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<document id="07A8769F18E118ABD12417EFC34A2D3A" ID-CLB-Dataset="21904" ID-DOI="10.5852/ejt.2016.199" ID-GBIF-Dataset="d88184f4-ceef-4392-a444-0ac1552778d3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840484" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590159499057" checkinUser="valdenar" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak &amp; Backeljau, Thierry" docDate="2016" docId="0385FA5C9617FFA1FDE48977DE26C157" docLanguage="en" docName="ejt-2016-199.pdf.imf" docOrigin="European Journal of Taxonomy 199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thyropygus undulatus Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau 2016, sp. nov." docType="treatment" docUuid="972296F7-CF7B-4AD4-A2AA-3DCE22596B3E" docUuidSource="ZooBank" docVersion="13" lastPageNumber="29" masterDocId="FFBC8224960DFFBDFF848A31DC7FC003" masterDocTitle="A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="27" updateTime="1701998209473" updateUser="plazi" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="29B843ADA14A25D279C44970306D3B40" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="11405D9FEEC4D2CEC4F3E2DF26FA0127">
<mods:title id="64D9E326D5E3EEDA0EACA77CB98D6732">A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="E66BC46FE2683F2CC6A1460520D83A8A" type="personal">
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<mods:namePart id="48AFDC84F49B93B8CF5DE451C8529554">Pimvichai, Piyatida</mods:namePart>
<mods:nameIdentifier id="7298DF508598208647CD5E59D6665EC7" type="LSID">4D882A89-E0C1-4F78-93E7-FA1B5A2650F2</mods:nameIdentifier>
<mods:affiliation id="4996632B120A5DFC00A055E9CD2411D7">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand. &amp; urn: lsid: zoobank. org: author: 4 D 882 A 89 - E 0 C 1 - 4 F 78 - 93 E 7 - FA 1 B 5 A 2650 F 2 &amp; Corresponding author: piyatida _ pimvichai @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="920690FDDAF1D705C3C772AE5FF59CDE" type="email">piyatida_pimvichai@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="B92B12A496511220130E4E76D9F9DF84">Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier id="2346792644152627FA29C043FA008300" type="LSID">FB09A817-000D-43C3-BCC4-2BC1E5373635</mods:nameIdentifier>
<mods:affiliation id="4EEDEED506B981393E9D8EB7C82AD6C7">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. &amp; Email: henghoff @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635</mods:affiliation>
<mods:nameIdentifier id="DEA98CA0181682A2E99E58D709418DCD" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
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<mods:role id="3526BDE4D91516999AD755B216C8D4BA">
<mods:roleTerm id="711CF6B51171EB496D1BC3900C8033C9">Author</mods:roleTerm>
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<mods:namePart id="99F457585410CD909B6CF378E612CBA2">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="38B69870DE5D14FBDB42AD37849EA229" type="LSID">AC935098-D901-4F35-A414-4B0D4FE44E79</mods:nameIdentifier>
<mods:affiliation id="2005ED43020131409103D4DD14478412">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Email: somsakp @ sc. chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79</mods:affiliation>
<mods:nameIdentifier id="4DB4DA02E37AEA261D321E6CFA6A02FA" type="email">somsakp@sc.chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="56C19FCEED0EC11399A6649B94818C52">Backeljau, Thierry</mods:namePart>
<mods:nameIdentifier id="39E4A180BA89705B80A3C705A6E6ED7C" type="LSID">E6F42575-36AE-4AD7-98C6-D083EF052568</mods:nameIdentifier>
<mods:affiliation id="2961C16FC4D455A605C8E6B506A1AE55">University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. &amp; Email: Thierry. Backeljau @ naturalsciences. be &amp; urn: lsid: zoobank. org: author: E 6 F 42575 - 36 AE- 4 AD 7 - 98 C 6 - D 083 EF 052568</mods:affiliation>
