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<mods:title id="05610F8392D4A2DB46CFCA121FC69A83">Description of two new species of Arcania Leach, 1817, from the western Pacific Ocean and redescriptions of A. undecimspinosa De Haan, 1841, and A. elongata Yokoya, 1933 (Crustacea: Brachyura: Leucosiidae)</mods:title>
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<taxonomicName id="4C354D3F381CD63B97B92EB0FDCECECA" ID-CoL="G7CC" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[151,620,691,717]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="undecimspinosa">
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<emphasis id="B941EAAE381CD63B97B92EB0FE14CECB" bold="true" box="[151,438,691,716]" italics="true" pageId="1" pageNumber="302">Arcania undecimspinosa</emphasis>
De Haan, 1841
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(
<figureCitation id="130E2A39381CD63B97B12ED4FEABCEF7" box="[159,265,727,752]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4. G 1 and G 2" captionStart-4="FIGURE 5. G 1" captionStart-5="FIGURE 6" captionStartId-0="1.[151,250,1920,1942]" captionStartId-1="2.[151,250,1689,1711]" captionStartId-2="3.[151,250,1316,1338]" captionStartId-3="4.[151,250,1820,1842]" captionStartId-4="5.[151,250,1769,1791]" captionStartId-5="6.[151,250,1363,1385]" captionTargetBox-0="[151,1436,1236,1898]" captionTargetBox-1="[151,1436,193,1667]" captionTargetBox-2="[151,1436,625,1293]" captionTargetBox-3="[198,1393,597,1786]" captionTargetBox-4="[325,1261,495,1747]" captionTargetBox-5="[193,1391,529,1336]" captionTargetId-0="figure@1.[151,1436,1236,1899]" captionTargetId-1="figure@2.[151,1436,193,1668]" captionTargetId-2="figure@3.[151,1436,625,1294]" captionTargetId-3="figure@4.[183,1404,589,1799]" captionTargetId-4="figure@5.[315,1272,483,1747]" captionTargetId-5="figure@6.[188,1398,517,1342]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionText-0="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." captionText-1="FIGURE 2. Arcania undecimspinosa De Haan, 1841. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view." captionText-2="FIGURE 3. Arcania undecimspinosa De Haan, 1841. ZRC 2002.0495, male, 31.0 × 29.8 mm, Qindao, China; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." captionText-3="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." captionText-4="FIGURE 5. G 1 of Arcania undecimspinosa De Haan, 1841. ZRC 2002.0495, male, 31.0 × 29.8 mm, Qindao, China; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventromesial view. Scales = 1 mm." captionText-5="FIGURE 6. Right vulva of Arcania undecimspinosa De Haan, 1841; a, NSMT-Cr 3839, 24.5 × 23.3 mm, Mikawa Bay, Japan; b, NSMT-Cr 4397, 28.2 × 27.0 mm, Kochi, Japan. Scale = 1 mm." httpUri-0="https://zenodo.org/record/228872/files/figure.png" httpUri-1="https://zenodo.org/record/228873/files/figure.png" httpUri-2="https://zenodo.org/record/228874/files/figure.png" httpUri-3="https://zenodo.org/record/228875/files/figure.png" httpUri-4="https://zenodo.org/record/228876/files/figure.png" httpUri-5="https://zenodo.org/record/228877/files/figure.png" pageId="1" pageNumber="302">Figs. 1–6</figureCitation>
)
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<taxonomicName id="4C354D3F381CD63B97B92F22FDD9CF31" ID-CoL="G7CC" authority="De Haan, 1841: 135" authorityName="De Haan" authorityPageNumber="135" authorityYear="1841" box="[151,635,800,822]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381CD63B97B92F22FE37CF31" box="[151,405,800,822]" italics="true" pageId="1" pageNumber="302">Arcania undecimspinosa</emphasis>
De Haan, 1841: 135
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, pl. 33, fig. 8.— 
<treatmentCitation id="0A9410AD381CD63B941F2F23FC7DCF31" author="Bell" box="[817,991,800,822]" page="367" pageId="1" pageNumber="302" year="1855">
<bibRefCitation id="EFA44B4D381CD63B941F2F23FC0BCF31" author="Bell" box="[817,937,800,822]" pageId="1" pageNumber="302" refString="Bell, T. (1855 a) Horae carcinologicae, or notices of Crustacea. I. A monograph of the Leucosiadae. Annals and Magazine of Natural History, 16, 361 - 367." type="journal article" year="1855" yearSuffix="a">Bell 1855a</bibRefCitation>
: 367
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; 1855b: 309; 1855c: 21.— 
<treatmentCitation id="0A9410AD381CD63B93D32F23FF42CF53" author="Herklots" page="28" pageId="1" pageNumber="302" year="1861">
<bibRefCitation id="EFA44B4D381CD63B93D32F23FA3ACF31" author="Herklots" box="[1277,1432,800,822]" pageId="1" pageNumber="302" refString="Herklots, J. A. (1861) Symbolae carcinologicae. Etudes sur la classe des Crustaces. I. Catalogue des Crustaces qui ont servi de base au systeme carcinologique de M. W. de Haan, redige d'apres la collection du Musee des Pays-Bas et les Crustaces de la faune du Japon. Tijdschrift voor Entomologie, 4, 116 - 156." type="journal article" year="1861">Herklots 1861</bibRefCitation>
: 28
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.— 
<treatmentCitation id="0A9410AD381CD63B97D12F3DFE34CF53" author="Ives" box="[255,406,830,852]" page="216" pageId="1" pageNumber="302" year="1891">
<bibRefCitation id="EFA44B4D381CD63B97D12F3DFEC4CF53" author="Ives" box="[255,358,830,852]" pageId="1" pageNumber="302" refString="Ives, J. E. (1891) Echinoderms and arthropods from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia, 43, 210 - 223, pls. 7 - 12." type="journal article" year="1891">Ives 1891</bibRefCitation>
: 216
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.— 
<treatmentCitation id="0A9410AD381CD63B969A2F3DFDF6CF53" author="Shen" box="[436,596,830,852]" page="107" pageId="1" pageNumber="302" year="1931">
<bibRefCitation id="EFA44B4D381CD63B969A2F3DFD86CF53" author="Shen" box="[436,548,830,852]" pageId="1" pageNumber="302" refString="Shen, C. J. (1931) The crabs of Hong Kong. Part I. The Hong Kong Naturalist, 2, 92 - 110, pls. 4 - 10." type="journal article" year="1931">Shen 1931</bibRefCitation>
: 107
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, pl. 10, fig. 1.— 
<treatmentCitation id="0A9410AD381CD63B95DD2F3DFC3FCF53" author="Sakai" box="[755,925,830,852]" page="123" pageId="1" pageNumber="302" year="1937">
<bibRefCitation id="EFA44B4D381CD63B95DD2F3DFCCECF53" author="Sakai" box="[755,876,830,852]" pageId="1" pageNumber="302" refString="Sakai, T. (1937) Studies on the crabs of Japan. II. Oxystomata. Science Reports of the Tokyo Bunrika Daigaku, Section B, Supplement No. 2, 3, 67 - 192, pls. 10 - 19." type="journal article" year="1937">Sakai 1937</bibRefCitation>
: 123
</treatmentCitation>
, fig. 15a, pl. 14, fig. 2; 1965: 40 (part?), pl. 16, fig. 2; 1976: 91, pl. 28, fig. 1.— 
<bibRefCitation id="EFA44B4D381CD63B96F22F5EFDD7CF74" author="Kamita" box="[476,629,861,883]" pageId="1" pageNumber="302" refString="Kamita, T. (1941 b) Studies of the Decapod Crustaceans of Chosen. Pt. I. Crabs. The Fisheries Society of Chosen, Keijo, 2 pls. + 2 + 2 + 14 + 289, 1 map." type="book" year="1941" yearSuffix="b">
<collectingCountry id="F322762C381CD63B96F22F5EFD8ECF74" box="[476,556,861,883]" name="Egypt" pageId="1" pageNumber="302">Kamita</collectingCountry>
1941b
</bibRefCitation>
, 40, fig. 13.— 
<treatmentCitation id="0A9410AD381CD63B942C2F5EFB81CF74" author="Holthuis" box="[770,1059,861,883]" page="119" pageId="1" pageNumber="302" year="1970">
<bibRefCitation id="EFA44B4D381CD63B942C2F5EFC50CF74" author="Holthuis" box="[770,1010,861,883]" pageId="1" pageNumber="302" refString="Holthuis, L. B. &amp; Sakai, T. (1970) Ph. F. von Siebold and Fauna Japonica: A history of early Japanese zoology. Academic Press of Japan, Tokyo, 323 pp + 38 color plates + 1 map." type="book" year="1970">Holthuis &amp; Sakai 1970</bibRefCitation>
: 119
</treatmentCitation>
, pl. 11, fig. 2.— 
<treatmentCitation id="0A9410AD381CD63B93E82F5EFADACF74" author="Miyake" box="[1222,1400,861,883]" page="60" pageId="1" pageNumber="302" year="1983">
<bibRefCitation id="EFA44B4D381CD63B93E82F5EFAF1CF74" author="Miyake" box="[1222,1363,861,883]" pageId="1" pageNumber="302" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color. Vol. II Brachyura (Crabs). Hoikusha, Osaka, 277 pp. [in Japanese]" type="book" year="1983">Miyake 1983</bibRefCitation>
: 60
</treatmentCitation>
, pl. 20, fig. 6.— 
<treatmentCitation id="0A9410AD381CD63B96112F7FFE5DCF95" author="Dai" box="[319,511,892,914]" page="66" pageId="1" pageNumber="302" year="1986">
<bibRefCitation id="EFA44B4D381CD63B96112F7FFE79CF95" author="Dai" box="[319,475,892,914]" pageId="1" pageNumber="302" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of Chinese Seas. China Ocean Press, Beijing, xvii + 642 pp., 74 pls. [in Chinese]" type="book" year="1986">
Dai 
<emphasis id="B941EAAE381CD63B96422F7EFE3CCF95" box="[364,414,892,914]" italics="true" pageId="1" pageNumber="302">et al.</emphasis>
1986
</bibRefCitation>
: 66
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, fig. 33-1, pl. 8, fig. 1.— 
<treatmentCitation id="0A9410AD381CD63B95DA2F7FFC08CF95" author="Huang" box="[756,938,892,914]" page="305" pageId="1" pageNumber="302" year="1989">
<bibRefCitation id="EFA44B4D381CD63B95DA2F7FFCDBCF95" author="Huang" box="[756,889,892,914]" pageId="1" pageNumber="302" refString="Huang, L. (1989) Oxystomata. In: Wei, C. &amp; Chen, Y. (Eds.), Fauna of Zhejiang. Crustacea. Zhejiang Science and Technology Publishing House, Hangzhou, pp. 294 - 324. [in Chinese]" type="book chapter" year="1989">Huang 1989</bibRefCitation>
: 305
</treatmentCitation>
, fig. 267.— 
<treatmentCitation id="0A9410AD381CD63B930F2F7FFB59CF95" author="Dai" box="[1057,1275,892,914]" page="73" pageId="1" pageNumber="302" year="1991">
<bibRefCitation id="EFA44B4D381CD63B930F2F7FFB75CF95" author="Dai" box="[1057,1239,892,914]" pageId="1" pageNumber="302" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China Sea. China Ocean Press, Beijing and Springer-Verlag, Berlin, Heidelberg, New York, Tokyo, 21 + 608 pp. + 74 pls." type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: 73
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, fig. 33-1, pl. 8, fig. 1.—Yamaguchi 1993: pl. 21c.— 
<treatmentCitation id="0A9410AD381CD63B95182F99FCD1CFB7" author="Yamaguchi" box="[566,883,922,944]" page="318" pageId="1" pageNumber="302" year="1993">
<bibRefCitation id="EFA44B4D381CD63B95182F99FCE3CFB7" author="Yamaguchi" box="[566,833,922,944]" pageId="1" pageNumber="302" refString="Yamaguchi, T. &amp; Baba, K. (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. In: Yamaguchi, T. (Ed.), Ph. F. von Siebold and Natural History of Japan, Crustacea. The Carcinological Society of Japan, Tokyo, pp. 145 - 570. [in Japanese with English summary and figure caption]" type="book chapter" year="1993">Yamaguchi &amp; Baba 1993</bibRefCitation>
: 318
</treatmentCitation>
, fig. 101; 2003: 33.— 
<treatmentCitation id="0A9410AD381CD63B937A2F99FB54CFB7" author="Galil" box="[1108,1270,922,944]" page="197" pageId="1" pageNumber="302" year="2001">
<bibRefCitation id="EFA44B4D381CD63B937A2F99FB64CFB7" author="Galil" box="[1108,1222,922,944]" pageId="1" pageNumber="302" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil 2001</bibRefCitation>
: 197
</treatmentCitation>
(part), figs. 3D, 7D (part).— 
<treatmentCitation id="0A9410AD381CD63B966E2FBAFD84CFC8" author="Takeda" box="[320,550,953,975]" page="37" pageId="1" pageNumber="302" year="2011">
<bibRefCitation id="EFA44B4D381CD63B966E2FBAFDA3CFC8" author="Takeda" box="[320,513,953,975]" pageId="1" pageNumber="302" refString="Takeda, M., Furuta, S., Miyanaga, T., Tamura, A. &amp; Wada, T. (2011) Crabs from the southwestern Sea of Japan along Tottori Prefecture and its vicinity, Japan. Bulletin of the Tottori Prefectural Museum, 48, 29 - 94. [in Japanese with English abstract]" type="journal article" year="2011">
Takeda 
<emphasis id="B941EAAE381CD63B96BD2FB9FE64CFC8" box="[403,454,953,975]" italics="true" pageId="1" pageNumber="302">et al.</emphasis>
2011
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: 37
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, fig. 9–26).
