by tightly pressing its flattened body against the mud-sand bottoms” (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0619F8BFFE47D657" author="Bishop" box="[274,498,1832,1858]" pageId="17" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2B3F7B0664F8BFFE16D654" box="[367,419,1832,1857]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
: 249), a behaviour perhaps not compatible with carrying behaviour. Pereiopods are not preserved in the small and highly sculptured
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B042AF8C7FC77D67C" ID-CoL="531cfc0b-40a9-4a55-b008-8e11b354cf51" box="[801,962,1872,1897]" class="Malacostraca" family="Dakoticancridae" genus="Tetracarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF2B3F7B042AF8C7FC77D67C" box="[801,962,1872,1897]" italics="true" pageId="17" pageNumber="18">Tetracarcinus</emphasis>
|
||
</taxonomicName>
|
||
and in the poorly known
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B020DF8C7FAD4D67C" box="[1286,1377,1872,1897]" class="Malacostraca" family="Dakoticancridae" genus="Seorsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF2B3F7B020DF8C7FAD4D67C" box="[1286,1377,1872,1897]" italics="true" pageId="17" pageNumber="18">Seorsus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B079CF8EFFEC6D687" author="Bishop" box="[151,371,1912,1938]" pageId="17" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF2B3F7B07F9F8EFFE90D684" box="[242,293,1912,1937]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C4C1CCEEF2B3F7B06A3F8EFFD21D684" ID-CoL="643ac661-6d3f-4be0-a28e-a0d2b5e4d4a2" box="[424,660,1912,1937]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF2B3F7B06A3F8EFFD21D684" box="[424,660,1912,1937]" italics="true" pageId="17" pageNumber="18">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B0624EF2B3F7B0597F8E0FD5BD684" box="[668,750,1911,1937]" pageId="17" pageNumber="18" rank="genus">
|
||
<emphasis id="B938BB5FEF2B3F7B0597F8E0FD5BD684" bold="true" box="[668,750,1911,1937]" pageId="17" pageNumber="18">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B938BB5FEF2B3F7B05F6F8E0FCF5D684" bold="true" box="[765,832,1911,1937]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
the reduction and dorsal position of the P4 and P5, which probably could have been positioned over the carapace, suggest the possibility of a carrying behaviour by both P4 and P5, similar to that found in extant Podotremata (
|
||
<bibRefCitation id="EFDD1ABCEF2B3F7B0496F85FFBCAD6F7" author="Guinot" box="[925,1151,1992,2018]" pageId="17" pageNumber="26" refString="Guinot, D., Doumenc, D. & Chintiroglou, C. C. (1995) A review of the carrying behaviour in brachyuran crabs, with additional information on the symbioses with sea anemones. The Raffles Bulletin of Zoology, 43, 377 - 416." type="journal article" year="1995">
|
||
Guinot
|
||
<emphasis id="B938BB5FEF2B3F7B04F1F85FFB9BD6F4" box="[1018,1070,1992,2017]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. 1995
|
||
</bibRefCitation>
|
||
). The reduced P5 could have only been involved in a carrying behaviour among dakoticancrids. This was suggested for
|
||
<taxonomicName id="4C4C1CCEEF283F7803F7FF0FFF42D1CC" ID-CoL="cdc0c084-90c4-482f-85b7-7807435b8e40" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF283F7803F7FF0FFF42D1CC" italics="true" pageId="18" pageNumber="19">Dakoticancer overana</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFDD1ABCEF283F78062FFF57FE44D1CF" author="Glaessner" box="[292,497,192,218]" pageId="18" pageNumber="26" refString="Glaessner, M. F. (1960) The fossil decapod Crustacea of New Zealand and the evolution of the Order Decapoda. New Zealand Geological Survey, Paleontological Bulletin, 31, 1 - 79." type="journal article" year="1960">Glaessner (1960)</bibRefCitation>