<mods:nameIdentifier id="F86274690E0B81CC097A82CBB5D353E4" type="email">ackeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="2011E45BCE4835403BF3A74F533D2CCA">European Journal of Taxonomy</mods:title>
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<mods:part id="BE49A69490C973880D78C5AABC546CDA">
<mods:date id="C8249C2D0D4B6CB9D7CF2411098DC56B">2016</mods:date>
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<mods:number id="4B9756A33A46DE084DF91AB9E6EC206B">2016-05-24</mods:number>
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<mods:number id="0BEE2CFA91FA9598D9D3B4257E31CA30">199</mods:number>
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<mods:identifier id="05E45EB6246F05E95905779546B24E78" type="DOI">10.5852/ejt.2016.199</mods:identifier>
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<mods:identifier id="554C14747187C7A79CD8A543E17373F9" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="629DA6E4CF03C00EECCBFC2D270023BC" type="Zenodo-Dep">3840484</mods:identifier>
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<treatment id="0385FA5C9617FFA1FDE48977DE26C157" ID-DOI="http://doi.org/10.5281/zenodo.3853326" ID-GBIF-Taxon="164249486" ID-Zenodo-Dep="3853326" LSID="urn:lsid:zoobank.org:act:972296F7-CF7B-4AD4-A2AA-3DCE22596B3E" httpUri="http://treatment.plazi.org/id/0385FA5C9617FFA1FDE48977DE26C157" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C33618C19617FFA7FDE48977D8B2C387" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B934B4A9617FFA7FDE48977DFACC362" blockId="26.[358,1229,838,935]" box="[608,979,838,865]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FDE48977DFACC362" box="[608,979,838,865]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="4C2C30C99617FFA7FDE48977DF0EC362" ID-CoL="56T4N" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[608,881,838,865]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undulatus" status="sp. nov.">
<emphasis id="B95897589617FFA7FDE48977DF0EC362" bold="true" box="[608,881,838,865]" italics="true" pageId="26" pageNumber="27">Thyropygus undulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239617FFA7FCFD8976DFACC362" box="[889,979,839,865]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B934B4A9617FFA7FEE28958D8B2C387" blockId="26.[358,1229,838,935]" box="[358,1229,873,900]" pageId="26" pageNumber="27">
<uri id="FFBD47489617FFA7FEE28958D8B2C387" box="[358,1229,873,900]" pageId="26" pageNumber="27">
urn:lsid:zoobank.org:act:
<uuid id="FF8A719F9617FFA7FD078958D8B2C387" box="[643,1229,873,900]" pageId="26" pageNumber="27">972296F7-CF7B-4AD4-A2AA-3DCE22596B3E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FD5989BCDF28C3A4" box="[733,855,909,935]" pageId="26" pageNumber="27" type="description">
<paragraph id="8B934B4A9617FFA7FD5989BCDF28C3A4" blockId="26.[358,1229,838,935]" box="[733,855,909,935]" pageId="26" pageNumber="27">
<figureCitation id="131757CF9617FFA7FD5989BCDF60C3A4" box="[733,799,909,935]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
A–E
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FF3989FFDD37C4A0" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8B934B4A9617FFA7FF3989FFDD49C3EB" blockId="26.[189,310,974,1000]" box="[189,310,974,1000]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FF3989FFDD49C3EB" bold="true" box="[189,310,974,1000]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B95897589617FFA7FF3989FFDD49C3EB" bold="true" box="[189,310,974,1000]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF3989CCDD37C4A0" blockId="26.[189,1399,1021,1187]" pageId="26" pageNumber="27">