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</paragraph>
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<emphasis id="B941EAAE381CD63B97B92FDAFE36CFE9" box="[151,404,985,1006]" italics="true" pageId="1" pageNumber="302">
<taxonomicName id="4C354D3F381CD63B97B92FDAFF4ACFE9" ID-CoL="7NQTF" box="[151,232,985,1006]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">Arcania</taxonomicName>
novemspinosa—
</emphasis>
<treatmentCitation id="0A9410AD381CD63B96BA2FDBFD8BCFE9" author="Hill" box="[404,553,984,1006]" page="201" pageId="1" pageNumber="302" year="1982">
<bibRefCitation id="EFA44B4D381CD63B96BA2FDBFE55CFE9" author="Hill" box="[404,503,984,1006]" pageId="1" pageNumber="302" refString="Hill, D. S. (1982) The Leucosiidae (Crustacea: Decapoda) of Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong, Hong Kong University Press, pp. 195 - 205." type="book chapter" year="1982">Hill 1982</bibRefCitation>
: 201
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, pl. 4D.— 
<treatmentCitation id="0A9410AD381CD63B95BF2FDBFCF1CFE9" author="Dai" box="[657,851,984,1006]" page="66" pageId="1" pageNumber="302" year="1986">
<bibRefCitation id="EFA44B4D381CD63B95BF2FDBFC8FCFE9" author="Dai" box="[657,813,984,1006]" pageId="1" pageNumber="302" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of Chinese Seas. China Ocean Press, Beijing, xvii + 642 pp., 74 pls. [in Chinese]" type="book" year="1986">
Dai 
<emphasis id="B941EAAE381CD63B95902FDAFD53CFE9" box="[702,753,984,1006]" italics="true" pageId="1" pageNumber="302">et al.</emphasis>
1986
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: 66
</treatmentCitation>
, fig. 33-2, 3; 
<treatmentCitation id="0A9410AD381CD63B94F52FDBFB14CFE9" author="Dai" box="[987,1206,984,1006]" page="74" pageId="1" pageNumber="302" year="1991">
<bibRefCitation id="EFA44B4D381CD63B94F52FDBFB32CFE9" author="Dai" box="[987,1168,984,1006]" pageId="1" pageNumber="302" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China Sea. China Ocean Press, Beijing and Springer-Verlag, Berlin, Heidelberg, New York, Tokyo, 21 + 608 pp. + 74 pls." type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: 74
</treatmentCitation>
, fig. 33-2, 3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32F6537381CD63997B92819FAE0CE3A" lastPageId="3" lastPageNumber="304" pageId="1" pageNumber="302" type="materials_examined">
<paragraph id="8B8A36BC381CD63B97B92819FED4C850" blockId="1.[151,1436,1050,1183]" pageId="1" pageNumber="302">
<emphasis id="B941EAAE381CD63B97B92819FE22C834" bold="true" box="[151,384,1050,1075]" pageId="1" pageNumber="302">Material examined.</emphasis>
<typeStatus id="548E881E381CD63B96A92818FE59C834" box="[391,507,1051,1075]" pageId="1" pageNumber="302" type="lectotype">Lectotype</typeStatus>
: 
<collectionCode id="ED24AE79381CD63B95282818FDC3C834" LSID="urn:lsid:biocol.org:col:34992" box="[518,609,1051,1075]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="302">RMNH</collectionCode>
. 
<collectionCode id="ED24AE79381CD63B954A2818FD13C834" box="[612,689,1051,1075]" pageId="1" pageNumber="302">CRUS</collectionCode>
.D.790, 
<specimenCount id="9D33FD35381CD63B942B2818FCF1C834" box="[773,851,1050,1075]" pageId="1" pageNumber="302" type="male">1 male</specimenCount>
, 16.5 × 
<quantity id="4CCD9B59381CD63B94802819FBB6C835" box="[942,1044,1050,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="302" unit="mm" value="16.5">16.5 mm</quantity>
, Nagasaki, Kyushu, 
<collectingCountry id="F322762C381CD63B93D52818FA9CC835" box="[1275,1342,1051,1074]" name="Japan" pageId="1" pageNumber="302">Japan</collectingCountry>
, coll. H. Bürger, 
<date id="FF8B107C381CD63B97DF283DFED1C850" box="[241,371,1086,1111]" pageId="1" pageNumber="302" value="1825" valueMax="1834">1825–1834</date>
.
</paragraph>
<paragraph id="8B8A36BC381CD63B97E92861FDF8C898" blockId="1.[151,1436,1050,1183]" pageId="1" pageNumber="302">
<typeStatus id="548E881E381CD63B97E92861FECDC87C" box="[199,367,1122,1147]" pageId="1" pageNumber="302" type="paralectotype">Paralectotypes</typeStatus>
: 
<collectionCode id="ED24AE79381CD63B96572861FE71C87C" LSID="urn:lsid:biocol.org:col:34992" box="[377,467,1122,1147]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="302">RMNH</collectionCode>
. 
<collectionCode id="ED24AE79381CD63B96F92861FD81C87C" box="[471,547,1122,1147]" pageId="1" pageNumber="302">CRUS</collectionCode>
.D.56126, 
<specimenCount id="9D33FD35381CD63B95BA2860FD5AC87C" box="[660,760,1122,1147]" pageId="1" pageNumber="302" type="female">1 female</specimenCount>
, 19.2 × 
<quantity id="4CCD9B59381CD63B947D2860FC18C87D" box="[851,954,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="1" pageNumber="302" unit="mm" value="19.2">19.2 mm</quantity>
, 3 individuals (sex unknown), 20.3 × 19.9, 21.1 × 
<quantity id="4CCD9B59381CD63B97CB2884FEE9C899" box="[229,331,1159,1183]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="1" pageNumber="302" unit="mm" value="21.1">21.1 mm</quantity>
, data same as 
<typeStatus id="548E881E381CD63B96C22885FDF5C898" box="[492,599,1158,1183]" pageId="1" pageNumber="302" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<caption id="DF4A6634381CD63B97B92B83FC0ACBB3" httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="1" pageNumber="302" targetBox="[151,1436,1236,1898]" targetPageId="1">
<paragraph id="8B8A36BC381CD63B97B92B83FC0ACBB3" blockId="1.[151,1436,1920,1972]" pageId="1" pageNumber="302">
<emphasis id="B941EAAE381CD63B97B92B83FEB5CB91" bold="true" box="[151,279,1920,1942]" pageId="1" pageNumber="302">FIGURE 1.</emphasis>
<taxonomicName id="4C354D3F381CD63B960E2B82FD6ECB91" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[288,716,1920,1942]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381CD63B960E2B82FDBCCB91" box="[288,542,1920,1942]" italics="true" pageId="1" pageNumber="302">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
. Lectotype. RMNH.CRUS.D.790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view.
</paragraph>
</caption>
<caption id="DF4A6634381FD63897B92A9AFD20CAC9" httpUri="https://zenodo.org/record/228873/files/figure.png" pageId="2" pageNumber="303" targetBox="[151,1436,193,1667]" targetPageId="2">
<paragraph id="8B8A36BC381FD63897B92A9AFD20CAC9" blockId="2.[151,1436,1689,1742]" pageId="2" pageNumber="303">
<emphasis id="B941EAAE381FD63897B92A9AFEB4CAA8" bold="true" box="[151,278,1689,1711]" pageId="2" pageNumber="303">FIGURE 2.</emphasis>
<taxonomicName id="4C354D3F381FD63896312A99FD6ACAA8" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[287,712,1689,1711]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381FD63896312A99FDBECAA8" box="[287,540,1689,1711]" italics="true" pageId="2" pageNumber="303">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view.
</paragraph>
</caption>
<paragraph id="8B8A36BC381FD63997E92AF9FA23CD47" blockId="2.[151,1437,1786,2027]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="304" pageId="2" pageNumber="303">
Additional material. 
<collectingCountry id="F322762C381FD63896812AF8FE51CB15" box="[431,499,1787,1810]" name="Japan" pageId="2" pageNumber="303">
<emphasis id="B941EAAE381FD63896812AF8FE51CB15" box="[431,499,1787,1810]" italics="true" pageId="2" pageNumber="303">Japan</emphasis>
</collectingCountry>
: 
<collectionCode id="ED24AE79381FD638952C2AF8FDF1CB14" LSID="urn:lsid:biocol.org:col:34871" box="[514,595,1787,1811]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="303">USNM</collectionCode>
26280, 
<specimenCount id="9D33FD35381FD63895832AF8FD59CB14" box="[685,763,1786,1811]" pageId="2" pageNumber="303" type="male">1 male</specimenCount>
, 21.1 × 
<quantity id="4CCD9B59381FD638947B2AF8FC19CB15" box="[853,955,1787,1811]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="2" pageNumber="303" unit="mm" value="21.1">21.1 mm</quantity>
, 
<specimenCount id="9D33FD35381FD63894E82AF8FB88CB14" box="[966,1066,1786,1811]" pageId="2" pageNumber="303" type="female">1 female</specimenCount>
, 21.2 × 
<quantity id="4CCD9B59381FD63893AD2AF8FB4BCB15" box="[1155,1257,1787,1811]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="2" pageNumber="303" unit="mm" value="21.2">21.2 mm</quantity>
, 1 ovig. female, 19.9 × 
<quantity id="4CCD9B59381FD63897C62B1CFEF2CB31" box="[232,336,1823,1847]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="2" pageNumber="303" unit="mm" value="20.3">20.3 mm</quantity>
, Nagasaki, Hizen, Kyushu, coll. 
