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<document id="A70A1F582F54423C59B644B9B2E2AF6D" ID-CLB-Dataset="21904" ID-DOI="10.5852/ejt.2016.199" ID-GBIF-Dataset="d88184f4-ceef-4392-a444-0ac1552778d3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840484" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590159499057" checkinUser="valdenar" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak &amp; Backeljau, Thierry" docDate="2016" docId="0385FA5C9611FFA0FDF88B4ADE26C5CA" docLanguage="en" docName="ejt-2016-199.pdf.imf" docOrigin="European Journal of Taxonomy 199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thyropygus ursus Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau 2016, sp. nov." docType="treatment" docUuid="D3E091F2-C075-458B-A9B5-DA35B673309B" docUuidSource="ZooBank" docVersion="12" lastPageNumber="30" masterDocId="FFBC8224960DFFBDFF848A31DC7FC003" masterDocTitle="A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="29" updateTime="1698841008082" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6379C081494C5B19E1C8BC477F8398F9">A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:name id="F0E7CD1702829685E4CBB287CBD946CA" type="personal">
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<mods:namePart id="E3B1AE80A0F9C930EAF1935B5C9DF058">Pimvichai, Piyatida</mods:namePart>
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<mods:affiliation id="41F095B482BE2BD3C1C7D746892019F8">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand. &amp; urn: lsid: zoobank. org: author: 4 D 882 A 89 - E 0 C 1 - 4 F 78 - 93 E 7 - FA 1 B 5 A 2650 F 2 &amp; Corresponding author: piyatida _ pimvichai @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="03CB1AF603E457D2705739CA83A4041F" type="email">piyatida_pimvichai@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="2666AD6EAC9AEAEBD7223162562B96A2">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="16C69EC3E3A50175D9BEDFD14DCE20DA">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. &amp; Email: henghoff @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635</mods:affiliation>
<mods:nameIdentifier id="8901FEE9311B7979C2B4D1911EFEAD29" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="CB8EB74DA980B3CB27F4C5ED992E5C02">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="4F754AFF1620F0E61972A8A7546629E9" type="LSID">AC935098-D901-4F35-A414-4B0D4FE44E79</mods:nameIdentifier>
<mods:affiliation id="72796940E097875B9F61AAEE43C9FDCB">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Email: somsakp @ sc. chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79</mods:affiliation>
<mods:nameIdentifier id="89EE97059E17EF67798E8FC4BAA074FA" type="email">somsakp@sc.chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="ECABFC24CB3EE525833B824746DB43BF">Backeljau, Thierry</mods:namePart>
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<mods:affiliation id="909B940645D5FDAE7AEBD10A3A19E106">University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. &amp; Email: Thierry. Backeljau @ naturalsciences. be &amp; urn: lsid: zoobank. org: author: E 6 F 42575 - 36 AE- 4 AD 7 - 98 C 6 - D 083 EF 052568</mods:affiliation>
<mods:nameIdentifier id="7A6D226A4BFE3BB06D2F925C29BF1DBE" type="email">ackeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="A41CFAC04DC98F637CDD552B14398B6A">European Journal of Taxonomy</mods:title>
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<mods:date id="74689D2A218C2D8197AC1E710968917B">2016</mods:date>
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<mods:number id="D25D26DEBDC9ADC30503730C9C51FAE5">2016-05-24</mods:number>
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<treatment id="0385FA5C9611FFA0FDF88B4ADE26C5CA" ID-DOI="http://doi.org/10.5281/zenodo.3853334" ID-GBIF-Taxon="164249488" ID-Zenodo-Dep="3853334" LSID="urn:lsid:zoobank.org:act:D3E091F2-C075-458B-A9B5-DA35B673309B" httpUri="http://treatment.plazi.org/id/0385FA5C9611FFA0FDF88B4ADE26C5CA" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C33618C19611FFA1FDF88B4AD8B8C1BA" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B934B4A9611FFA1FDF88B4ADFC7C195" blockId="28.[365,1223,379,476]" box="[636,952,379,406]" pageId="28" pageNumber="29">