A species of the
<taxonomicName id="4C2C30C99617FFA7FEF389CFDDA3C414" authorityName="Arw" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[375,476,1022,1047]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="B95897589617FFA7FEF389CFDDA3C414" box="[375,476,1022,1047]" italics="true" pageId="26" pageNumber="27">opinatus</emphasis>
</taxonomicName>
subgroup. Lateral process (
<emphasis id="B95897589617FFA7FC9F89CCDF3FC414" box="[795,832,1021,1047]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
) flattened, slightly curved, its laterodistal margin coarsely dentate. Similar in this respect to
<taxonomicName id="4C2C30C99617FFA7FD2F8E10DF3EC439" box="[683,833,1056,1082]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cristagalli">
<emphasis id="B95897589617FFA7FD2F8E10DF3EC439" box="[683,833,1056,1082]" italics="true" pageId="26" pageNumber="27">T. cristagalli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C30C99617FFA7FCCA8E10DF9BC439" baseAuthorityName="Demange" baseAuthorityYear="1961" box="[846,996,1056,1082]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="implicatus">
<emphasis id="B95897589617FFA7FCCA8E10DF9BC439" box="[846,996,1056,1082]" italics="true" pageId="26" pageNumber="27">T. implicatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C30C99617FFA7FB988E10D802C439" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[1052,1149,1056,1082]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="culter" status="sp. nov.">
<emphasis id="B95897589617FFA7FB988E10D802C439" box="[1052,1149,1056,1082]" italics="true" pageId="26" pageNumber="27">T. culter</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239617FFA7FB008E11D8A1C439" box="[1156,1246,1056,1082]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
Differs from these species by having the mesal process of the posterior coxal fold (
<emphasis id="B95897589617FFA7FB968E75D83AC45E" box="[1042,1093,1092,1117]" italics="true" pageId="26" pageNumber="27">pmp</emphasis>
) with two rounded distal lobes, visible in anterior view between
<emphasis id="B95897589617FFA7FD0B8E57DECBC483" box="[655,692,1126,1152]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
and
<emphasis id="B95897589617FFA7FD6B8E56DF56C483" box="[751,809,1127,1152]" italics="true" pageId="26" pageNumber="27">amp,</emphasis>
and by having the tibial spine (
<emphasis id="B95897589617FFA7FB268E56D8CDC483" box="[1186,1202,1127,1152]" italics="true" pageId="26" pageNumber="27">ti</emphasis>
) not ending in a sharp point.
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FF398EFADE37C534" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8B934B4A9617FFA7FF398EFADD3BC4E6" blockId="26.[189,324,1227,1253]" box="[189,324,1227,1253]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FF398EFADD3BC4E6" bold="true" box="[189,324,1227,1253]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B95897589617FFA7FF398EFADD3BC4E6" bold="true" box="[189,324,1227,1253]" pageId="26" pageNumber="27">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398EC8DE37C534" blockId="26.[189,1398,1273,1335]" pageId="26" pageNumber="27">The name is a Latin adjective, referring to the undulate/coarsely dentate laterodistal margin of the lateral process of the anterior coxal fold.</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FF398F6FDFFEC676" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B934B4A9617FFA7FF398F6FDDD5C57B" blockId="26.[189,426,1374,1400]" box="[189,426,1374,1400]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FF398F6FDDD5C57B" bold="true" box="[189,426,1374,1400]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B95897589617FFA7FF398F6FDDD5C57B" bold="true" box="[189,426,1374,1400]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B4441179617FFA7FF6E8FBCDDEAC613" ID-GBIF-Occurrence="2625222301" collectingDate="2009-01-16" collectionCode="CUMZ-D" collectorName="P. Pimvichai &amp; Pimvichai, C &amp; Animal Systematics Research Unit" country="Thailand" latitude="8.275" location="Khao Phanom Bencha" longLatPrecision="21" longitude="98.924995" municipality="Muang District" pageId="26" pageNumber="27" specimenCode="CUMZ-D00087" specimenCount="1" stateProvince="Krabi" typeStatus="holotype">