<collectingCountry id="F322762C381FD63895E92B1DFCB0CB30" box="[711,786,1822,1847]" name="Jordan" pageId="2" pageNumber="303">Jordan</collectingCountry>
&amp; Snyder, 1900.— 
<collectionCode id="ED24AE79381FD63894C02B1CFBE0CB30" LSID="urn:lsid:biocol.org:col:34871" box="[1006,1090,1823,1847]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="303">USNM</collectionCode>
45833, 1 ovig. female, 19.7 × 
<quantity id="4CCD9B59381FD63897B92B40FF5CCB5D" box="[151,254,1859,1883]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="2" pageNumber="303" unit="mm" value="20.2">20.2 mm</quantity>
, Nagasaki, Kyushu, exchange with Imperial Tokyo University.—NSMT-Cr 8371, 
<specimenCount id="9D33FD35381FD63893892B40FAAECB5C" box="[1191,1292,1858,1883]" pageId="2" pageNumber="303" type="female">1 female</specimenCount>
, 16.7 × 
<quantity id="4CCD9B59381FD63892452B41FF67CB79" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="303" unit="mm" value="16.2">16.2 mm</quantity>
, Ariake Sea, Kyushu, coll. 
<date id="FF8B107C381FD638952D2B64FDD0CB78" box="[515,626,1894,1919]" pageId="2" pageNumber="303" value="1958-09">Sep. 1958</date>
.—NSMT-Cr 8200, 
<specimenCount id="9D33FD35381FD63894752B64FC09CB78" box="[859,939,1894,1919]" pageId="2" pageNumber="303" type="male">1 male</specimenCount>
, 19.9 × 
<quantity id="4CCD9B59381FD63893222B64FBD7CB79" box="[1036,1141,1895,1919]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="2" pageNumber="303" unit="mm" value="19.1">19.1 mm</quantity>
, Ariake, Kyushu, coll. N. Yoshikawa, 
<date id="FF8B107C381FD638960D2B89FE1ECBA4" box="[291,444,1930,1955]" pageId="2" pageNumber="303" value="1982-06-15">15 June 1982</date>
.—NSMT-Cr 8009, 
<specimenCount id="9D33FD35381FD638958B2B88FD54CBA4" box="[677,758,1930,1955]" pageId="2" pageNumber="303" type="male">1 male</specimenCount>
, 16.5 × 
<quantity id="4CCD9B59381FD63894772B89FC60CBA5" box="[857,962,1930,1955]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="2" pageNumber="303" unit="mm" value="15.6">15.6 mm</quantity>
, off Jogajima, coll. E. Tsuchida, 
<date id="FF8B107C381FD638926D2B88FF6CCBC0" pageId="2" pageNumber="303" value="1982-03-11">11 Mar. 1982</date>
.— 
<collectionCode id="ED24AE79381FD63897C12BACFEE1CBC0" LSID="urn:lsid:biocol.org:col:34871" box="[239,323,1967,1991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="303">USNM</collectionCode>
1199103, 
<specimenCount id="9D33FD35381FD63896962BACFDA4CBC0" box="[440,518,1966,1991]" pageId="2" pageNumber="303" type="male">1 male</specimenCount>
, Shizuoka, Suruga Bay, Seno Umi, 
<emphasis id="B941EAAE381FD63894BA2BADFC5CCBC0" box="[916,1022,1966,1991]" italics="true" pageId="2" pageNumber="303">Albatross</emphasis>
, stn. D. 3702, 
<date id="FF8B107C381FD638938F2BACFA85CBC0" box="[1185,1319,1967,1991]" pageId="2" pageNumber="303" value="1900-05-07">7 May 1900</date>
.— 
<collectionCode id="ED24AE79381FD63892662BACFA3ECBC0" LSID="urn:lsid:biocol.org:col:34871" box="[1352,1436,1967,1991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="303">USNM</collectionCode>
63676, 1 ovig. female, 26.3 × 
<quantity id="4CCD9B59381FD63896C72BD1FDEDCBED" box="[489,591,2002,2027]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="2" pageNumber="303" unit="mm" value="25.9">25.9 mm</quantity>
, Misaki, 
<collectingCountry id="F322762C381FD63895992BD0FD58CBED" box="[695,762,2003,2026]" name="Japan" pageId="2" pageNumber="303">Japan</collectingCountry>
, coll. A.S. Pearse (bought from Kuma-san), 1930.—NSMT- Cr 3839, 
<specimenCount id="9D33FD35381ED639962A2C9BFEC9CCB7" box="[260,363,151,176]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 24.5 × 
<quantity id="4CCD9B59381ED63996E22C9BFD96CCA8" box="[460,564,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="304" unit="mm" value="23.3">23.3 mm</quantity>
, Mikawa-Isshiki, Mikawa Bay, coll. 
<date id="FF8B107C381ED63994F32C94FBD2CCB7" box="[989,1136,151,176]" pageId="3" pageNumber="304" value="1966-01-15">15 Jan. 1966</date>
.— 
<collectionCode id="ED24AE79381ED63993BF2C94FB47CCB7" LSID="urn:lsid:biocol.org:col:34871" box="[1169,1253,151,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
134198, 19.6 × 
<quantity id="4CCD9B59381ED63997B92CBFFF5CCCD3" box="[151,254,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="304" unit="mm" value="18.5">18.5 mm</quantity>
, 
<quantity id="4CCD9B59381ED63996252CBFFEF1CCD2" box="[267,339,188,213]" metricMagnitude="4" metricUnit="m" metricValue="1.41622272" pageId="3" pageNumber="304" unit="mi" value="8.8">8.8 mi</quantity>
off Nomasaki, Honshu Island, stn. Alb. D-3725, 13 faths. (ca. 
<quantity id="4CCD9B59381ED639930F2CBFFBD6CCD3" box="[1057,1140,188,212]" metricMagnitude="1" metricUnit="m" metricValue="2.38" pageId="3" pageNumber="304" unit="m" value="23.8">23.8 m</quantity>
), 
<date id="FF8B107C381ED63993A72CBFFA82CCD3" box="[1161,1312,188,213]" pageId="3" pageNumber="304" value="1900-05-15">15 May 1900</date>
.—RUMF- ZC-2574, 
<specimenCount id="9D33FD35381ED63996222CE3FED1CCFF" box="[268,371,223,248]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 12.6 × 
<quantity id="4CCD9B59381ED63996FB2CE3FD92CCF0" box="[469,560,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="304" unit="mm" value="9.9">9.9 mm</quantity>
, Takehara City, Hiroshima, 
<quantity id="4CCD9B59381ED639945A2CDCFC0ACCFF" box="[884,936,223,248]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="304" unit="m" value="15.0">15m</quantity>
, coll. D. Uyeno, 
<date id="FF8B107C381ED639935C2CE3FAAACCFF" box="[1138,1288,224,248]" pageId="3" pageNumber="304" value="2007-10-14">14 Oct. 2007</date>
.—NSMT-Cr 4397, 
<specimenCount id="9D33FD35381ED63997CD2D07FEE0CD1A" box="[227,322,260,285]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
, ca 25.9 × 
<quantity id="4CCD9B59381ED63996FA2D07FDE2CD1B" box="[468,576,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="3" pageNumber="304" unit="mm" value="25.1">25.1 mm</quantity>
, 
<specimenCount id="9D33FD35381ED639957F2D07FD19CD1A" box="[593,699,260,285]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 28.2 × 
<quantity id="4CCD9B59381ED63994092D07FC31CD1B" box="[807,915,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="3" pageNumber="304" unit="mm" value="27.2">27.2 mm</quantity>
, Mimase, 
<collectingCountry id="F322762C381ED639933D2D07FBF8CD1A" box="[1043,1114,260,285]" name="India" pageId="3" pageNumber="304">Kochi</collectingCountry>
, coll. Y. Koyama, 
<date id="FF8B107C381ED639926F2D07FF6FCD47" pageId="3" pageNumber="304" value="1973-05-11">11 May 1973</date>
.— 
<collectionCode id="ED24AE79381ED63997C02D2BFEE0CD47" LSID="urn:lsid:biocol.org:col:34871" box="[238,322,296,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
18873, 
<specimenCount id="9D33FD35381ED63996B02D2BFDA0CD47" box="[414,514,295,320]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 13.2 × 
<quantity id="4CCD9B59381ED63995722D2BFD60CD38" box="[604,706,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="304" unit="mm" value="13.1">13.1 mm</quantity>
, 
<collectingCountry id="F322762C381ED63995E32D2BFCB2CD38" box="[717,784,296,319]" name="Japan" pageId="3" pageNumber="304">Japan</collectingCountry>
, coll. Mr. Sakamoto (passed through Garrett Droppers).
</paragraph>
<paragraph id="8B8A36BC381ED63997E92D4EFC03CDAA" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="304">
<collectingCountry id="F322762C381ED63997E92D4EFEAECD63" box="[199,268,333,356]" name="South Korea" pageId="3" pageNumber="304">
<emphasis id="B941EAAE381ED63997E92D4EFEAECD63" box="[199,268,333,356]" italics="true" pageId="3" pageNumber="304">Korea</emphasis>
</collectingCountry>
: 
<collectionCode id="ED24AE79381ED63996332D4FFE28CD63" box="[285,394,332,356]" pageId="3" pageNumber="304">MADBK</collectionCode>
171803-017, 
<specimenCount id="9D33FD35381ED63995332D4FFDCFCD62" box="[541,621,332,357]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
, 30.3 × 
<quantity id="4CCD9B59381ED63995E12D4FFC95CD63" box="[719,823,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="3" pageNumber="304" unit="mm" value="28.7">28.7 mm</quantity>
, 
<specimenCount id="9D33FD35381ED639946B2D4FFC09CD62" box="[837,939,332,357]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 22.0 × 
<quantity id="4CCD9B59381ED63993222D4FFBD7CD63" box="[1036,1141,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="3" pageNumber="304" unit="mm" value="21.9">21.9 mm</quantity>
, Jinhae-gu, Changwon-si, Gyeongsangnam-do, 
<date id="FF8B107C381ED63996A62D73FDB4CD8F" box="[392,534,367,392]" pageId="3" pageNumber="304" value="1968-08-01">1 Aug. 1968</date>
.— 
<collectionCode id="ED24AE79381ED63995192D6CFD06CD8F" box="[567,676,367,392]" pageId="3" pageNumber="304">MADBK</collectionCode>
171803-016, 
<specimenCount id="9D33FD35381ED63994172D73FC2BCD8F" box="[825,905,367,392]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
, 30.1 × 
<quantity id="4CCD9B59381ED63994C72D73FBF2CD80" box="[1001,1104,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" pageId="3" pageNumber="304" unit="mm" value="30.8">30.8 mm</quantity>
, 2 ovig. female, 29.8 × 30.2, 30.8 × 
<quantity id="4CCD9B59381ED63997CB2D97FEE9CDAB" box="[229,331,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="3" pageNumber="304" unit="mm" value="30.2">30.2 mm</quantity>
, Namcheon-dong, Suyeong-gu, Busan, 
<date id="FF8B107C381ED63994272D97FC3FCDAA" box="[777,925,404,429]" pageId="3" pageNumber="304" value="1968-07-10">10 July 1968</date>
.