<heading id="D0DBFC269611FFA1FDF88B4ADFC7C195" box="[636,952,379,406]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4C2C30C99611FFA1FDF88B4ADF29C195" ID-CoL="56T4Q" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[636,854,379,406]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B95897589611FFA1FDF88B4ADF29C195" bold="true" box="[636,854,379,406]" italics="true" pageId="28" pageNumber="29">Thyropygus ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239611FFA1FCD98B4DDFC7C195" box="[861,952,380,406]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FFBD47489611FFA1FEE98BAFD8B8C1BA" box="[365,1223,414,441]" pageId="28" pageNumber="29">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33618C19611FFA1FD508BF3DF1FC1DF" box="[724,864,450,476]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B934B4A9611FFA1FD508BF3DF1FC1DF" blockId="28.[365,1223,379,476]" box="[724,864,450,476]" pageId="28" pageNumber="29">
<figureCitation id="131757CF9611FFA1FD508BF3DF5BC1DF" box="[724,804,450,476]" captionStart="Fig" captionStartId="28.[189,232,1930,1956]" captionTargetBox="[289,1281,695,1897]" captionTargetId="figure@28.[277,1311,693,1910]" captionTargetPageId="28" captionText="Fig. 10. Thyropygus ursus sp. nov., from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="28" pageNumber="29">Fig. 10</figureCitation>
AD
</paragraph>
</subSubSection>
<subSubSection id="C33618C19611FFA1FF398832D851C290" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B934B4A9611FFA1FF398832DD49C21E" blockId="28.[189,310,515,541]" box="[189,310,515,541]" pageId="28" pageNumber="29">
<heading id="D0DBFC269611FFA1FF398832DD49C21E" bold="true" box="[189,310,515,541]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B95897589611FFA1FF398832DD49C21E" bold="true" box="[189,310,515,541]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9611FFA1FF398803D851C290" blockId="28.[189,1399,562,659]" pageId="28" pageNumber="29">
A species of the
<taxonomicName id="4C2C30C99611FFA1FEFB8802DD9BC24F" authorityName="Arw" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[383,484,563,588]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="B95897589611FFA1FEFB8802DD9BC24F" box="[383,484,563,588]" italics="true" pageId="28" pageNumber="29">opinatus</emphasis>
</taxonomicName>
subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold (
<emphasis id="B95897589611FFA1FDD88864DEFEC26C" box="[604,641,597,623]" italics="true" pageId="28" pageNumber="29">alp</emphasis>
) flattened, broad, apically curved caudad and ending in a short spine, as well as by the first femoral spine (
<emphasis id="B95897589611FFA1FD3D8849DE9AC291" box="[697,741,632,658]" italics="true" pageId="28" pageNumber="29">fe 1</emphasis>
) being very short and erect.
</paragraph>
</subSubSection>
<caption id="DF531BC29611FFA1FF398DBBDFEBC7E8" ID-DOI="http://doi.org/10.5281/zenodo.3840506" ID-Zenodo-Dep="3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1930,1956]" targetBox="[289,1281,695,1897]" targetPageId="28">
<paragraph id="8B934B4A9611FFA1FF398DBBDFEBC7E8" blockId="28.[189,1399,1929,2027]" pageId="28" pageNumber="29">
<emphasis id="B95897589611FFA1FF398DBBDD62C7A7" bold="true" box="[189,285,1930,1956]" pageId="28" pageNumber="29">Fig. 10.</emphasis>
<taxonomicName id="4C2C30C99611FFA1FEAC8DBBDD83C7A7" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[296,508,1930,1956]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B95897589611FFA1FEAC8DBBDD83C7A7" box="[296,508,1930,1956]" italics="true" pageId="28" pageNumber="29">Thyropygus ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239611FFA1FD838DBADE18C7A6" box="[519,615,1931,1957]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods.
<emphasis id="B95897589611FFA1FF398D9CDCADC7C4" bold="true" box="[189,210,1965,1991]" pageId="28" pageNumber="29">A</emphasis>
. Anterior view, left telopodite removed.