<paragraph id="8B934B4A9617FFA7FF6E8FBCDD23C5A4" blockId="26.[189,1399,1421,1552]" box="[234,348,1421,1447]" pageId="26" pageNumber="27">
<emphasis id="B95897589617FFA7FF6E8FBCDD23C5A4" bold="true" box="[234,348,1421,1447]" pageId="26" pageNumber="27">
<typeStatus id="5497F5E89617FFA7FF6E8FBCDD23C5A4" box="[234,348,1421,1447]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398F81DDEAC613" blockId="26.[189,1399,1421,1552]" pageId="26" pageNumber="27">
<collectingCountry id="F33B0BDA9617FFA7FF398F81DD26C5C9" box="[189,345,1456,1482]" name="Thailand" pageId="26" pageNumber="27">THAILAND</collectingCountry>
: ³,
<collectingRegion id="49E885A89617FFA7FE158F81DE35C5C9" box="[401,586,1456,1482]" country="Thailand" name="Krabi" pageId="26" pageNumber="27">Krabi Province</collectingRegion>
,
<collectingMunicipality id="6BF7D1309617FFA7FDDE8F81DF6CC5C9" box="[602,787,1456,1482]" pageId="26" pageNumber="27">Muang District</collectingMunicipality>
,
<location id="8EF31D919617FFA7FCA78F81D84BC5C9" LSID="urn:lsid:plazi:treatment:0385FA5C9617FFA1FDE48977DE26C157:8EF31D919617FFA7FCA78F81D84BC5C9" box="[803,1076,1456,1482]" country="Thailand" latitude="8.275" longLatPrecision="21" longitude="98.924995" municipality="Muang District" name="Khao Phanom Bencha" pageId="26" pageNumber="27" stateProvince="Krabi">Khao Phanom Bencha</location>
,
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,
<geoCoordinate id="EE182D8D9617FFA7FB598F81D90EC5C9" box="[1245,1393,1456,1483]" degrees="98" direction="east" minutes="55" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="30" value="98.924995">98°55'30&quot; E</geoCoordinate>
,
<date id="FF926D8A9617FFA7FF398FE2DD24C5EE" box="[189,347,1491,1518]" pageId="26" pageNumber="27" value="2009-01-16">
<collectingDate id="EFD694629617FFA7FF398FE2DD24C5EE" box="[189,347,1491,1518]" pageId="26" pageNumber="27" value="2009-01-16">16 Jan. 2009</collectingDate>
</date>
, leg.
<collectorName id="26D92E9C9617FFA7FE1A8FE5DE49C5EE" box="[414,566,1491,1518]" pageId="26" pageNumber="27">P. Pimvichai</collectorName>
, C. Sutcharit and members of the
<collectorName id="26D92E9C9617FFA7FC518FE2D909C5EE" box="[981,1398,1491,1517]" pageId="26" pageNumber="27">Animal Systematics Research Unit</collectorName>
(
<specimenCode id="DB8AE3319617FFA7FF418FC7DDF5C613" box="[197,394,1526,1552]" collectionCode="CUMZ-D" pageId="26" pageNumber="27">CUMZ-D00087</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8B934B4A9617FFA7FF6E8C09DD18C651" blockId="26.[189,897,1592,1653]" box="[234,359,1592,1618]" pageId="26" pageNumber="27">
<emphasis id="B95897589617FFA7FF6E8C09DD18C651" bold="true" box="[234,359,1592,1618]" pageId="26" pageNumber="27">
<typeStatus id="5497F5E89617FFA7FF6E8C09DD18C651" box="[234,359,1592,1618]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398C6ADFFEC676" blockId="26.[189,897,1592,1653]" box="[189,897,1627,1653]" pageId="26" pageNumber="27">
<collectingCountry id="F33B0BDA9617FFA7FF398C6ADD26C676" box="[189,345,1627,1653]" name="Thailand" pageId="26" pageNumber="27">THAILAND</collectingCountry>
:
<specimenCount id="9D2A80C39617FFA7FEE08C6ADDEFC676" box="[356,400,1627,1653]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
, same data as