</paragraph>
<paragraph id="8B8A36BC381ED63997E92DB4FAB1CE1F" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="304">
<collectingCountry id="F322762C381ED63997E92DB4FEA9CDD7" box="[199,267,439,464]" name="China" pageId="3" pageNumber="304">
<emphasis id="B941EAAE381ED63997E92DB4FEA9CDD7" box="[199,267,439,464]" italics="true" pageId="3" pageNumber="304">China</emphasis>
</collectingCountry>
: 
<collectionCode id="ED24AE79381ED63996342DBBFEF3CDD7" box="[282,337,440,464]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="304">ZRC</collectionCode>
2002.0495, 
<specimenCount id="9D33FD35381ED63996F32DBBFD94CDD7" box="[477,566,439,464]" pageId="3" pageNumber="304" type="male">4 males</specimenCount>
, 27.2 × 26.7—31.0 × 
<quantity id="4CCD9B59381ED63994022DBBFC30CDC8" box="[812,914,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="3" pageNumber="304" unit="mm" value="29.8">29.8 mm</quantity>
, 1 ovig. female, 31.6 × 
<quantity id="4CCD9B59381ED63993B52DBBFAA3CDC8" box="[1179,1281,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="3" pageNumber="304" unit="mm" value="31.8">31.8 mm</quantity>
, Tuandao, off Qindao, coll. P.K.L. Ng via fishermen, 
<date id="FF8B107C381ED63995622DDFFCB2CDF3" box="[588,784,476,500]" pageId="3" pageNumber="304" value="2002-08-23" valueMax="2002-08-28" valueMin="2002-08-23">23–28 Aug. 2002</date>
.— 
<collectionCode id="ED24AE79381ED639941F2DDFFC26CDF3" LSID="urn:lsid:biocol.org:col:34871" box="[817,900,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
57771, 
<specimenCount id="9D33FD35381ED63994F02DDFFB94CDF2" box="[990,1078,476,501]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
, 23.9 × 24.8, 31.7 × 
<quantity id="4CCD9B59381ED63992332DDFFA21CDF3" box="[1309,1411,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="3" pageNumber="304" unit="mm" value="31.1">31.1 mm</quantity>
, 
<specimenCount id="9D33FD35381ED63992A02DDFFF44CE1F" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
, 32.1 × 
<quantity id="4CCD9B59381ED639966E2DFCFE04CE10" box="[320,422,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="3" pageNumber="304" unit="mm" value="35.1">35.1 mm</quantity>
, 
<specimenCount id="9D33FD35381ED639969F2E03FE52CE1F" box="[433,496,511,536]" pageId="3" pageNumber="304" type="generic">1 juv.</specimenCount>
female (damaged), Tsimei (=Jimei, Fujian), coll. S.F. Light, 
<date id="FF8B107C381ED63993B72E03FAADCE1F" box="[1177,1295,512,536]" pageId="3" pageNumber="304" value="1923-06">June 1923</date>
.
</paragraph>
<paragraph id="8B8A36BC381ED63997E92E26FAE0CE3A" blockId="3.[151,1437,151,573]" box="[199,1346,548,573]" pageId="3" pageNumber="304">
<collectingCountry id="F322762C381ED63997E92E26FEBBCE3B" box="[199,281,549,572]" name="Taiwan" pageId="3" pageNumber="304">
<emphasis id="B941EAAE381ED63997E92E26FEBBCE3B" box="[199,281,549,572]" italics="true" pageId="3" pageNumber="304">Taiwan</emphasis>
</collectingCountry>
: 
<collectionCode id="ED24AE79381ED63996092E27FEFFCE3B" box="[295,349,548,572]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="304">ZRC</collectionCode>
1997.0382, 
<specimenCount id="9D33FD35381ED63996C42E27FD95CE3A" box="[490,567,548,573]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
, 18.3 × 
<quantity id="4CCD9B59381ED63995BC2E27FD5ACE3B" box="[658,760,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="304" unit="mm" value="16.7">16.7 mm</quantity>
, Donggang, Pingtung, coll. P.K.L. Ng, 
<date id="FF8B107C381ED63993812E27FA9FCE3A" box="[1199,1341,548,573]" pageId="3" pageNumber="304" value="1996-08-05">5 Aug. 1996</date>
.
</paragraph>
</subSubSection>
<caption id="DF4A6634381ED63997B92927FD4CC95F" httpUri="https://zenodo.org/record/228874/files/figure.png" pageId="3" pageNumber="304" targetBox="[151,1436,625,1293]" targetPageId="3">
<paragraph id="8B8A36BC381ED63997B92927FD4CC95F" blockId="3.[151,1435,1316,1368]" pageId="3" pageNumber="304">
<emphasis id="B941EAAE381ED63997B92927FEB9C93D" bold="true" box="[151,283,1316,1338]" pageId="3" pageNumber="304">FIGURE 3.</emphasis>
<taxonomicName id="4C354D3F381ED63996052927FD49C93D" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[299,747,1316,1338]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381ED63996052927FD8DC93D" box="[299,559,1316,1338]" italics="true" pageId="3" pageNumber="304">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
. ZRC 2002.0495, male, 31.0 × 29.8 mm, Qindao, China; a, cephalothorax, dorsal view; b, cephalothorax, ventral view.
</paragraph>
</caption>
<subSubSection id="C32F6537381ED63F97E92987FE21CDAA" lastPageId="5" lastPageNumber="306" pageId="3" pageNumber="304" type="description">
<paragraph id="8B8A36BC381ED63997E92987FD63CB02" blockId="3.[151,1437,1412,2014]" pageId="3" pageNumber="304">
<emphasis id="B941EAAE381ED63997E92987FED6C99A" bold="true" box="[199,372,1412,1437]" pageId="3" pageNumber="304">Redescription.</emphasis>
Carapace rounded, dorsal surface covered somewhat sparsely by columnar granules, granules larger at gastric to intestinal regions (
<figureCitation id="130E2A39381ED639957B29AAFD07C9C6" box="[597,677,1449,1474]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="3" pageNumber="304">Figs. 1</figureCitation>
a, 2, 3a); pterygostomial, subepatic regions evenly covered with rounded granules (
<figureCitation id="130E2A39381ED639965F29CFFE1BC9E2" box="[369,441,1484,1509]" captionStart="FIGURE 2" captionStartId="2.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1667]" captionTargetId="figure@2.[151,1436,193,1668]" captionTargetPageId="2" captionText="FIGURE 2. Arcania undecimspinosa De Haan, 1841. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view." httpUri="https://zenodo.org/record/228873/files/figure.png" pageId="3" pageNumber="304">Fig. 2</figureCitation>
b). Front divided into 2 triangular lobes by obtuse triangular gap, lobes not strongly produced, weakly rounded in some large individuals (
<figureCitation id="130E2A39381ED63995C129F2FC98CA0E" box="[751,826,1521,1546]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="3" pageNumber="304">Figs. 1</figureCitation>
a, 2a). Carapace with subhepatic, anterolateral, lateral, posterolateral, posterior, intestinal spines. Anterolateral spines smallest, followed in size by subhepatic spine, other spines of almost same length. Only branchiocardiac groove distinct. Antennular fossae oblique, antenna excluded from antenular fossa by plate extended from posterior margin of antennullar basal segment, plate with triangular lobe anteriorly, seta posteriorly. Mesial end of infraorbital margin forming dorsoventrally compressed, sharp, triangular spine. Anteromesial corner of pterygostomial region, anterolateral margin of buccal cavern forming laterally compressed triangular spines fitting distal part of exopod of closed mxp3; distance between spines relatively narrow due to slender exopod of mxp3.
</paragraph>
<paragraph id="8B8A36BC381ED63997E92B11FCC6CB2D" blockId="3.[151,1437,1412,2014]" box="[199,868,1809,1834]" pageId="3" pageNumber="304">Eyes fitting within orbit, only corneal part slightly exposed.</paragraph>
<paragraph id="8B8A36BC381ED63997E92B36FBC2CBBE" blockId="3.[151,1437,1412,2014]" pageId="3" pageNumber="304">
Mxp3 (
<figureCitation id="130E2A39381ED63996312B36FED1CB49" box="[287,371,1845,1870]" captionStart="FIGURE 2" captionStartId="2.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1667]" captionTargetId="figure@2.[151,1436,193,1668]" captionTargetPageId="2" captionText="FIGURE 2. Arcania undecimspinosa De Haan, 1841. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view." httpUri="https://zenodo.org/record/228873/files/figure.png" pageId="3" pageNumber="304">Figs. 2</figureCitation>
b, 3b) sparsely covered with rounded granules; merus length about two-fifths of ischium; ischium, merus each with longitudinal groove on mesial third, half; female merus with longitudinal row of setae along longitudinal groove. Exopod narrowed at level of proximal two-fifths of merus, gap between narrowed part of exopod, merus thus fitting triangular spine of anterolateral margin of buccal cavern.
</paragraph>
<paragraph id="8B8A36BC381ED63E97E92BC6FE08CD1A" blockId="3.[151,1437,1412,2014]" lastBlockId="4.[151,1437,151,536]" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304">
Thoracic sternites 4–7 (
<figureCitation id="130E2A39381ED63996CF2BC6FD87CBD9" box="[481,549,1989,2014]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="3" pageNumber="304">Fig. 1</figureCitation>
b) covered with rounded granules; granulation sparser than pterygostomial region. Sternal button of abdominal holding on anterior end of thoracic sternite 5 at lateral slope of thoracic cavity, button fitting proximolateral cup of somite 6. Penis coxal. Vulva on mesial end of thoracic suture 5/6, on somite 6 but extending mesial end of suture 5/6 anteriorly (
<figureCitation id="130E2A393819D63E95FF2CDCFCB5CCFF" box="[721,791,223,248]" captionStart="FIGURE 6" captionStartId="6.[151,250,1363,1385]" captionTargetBox="[193,1391,529,1336]" captionTargetId="figure@6.[188,1398,517,1342]" captionTargetPageId="6" captionText="FIGURE 6. Right vulva of Arcania undecimspinosa De Haan, 1841; a, NSMT-Cr 3839, 24.5 × 23.3 mm, Mikawa Bay, Japan; b, NSMT-Cr 4397, 28.2 × 27.0 mm, Kochi, Japan. Scale = 1 mm." httpUri="https://zenodo.org/record/228877/files/figure.png" pageId="4" pageNumber="305">Fig. 6</figureCitation>
); posterolateral part of vulva swollen; opening narrowed laterally, entire mesially.
</paragraph>
<paragraph id="8B8A36BC3819D63E97E92D24FDAACE10" blockId="4.[151,1437,151,536]" pageId="4" pageNumber="305">
Chelipeds (
<figureCitation id="130E2A393819D63E96622D24FE37CD47" box="[332,405,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1667]" captionTargetId="figure@2.[151,1436,193,1668]" captionTargetPageId="2" captionText="FIGURE 2. Arcania undecimspinosa De Haan, 1841. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view." httpUri="https://zenodo.org/record/228873/files/figure.png" pageId="4" pageNumber="305">Fig. 2</figureCitation>
a) moderately long, subequal. Merus slender, shorter than chela, entirely covered with rounded granules, granules smaller on lower surface. Carpus, palm sparsely covered with minute granules, carpus smooth without knob on external margin. Fingers slender, gape absent when closed, with 14–18 almost equidistantly placed, small teeth with much smaller teeth throughout. Ambulatory legs slender; merus longest of articles, covered with minute rounded granules. Carpus, propodus covered with minute granules. Dactylus mesiolateraly depressed in cross-section, with sharp keels on mid-lines of mesial, lateral surfaces, keel on lateral surface flanked by rows of setae.
</paragraph>
<caption id="DF4A66343819D63E97B92B1FFB52CB56" httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="4" pageNumber="305" targetBox="[198,1393,597,1786]" targetPageId="4">
<paragraph id="8B8A36BC3819D63E97B92B1FFB52CB56" blockId="4.[151,1435,1820,1873]" pageId="4" pageNumber="305">
<emphasis id="B941EAAE3819D63E97B92B1FFEB0CB35" bold="true" box="[151,274,1820,1842]" pageId="4" pageNumber="305">FIGURE 4.</emphasis>
G1 and G2 of 
<taxonomicName id="4C354D3F3819D63E96892B1EFCE3CB35" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[423,833,1820,1842]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE3819D63E96892B1EFD02CB35" box="[423,672,1820,1842]" italics="true" pageId="4" pageNumber="305">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
. Lectotype. RMNH.CRUS.D.790, male, 16.5 × 16.5 mm; a, left G1, mesial view; b, left G1, ventral view; c, distal part of left G1, ventral view; d, left G2. Scales = 1 mm.