<emphasis id="B95897589611FFA1FD2F8D9FDEC0C7CB" bold="true" box="[683,703,1966,1992]" pageId="28" pageNumber="29">B</emphasis>
. Posterior view, left telopodite removed.
<emphasis id="B95897589611FFA1FB1B8D9CD8CBC7C4" bold="true" box="[1183,1204,1965,1991]" pageId="28" pageNumber="29">C</emphasis>
. Left telopodite, posterior-mesal view.
<emphasis id="B95897589611FFA1FE448DE0DDAAC7E8" bold="true" box="[448,469,2001,2027]" pageId="28" pageNumber="29">D</emphasis>
. Left telopodite, anterior-lateral view.
</paragraph>
</caption>
<subSubSection id="C33618C19610FFA0FF398B3ADF08C174" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B934B4A9610FFA0FF398B3ADD3BC126" blockId="29.[189,324,267,293]" box="[189,324,267,293]" pageId="29" pageNumber="30">
<heading id="D0DBFC269610FFA0FF398B3ADD3BC126" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B95897589610FFA0FF398B3ADD3BC126" bold="true" box="[189,324,267,293]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398B08DF08C174" blockId="29.[189,1399,313,375]" pageId="29" pageNumber="30">
The name is a Latin noun in apposition, meaning “bear”, and refers to the (somewhat) bearhead-like profile of the lateral process of the anterior coxal fold (
<emphasis id="B95897589610FFA0FCC48B6CDF1AC174" box="[832,869,349,375]" italics="true" pageId="29" pageNumber="30">alp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19610FFA0FF398BAFDF5BC291" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8B934B4A9610FFA0FF398BAFDDD5C1BB" blockId="29.[189,426,414,440]" box="[189,426,414,440]" pageId="29" pageNumber="30">
<heading id="D0DBFC269610FFA0FF398BAFDDD5C1BB" bold="true" box="[189,426,414,440]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B95897589610FFA0FF398BAFDDD5C1BB" bold="true" box="[189,426,414,440]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B4441179610FFA0FF6E8BFCDF3FC22D" ID-GBIF-Occurrence="2625222304" collectingDate="2009-01-02" collectorName="B. Daubl &amp; E. Haring" country="Thailand" latitude="7.6555557" location="Lanta Islands" longLatPrecision="21" longitude="99.04195" municipality="Koh Lanta District" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Krabi" typeStatus="holotype">
<paragraph id="8B934B4A9610FFA0FF6E8BFCDD23C1E4" blockId="29.[189,1399,461,558]" box="[234,348,461,487]" pageId="29" pageNumber="30">
<emphasis id="B95897589610FFA0FF6E8BFCDD23C1E4" bold="true" box="[234,348,461,487]" pageId="29" pageNumber="30">
<typeStatus id="5497F5E89610FFA0FF6E8BFCDD23C1E4" box="[234,348,461,487]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398BC1DF3FC22D" blockId="29.[189,1399,461,558]" pageId="29" pageNumber="30">
<collectingCountry id="F33B0BDA9610FFA0FF398BC1DD26C209" box="[189,345,496,522]" name="Thailand" pageId="29" pageNumber="30">THAILAND</collectingCountry>
: ³,
<collectingRegion id="49E885A89610FFA0FE0A8BC1DE3AC209" box="[398,581,496,522]" country="Thailand" name="Krabi" pageId="29" pageNumber="30">Krabi Province</collectingRegion>
,
<collectingMunicipality id="6BF7D1309610FFA0FDD08BC1DF47C209" box="[596,824,496,522]" pageId="29" pageNumber="30">Koh Lanta District</collectingMunicipality>
,
<location id="8EF31D919610FFA0FCC28BC1DF96C209" LSID="urn:lsid:plazi:treatment:0385FA5C9611FFA0FDF88B4ADE26C5CA:8EF31D919610FFA0FCC28BC1DF96C209" box="[838,1001,496,522]" country="Thailand" latitude="7.6555557" longLatPrecision="21" longitude="99.04195" municipality="Koh Lanta District" name="Lanta Islands" pageId="29" pageNumber="30" stateProvince="Krabi">Lanta Islands</location>
,
<geoCoordinate id="EE182D8D9610FFA0FC738BC1D800C209" box="[1015,1151,496,522]" degrees="7" direction="north" minutes="39" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="20" value="7.6555557">7°39'20&quot; N</geoCoordinate>
,
<geoCoordinate id="EE182D8D9610FFA0FB098BC1D95FC209" box="[1165,1312,496,522]" degrees="99" direction="east" minutes="02" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="31" value="99.04195">99°02'31&quot; E</geoCoordinate>
,
<date id="FF926D8A9610FFA0FAAB8BC1DC84C22E" pageId="29" pageNumber="30" value="2009-01-02">
<collectingDate id="EFD694629610FFA0FAAB8BC1DC84C22E" pageId="29" pageNumber="30" value="2009-01-02">2 Jan. 2009</collectingDate>
</date>
, leg.
<collectorName id="26D92E9C9610FFA0FEBD8822DDDCC22E" box="[313,419,531,557]" pageId="29" pageNumber="30">B. Däubl</collectorName>
and
<collectorName id="26D92E9C9610FFA0FE588822DE32C22E" box="[476,589,531,557]" pageId="29" pageNumber="30">E. Haring</collectorName>
(NMHW-Inv.7855).
</paragraph>
</materialsCitation>
<paragraph id="8B934B4A9610FFA0FF6E8864DD18C26C" blockId="29.[189,804,597,658]" box="[234,359,597,623]" pageId="29" pageNumber="30">
<emphasis id="B95897589610FFA0FF6E8864DD18C26C" bold="true" box="[234,359,597,623]" pageId="29" pageNumber="30">
<typeStatus id="5497F5E89610FFA0FF6E8864DD18C26C" box="[234,359,597,623]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398849DF5BC291" blockId="29.[189,804,597,658]" box="[189,804,632,658]" pageId="29" pageNumber="30">
<collectingCountry id="F33B0BDA9610FFA0FF398849DD26C291" box="[189,345,632,658]" name="Thailand" pageId="29" pageNumber="30">THAILAND</collectingCountry>
:
<specimenCount id="9D2A80C39610FFA0FEE08849DDEFC291" box="[356,400,632,658]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
, same data as
<typeStatus id="5497F5E89610FFA0FDBE8849DEDEC291" box="[570,673,632,658]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3DD38F9610FFA0FD348849DF68C291" box="[688,791,632,658]" pageId="29" pageNumber="30">NMHW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19610FFA0FF398888DE26C5CA" pageId="29" pageNumber="30" type="description">
<paragraph id="8B934B4A9610FFA0FF398888DD31C2D0" blockId="29.[189,334,697,723]" box="[189,334,697,723]" pageId="29" pageNumber="30">
<heading id="D0DBFC269610FFA0FF398888DD31C2D0" bold="true" box="[189,334,697,723]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B95897589610FFA0FF398888DD31C2D0" bold="true" box="[189,334,697,723]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9610FFA0FF3988D6DDF6C368" blockId="29.[189,1398,743,875]" pageId="29" pageNumber="30">
Adult male with 62 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9610FFA0FC0B88D9DFA4C301" box="[911,987,744,770]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="cm" value="11.0">11 cm</quantity>
, width
<quantity id="4CD4E6AF9610FFA0FBB088D6D8EDC301" box="[1076,1170,743,770]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="29" pageNumber="30" unit="mm" value="6.4">6.4 mm</quantity>
. Adult female with 61 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9610FFA0FD43893ADF6BC326" box="[711,788,779,805]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="cm" value="12.0">12 cm</quantity>
, width
<quantity id="4CD4E6AF9610FFA0FCEA893ADFB2C326" box="[878,973,779,805]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="29" pageNumber="30" unit="mm" value="7.4">7.4 mm</quantity>
. Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown.
</paragraph>
<paragraph id="8B934B4A9610FFA0FF3989A2D8F7C4C4" blockId="29.[189,1399,915,1223]" pageId="29" pageNumber="30">
GONOPODS (
<figureCitation id="131757CF9610FFA0FEC689A2DDEDC3AE" box="[322,402,915,941]" captionStart="Fig" captionStartId="28.[189,232,1930,1956]" captionTargetBox="[289,1281,695,1897]" captionTargetId="figure@28.[277,1311,693,1910]" captionTargetPageId="28" captionText="Fig. 10. Thyropygus ursus sp. nov., from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="29" pageNumber="30">Fig. 10</figureCitation>
AD).Anterior coxal fold (
<emphasis id="B95897589610FFA0FD4F89A2DE98C3AF" box="[715,743,915,940]" italics="true" pageId="29" pageNumber="30">ac</emphasis>
;
<figureCitation id="131757CF9610FFA0FD7789A2DF20C3AE" box="[755,863,915,941]" captionStart="Fig" captionStartId="28.[189,232,1930,1956]" captionTargetBox="[289,1281,695,1897]" captionTargetId="figure@28.[277,1311,693,1910]" captionTargetPageId="28" captionText="Fig. 10. Thyropygus ursus sp. nov., from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="29" pageNumber="30">Fig. 10A</figureCitation>
): lateral process (
<emphasis id="B95897589610FFA0FBA189A2D835C3AE" box="[1061,1098,915,941]" italics="true" pageId="29" pageNumber="30">alp</emphasis>
) flattened, broad, apically curved caudad, tip with short spine, lateral margin slightly folded; mesal process (
<emphasis id="B95897589610FFA0FBF28986D8D6C3D3" box="[1142,1193,951,976]" italics="true" pageId="29" pageNumber="30">amp</emphasis>
) shorter than
<emphasis id="B95897589610FFA0FACE8987D910C3D3" box="[1354,1391,950,976]" italics="true" pageId="29" pageNumber="30">alp</emphasis>
, slender, straight, directed distad, pointed. Posterior coxal fold (
<emphasis id="B95897589610FFA0FC3489EBDFB3C3F0" box="[944,972,986,1011]" italics="true" pageId="29" pageNumber="30">pc</emphasis>
;
<figureCitation id="131757CF9610FFA0FC5989E8D839C3F0" box="[989,1094,985,1011]" captionStart="Fig" captionStartId="28.[189,232,1930,1956]" captionTargetBox="[289,1281,695,1897]" captionTargetId="figure@28.[277,1311,693,1910]" captionTargetPageId="28" captionText="Fig. 10. Thyropygus ursus sp. nov., from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="29" pageNumber="30">Fig. 10B</figureCitation>
) basally with moderately high paracoxites (
<emphasis id="B95897589610FFA0FE1489CCDDD3C415" box="[400,428,1021,1046]" italics="true" pageId="29" pageNumber="30">px</emphasis>
), distally truncate, forming shelf to accommodate telopodite, mesal process (
<emphasis id="B95897589610FFA0FABE89CCD912C415" box="[1338,1389,1021,1046]" italics="true" pageId="29" pageNumber="30">pmp</emphasis>
) very small, directed distolaterad. Telopodite (
<figureCitation id="131757CF9610FFA0FD538E2EDF53C439" box="[727,812,1055,1082]" captionStart="Fig" captionStartId="28.[189,232,1930,1956]" captionTargetBox="[289,1281,695,1897]" captionTargetId="figure@28.[277,1311,693,1910]" captionTargetPageId="28" captionText="Fig. 10. Thyropygus ursus sp. nov., from Lanta Islands, holotype (NMHW-Inv.7855), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840506" httpUri="https://zenodo.org/record/3840506/files/figure.png" pageId="29" pageNumber="30">Fig. 10</figureCitation>
CD) leaving coxite over shelf of posterior coxal fold; femoral spine double,
<emphasis id="B95897589610FFA0FD8F8E72DE46C45E" box="[523,569,1091,1117]" italics="true" pageId="29" pageNumber="30">fe 1</emphasis>
very short, erect, situated above
<emphasis id="B95897589610FFA0FC4E8E72DF87C45E" box="[970,1016,1091,1117]" italics="true" pageId="29" pageNumber="30">fe 2</emphasis>
,
<emphasis id="B95897589610FFA0FB8C8E72D849C45E" box="[1032,1078,1091,1117]" italics="true" pageId="29" pageNumber="30">fe 2</emphasis>
very long, slender, curved downward,
<emphasis id="B95897589610FFA0FECD8E56DDEDC483" box="[329,402,1127,1152]" italics="true" pageId="29" pageNumber="30">in situ</emphasis>
resting behind
<emphasis id="B95897589610FFA0FDC88E57DE0EC483" box="[588,625,1126,1152]" italics="true" pageId="29" pageNumber="30">alp</emphasis>
; tibial spine (
<emphasis id="B95897589610FFA0FC9C8E56DF57C483" box="[792,808,1127,1152]" italics="true" pageId="29" pageNumber="30">ti</emphasis>
) long, slender, curving in horizontal plane, its tip close to base of
<emphasis id="B95897589610FFA0FEFD8EB8DDF1C4A0" box="[377,398,1161,1187]" italics="true" pageId="29" pageNumber="30">fe</emphasis>
; apical part: lamellar lobe (
<emphasis id="B95897589610FFA0FD578EB8DE9CC4A0" box="[723,739,1161,1187]" italics="true" pageId="29" pageNumber="30">ll</emphasis>
) broad, bent down; palette (
<emphasis id="B95897589610FFA0FBAB8EBBD833C4A0" box="[1071,1100,1162,1187]" italics="true" pageId="29" pageNumber="30">pa</emphasis>
) simple, gutter-like, with small crest (
<emphasis id="B95897589610FFA0FEC88E9CDD1BC4C5" box="[332,356,1197,1222]" italics="true" pageId="29" pageNumber="30">cr</emphasis>
) near tip; distally with about twelve brownish blepharochaetae (
<emphasis id="B95897589610FFA0FBDD8E9DD809C4C5" box="[1113,1142,1196,1222]" italics="true" pageId="29" pageNumber="30">bp</emphasis>
).
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398EDCDD19C50B" blockId="29.[189,358,1261,1288]" box="[189,358,1261,1288]" pageId="29" pageNumber="30">
<heading id="D0DBFC269610FFA0FF398EDCDD19C50B" bold="true" box="[189,358,1261,1288]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B95897589610FFA0FF398EDCDD19C50B" bold="true" box="[189,358,1261,1288]" pageId="29" pageNumber="30">DNA barcode</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398F2DDD49C559" blockId="29.[189,1398,1308,1370]" pageId="29" pageNumber="30">
The GenBank accession number of the barcode of the
<typeStatus id="5497F5E89610FFA0FCC38F2DDFD1C535" box="[839,942,1308,1334]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
is
<accessionNumber id="947FD6A99610FFA0FC568F2DD82AC534" box="[978,1109,1308,1335]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU306523" pageId="29" pageNumber="30">KU306523</accessionNumber>
(voucher code NMHW- Inv.7855).
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398FB0DDBBC598" blockId="29.[189,452,1409,1435]" box="[189,452,1409,1435]" pageId="29" pageNumber="30">
<emphasis id="B95897589610FFA0FF398FB0DD29C598" bold="true" box="[189,342,1409,1435]" pageId="29" pageNumber="30">Distribution</emphasis>
(
<figureCitation id="131757CF9610FFA0FEE28FB0DDC4C598" box="[358,443,1409,1435]" captionStart="Fig" captionStartId="30.[189,232,1998,2024]" captionTargetBox="[189,1398,829,1964]" captionTargetId="figure@30.[189,1398,829,1964]" captionTargetPageId="30" captionText="Fig. 12. Known distribution of the species of the T. opinatus subgroup." figureDoi="http://doi.org/10.5281/zenodo.3840510" httpUri="https://zenodo.org/record/3840510/files/figure.png" pageId="29" pageNumber="30">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8B934B4A9610FFA0FF398F81DE26C5CA" blockId="29.[189,601,1455,1482]" box="[189,601,1455,1482]" pageId="29" pageNumber="30">
Known only from the
<typeStatus id="5497F5E89610FFA0FE468F81DD8AC5C9" box="[450,501,1456,1482]" pageId="29" pageNumber="30">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>