<typeStatus id="5497F5E89617FFA7FDBE8C6ADEDEC676" box="[570,673,1627,1653]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3DD38F9617FFA7FD348C6ADF56C676" box="[688,809,1627,1653]" pageId="26" pageNumber="27">CUMZ-D</collectionCode>
00099).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA1FF398CADDE26C157" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="8B934B4A9617FFA7FF398CADDD31C6B5" blockId="26.[189,334,1692,1718]" box="[189,334,1692,1718]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FF398CADDD31C6B5" bold="true" box="[189,334,1692,1718]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B95897589617FFA7FF398CADDD31C6B5" bold="true" box="[189,334,1692,1718]" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398CFADF6AC74D" blockId="26.[189,1398,1739,1871]" pageId="26" pageNumber="27">
Adult male with 63 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9617FFA7FC0A8CFADFA4C6E6" box="[910,987,1739,1766]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="cm" value="15.0">15 cm</quantity>
, width
<quantity id="4CD4E6AF9617FFA7FBB08CFAD8ECC6E6" box="[1076,1171,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="26" pageNumber="27" unit="mm" value="8.7">8.7 mm</quantity>
. Adult female with 64 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9617FFA7FD438CDFDF6BC70A" box="[711,788,1774,1801]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="27" unit="cm" value="14.0">14 cm</quantity>
, width
<quantity id="4CD4E6AF9617FFA7FCEA8CDFDFB2C70A" box="[878,973,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="26" pageNumber="27" unit="mm" value="8.5">8.5 mm</quantity>
. Colour in life unknown; preserved specimens with head, legs, middorsal metazona, epiproct, paraprocts and hypoproct brown; antennae and metazona dark brown; prozona whitish-brown.
</paragraph>
<paragraph id="8B934B4A9617FFA6FF398D47D8E2C21F" blockId="26.[189,1398,1910,2007]" lastBlockId="27.[189,1399,267,540]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
GONOPODS (
<figureCitation id="131757CF9617FFA7FEC08D47DDF8C793" box="[324,391,1910,1936]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
A–E). Anterior coxal fold (
<emphasis id="B95897589617FFA7FD418D46DE9EC793" box="[709,737,1911,1936]" italics="true" pageId="26" pageNumber="27">ac</emphasis>
;
<figureCitation id="131757CF9617FFA7FD6B8D47DF32C793" box="[751,845,1910,1936]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
): lateral process (
<emphasis id="B95897589617FFA7FB9E8D47D840C793" box="[1050,1087,1910,1936]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
) flattened, curving mesad, its laterodistal margin coarsely dentate, terminating in a short spine, tip curving against tip of opposite side, lateral margins with serrate crest (
<figureCitation id="131757CF9617FFA7FD1D8D8DDE88C7D5" box="[665,759,1980,2006]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="26" pageNumber="27">Fig. 9C</figureCitation>
, arrow); mesal process (
<emphasis id="B95897589617FFA7FBA48D8CD82CC7D5" box="[1056,1107,1981,2006]" italics="true" pageId="26" pageNumber="27">amp</emphasis>
) much shorter than
<emphasis id="B95897589617FFA7FACE8D8DD910C7D5" box="[1354,1391,1980,2006]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
, slender, curving mesad. Posterior coxal fold (
<emphasis id="B95897589616FFA6FD478B3DDEA0C126" box="[707,735,268,293]" italics="true" pageId="27" pageNumber="28">pc</emphasis>
;
<figureCitation id="131757CF9616FFA6FD688B3ADF3DC126" box="[748,834,267,293]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="27" pageNumber="28">Fig. 9B</figureCitation>
) basally with moderately high lateral paracoxites (
<emphasis id="B95897589616FFA6FF438B1EDC9CC14B" box="[199,227,303,328]" italics="true" pageId="27" pageNumber="28">px</emphasis>
); mesal process (
<emphasis id="B95897589616FFA6FE288B1EDDA0C14B" box="[428,479,303,328]" italics="true" pageId="27" pageNumber="28">pmp</emphasis>
) distally with two rounded lobes, visible in anterior view between
<emphasis id="B95897589616FFA6FB648B1ED97AC14A" box="[1248,1285,303,329]" italics="true" pageId="27" pageNumber="28">alp</emphasis>
and
<emphasis id="B95897589616FFA6FAB88B1ED910C14B" box="[1340,1391,303,328]" italics="true" pageId="27" pageNumber="28">amp</emphasis>
; lateral process (
<emphasis id="B95897589616FFA6FEFA8B63DDDCC16F" box="[382,419,338,364]" italics="true" pageId="27" pageNumber="28">plp</emphasis>
) erect, digitiform. Telopodite (
<figureCitation id="131757CF9616FFA6FC9A8B63DF19C16F" box="[798,870,338,364]" captionStart="Fig" captionStartId="27.[189,232,1930,1956]" captionTargetBox="[257,1362,777,1891]" captionTargetId="figure@27.[250,1368,770,1901]" captionTargetPageId="27" captionText="Fig. 9. Thyropygus undulatus sp. nov., from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="27" pageNumber="28">Fig. 9</figureCitation>
D–E) leaving coxite over shelf of posterior coxal fold; femoral spine (
<emphasis id="B95897589616FFA6FE748B44DE7AC18C" box="[496,517,373,399]" italics="true" pageId="27" pageNumber="28">fe</emphasis>
) duplicated,
<emphasis id="B95897589616FFA6FD1A8B44DEB7C18C" box="[670,712,373,399]" italics="true" pageId="27" pageNumber="28">fe 1</emphasis>
relatively long, projecting above top of telopodite, curving backward, ending in slender barb,
<emphasis id="B95897589616FFA6FDD38BA9DEFCC1B1" box="[599,643,408,434]" italics="true" pageId="27" pageNumber="28">fe 2</emphasis>
basally broad, situated under
<emphasis id="B95897589616FFA6FC6C8BA9D86BC1B1" box="[1000,1044,408,434]" italics="true" pageId="27" pageNumber="28">fe 1</emphasis>
, curving backward, ending in slender barb; tibial spine (
<emphasis id="B95897589616FFA6FE738B8DDE78C1D6" box="[503,519,444,469]" italics="true" pageId="27" pageNumber="28">ti</emphasis>
) long, curving in horizontal plane, not ending in sharp spine; apical part: spatulate lobe (
<emphasis id="B95897589616FFA6FEF58BEEDDFAC1FA" box="[369,389,479,505]" italics="true" pageId="27" pageNumber="28">sl</emphasis>
) rounded, with slender long spine (
<emphasis id="B95897589616FFA6FCAF8BEEDF35C1FA" box="[811,842,479,505]" italics="true" pageId="27" pageNumber="28">sls</emphasis>
) at base of apical part, opposite origin of tibial spine; palette (
<emphasis id="B95897589616FFA6FEEE8832DDF8C21F" box="[362,391,515,540]" italics="true" pageId="27" pageNumber="28">pa</emphasis>
) simple, distally with about eleven brownish blepharochaetae (
<emphasis id="B95897589616FFA6FBEB8833D8F3C21F" box="[1135,1164,514,540]" italics="true" pageId="27" pageNumber="28">bp</emphasis>
).
</paragraph>
<paragraph id="8B934B4A9616FFA6FF398872DD19C25E" blockId="27.[189,358,579,605]" box="[189,358,579,605]" pageId="27" pageNumber="28">
<heading id="D0DBFC269616FFA6FF398872DD19C25E" bold="true" box="[189,358,579,605]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B95897589616FFA6FF398872DD19C25E" bold="true" box="[189,358,579,605]" pageId="27" pageNumber="28">DNA barcode</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9616FFA6FF398843DDF3C2AC" blockId="27.[189,1398,626,687]" pageId="27" pageNumber="28">
The GenBank accession number of the barcode of the
<typeStatus id="5497F5E89616FFA6FC118843DF83C28F" box="[917,1020,626,652]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
is
<accessionNumber id="947FD6A99616FFA6FBB68843D8CBC28E" box="[1074,1204,626,653]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU306520" pageId="27" pageNumber="28">KU306520</accessionNumber>
(voucher code CUMZ-D00087).
</paragraph>
<caption id="DF531BC29616FFA6FF398DBBD949C7E9" ID-DOI="http://doi.org/10.5281/zenodo.3840502" ID-Zenodo-Dep="3840502" httpUri="https://zenodo.org/record/3840502/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1930,1956]" targetBox="[257,1362,777,1891]" targetPageId="27">
<paragraph id="8B934B4A9616FFA6FF398DBBD949C7E9" blockId="27.[189,1399,1928,2027]" pageId="27" pageNumber="28">
<emphasis id="B95897589616FFA6FF398DBBDD71C7A7" bold="true" box="[189,270,1930,1956]" pageId="27" pageNumber="28">Fig. 9.</emphasis>
<taxonomicName id="4C2C30C99616FFA6FE9E8DBBDE5CC7A7" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[282,547,1930,1956]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undulatus" status="sp. nov.">
<emphasis id="B95897589616FFA6FE9E8DBBDE5CC7A7" box="[282,547,1930,1956]" italics="true" pageId="27" pageNumber="28">Thyropygus undulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239616FFA6FDAA8DBBDEF0C7A7" box="[558,655,1930,1956]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, from Khao Phanom Bencha, holotype (CUMZ-D00087), ³, gonopods.
<emphasis id="B95897589616FFA6FEC18D9CDD25C7C4" bold="true" box="[325,346,1965,1991]" pageId="27" pageNumber="28">A</emphasis>
. Anterior view, left telopodite removed.
<emphasis id="B95897589616FFA6FCE58D9CDF0AC7C4" bold="true" box="[865,885,1965,1991]" pageId="27" pageNumber="28">B</emphasis>
. Posterior view, left telopodite removed.
<emphasis id="B95897589616FFA6FF398DE1DCADC7E9" bold="true" box="[189,210,2000,2026]" pageId="27" pageNumber="28">C</emphasis>
. Lateral view.
<emphasis id="B95897589616FFA6FE058DE1DDE9C7E9" bold="true" box="[385,406,2000,2026]" pageId="27" pageNumber="28">D</emphasis>
. Left telopodite, posterior-mesal view.
<emphasis id="B95897589616FFA6FCE08DE1DF07C7E9" bold="true" box="[868,888,2000,2026]" pageId="27" pageNumber="28">E</emphasis>
. Left telopodite, anterior-lateral view.
</paragraph>
</caption>
<paragraph id="8B934B4A9611FFA1FF398B3ADDBBC126" blockId="28.[189,452,267,293]" box="[189,452,267,293]" pageId="28" pageNumber="29">
<emphasis id="B95897589611FFA1FF398B3ADD29C126" bold="true" box="[189,342,267,293]" pageId="28" pageNumber="29">Distribution</emphasis>
(
<figureCitation id="131757CF9611FFA1FEE28B3ADDC4C126" box="[358,443,267,293]" captionStart="Fig" captionStartId="30.[189,232,1998,2024]" captionTargetBox="[189,1398,829,1964]" captionTargetId="figure@30.[189,1398,829,1964]" captionTargetPageId="30" captionText="Fig. 12. Known distribution of the species of the T. opinatus subgroup." figureDoi="http://doi.org/10.5281/zenodo.3840510" httpUri="https://zenodo.org/record/3840510/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8B934B4A9611FFA1FF398B0BDE26C157" blockId="28.[189,601,314,340]" box="[189,601,314,340]" pageId="28" pageNumber="29">
Known only from the
<typeStatus id="5497F5E89611FFA1FE468B0BDD8AC157" box="[450,501,314,340]" pageId="28" pageNumber="29">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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