</paragraph>
</caption>
<paragraph id="8B8A36BC3819D63F97E92B7DFB31CCD3" blockId="4.[151,1436,1918,2015]" lastBlockId="5.[151,1437,151,429]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
Abdomen (
<figureCitation id="130E2A393819D63E96692B7DFE25CB90" box="[327,391,1918,1943]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="4" pageNumber="305">Fig. 1</figureCitation>
b) with sparse, rounded granules; somites 3–5 fused, with proximolateral protuberances on external surface, each protuberance not distinctly larger than central part between two protuberances. Lateral margins of somite 6 convex in large individuals. G1 (
<figureCitation id="130E2A393819D63E95C72BC5FC95CBD8" box="[745,823,1990,2015]" captionStart="FIGURE 4. G 1 and G 2" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[198,1393,597,1786]" captionTargetId="figure@4.[183,1404,589,1799]" captionTargetPageId="4" captionText="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="4" pageNumber="305">Figs. 4</figureCitation>
a–c, 5) slender, nearly straight, slightly curved dorsally in mesial view; distal part only slightly upcurved in mesial view, lateral margin wrapping onto mesial margin submedially, forming pipe-like opening. G2 (
<figureCitation id="130E2A393818D63F95BF2CBFFD76CCD2" box="[657,724,188,213]" captionStart="FIGURE 4. G 1 and G 2" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[198,1393,597,1786]" captionTargetId="figure@4.[183,1404,589,1799]" captionTargetPageId="4" captionText="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
d) short, opening facing dorsally 
<emphasis id="B941EAAE3818D63F93692CBEFB2ECCD3" box="[1095,1164,189,212]" italics="true" pageId="5" pageNumber="306">in situ</emphasis>
.
</paragraph>
<paragraph id="8B8A36BC3818D63F97E92CDCFD1DCD1A" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="306">
<emphasis id="B941EAAE3818D63F97E92CDCFEEDCCFF" bold="true" box="[199,335,223,248]" pageId="5" pageNumber="306">Coloration.</emphasis>
Body and cheliped merus pinkish dorsally, with a pair of reddish lines and semicircular marks on the dorsal surface (
<bibRefCitation id="EFA44B4D3818D63F96442D07FD9CCD1B" author="Takeda" box="[362,574,260,285]" pageId="5" pageNumber="306" refString="Takeda, M., Furuta, S., Miyanaga, T., Tamura, A. &amp; Wada, T. (2011) Crabs from the southwestern Sea of Japan along Tottori Prefecture and its vicinity, Japan. Bulletin of the Tottori Prefectural Museum, 48, 29 - 94. [in Japanese with English abstract]" type="journal article" year="2011">
Takeda 
<emphasis id="B941EAAE3818D63F96EA2D06FE5ECD1A" box="[452,508,260,285]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
2011
</bibRefCitation>
: fig. 9–26).
</paragraph>
<paragraph id="8B8A36BC3818D63F97E92D24FE21CDAA" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="306">
<emphasis id="B941EAAE3818D63F97E92D24FDA6CD47" bold="true" box="[199,516,295,320]" pageId="5" pageNumber="306">Geographical distribution.</emphasis>
<collectingCountry id="F322762C3818D63F95252D2BFDE9CD38" box="[523,587,296,319]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
(Pacific coast: from Sagami Bay to 
<collectingCountry id="F322762C3818D63F94CE2D24FB8ACD47" box="[992,1064,295,320]" name="India" pageId="5" pageNumber="306">Kochi</collectingCountry>
; Kyushu: Ariake, Nagasaki [
<typeStatus id="548E881E3818D63F92472D2BFA3ECD47" box="[1385,1436,296,320]" pageId="5" pageNumber="306">type</typeStatus>
locality]); 
<collectingCountry id="F322762C3818D63F96232D4EFEF3CD63" box="[269,337,333,356]" name="South Korea" pageId="5" pageNumber="306">Korea</collectingCountry>
(Namcheon-dong, Jinhae-gu); 
<collectingCountry id="F322762C3818D63F959C2D4FFD57CD62" box="[690,757,332,357]" name="China" pageId="5" pageNumber="306">China</collectingCountry>
(Qingdao; Tsimei; Fukien; Hangzhou; Kwangtung; Cheung Chau Island, 
<collectingCountry id="F322762C3818D63F961F2D73FE1FCD8F" box="[305,445,368,392]" name="Hong Kong" pageId="5" pageNumber="306">Hong Kong</collectingCountry>
); 
<collectingCountry id="F322762C3818D63F96FD2D6CFD87CD8F" box="[467,549,367,392]" name="Taiwan" pageId="5" pageNumber="306">Taiwan</collectingCountry>
(Donggang). (De Haan 1844; 
<bibRefCitation id="EFA44B4D3818D63F94A42D6CFBA5CD8F" author="Shen" box="[906,1031,367,392]" pageId="5" pageNumber="306" refString="Shen, C. J. (1931) The crabs of Hong Kong. Part I. The Hong Kong Naturalist, 2, 92 - 110, pls. 4 - 10." type="journal article" year="1931">Shen 1931</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D3818D63F93382D6CFB3BCD8F" author="Sakai" box="[1046,1177,367,392]" pageId="5" pageNumber="306" refString="Sakai, T. (1937) Studies on the crabs of Japan. II. Oxystomata. Science Reports of the Tokyo Bunrika Daigaku, Section B, Supplement No. 2, 3, 67 - 192, pls. 10 - 19." type="journal article" year="1937">Sakai 1937</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D3818D63F93862D6CFAE1CD8F" author="Miyake" box="[1192,1347,367,392]" pageId="5" pageNumber="306" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color. Vol. II Brachyura (Crabs). Hoikusha, Osaka, 277 pp. [in Japanese]" type="book" year="1983">Miyake 1983</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D3818D63F927C2D73FF71CDAB" author="Huang" pageId="5" pageNumber="306" refString="Huang, L. (1989) Oxystomata. In: Wei, C. &amp; Chen, Y. (Eds.), Fauna of Zhejiang. Crustacea. Zhejiang Science and Technology Publishing House, Hangzhou, pp. 294 - 324. [in Chinese]" type="book chapter" year="1989">Huang 1989</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<caption id="DF4A66343818D63F97B92AEAFBC8CB19" httpUri="https://zenodo.org/record/228876/files/figure.png" pageId="5" pageNumber="306" targetBox="[325,1261,495,1747]" targetPageId="5">
<paragraph id="8B8A36BC3818D63F97B92AEAFBC8CB19" blockId="5.[151,1436,1769,1822]" pageId="5" pageNumber="306">
<emphasis id="B941EAAE3818D63F97B92AEAFEB0CAF8" bold="true" box="[151,274,1769,1791]" pageId="5" pageNumber="306">FIGURE 5.</emphasis>
G1 of 
<taxonomicName id="4C354D3F3818D63F96772AE9FD56CAF8" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[345,756,1769,1791]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE3818D63F96772AE9FDF0CAF8" box="[345,594,1769,1791]" italics="true" pageId="5" pageNumber="306">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
. ZRC 2002.0495, male, 31.0 × 29.8 mm, Qindao, China; a, left G1, mesial view; b, left G1, ventral view; c, distal part of left G1, ventromesial view. Scales = 1 mm.
</paragraph>
</caption>
<subSubSection id="C32F65373818D63C97E92B49FEEACDD7" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306" type="discussion">
<paragraph id="8B8A36BC3818D63C97E92B49FEEACDD7" blockId="5.[151,1436,1866,2035]" lastBlockId="6.[151,1436,151,464]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<emphasis id="B941EAAE3818D63F97E92B49FE99CB64" bold="true" box="[199,315,1866,1891]" pageId="5" pageNumber="306">Remarks.</emphasis>
<taxonomicName id="4C354D3F3818D63F96662B48FC8ACB64" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[328,808,1866,1891]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE3818D63F96662B48FDC1CB64" box="[328,611,1866,1891]" italics="true" pageId="5" pageNumber="306">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
, was described from material collected at Nagasaki, western Kyushu, 
<collectingCountry id="F322762C3818D63F964E2B73FE01CB80" box="[352,419,1904,1927]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
. 
<bibRefCitation id="EFA44B4D3818D63F969F2B6DFD45CB80" author="Yamaguchi" box="[433,743,1902,1927]" pageId="5" pageNumber="306" refString="Yamaguchi, T. &amp; Baba, K. (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. In: Yamaguchi, T. (Ed.), Ph. F. von Siebold and Natural History of Japan, Crustacea. The Carcinological Society of Japan, Tokyo, pp. 145 - 570. [in Japanese with English summary and figure caption]" type="book chapter" year="1993">Yamaguchi &amp; Baba (1993)</bibRefCitation>
and 
<bibRefCitation id="EFA44B4D3818D63F940A2B73FBB4CB80" box="[804,1046,1902,1927]" pageId="5" pageNumber="306" refString="Fransen, C. H. J. M., Holthuis, L. B. &amp; Adema, J. P. H. M. (1997) Type-catalogue of the decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, 311, i - xvi, 1 - 344." type="journal article">
Fransen 
<emphasis id="B941EAAE3818D63F94A92B73FC60CB80" box="[903,962,1902,1927]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
(1997)
</bibRefCitation>
listed 15 
<typeStatus id="548E881E3818D63F93A32B73FB50CB8F" box="[1165,1266,1904,1928]" pageId="5" pageNumber="306" type="syntype">syntypes</typeStatus>
, of which five specimens are preserved in ethanol and the rest are dried. The ethanol-preserved specimens comprise three males (two of which are confirmed by the presence of fragmented male thoracic sternites), one female, and one individual (sex unknown). 
<bibRefCitation id="EFA44B4D3818D63F967B2BD9FD33CBF4" author="Yamaguchi" box="[341,657,2010,2035]" pageId="5" pageNumber="306" refString="Yamaguchi, T. &amp; Baba, K. (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. In: Yamaguchi, T. (Ed.), Ph. F. von Siebold and Natural History of Japan, Crustacea. The Carcinological Society of Japan, Tokyo, pp. 145 - 570. [in Japanese with English summary and figure caption]" type="book chapter" year="1993">Yamaguchi &amp; Baba (1993)</bibRefCitation>
indicated that the ethanol-preserved lot contained a 
<typeStatus id="548E881E3818D63F922F2BD9FACACBF4" box="[1281,1384,2010,2035]" pageId="5" pageNumber="306" type="lectotype">lectotype</typeStatus>
and 
<typeStatus id="548E881E381BD63C97B92C94FE9ECCB7" box="[151,316,151,176]" pageId="6" pageNumber="307" type="paralectotype">paralectotypes</typeStatus>
, but they did not note nor indicate which individual was selected as 
<typeStatus id="548E881E381BD63C936A2C94FB0CCCB7" box="[1092,1198,151,176]" pageId="6" pageNumber="307" type="lectotype">lectotype</typeStatus>
. Since Yamaguchi &amp; Baba’s (1993) 
<typeStatus id="548E881E381BD63C96112CBFFE07CCD2" box="[319,421,188,213]" pageId="6" pageNumber="307" type="lectotype">lectotype</typeStatus>
designation is invalid, the 
<typeStatus id="548E881E381BD63C95F42CBFFCE3CCD2" box="[730,833,188,213]" pageId="6" pageNumber="307" type="lectotype">lectotype</typeStatus>
is here designated for a male (RMNH.CRUS.D.790; 16.5 × 
<quantity id="4CCD9B59381BD63C97CB2CDCFEE8CCFF" box="[229,330,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="307" unit="mm" value="16.5">16.5 mm</quantity>
) form the ethanol-preserved specimens (
<figureCitation id="130E2A39381BD63C943C2CDCFCFDCCFF" box="[786,863,223,248]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="6" pageNumber="307">Figs. 1</figureCitation>
, 
<figureCitation id="130E2A39381BD63C94432CE3FCDCCCFF" box="[877,894,224,248]" captionStart="FIGURE 4. G 1 and G 2" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[198,1393,597,1786]" captionTargetId="figure@4.[183,1404,589,1799]" captionTargetPageId="4" captionText="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="6" pageNumber="307">4</figureCitation>
). The 
<typeStatus id="548E881E381BD63C94EB2CDCFB8ECCFF" box="[965,1068,223,248]" pageId="6" pageNumber="307" type="lectotype">lectotype</typeStatus>
is in a relatively good condition, allowing the observation of the G1 (
<figureCitation id="130E2A39381BD63C95142D07FDDDCD1B" box="[570,639,260,285]" captionStart="FIGURE 4. G 1 and G 2" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[198,1393,597,1786]" captionTargetId="figure@4.[183,1404,589,1799]" captionTargetPageId="4" captionText="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="6" pageNumber="307">Fig. 4</figureCitation>
a–c). 
<taxonomicName id="4C354D3F381BD63C95912D07FC74CD1A" box="[703,982,260,285]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C95912D07FC74CD1A" box="[703,982,260,285]" italics="true" pageId="6" pageNumber="307">Arcania undecimspinosa</emphasis>
</taxonomicName>
is characterized by a rounded carapace (
<figureCitation id="130E2A39381BD63C97B12D24FF49CD47" box="[159,235,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="6" pageNumber="307">Figs. 1</figureCitation>
a, 2, 3a), front with triangular lobes with an almost regular triangular gap between the lobes (
<figureCitation id="130E2A39381BD63C923F2D24FAFECD47" box="[1297,1372,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="6" pageNumber="307">Figs. 1</figureCitation>
a, 2a, 3a), distinct marginal spines in the carapace (
<figureCitation id="130E2A39381BD63C95BB2D4FFD43CD63" box="[661,737,332,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="6" pageNumber="307">Figs. 1</figureCitation>
a, 2a, 3a), dorsal surface of the carapace being rather sparsely covered by columnar granules (granules larger from gastric to intestinal regions) (
<figureCitation id="130E2A39381BD63C93022D6CFBD2CD8F" box="[1068,1136,367,392]" captionStart="FIGURE 2" captionStartId="2.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1667]" captionTargetId="figure@2.[151,1436,193,1668]" captionTargetPageId="2" captionText="FIGURE 2. Arcania undecimspinosa De Haan, 1841. NSMT-Cr 4397, male, ca. 25.9 × 25.1 mm (front damaged), Kochi, Japan; a, habitus; b, cephalothorax, frontal view." httpUri="https://zenodo.org/record/228873/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
b), almost straight G1 with pipe-like distal end (
<figureCitation id="130E2A39381BD63C96A62D97FE78CDAB" box="[392,474,404,429]" captionStart="FIGURE 4. G 1 and G 2" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[198,1393,597,1786]" captionTargetId="figure@4.[183,1404,589,1799]" captionTargetPageId="4" captionText="FIGURE 4. G 1 and G 2 of Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm; a, left G 1, mesial view; b, left G 1, ventral view; c, distal part of left G 1, ventral view; d, left G 2. Scales = 1 mm." httpUri="https://zenodo.org/record/228875/files/figure.png" pageId="6" pageNumber="307">Figs. 4</figureCitation>
a–c, 5) and a vulva consisting of a narrowed lateral end and with an entire mesial margin (
<figureCitation id="130E2A39381BD63C97DB2DB4FE98CDD7" box="[245,314,439,464]" captionStart="FIGURE 6" captionStartId="6.[151,250,1363,1385]" captionTargetBox="[193,1391,529,1336]" captionTargetId="figure@6.[188,1398,517,1342]" captionTargetPageId="6" captionText="FIGURE 6. Right vulva of Arcania undecimspinosa De Haan, 1841; a, NSMT-Cr 3839, 24.5 × 23.3 mm, Mikawa Bay, Japan; b, NSMT-Cr 4397, 28.2 × 27.0 mm, Kochi, Japan. Scale = 1 mm." httpUri="https://zenodo.org/record/228877/files/figure.png" pageId="6" pageNumber="307">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4A6634381BD63C97B92950FC8FC98F" httpUri="https://zenodo.org/record/228877/files/figure.png" pageId="6" pageNumber="307" targetBox="[193,1391,529,1336]" targetPageId="6">
<paragraph id="8B8A36BC381BD63C97B92950FC8FC98F" blockId="6.[151,1436,1363,1416]" pageId="6" pageNumber="307">
<emphasis id="B941EAAE381BD63C97B92950FEB1C96E" bold="true" box="[151,275,1363,1385]" pageId="6" pageNumber="307">FIGURE 6.</emphasis>
Right vulva of 
<taxonomicName id="4C354D3F381BD63C969D2957FCEDC96E" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[435,847,1363,1385]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C969D2957FD0CC96E" box="[435,686,1363,1385]" italics="true" pageId="6" pageNumber="307">Arcania undecimspinosa</emphasis>
De Haan, 1841
</taxonomicName>
; a, NSMT-Cr 3839, 24.5 × 23.3 mm, Mikawa Bay, Japan; b, NSMT-Cr 4397, 28.2 × 27.0 mm, Kochi, Japan. Scale = 1 mm.
</paragraph>
</caption>
<subSubSection id="C32F6537381BD63D97E929B3FA9BCF5A" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307" type="reference_group">
<paragraph id="8B8A36BC381BD63C97E929B3FADDCAC3" blockId="6.[151,1437,1455,2020]" pageId="6" pageNumber="307">
<taxonomicName id="4C354D3F381BD63C97E929B3FE43C9CF" ID-CoL="G7CC" box="[199,481,1455,1480]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C97E929B3FE43C9CF" box="[199,481,1455,1480]" italics="true" pageId="6" pageNumber="307">Arcania undecimspinosa</emphasis>
</taxonomicName>
is morphologically most similar to 
<taxonomicName id="4C354D3F381BD63C94BD29B3FB84C9C0" ID-CoL="G7BF" box="[915,1062,1456,1479]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="B941EAAE381BD63C94BD29B3FB84C9C0" box="[915,1062,1456,1479]" italics="true" pageId="6" pageNumber="307">A. cornigera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C931D29ACFBD5C9CF" box="[1075,1143,1455,1480]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B941EAAE381BD63C931D29ACFBD5C9CF" bold="true" box="[1075,1143,1455,1480]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
(see 
<emphasis id="B941EAAE381BD63C939029ACFA89C9CF" bold="true" box="[1214,1323,1455,1480]" pageId="6" pageNumber="307">Remarks</emphasis>
under 
<taxonomicName id="4C354D3F381BD63C92AA29B3FEA4C9EB" ID-CoL="G7BF" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="B941EAAE381BD63C92AA29B3FEA4C9EB" italics="true" pageId="6" pageNumber="307">A. cornigera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C962329D0FEEEC9EB" box="[269,332,1491,1516]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B941EAAE381BD63C962329D0FEEEC9EB" bold="true" box="[269,332,1491,1516]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
). It should be noted that 
<taxonomicName id="4C354D3F381BD63C954529D7FCE2C9EB" ID-CoL="G7CC" box="[619,832,1491,1516]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C954529D7FCE2C9EB" box="[619,832,1491,1516]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C354D3F381BD63C945929D7FBA4C9EB" ID-CoL="G7BF" box="[887,1030,1492,1516]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="B941EAAE381BD63C945929D7FBA4C9EB" box="[887,1030,1492,1516]" italics="true" pageId="6" pageNumber="307">A. cornigera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C932329D0FBEFC9EB" box="[1037,1101,1491,1516]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B941EAAE381BD63C932329D0FBEFC9EB" bold="true" box="[1037,1101,1491,1516]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
have been collected from the same areas (e.g. 
<collectingCountry id="F322762C381BD63C964B29F4FE0ECA17" box="[357,428,1527,1552]" name="India" pageId="6" pageNumber="307">Kochi</collectingCountry>
, Pacific coast of Shikoku, 
<collectingCountry id="F322762C381BD63C95D929FBFC98CA08" box="[759,826,1528,1551]" name="Japan" pageId="6" pageNumber="307">Japan</collectingCountry>
; Donggang, southwestern 
<collectingCountry id="F322762C381BD63C935929F4FB6CCA17" box="[1143,1230,1527,1552]" name="Taiwan" pageId="6" pageNumber="307">Taiwan</collectingCountry>
). This superficial similarity and overlapping distribution may have led to misidentifications. Among the 
<taxonomicName id="4C354D3F381BD63C93EE2A1FFA3ECA33" ID-CoL="G7CC" box="[1216,1436,1563,1588]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C93EE2A1FFA3ECA33" box="[1216,1436,1563,1588]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
specimens recorded by 
<bibRefCitation id="EFA44B4D381BD63C96832A3CFDF9CA5F" author="Yokoya" box="[429,603,1599,1624]" pageId="6" pageNumber="307" refString="Yokoya, Y. (1933) On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, chiefly based upon materials collected by s. s. Soyo-Maru during the years 1923 - 1930. Journal of the College of Agriculture, Tokyo Imperial University, 12 (1), 1 - 226." type="journal article" year="1933">Yokoya (1933)</bibRefCitation>
, the specimens from Kyo-gasaki (KMNH IvR 400,102; st. 540), Goto Islands (KMNH IvR 400,104; st. 451) and Jeju Island (KMNH IvR 400,103; labelled as Saishu-to, st. 454) are indeed 
<taxonomicName id="4C354D3F381BD63C97C42A8BFED8CA98" ID-CoL="G7BF" box="[234,378,1672,1695]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="B941EAAE381BD63C97C42A8BFED8CA98" box="[234,378,1672,1695]" italics="true" pageId="6" pageNumber="307">A. cornigera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C96AC2A84FE60CAA7" box="[386,450,1671,1696]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B941EAAE381BD63C96AC2A84FE60CAA7" bold="true" box="[386,450,1671,1696]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
<bibRefCitation id="EFA44B4D381BD63C96E52A84FDFECAA7" author="Sakai" box="[459,604,1671,1696]" pageId="6" pageNumber="307" refString="Sakai, T. (1965) The crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen, Tokyo, 206 pp. + 100 pls. + 92 pp., in Japanese + 32 pp. of bibiliography and index pp." type="book" year="1965">Sakai (1965)</bibRefCitation>
also figured the anterior part of the cephalothorax of 
<taxonomicName id="4C354D3F381BD63C93EA2A8BFAF0CA98" ID-CoL="G7BF" box="[1220,1362,1672,1695]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="B941EAAE381BD63C93EA2A8BFAF0CA98" box="[1220,1362,1672,1695]" italics="true" pageId="6" pageNumber="307">A. cornigera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C92752A84FA3ECAA7" box="[1371,1436,1671,1696]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B941EAAE381BD63C92752A84FA3ECAA7" bold="true" box="[1371,1436,1671,1696]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
(
<bibRefCitation id="EFA44B4D381BD63C97B02AA8FED6CAC3" author="Sakai" box="[158,372,1707,1732]" pageId="6" pageNumber="307" refString="Sakai, T. (1965) The crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen, Tokyo, 206 pp. + 100 pls. + 92 pp., in Japanese + 32 pp. of bibiliography and index pp." type="book" year="1965" yearSuffix="a">Sakai 1965: fig. 6a</bibRefCitation>
) and the habitus of 
<taxonomicName id="4C354D3F381BD63C957D2AAFFC85CAC3" ID-CoL="G7CC" box="[595,807,1707,1732]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" sensu="stricto" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C957D2AAFFC85CAC3" box="[595,807,1707,1732]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381BD63C94032AAFFCF3CAC4" box="[813,849,1708,1731]" pageId="6" pageNumber="307" sensu="stricto">s.s.</taxonomicNameLabel>
as 
<taxonomicName id="4C354D3F381BD63C94582AAFFBE8CAC3" ID-CoL="G7CC" box="[886,1098,1707,1732]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C94582AAFFBE8CAC3" box="[886,1098,1707,1732]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA44B4D381BD63C93772AA8FAD0CAC3" author="Sakai" box="[1113,1394,1707,1732]" pageId="6" pageNumber="307" refString="Sakai, T. (1965) The crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen, Tokyo, 206 pp. + 100 pls. + 92 pp., in Japanese + 32 pp. of bibiliography and index pp." type="book" year="1965">Sakai 1965: pl. 16, fig. 2</bibRefCitation>
).
</paragraph>
<paragraph id="8B8A36BC381BD63D97E92ACCFC66CD47" blockId="6.[151,1437,1455,2020]" lastBlockId="7.[151,1436,151,861]" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307">
<bibRefCitation id="EFA44B4D381BD63C97E92ACCFEF8CAEF" author="Miers" box="[199,346,1743,1768]" pageId="6" pageNumber="307" refString="Miers, E. J. (1877) Notes upon the Oxystomatous Crustacea. Transactions of the Linnean Society of London, Series 2, 1 (5), 235 - 249, pls. 38 - 40." type="journal article" year="1877">Miers (1877)</bibRefCitation>
described 
<taxonomicName id="4C354D3F381BD63C96FA2AD3FDC7CAEF" ID-CoL="G7BT" box="[468,613,1743,1768]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="granulosa">
<emphasis id="B941EAAE381BD63C96FA2AD3FDC7CAEF" box="[468,613,1743,1768]" italics="true" pageId="6" pageNumber="307">A. granulosa</emphasis>
</taxonomicName>
from Moreton Bay, 
<collectingCountry id="F322762C381BD63C94652ACCFC16CAEF" box="[843,948,1743,1768]" name="Australia" pageId="6" pageNumber="307">Australia</collectingCountry>
, which was cited by 
<bibRefCitation id="EFA44B4D381BD63C938E2ACCFAE5CAEF" author="Haswell" box="[1184,1351,1743,1768]" pageId="6" pageNumber="307" refString="Haswell, W. A. (1880) Contributions to a monograph of Australian Leucosiidae. Proceedings of the Linnean Society of New South Wales, 4 (1), 44 - 60, pls. 5, 6." type="journal article" year="1880">Haswell (1880</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D381BD63C927D2AD3FA32CAEF" author="Haswell" box="[1363,1424,1744,1768]" pageId="6" pageNumber="307" refString="Haswell, W. A. (1882) Catalogue of the Australian Stalk- and Sessile-Eyed Crustacea. Australian Museum, Sydney, 327 pp + 4 pls." type="book" year="1882">1882</bibRefCitation>
). 
<bibRefCitation id="EFA44B4D381BD63C97B92AF0FE89CB0B" author="Miers" box="[151,299,1779,1804]" pageId="6" pageNumber="307" refString="Miers, E. J. (1879) On a collection of Crustacea made by Capt. H. C. St. John, R. N., in the Corean and Japanese Seas. Part I. Podophthalmia. With an appendix by Capt. H. C. St. John. Proceedings of the Scientific Meetings of the Zoological Society of London, 1879, 18 - 61." type="book chapter" year="1879">Miers (1879)</bibRefCitation>
subsequently synonymized 
<taxonomicName id="4C354D3F381BD63C95422AF7FD5DCB0B" ID-CoL="G7BT" box="[620,767,1779,1804]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="granulosa">
<emphasis id="B941EAAE381BD63C95422AF7FD5DCB0B" box="[620,767,1779,1804]" italics="true" pageId="6" pageNumber="307">A. granulosa</emphasis>
</taxonomicName>
under 
<emphasis id="B941EAAE381BD63C947E2AF7FB8FCB0B" box="[848,1069,1779,1804]" italics="true" pageId="6" pageNumber="307">
<taxonomicName id="4C354D3F381BD63C947E2AF7FB8BCB0B" ID-CoL="G7CC" box="[848,1065,1779,1804]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">A. undecimspinosa</taxonomicName>
.
</emphasis>
Subsequent authors (e.g. 
<bibRefCitation id="EFA44B4D381BD63C927A2AF0FF71CB37" author="Miers" pageId="6" pageNumber="307" refString="Miers, E. J. (1884) Crustacea (Brachyura). In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the voyage of H. M. S. ' Alert' 1881 - 1882. Part II. The collections from the Western Indian Ocean, 8 (2), 513 - 575, pls. 46 - 52. [British Museum (Natural History), London]" type="journal article" year="1884">Miers, 1884</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D381BD63C97F32B14FE3DCB37" author="Henderson" box="[221,415,1815,1840]" pageId="6" pageNumber="307" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, Series 2 Zoology, 5 (10), 325 - 458, pls 36 - 40." type="journal article" year="1893">Henderson, 1893</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D381BD63C96842B14FDFACB37" author="Campbell" box="[426,600,1815,1840]" pageId="6" pageNumber="307" refString="Campbell, B. M. (1971) New records and new species of crabs (Crustacea: Brachyura) trawled off southern Queensland: Dromiacea, Homolidea, Gymnopleura, Corystoidea, and Oxystomata). Memoirs of the Queensland Museum, 16 (1), 27 - 48, pls. 2, 3." type="journal article" year="1971">Campbell 1971</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D381BD63C954D2B14FCE7CB37" author="Davie" box="[611,837,1815,1840]" pageId="6" pageNumber="307" refString="Davie, P. J. F. &amp; Short, J. W. (1989) Deepwater Brachyura (Crustacea: Decapoda) from southern Queensland, Australia with description of four new species. Memoirs of the Queensland Museum, 27 (2), 157 - 187." type="journal article" year="1989">Davie &amp; Short 1989</bibRefCitation>
) recorded “ 
<taxonomicName id="4C354D3F381BD63C94E62B1BFB38CB37" ID-CoL="G7CC" box="[968,1178,1815,1840]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C94E62B1BFB38CB37" box="[968,1178,1815,1840]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
” from Moreton Bay as well. Miers’s (1877) specimen is, however, only about 1/ 
<quantity id="4CCD9B59381BD63C94362B3FFCFCCB53" box="[792,862,1851,1876]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="307" unit="in" value="3.0">3 inch</quantity>
carapace length by 1/ 
<quantity id="4CCD9B59381BD63C937A2B3FFB3ECB53" box="[1108,1180,1851,1876]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="307" unit="in" value="4.0">4 inch</quantity>
carapace width (about 8.5 × 
<quantity id="4CCD9B59381BD63C97F62B5CFE90CB7F" box="[216,306,1887,1912]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="307" unit="mm" value="6.4">6.4 mm</quantity>
), which is too small to allow the correct identification of 
<taxonomicName id="4C354D3F381BD63C949B2B63FB2BCB7F" ID-CoL="G7CC" box="[949,1161,1887,1912]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C949B2B63FB2BCB7F" box="[949,1161,1887,1912]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
and allied species. 
<bibRefCitation id="EFA44B4D381BD63C924A2B5CFF43CB9B" author="Galil" pageId="6" pageNumber="307" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil (2001)</bibRefCitation>
identified Campbell &amp; Stephenson’s (1970) 
<taxonomicName id="4C354D3F381BD63C95C22B87FC0FCB9B" ID-CoL="G7BZ" box="[748,941,1924,1948]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="novemspinosa">
<emphasis id="B941EAAE381BD63C95C22B87FC0FCB9B" box="[748,941,1924,1948]" italics="true" pageId="6" pageNumber="307">A. novemspinosa</emphasis>
</taxonomicName>
from Moreton Bay as 
<taxonomicName id="4C354D3F381BD63C93902B87FA36CB9B" ID-CoL="G7CC" box="[1214,1428,1923,1948]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381BD63C93902B87FA36CB9B" box="[1214,1428,1923,1948]" italics="true" pageId="6" pageNumber="307">A. undecimspinosa</emphasis>
</taxonomicName>
. Campbell &amp; Stephenson’s drawing (1970: fig. 13), however, shows a distally curved G1 with an oblique line of setae, proportionally too broad posterior teeth and almost indiscernible anterolateral teeth of the carapace. This is clearly neither 
<taxonomicName id="4C354D3F381AD63D96102C9BFDB3CCB7" ID-CoL="G7CC" box="[318,529,151,176]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D96102C9BFDB3CCB7" box="[318,529,151,176]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
(
<figureCitation id="130E2A39381AD63D950F2C94FDCECCB7" box="[545,620,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1920,1942]" captionTargetBox="[151,1436,1236,1898]" captionTargetId="figure@1.[151,1436,1236,1899]" captionTargetPageId="1" captionText="FIGURE 1. Arcania undecimspinosa De Haan, 1841. Lectotype. RMNH. CRUS. D. 790, male, 16.5 × 16.5 mm, Nagasaki, Kyushu, Japan; a, cephalothorax, dorsal view; b, cephalothorax, ventral view." httpUri="https://zenodo.org/record/228872/files/figure.png" pageId="7" pageNumber="308">Figs. 1</figureCitation>
a, 2a, 3a, 4a–c, 5) nor 
<taxonomicName id="4C354D3F381AD63D94492C9BFB87CCB7" ID-CoL="G7BZ" box="[871,1061,152,176]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="novemspinosa">
<emphasis id="B941EAAE381AD63D94492C9BFB87CCB7" box="[871,1061,152,176]" italics="true" pageId="7" pageNumber="308">A. novemspinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA44B4D381AD63D931D2C94FB0DCCB7" author="Galil" box="[1075,1199,151,176]" pageId="7" pageNumber="308" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil 2001</bibRefCitation>
, figs. 2F, 6C; 
<bibRefCitation id="EFA44B4D381AD63D92642C94FE8DCCD3" author="Fujii" pageId="7" pageNumber="308" refString="Fujii, T. &amp; Naruse, T. (2013) First record of Arcania novemspinosa (Crustacea: Decapoda: Leucosiidae) from Okinawa Island, Japan. Fauna Ryukyuana, 3, 1 - 6. [in Japanese]" type="journal article" year="2013">Fujii &amp; Naruse 2013</bibRefCitation>
: figs. 1, 2A). The condition of the G1 and the posterior teeth of the carapace of Campbell &amp; Stephenson’s (1970) “ 
<taxonomicName id="4C354D3F381AD63D96BA2CE3FDF6CCFF" ID-CoL="G7BZ" box="[404,596,224,248]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="novemspinosa">
<emphasis id="B941EAAE381AD63D96BA2CE3FDF6CCFF" box="[404,596,224,248]" italics="true" pageId="7" pageNumber="308">A. novemspinosa</emphasis>
</taxonomicName>
” are rather closer to 
<taxonomicName id="4C354D3F381AD63D94652CE3FBFBCCFF" ID-CoL="G7BM" authority="Galil, 2001" authorityName="Galil" authorityYear="2001" box="[843,1113,223,248]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="foliolata">
<emphasis id="B941EAAE381AD63D94652CE3FC69CCFF" box="[843,971,223,248]" italics="true" pageId="7" pageNumber="308">A. foliolata</emphasis>
<bibRefCitation id="EFA44B4D381AD63D94FB2CDCFBFBCCFF" author="Galil" box="[981,1113,223,248]" pageId="7" pageNumber="308" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil, 2001</bibRefCitation>
</taxonomicName>
. The taxonomic status of 
<taxonomicName id="4C354D3F381AD63D92AA2CE3FEABCD1A" ID-CoL="G7BT" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="granulosa">
<emphasis id="B941EAAE381AD63D92AA2CE3FEABCD1A" italics="true" pageId="7" pageNumber="308">A. granulosa</emphasis>
</taxonomicName>
as well as the 
<taxonomicName id="4C354D3F381AD63D96822D07FDA4CD1C" ID-CoL="7NQTF" box="[428,518,260,283]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B941EAAE381AD63D96822D07FDA4CD1C" box="[428,518,260,283]" italics="true" pageId="7" pageNumber="308">Arcania</emphasis>
</taxonomicName>
species collected from Moreton Bay, including “ 
<taxonomicName id="4C354D3F381AD63D93052D07FB0ECD1A" ID-CoL="G7BK" box="[1067,1196,260,285]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B941EAAE381AD63D93052D07FB0ECD1A" box="[1067,1196,260,285]" italics="true" pageId="7" pageNumber="308">A. elongata</emphasis>
</taxonomicName>
” of 
<bibRefCitation id="EFA44B4D381AD63D93F32D07FA3ECD1B" author="Campbell" box="[1245,1436,260,285]" pageId="7" pageNumber="308" refString="Campbell, B. M. (1971) New records and new species of crabs (Crustacea: Brachyura) trawled off southern Queensland: Dromiacea, Homolidea, Gymnopleura, Corystoidea, and Oxystomata). Memoirs of the Queensland Museum, 16 (1), 27 - 48, pls. 2, 3." type="journal article" year="1971">Campbell (1971)</bibRefCitation>
(see 
<emphasis id="B941EAAE381AD63D97E52D24FE9ACD47" bold="true" box="[203,312,295,320]" pageId="7" pageNumber="308">Remarks</emphasis>
for 
<taxonomicName id="4C354D3F381AD63D96482D2BFE45CD47" ID-CoL="G7BK" box="[358,487,295,320]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B941EAAE381AD63D96482D2BFE45CD47" box="[358,487,295,320]" italics="true" pageId="7" pageNumber="308">A. elongata</emphasis>
</taxonomicName>
), should be reconsidered in future studies.
</paragraph>
<paragraph id="8B8A36BC381AD63D97E92D4FFAEECD8F" blockId="7.[151,1436,151,861]" pageId="7" pageNumber="308">
<bibRefCitation id="EFA44B4D381AD63D97E92D4FFE0ECD62" author="Alcock" box="[199,428,332,357]" pageId="7" pageNumber="308" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystomata. Journal of the Asiatic Society of Bengal, 65 (2), 134 - 296, pls. 6 - 8." type="journal article" year="1896">Alcock (1896: 267)</bibRefCitation>
and 
<bibRefCitation id="EFA44B4D381AD63D96C22D4EFDC5CD62" author="Tan" box="[492,615,332,357]" pageId="7" pageNumber="308" refString="Tan, C. G. S. (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30, 1021 - 1058. http: // dx. doi. org / 10.1080 / 00222939600770551" type="journal article" year="1996">Tan (1996</bibRefCitation>
: 1028) mentioned that 
<taxonomicName id="4C354D3F381AD63D94522D4FFB9CCD63" ID-CoL="G7BZ" box="[892,1086,332,356]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="novemspinosa">
<emphasis id="B941EAAE381AD63D94522D4FFB9CCD63" box="[892,1086,332,356]" italics="true" pageId="7" pageNumber="308">A. novemspinosa</emphasis>
</taxonomicName>
may be synonymous with 
<taxonomicName id="4C354D3F381AD63D92AA2D4FFEE9CD8F" ID-CoL="G7CC" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D92AA2D4FFEE9CD8F" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
, but they are clearly different species (see 
<bibRefCitation id="EFA44B4D381AD63D94042D6CFC06CD8F" author="Galil" box="[810,932,367,392]" pageId="7" pageNumber="308" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil 2001</bibRefCitation>
; 
<bibRefCitation id="EFA44B4D381AD63D949E2D6CFB38CD8F" author="Fujii" box="[944,1178,367,392]" pageId="7" pageNumber="308" refString="Fujii, T. &amp; Naruse, T. (2013) First record of Arcania novemspinosa (Crustacea: Decapoda: Leucosiidae) from Okinawa Island, Japan. Fauna Ryukyuana, 3, 1 - 6. [in Japanese]" type="journal article" year="2013">Fujii &amp; Naruse 2013</bibRefCitation>
; present study).
</paragraph>
<paragraph id="8B8A36BC381AD63D97E92D97FDAACE10" blockId="7.[151,1436,151,861]" pageId="7" pageNumber="308">
<bibRefCitation id="EFA44B4D381AD63D97E92D97FE6DCDAB" author="Hill" box="[199,463,404,429]" pageId="7" pageNumber="308" refString="Hill, D. S. (1982) The Leucosiidae (Crustacea: Decapoda) of Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong, Hong Kong University Press, pp. 195 - 205." type="book chapter" year="1982">Hill (1982: 201, pl. 4D)</bibRefCitation>
recorded “ 
<taxonomicName id="4C354D3F381AD63D95642D97FCAACDAB" ID-CoL="G7BZ" box="[586,776,404,428]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="novemspinosa">
<emphasis id="B941EAAE381AD63D95642D97FCAACDAB" box="[586,776,404,428]" italics="true" pageId="7" pageNumber="308">A. novemspinosa</emphasis>
</taxonomicName>
” from 
<collectingCountry id="F322762C381AD63D94762D96FC7FCDAA" box="[856,989,405,429]" name="Hong Kong" pageId="7" pageNumber="308">Hong Kong</collectingCountry>
, but its less produced and obtuse frontal lobes indicate that it is 
<taxonomicName id="4C354D3F381AD63D968D2DBBFDDACDD7" ID-CoL="G7CC" box="[419,632,439,464]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D968D2DBBFDDACDD7" box="[419,632,439,464]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
. He also noted the body color of the specimen as “uniformly pinkishbrown with two faint broad darker stripes dorsally down the carapace” (
<bibRefCitation id="EFA44B4D381AD63D94F42DDFFB2ACDF3" author="Hill" box="[986,1160,476,501]" pageId="7" pageNumber="308" refString="Hill, D. S. (1982) The Leucosiidae (Crustacea: Decapoda) of Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong, Hong Kong University Press, pp. 195 - 205." type="book chapter" year="1982">Hill 1982: 201</bibRefCitation>
), which agrees with the coloration of 
<taxonomicName id="4C354D3F381AD63D96032E03FDA2CE1F" ID-CoL="G7CC" box="[301,512,511,536]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D96032E03FDA2CE1F" box="[301,512,511,536]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8A36BC381AD63D97E92E27FA9BCF5A" blockId="7.[151,1436,151,861]" pageId="7" pageNumber="308">
<bibRefCitation id="EFA44B4D381AD63D97E92E27FEF5CE3B" author="Galil" box="[199,343,548,573]" pageId="7" pageNumber="308" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil (2001)</bibRefCitation>
listed a record of 
<taxonomicName id="4C354D3F381AD63D956C2E27FCA9CE3B" ID-CoL="G7BL" box="[578,779,548,572]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="erinacea">
<emphasis id="B941EAAE381AD63D956C2E27FCA9CE3B" box="[578,779,548,572]" italics="true" pageId="7" pageNumber="308">Arcania erinacea</emphasis>
</taxonomicName>
from 
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by 
<bibRefCitation id="EFA44B4D381AD63D94DF2E27FAEBCE3B" author="Tirmizi" box="[1009,1353,548,573]" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi (1988: 72)</bibRefCitation>
in the synonymy list of 
<taxonomicName id="4C354D3F381AD63D964C2E4BFD95CE67" ID-CoL="G7CC" box="[354,567,583,608]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D964C2E4BFD95CE67" box="[354,567,583,608]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
. The drawing by 
<bibRefCitation id="EFA44B4D381AD63D94262E44FB20CE67" author="Tirmizi" box="[776,1154,583,608]" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi (1988: fig. 20)</bibRefCitation>
, however, clearly shows serrate meri of the cheliped and fourth ambulatory leg, which are not observed in 
<taxonomicName id="4C354D3F381AD63D936B2E6FFAB8CE82" ID-CoL="G7CC" box="[1093,1306,620,645]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D936B2E6FFAB8CE82" box="[1093,1306,620,645]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFA44B4D381AD63D92042E6FFE19CEAF" author="Tirmizi" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi (1988: 76, fig. 22)</bibRefCitation>
provisionally identified other specimens from Karachi, 
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as 
<taxonomicName id="4C354D3F381AD63D93EE2E93FA36CEAF" ID-CoL="G7CC" box="[1216,1428,655,680]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D93EE2E93FA36CEAF" box="[1216,1428,655,680]" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
. As 
<bibRefCitation id="EFA44B4D381AD63D97E82EB7FE56CECB" author="Tirmizi" box="[198,500,692,717]" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi (1988)</bibRefCitation>
noted, the G1 of the 
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specimen differs markedly from that of 
<taxonomicName id="4C354D3F381AD63D92AA2EB7FEE9CEF7" ID-CoL="G7CC" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" sensu="stricto" species="undecimspinosa">
<emphasis id="B941EAAE381AD63D92AA2EB7FEE9CEF7" italics="true" pageId="7" pageNumber="308">A. undecimspinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27257D5381AD63D96782EDBFED8CEE8" box="[342,378,728,751]" pageId="7" pageNumber="308" sensu="stricto">s.s.</taxonomicNameLabel>
in its strong distal curvature over the distal third (
<bibRefCitation id="EFA44B4D381AD63D94FC2ED4FAFBCEF7" author="Tirmizi" box="[978,1369,727,752]" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi 1988: fig. 22G</bibRefCitation>
). The distally bent G1 of the 
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specimen may be similar to that of 
<taxonomicName id="4C354D3F381AD63D94A72EFFFB00CF13" ID-CoL="67PSV" authority="Chen, 1989" authorityName="Chen" authorityYear="1989" box="[905,1186,764,789]" class="Malacostraca" family="Leucosiidae" genus="Arcania" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brevifrons">
<emphasis id="B941EAAE381AD63D94A72EFFFBBBCF12" box="[905,1049,764,789]" italics="true" pageId="7" pageNumber="308">A. brevifrons</emphasis>
Chen, 1989
</taxonomicName>
, but the former differs from the latter by “half tube bent portion” of the distal part of the G1 (
<bibRefCitation id="EFA44B4D381AD63D94E72F1CFAA2CF3F" author="Tirmizi" box="[969,1280,799,824]" pageId="7" pageNumber="308" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1988) Crustacea: Brachyura (Dromiacea, Archaeobrachyura, Oxystomata, Oxyrhyncha). Marine Fauna of Pakistan, 4, i - viii, 2 unnumbered color plates, 1 - 244, 1 unnumbered plate." type="journal article" year="1988">Tirmizi &amp; Kazmi 1988: 79</bibRefCitation>
) (flattened in Chen 1989: fig. 32e; 
<bibRefCitation id="EFA44B4D381AD63D96AA2F47FE5DCF5B" author="Galil" box="[388,511,836,861]" pageId="7" pageNumber="308" refString="Galil, B. S. (2001) A revision of the genus Arcania Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 169 - 205." type="journal article" year="2001">Galil 2001</bibRefCitation>
: fig. 4A). The identities of the two 
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species remain questionable.
</paragraph>
</subSubSection>
</treatment>
</document>