|
||
. The thickness of the body, and the fact that the P4 and P5 are reduced, indicate that the
|
||
<taxonomicName id="4C4C1CCEEF283F7807FEFF7FFE28D017" ID-CoL="629869d8-410d-4b3e-8aea-a8f9c635e382" box="[245,413,232,258]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
|
||
<emphasis id="B938BB5FEF283F7806AFFF70FE4ED014" bold="true" box="[420,507,231,257]" pageId="18" pageNumber="19">n. fam.</emphasis>
|
||
may have had a different mode of life than the
|
||
<taxonomicName id="4C4C1CCEEF283F780323FF7FFB59D017" ID-CoL="377b5adf-9ed1-498f-8aef-7a7c4133e597" box="[1064,1260,232,258]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF283F7807CEFE87FDBAD37F" blockId="18.[151,1437,152,618]" pageId="18" pageNumber="19">
|
||
The Dakoticancroidea is, to our knowledge, one of the first fossil brachyuran groups in which a carrying behaviour may be inferred from reasonably certain morphological characters, that is, the last pereiopod(s) that are nearly entirely preserved. Fossil crabs with preserved last pereiopod(s) are rare and only a few homolids have so far been recorded (see
|
||
<bibRefCitation id="EFDD1ABCEF283F78051AFE1FFD12D0B7" author="Larghi" box="[529,679,392,418]" pageId="18" pageNumber="26" refString="Larghi, C. (2004) Brachyuran decapod Crustacea from the Upper Cretaceous of Lebanon. Journal of Paleontology, 78, 528 - 541." type="journal article" year="2004">Larghi 2004</bibRefCitation>
|
||
: 529). It is still a matter of conjecture to suppose that carrying behaviour was a regular phenomenon in fossil Homoloidea, as in extant representatives (as well as in extant homolodromioids) since the last pereiopods are very poorly preserved in fossil record. The Cenomanian
|
||
<taxonomicName id="4C4C1CCEEF283F78079CFD97FDA4D30F" authority="Larghi, 2004" authorityName="Larghi" authorityYear="2004" box="[151,529,512,538]" class="Malacostraca" family="Necrocarcinidae" genus="Corazzatocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF283F78079CFD97FEC6D30C" box="[151,371,512,537]" italics="true" pageId="18" pageNumber="19">Corazzatocarcinus</emphasis>
|
||
<bibRefCitation id="EFDD1ABCEF283F780671FD97FDA4D30F" author="Larghi" box="[378,529,512,538]" pageId="18" pageNumber="26" refString="Larghi, C. (2004) Brachyuran decapod Crustacea from the Upper Cretaceous of Lebanon. Journal of Paleontology, 78, 528 - 541." type="journal article" year="2004">Larghi, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDD1ABCEF283F780514FD97FCD1D30F" author="Larghi" box="[543,868,512,538]" pageId="18" pageNumber="26" refString="Larghi, C. (2004) Brachyuran decapod Crustacea from the Upper Cretaceous of Lebanon. Journal of Paleontology, 78, 528 - 541." type="journal article" year="2004">Larghi 2004: 530, figs. 2–4</bibRefCitation>
|
||
), assumed to belong to the Podotremata (
|
||
<bibRefCitation id="EFDD1ABCEF283F780248FD97FF48D357" author="Vega" pageId="18" pageNumber="27" refString="Vega, F. J., Nyborg, T., Rojas, A., Patarroyo, P., Luque, J., Porras Muzquiz, H. & Stinnesbeck, W. (2007) Upper Cretaceous Crustacea from Mexico and Colombia: similar faunas and evironments during Turonian times. Revista Mexicana de Ciencias Geologicas, 24, 403 - 422." type="journal article" year="2007">
|
||
Vega
|
||
<emphasis id="B938BB5FEF283F78028CFD97FF1BD354" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
), also shows clearly visible, subdorsal and reduced P4 and P5. Its assignment to a particular podotreme family is still problematic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |