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<document id="81A40C742DBFCD3BF58803B0358D3D66" ID-DOI="10.5852/ejt.2020.722.1139" ID-Zenodo-Dep="4239287" ID-ZooBank="90B3B475-49D9-4869-8F5F-A788DB880356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1604417496755" checkinUser="valdenar" docAuthor="Gilasian, Ebrahim, Steenis, Jeroen van &amp; Parchami-Araghi, Mehrdad" docDate="2020" docId="03855E28D767FFE1FDD8C670FEFD403B" docLanguage="en" docName="ejt-722_gilasian_van-steenis_parchami-araghi.pdf.imf" docOrigin="European Journal of Taxonomy 722" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eumerus coeruleus" docType="treatment" docVersion="5" lastPageNumber="122" masterDocId="FFBC2650D777FFF1FF91C779FFF44512" masterDocTitle="Review of the Eumerus tricolor species group (Diptera: Syrphidae) in Iran, with description of six new species" masterLastPageNumber="152" masterPageNumber="106" pageNumber="122" updateTime="1698793012035" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8BC72E552CBA3DC03EAA3D8F5B1D11BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="447F52689C96DE10E364EBB0BE0F87D2">
<mods:title id="62FD18759ED399D34AE72F0B8F231E99">Review of the Eumerus tricolor species group (Diptera: Syrphidae) in Iran, with description of six new species</mods:title>
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<mods:name id="C5A7E59BA0C8AF0883DF4C76B248A0E7" type="personal">
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<mods:namePart id="BB7B373A0670BAB68A4F9940CD7C2DC1">Gilasian, Ebrahim</mods:namePart>
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<mods:affiliation id="E63F91866D5C996AFFBFB36AFD90EC86">Insect Taxonomy Research Department, Iranian Research Institute of Plant Protection (IRIPP), Agricultural Research, Education and Extension Organization (AREEO), Tehran, 19395 - 1454, Iran.</mods:affiliation>
<mods:nameIdentifier id="E205C0D2C8E2BF673ABD606D725F94DE" type="email">ebrahimgilasian@gmail.com;gilasian@iripp.ir</mods:nameIdentifier>
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<mods:namePart id="DE34B011A444C213BE815B8D221DA1F1">Steenis, Jeroen van</mods:namePart>
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<mods:affiliation id="40A698F633D9CE3F7E603415D0D40280">Research Associate NBC-Naturalis, Leiden. % Hof der Toekomst 48, 3823 HX Amersfoort, The Netherlands.</mods:affiliation>
<mods:nameIdentifier id="DDBE98AE30153F4AE0578AA8FA16C66B" type="email">jvansteenis@syrphidaeintrees.com</mods:nameIdentifier>
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<mods:namePart id="8EBF80E3A77B175FA32EF78157B049D2">Parchami-Araghi, Mehrdad</mods:namePart>
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<mods:affiliation id="C7405345BE79B82FA58F9834B9D47E5C">Insect Taxonomy Research Department, Iranian Research Institute of Plant Protection (IRIPP), Agricultural Research, Education and Extension Organization (AREEO), Tehran, 19395 - 1454, Iran.</mods:affiliation>
<mods:nameIdentifier id="6FE3CF6C8C659F48D4D9234A8BBFF783" type="email">maraghi20@yahoo.ca</mods:nameIdentifier>
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<mods:title id="5EC14E165F35559C0814B0F1A6FF4040">European Journal of Taxonomy</mods:title>
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<mods:date id="E9E29FD5F45C4473AFD3D106A27F4EFC">2020</mods:date>
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<mods:number id="689F5464C9C7B90B55573F2DF67444B0">2020-11-02</mods:number>
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<mods:identifier id="CC4E58E0FA56DB03C74489A4957CDBD9" type="DOI">10.5852/ejt.2020.722.1139</mods:identifier>
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<treatment id="03855E28D767FFE1FDD8C670FEFD403B" LSID="urn:lsid:plazi:treatment:03855E28D767FFE1FDD8C670FEFD403B" httpUri="http://treatment.plazi.org/id/03855E28D767FFE1FDD8C670FEFD403B" lastPageNumber="122" pageId="16" pageNumber="122">
<subSubSection id="C336BCB5D767FFE1FDD8C670FC1E4431" box="[585,1002,264,291]" pageId="16" pageNumber="122" type="nomenclature">
<paragraph id="8B93EF3ED767FFE1FDD8C670FC1E4431" blockId="16.[479,1109,264,327]" box="[585,1002,264,291]" pageId="16" pageNumber="122">
<heading id="D0DB5852D767FFE1FDD8C670FC1E4431" box="[585,1002,264,291]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="122" reason="2">
<taxonomicName id="4C2C94BDD767FFE1FDD8C670FC1E4431" ID-CoL="6H8RH" authority="(Becker, 1913)" baseAuthorityName="Becker" baseAuthorityYear="1913" box="[585,1002,264,291]" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="122" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="B958332CD767FFE1FDD8C670FCC04431" bold="true" box="[585,820,264,291]" italics="true" pageId="16" pageNumber="122">Eumerus coeruleus</emphasis>
(
<bibRefCitation id="EFBD92CFD767FFE1FCD5C670FC164431" author="Becker T." box="[836,994,265,291]" pageId="16" pageNumber="122" pagination="503 - 654" refId="ref17880" refString="Becker T. 1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musee zoologique de l'Academie des sciences de St. Petersbourg 17 (1912): 503 - 654. https: // doi. org / 10.5962 / bhl. title. 8570" type="journal article" year="1913">Becker, 1913</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C336BCB5D767FFE1FE4EC655FBA14454" box="[479,1109,300,327]" pageId="16" pageNumber="122" type="description">
<paragraph id="8B93EF3ED767FFE1FE4EC655FBA14454" blockId="16.[479,1109,264,327]" box="[479,1109,300,327]" pageId="16" pageNumber="122">
<figureCitation id="1317F3BBD767FFE1FE4EC655FDC84454" box="[479,572,300,326]" captionStart="Fig" captionStartId="7.[189,232,1812,1838]" captionTargetBox="[289,1297,265,1777]" captionTargetId="figure-18@7.[289,1297,265,1777]" captionTargetPageId="7" captionText="Fig. 3. Adult habitus, ♂, dorsal view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. falsus Becker, 1922, Chekab Valley, Iran (HMIM). D. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). E. E. coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). F. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). G. E. kazanovzkyae Paramonov, 1927, abdomen, Paveh, Iran (HMIM). H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239303" httpUri="https://zenodo.org/record/4239303/files/figure.png" pageId="16" pageNumber="122">Figs 3E</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FDD9C655FD984455" box="[584,620,300,327]" captionStart="Fig" captionStartId="9.[189,232,1916,1942]" captionTargetBox="[189,1398,836,1873]" captionTargetId="figure-149@9.[189,1398,836,1873]" captionTargetPageId="9" captionText="Fig. 5. Adult habitus, ♀, dorsal view. A. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). B. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). C. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). D. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239315" httpUri="https://zenodo.org/record/4239315/files/figure.png" pageId="16" pageNumber="122">5B</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FDE8C655FD684454" box="[633,668,300,326]" captionStart="Fig" captionStartId="13.[189,232,1814,1840]" captionTargetBox="[290,1294,268,1774]" captionTargetId="figure-19@13.[290,1294,268,1774]" captionTargetPageId="13" captionText="Fig. 6. Adult habitus, ♂, lateral view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. falsus Becker, 1922, Chekab Valley, Iran (HMIM). D. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). E. E. coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). F. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). G. E. kazanovzkyae Paramonov, 1927, Paveh, Iran (HMIM). H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239319" httpUri="https://zenodo.org/record/4239319/files/figure.png" pageId="16" pageNumber="122">6E</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FD39C655FD394454" box="[680,717,300,326]" captionStart="Fig" captionStartId="15.[189,232,1812,1838]" captionTargetBox="[189,1398,336,1746]" captionTargetId="figure-19@15.[189,1399,336,1746]" captionTargetPageId="15" captionText="Fig. 8. A–E. Adult habitus, lateral view. F. Basicosta. A–D. Female. E–F. Male. A. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). B. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). C. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). D. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). E. E. tricolor (Fabricius, 1798), Paleochori, Greece (JSA). F. E. tadzhikorum Stackelberg, 1949, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm. Abbreviations: ac = anterodorsal costal setae; pc = posterodorsal costal setae." figureDoi="http://doi.org/10.5281/zenodo.4239333" httpUri="https://zenodo.org/record/4239333/files/figure.png" pageId="16" pageNumber="122">8B</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FD48C655FD014454" box="[729,757,300,326]" captionStart="Fig" captionStartId="26.[189,232,1882,1908]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure-82@26.[189,1399,596,1846]" captionTargetPageId="26" captionText="Fig. 14. Head, ♀, dorsal view (A, C, E, G) and lateral view (B, D, F, H). A–B. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). C–D. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). E–F. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). G–H. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4239361" httpUri="https://zenodo.org/record/4239361/files/figure.png" pageId="16" pageNumber="122">14</figureCitation>
C–D, 15A–B, 18D, 20D, 22D
</paragraph>
</subSubSection>
<subSubSection id="C336BCB5D767FFE1FF2CC617FD8E4495" box="[189,634,365,392]" pageId="16" pageNumber="122" type="reference_group">
<paragraph id="8B93EF3ED767FFE1FF2CC617FD8E4495" blockId="16.[189,634,365,392]" box="[189,634,365,392]" pageId="16" pageNumber="122">
<treatmentCitationGroup id="AB3CC810D767FFE1FF2CC617FD8E4495" box="[189,634,365,392]" pageId="16" pageNumber="122">
<treatmentCitation id="0A8DC92FD767FFE1FF2CC617FD824495" author="Becker T." box="[189,630,365,392]" page="602" pageId="16" pageNumber="122" year="1913">
<taxonomicName id="4C2C94BDD767FFE1FF2CC617FD824495" authority="Becker, 1913: 602" authorityName="Becker" authorityPageNumber="602" authorityYear="1913" box="[189,630,365,392]" class="Magnoliopsida" family="Rutaceae" genus="Lampetia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="122" phylum="Tracheophyta" rank="species" species="coerulea">
<emphasis id="B958332CD767FFE1FF2CC617FE63449A" box="[189,407,366,392]" italics="true" pageId="16" pageNumber="122">Lampetia coerulea</emphasis>
<bibRefCitation id="EFBD92CFD767FFE1FE0FC617FD824495" author="Becker T." box="[414,630,365,392]" pageId="16" pageNumber="122" pagination="503 - 654" refId="ref17880" refString="Becker T. 1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musee zoologique de l'Academie des sciences de St. Petersbourg 17 (1912): 503 - 654. https: // doi. org / 10.5962 / bhl. title. 8570" type="journal article" year="1913">Becker, 1913: 602</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C336BCB5D767FFE1FF2CC6D6FC6E47D9" pageId="16" pageNumber="122" type="diagnosis">
<paragraph id="8B93EF3ED767FFE1FF2CC6D6FEC244DB" blockId="16.[189,310,431,457]" box="[189,310,431,457]" pageId="16" pageNumber="122">
<heading id="D0DB5852D767FFE1FF2CC6D6FEC244DB" bold="true" box="[189,310,431,457]" fontSize="11" level="3" pageId="16" pageNumber="122" reason="3">
<emphasis id="B958332CD767FFE1FF2CC6D6FEC244DB" bold="true" box="[189,310,431,457]" pageId="16" pageNumber="122">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B93EF3ED767FFE1FF2CC6A7FC6E47D9" blockId="16.[189,1399,477,715]" pageId="16" pageNumber="122">
Eyes pilose (
<figureCitation id="1317F3BBD767FFE1FEC6C6A4FE5944EA" box="[343,429,477,504]" captionStart="Fig" captionStartId="26.[189,232,1882,1908]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure-82@26.[189,1399,596,1846]" captionTargetPageId="26" captionText="Fig. 14. Head, ♀, dorsal view (A, C, E, G) and lateral view (B, D, F, H). A–B. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). C–D. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). E–F. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). G–H. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4239361" httpUri="https://zenodo.org/record/4239361/files/figure.png" pageId="16" pageNumber="122">Fig. 14</figureCitation>
C–D), males dichoptic, distance between eyes in males less than width of ocellar triangle (
<figureCitation id="1317F3BBD767FFE1FEB9C578FE88470E" box="[296,380,513,540]" captionStart="Fig" captionStartId="27.[189,232,1882,1908]" captionTargetBox="[190,1398,651,1842]" captionTargetId="figure-70@27.[190,1400,651,1845]" captionTargetPageId="27" captionText="Fig. 15. Head, ♂, dorsal view (A, C, E, G) and lateral view (B, D, F, H). A–B. Eumerus coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). C–D. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). E–F. E. kazanovzkyae Paramonov, 1927, Paveh, Iran (HMIM). G–H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4239367" httpUri="https://zenodo.org/record/4239367/files/figure.png" pageId="16" pageNumber="122">Fig. 15</figureCitation>
A–B); face with whitish pile; ocellar triangle equilateral (
<figureCitation id="1317F3BBD767FFE1FBB8C578FB6C470E" box="[1065,1176,513,540]" captionStart="Fig" captionStartId="27.[189,232,1882,1908]" captionTargetBox="[190,1398,651,1842]" captionTargetId="figure-70@27.[190,1400,651,1845]" captionTargetPageId="27" captionText="Fig. 15. Head, ♂, dorsal view (A, C, E, G) and lateral view (B, D, F, H). A–B. Eumerus coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). C–D. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). E–F. E. kazanovzkyae Paramonov, 1927, Paveh, Iran (HMIM). G–H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4239367" httpUri="https://zenodo.org/record/4239367/files/figure.png" pageId="16" pageNumber="122">Fig. 15A</figureCitation>
); basoflagellomere orange; scutum and scutellum with dense and long whitish pile (
<figureCitation id="1317F3BBD767FFE1FC21C55DFBFB472C" box="[944,1039,548,574]" captionStart="Fig" captionStartId="13.[189,232,1814,1840]" captionTargetBox="[290,1294,268,1774]" captionTargetId="figure-19@13.[290,1294,268,1774]" captionTargetPageId="13" captionText="Fig. 6. Adult habitus, ♂, lateral view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. falsus Becker, 1922, Chekab Valley, Iran (HMIM). D. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). E. E. coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). F. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). G. E. kazanovzkyae Paramonov, 1927, Paveh, Iran (HMIM). H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239319" httpUri="https://zenodo.org/record/4239319/files/figure.png" pageId="16" pageNumber="122">Figs 6E</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FB8AC55DFBB4472C" box="[1051,1088,548,574]" captionStart="Fig" captionStartId="15.[189,232,1812,1838]" captionTargetBox="[189,1398,336,1746]" captionTargetId="figure-19@15.[189,1399,336,1746]" captionTargetPageId="15" captionText="Fig. 8. A–E. Adult habitus, lateral view. F. Basicosta. A–D. Female. E–F. Male. A. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). B. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). C. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). D. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). E. E. tricolor (Fabricius, 1798), Paleochori, Greece (JSA). F. E. tadzhikorum Stackelberg, 1949, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm. Abbreviations: ac = anterodorsal costal setae; pc = posterodorsal costal setae." figureDoi="http://doi.org/10.5281/zenodo.4239333" httpUri="https://zenodo.org/record/4239333/files/figure.png" pageId="16" pageNumber="122">8B</figureCitation>
), weakly pollinose; pleura above metacoxa bare; legs long pilose; tarsi and basal half of tibiae orange; metafemur enlarged; metatarsus without dense thick black pile fringe; wing hyaline, entirely covered with microtrichia; vein R
<subScript id="17A8ED7BD767FFE1FF40C5D9FF1F47BC" attach="left" box="[209,235,672,686]" fontSize="6" pageId="16" pageNumber="122">4+5</subScript>
deeply sinuate (
<figureCitation id="1317F3BBD767FFE1FE3FC5F4FDFF47BA" box="[430,523,653,680]" captionStart="Fig" captionStartId="9.[189,232,1916,1942]" captionTargetBox="[189,1398,836,1873]" captionTargetId="figure-149@9.[189,1398,836,1873]" captionTargetPageId="9" captionText="Fig. 5. Adult habitus, ♀, dorsal view. A. Eumerus brevipilosus Gilasian &amp; van Steenis sp. nov., paratype (HMIM). B. E. coeruleus (Becker, 1913), Ghoochan, Iran (HMIM). C. E. longitarsis Peck, 1979, Mian Jangal, Iran (JSA). D. E. persarum Stackelberg, 1961, Bazman, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239315" httpUri="https://zenodo.org/record/4239315/files/figure.png" pageId="16" pageNumber="122">Fig. 5B</figureCitation>
); abdominal terga II–IV orange laterally; terga II–III with distinct median black vitta (
<figureCitation id="1317F3BBD767FFE1FED9C5C8FE5747D9" box="[328,419,689,715]" captionStart="Fig" captionStartId="7.[189,232,1812,1838]" captionTargetBox="[289,1297,265,1777]" captionTargetId="figure-18@7.[289,1297,265,1777]" captionTargetPageId="7" captionText="Fig. 3. Adult habitus, ♂, dorsal view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. falsus Becker, 1922, Chekab Valley, Iran (HMIM). D. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). E. E. coeruleus (Becker, 1913), Chekab Valley, Iran (JSA). F. E. jacobsoni Becker, 1913, Chekab Valley, Iran (JSA). G. E. kazanovzkyae Paramonov, 1927, abdomen, Paveh, Iran (HMIM). H. E. longitarsis Peck, 1979, Chekab Valley, Iran (JSA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4239303" httpUri="https://zenodo.org/record/4239303/files/figure.png" pageId="16" pageNumber="122">Fig. 3E</figureCitation>
); male genitalia as in
<figureCitation id="1317F3BBD767FFE1FD32C5C8FCE747D9" box="[675,787,689,715]" captionStart="Fig" captionStartId="31.[189,232,1850,1876]" captionTargetBox="[301,1285,286,1803]" captionTargetId="figure-18@31.[298,1289,263,1811]" captionTargetPageId="31" captionText="Fig. 18. Epandrium, cercus and surstylus, lateral view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). D. E. coeruleus (Becker, 1913), Chekab Valley, Iran (HMIM). E. E. ovoformus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). F. E. richteri Stackelberg, 1960, Lar, Iran (HMIM). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4239387" httpUri="https://zenodo.org/record/4239387/files/figure.png" pageId="16" pageNumber="122">Figs 18D</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FC8FC5C8FCA047D9" box="[798,852,689,715]" captionStart="Fig" captionStartId="35.[189,232,1848,1874]" captionTargetBox="[306,1283,308,1705]" captionTargetId="figure-19@35.[300,1288,265,1809]" captionTargetPageId="35" captionText="Fig. 20. Right surstylus, dorsal view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. chekabicus Gilasian &amp; van Steenis. sp. nov., holotype (HMIM). D. E. coeruleus (Becker, 1913), Chekab Valley, Iran (HMIM). E. E. ovoformus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). F. E. richteri Stackelberg, 1960, Lar, Iran (HMIM). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4239395" httpUri="https://zenodo.org/record/4239395/files/figure.png" pageId="16" pageNumber="122">20D</figureCitation>
,
<figureCitation id="1317F3BBD767FFE1FCF1C5C8FC6147D9" box="[864,917,689,715]" captionStart="Fig" captionStartId="37.[189,232,1847,1873]" captionTargetBox="[213,1377,308,1786]" captionTargetId="figure-18@37.[205,1381,265,1808]" captionTargetPageId="37" captionText="Fig. 22. Hypandrium, lateral view. A. Eumerus atricolorus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). B. E. brevipilosus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). C. E. chekabicus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). D. E. coeruleus (Becker, 1913), Chekab Valley, Iran (HMIM). E. E. ovoformus Gilasian &amp; van Steenis sp. nov., holotype (HMIM). F. E. richteri Stackelberg, 1960, Lar, Iran (HMIM). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4239401" httpUri="https://zenodo.org/record/4239401/files/figure.png" pageId="16" pageNumber="122">22D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336BCB5D767FFE1FF2CC58BFD814184" pageId="16" pageNumber="122" type="materials_examined">
<paragraph id="8B93EF3ED767FFE1FF2CC58BFE5E461E" blockId="16.[189,426,754,780]" box="[189,426,754,780]" pageId="16" pageNumber="122">
<heading id="D0DB5852D767FFE1FF2CC58BFE5E461E" bold="true" box="[189,426,754,780]" fontSize="11" level="3" pageId="16" pageNumber="122" reason="3">
<emphasis id="B958332CD767FFE1FF2CC58BFE5E461E" bold="true" box="[189,426,754,780]" pageId="16" pageNumber="122">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B44E563D767FFE1FF7BC458FD4B4693" collectingDate="1901-05-13" collectionCode="ZISP" collectorName="P. Beluchist." country="Iran" determinerName="Becker" location="IRAN" pageId="16" pageNumber="122" specimenCount="1" typeStatus="holotype">
<paragraph id="8B93EF3ED767FFE1FF7BC458FEA84629" blockId="16.[189,1398,801,898]" box="[234,348,801,827]" pageId="16" pageNumber="122">
<emphasis id="B958332CD767FFE1FF7BC458FEA84629" bold="true" box="[234,348,801,827]" pageId="16" pageNumber="122">
<typeStatus id="5497519CD767FFE1FF7BC458FEA84629" box="[234,348,801,827]" pageId="16" pageNumber="122" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B93EF3ED767FFE1FF2CC43DFD4B4693" blockId="16.[189,1398,801,898]" pageId="16" pageNumber="122">
<collectingCountry id="F33BAFAED767FFE1FF2CC43DFEF1464C" box="[189,261,836,862]" name="Iran" pageId="16" pageNumber="122">IRAN</collectingCountry>
•
<specimenCount id="9D2A24B7D767FFE1FE8EC43BFEC2464C" box="[287,310,834,862]" pageId="16" pageNumber="122" type="male">♂</specimenCount>
; “P. Beluchist. /
<date id="FF92C9FED767FFE1FE6DC43DFD80464C" box="[508,628,836,862]" pageId="16" pageNumber="122" value="1901-05-13">
<collectingDate id="EFD63016D767FFE1FE6DC43DFD80464C" box="[508,628,836,862]" pageId="16" pageNumber="122" value="1901-05-13">13.V.1901</collectingDate>
</date>
//
<taxonomicName id="4C2C94BDD767FFE1FD04C43CFC4A464C" authority="Beck." authorityName="Becker" authorityYear="1913" box="[661,958,836,862]" class="Magnoliopsida" family="Rutaceae" genus="Lampetia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="122" phylum="Tracheophyta" rank="species" species="coerulea">
<emphasis id="B958332CD767FFE1FD04C43CFC86464C" box="[661,882,836,862]" italics="true" pageId="16" pageNumber="122">Lampetia coerulea</emphasis>
Beck.
</taxonomicName>
/ det.
<determinerName id="7E4F03FAD767FFE1FB98C43DFBA8464C" box="[1033,1116,836,862]" pageId="16" pageNumber="122">Becker</determinerName>
//
<typeStatus id="5497519CD767FFE1FBEDC43DFB02464C" box="[1148,1270,836,862]" pageId="16" pageNumber="122" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C2C94BDD767FFE1FB6FC43CFA92464C" authorityName="Meigen" authorityYear="1822" box="[1278,1382,837,862]" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B958332CD767FFE1FB6FC43CFA92464C" box="[1278,1382,837,862]" italics="true" pageId="16" pageNumber="122">Eumerus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C2C94BDD767FFE1FF2CC41EFEDB4693" baseAuthorityName="Becker" baseAuthorityYear="1913" box="[189,303,871,897]" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="122" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="B958332CD767FFE1FF2CC41EFEDB4693" box="[189,303,871,897]" italics="true" pageId="16" pageNumber="122">coeruleus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFBD92CFD767FFE1FED7C41EFE154693" author="Becker T." box="[326,481,871,897]" pageId="16" pageNumber="122" pagination="503 - 654" refId="ref17880" refString="Becker T. 1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musee zoologique de l'Academie des sciences de St. Petersbourg 17 (1912): 503 - 654. https: // doi. org / 10.5962 / bhl. title. 8570" type="journal article" year="1913">Becker, 1913</bibRefCitation>
[red label]”;
<collectionCode id="ED3D77FBD767FFE1FDEFC41EFD4F4693" box="[638,699,871,897]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="16" pageNumber="122">ZISP</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B93EF3ED767FFE1FF7BC4D1FE5246D0" blockId="16.[189,1399,936,1174]" box="[234,422,936,962]" pageId="16" pageNumber="122">
<emphasis id="B958332CD767FFE1FF7BC4D1FE5246D0" bold="true" box="[234,422,936,962]" pageId="16" pageNumber="122">Other material</emphasis>
</paragraph>
<paragraph id="8B93EF3ED767FFE1FF2CC4B5FD814184" blockId="16.[189,1399,936,1174]" pageId="16" pageNumber="122">
<materialsCitation id="3B44E563D767FFE1FF2CC4B5FB844118" collectingDate="2006-05-30" collectionCode="HMIM" collectorName="E. Gilasian" country="Iran" county="Ghoochan" elevation="1746" latitude="37.428917" location="Chovinli" longLatPrecision="1" longitude="58.53258" municipality="Emamgholi" pageId="16" pageNumber="122" specimenCount="1" specimenCount-female="1" stateProvince="Khorasan prov.">
<collectingCountry id="F33BAFAED767FFE1FF2CC4B5FEF146F4" box="[189,261,972,998]" name="Iran" pageId="16" pageNumber="122">IRAN</collectingCountry>
•
<specimenCount id="9D2A24B7D767FFE1FE8AC4B5FEB346F4" box="[283,327,972,998]" pageId="16" pageNumber="122" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="F33BAFAED767FFE1FECFC4B5FE5946F4" box="[350,429,972,998]" name="Iran" pageId="16" pageNumber="122">IRAN</collectingCountry>
:
<collectingRegion id="49E821DCD767FFE1FE26C4B5FD9F46F4" box="[439,619,972,998]" country="Iran" name="Khorasan-e Shomali" pageId="16" pageNumber="122">Khorasan prov.</collectingRegion>
,
<collectingCounty id="62F297B2D767FFE1FDE7C4B5FD1B46F4" box="[630,751,972,998]" pageId="16" pageNumber="122">Ghoochan</collectingCounty>
/
<collectingMunicipality id="6BF77544D767FFE1FC93C4B5FC7A46F4" box="[770,910,972,998]" pageId="16" pageNumber="122">Emamgholi</collectingMunicipality>
,
<location id="8EF3B9E5D767FFE1FC09C4B5FC0B46F4" LSID="urn:lsid:plazi:treatment:03855E28D767FFE1FDD8C670FEFD403B:8EF3B9E5D767FFE1FC09C4B5FC0B46F4" box="[920,1023,972,998]" country="Iran" county="Ghoochan" latitude="37.428917" longLatPrecision="1" longitude="58.53258" municipality="Emamgholi" name="Chovinli" pageId="16" pageNumber="122" stateProvince="Khorasan prov.">Chovinli</location>
/
<geoCoordinate id="EE1889F9D767FFE1FB82C4B3FB4C46F4" box="[1043,1208,970,998]" degrees="37" direction="north" minutes="25" orientation="latitude" pageId="16" pageNumber="122" precision="1" seconds="44.1" value="37.428917">37°25′44.1″N</geoCoordinate>
,
<geoCoordinate id="EE1889F9D767FFE1FB53C4B3FA8646F4" box="[1218,1394,970,998]" degrees="058" direction="east" minutes="31" orientation="longitude" pageId="16" pageNumber="122" precision="1" seconds="57.3" value="58.53258">058°31′57.3″E</geoCoordinate>
, /
<quantity id="4CD442DBD767FFE1FF5AC496FEDD4118" box="[203,297,1007,1034]" metricMagnitude="3" metricUnit="m" metricValue="1.746" pageId="16" pageNumber="122" unit="m" value="1746.0">
<elevation id="0001080DD767FFE1FF5AC496FEDD4118" box="[203,297,1007,1034]" metricMagnitude="3" metricUnit="m" metricValue="1.746" pageId="16" pageNumber="122" unit="m" value="1746.0">1746 m</elevation>
</quantity>
.;
<date id="FF92C9FED767FFE1FEA8C496FE58411B" box="[313,428,1007,1033]" pageId="16" pageNumber="122" value="2006-05-30">
<collectingDate id="EFD63016D767FFE1FEA8C496FE58411B" box="[313,428,1007,1033]" pageId="16" pageNumber="122" value="2006-05-30">30.v.2006</collectingDate>
</date>
/
<collectorName id="26D98AE8D767FFE1FE50C489FDB6411B" box="[449,578,1007,1034]" pageId="16" pageNumber="122">E. Gilasian</collectorName>
//
<taxonomicName id="4C2C94BDD767FFE1FDCEC489FCCB411B" baseAuthorityName="Becker" baseAuthorityYear="1913" box="[607,831,1007,1033]" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="122" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="B958332CD767FFE1FDCEC489FCCB411B" box="[607,831,1007,1033]" italics="true" pageId="16" pageNumber="122">Eumerus coeruleus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFBD92CFD767FFE1FCCDC496FC09411B" author="Becker T." box="[860,1021,1007,1033]" pageId="16" pageNumber="122" pagination="503 - 654" refId="ref17880" refString="Becker T. 1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musee zoologique de l'Academie des sciences de St. Petersbourg 17 (1912): 503 - 654. https: // doi. org / 10.5962 / bhl. title. 8570" type="journal article" year="1913">Becker, 1913</bibRefCitation>
)”;
<collectionCode id="ED3D77FBD767FFE1FB8CC489FB844118" box="[1053,1136,1008,1034]" pageId="16" pageNumber="122">HMIM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B44E563D767FFE1FB16C496FE0F413E" collectingDate="2006-05-30" collectionCode="JSA" collectorName="E. Gilasian" country="Iran" county="Ghoochan" elevation="1746" latitude="37.428917" location="Chovinli" longLatPrecision="1" longitude="58.53258" municipality="Emamgholi" pageId="16" pageNumber="122" specimenCount="1" specimenCount-female="1" stateProvince="Khorasan-e Shomali">
<specimenCount id="9D2A24B7D767FFE1FB16C496FB474118" box="[1159,1203,1007,1034]" pageId="16" pageNumber="122" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED3D77FBD767FFE1FE5BC36BFE0F413E" box="[458,507,1042,1068]" pageId="16" pageNumber="122">JSA</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B44E563D767FFE1FD87C36BFB374161" collectedFrom="near pool" collectingDate="2016-05-25" collectingDateMax="2016-06-15" collectingDateMin="2016-05-25" collectingMethod="Malaise trap" collectionCode="HMIM" collectorName="E. Gilasian" country="Iran" county="Haftad" elevation="2219" latitude="34.116802" location="Chekab Valley" longLatPrecision="1" longitude="50.273693" municipality="Qolleh Protected Area" pageId="16" pageNumber="122" specimenCount="82" specimenCount-female="59" stateProvince="Markazi prov.">
<specimenCount id="9D2A24B7D767FFE1FD87C36BFD9E413E" box="[534,618,1040,1068]" pageId="16" pageNumber="122">23 ♂♂</specimenCount>
,
<specimenCount id="9D2A24B7D767FFE1FDEBC36BFD3A413F" box="[634,718,1042,1069]" pageId="16" pageNumber="122" type="female">59 ♀♀</specimenCount>
; “
<collectingCountry id="F33BAFAED767FFE1FD78C36BFCCC413E" box="[745,824,1042,1068]" name="Iran" pageId="16" pageNumber="122">IRAN</collectingCountry>
:
<collectingRegion id="49E821DCD767FFE1FCD5C36BFC1A413F" box="[836,1006,1042,1069]" country="Iran" name="Markazi" pageId="16" pageNumber="122">Markazi prov.</collectingRegion>
,
<collectingCounty id="62F297B2D767FFE1FC6AC36BFBBF413E" box="[1019,1099,1042,1068]" pageId="16" pageNumber="122">Haftad</collectingCounty>
/
<collectingMunicipality id="6BF77544D767FFE1FBF7C36BFA86413E" box="[1126,1394,1042,1068]" pageId="16" pageNumber="122">Qolleh Protected Area</collectingMunicipality>
, /
<location id="8EF3B9E5D767FFE1FF5EC34CFE8A415D" LSID="urn:lsid:plazi:treatment:03855E28D767FFE1FDD8C670FEFD403B:8EF3B9E5D767FFE1FF5EC34CFE8A415D" box="[207,382,1077,1103]" country="Iran" county="Haftad" latitude="34.116802" longLatPrecision="1" longitude="50.273693" municipality="Qolleh Protected Area" name="Chekab Valley" pageId="16" pageNumber="122" stateProvince="Markazi prov.">Chekab Valley</location>
,
<quantity id="4CD442DBD767FFE1FE1DC34CFE1D4142" box="[396,489,1077,1104]" metricMagnitude="3" metricUnit="m" metricValue="2.219" pageId="16" pageNumber="122" unit="m" value="2219.0">
<elevation id="0001080DD767FFE1FE1DC34CFE1D4142" box="[396,489,1077,1104]" metricMagnitude="3" metricUnit="m" metricValue="2.219" pageId="16" pageNumber="122" unit="m" value="2219.0">2219 m</elevation>
</quantity>
, /
<geoCoordinate id="EE1889F9D767FFE1FD9BC34DFD544142" box="[522,672,1076,1104]" degrees="34" direction="north" minutes="07" orientation="latitude" pageId="16" pageNumber="122" precision="1" seconds="0.5" value="34.116802">34°07′0.5″N</geoCoordinate>
,
<geoCoordinate id="EE1889F9D767FFE1FD3FC34DFCAA4142" box="[686,862,1076,1104]" degrees="050" direction="east" minutes="16" orientation="longitude" pageId="16" pageNumber="122" precision="1" seconds="25.3" value="50.273693">050°16′25.3″E</geoCoordinate>
, /
<date id="FF92C9FED767FFE1FCEFC34CFBB44142" box="[894,1088,1077,1104]" pageId="16" pageNumber="122" value="2016-05-25" valueMax="2016-06-15" valueMin="2016-05-25">
<collectingDate id="EFD63016D767FFE1FCEFC34CFBB44142" box="[894,1088,1077,1104]" pageId="16" pageNumber="122" value="2016-05-25" valueMax="2016-06-15" valueMin="2016-05-25">25.v–15.vi.2016</collectingDate>
</date>
,
<collectingMethod id="526D9729D767FFE1FBDEC34CFB174142" box="[1103,1251,1077,1104]" pageId="16" pageNumber="122">Malaise trap</collectingMethod>
/
<collectedFrom id="3830ACC3D767FFE1FB6EC34FFA86415D" box="[1279,1394,1077,1104]" pageId="16" pageNumber="122">near pool</collectedFrom>
, E. Gilasian &amp; M. / Parchami-Araghi //
<taxonomicName id="4C2C94BDD767FFE1FD0DC320FC754160" baseAuthorityName="Becker" baseAuthorityYear="1913" box="[668,897,1112,1138]" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="122" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="B958332CD767FFE1FD0DC320FC754160" box="[668,897,1112,1138]" italics="true" pageId="16" pageNumber="122">Eumerus coeruleus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFBD92CFD767FFE1FC37C321FBB84161" author="Becker T." box="[934,1100,1112,1139]" pageId="16" pageNumber="122" pagination="503 - 654" refId="ref17880" refString="Becker T. 1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musee zoologique de l'Academie des sciences de St. Petersbourg 17 (1912): 503 - 654. https: // doi. org / 10.5962 / bhl. title. 8570" type="journal article" year="1913">Becker, 1913</bibRefCitation>
)”;
<collectionCode id="ED3D77FBD767FFE1FBE1C320FB374161" box="[1136,1219,1113,1139]" pageId="16" pageNumber="122">HMIM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B44E563D767FFE1FB73C320FD864184" collectedFrom="near pool" collectingDate="2016-05-25" collectingDateMax="2016-06-15" collectingDateMin="2016-05-25" collectingMethod="Malaise trap" collectionCode="JSA" collectorName="E. Gilasian" country="Iran" county="Haftad" elevation="2219" latitude="34.116802" location="Chekab Valley" longLatPrecision="1" longitude="50.273693" municipality="Qolleh Protected Area" pageId="16" pageNumber="122" specimenCount="2" specimenCount-male="2" stateProvince="Markazi">
<specimenCount id="9D2A24B7D767FFE1FB73C320FADC4160" box="[1250,1320,1110,1139]" pageId="16" pageNumber="122" type="male">2 ♂♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED3D77FBD767FFE1FDACC305FD864184" box="[573,626,1148,1174]" pageId="16" pageNumber="122">JSA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336BCB5D767FFE1FF2CC3C4FEFD403B" pageId="16" pageNumber="122" type="distribution">
<paragraph id="8B93EF3ED767FFE1FF2CC3C4FEA241C5" blockId="16.[189,342,1213,1239]" box="[189,342,1213,1239]" pageId="16" pageNumber="122">
<heading id="D0DB5852D767FFE1FF2CC3C4FEA241C5" bold="true" box="[189,342,1213,1239]" fontSize="11" level="3" pageId="16" pageNumber="122" reason="3">
<emphasis id="B958332CD767FFE1FF2CC3C4FEA241C5" bold="true" box="[189,342,1213,1239]" pageId="16" pageNumber="122">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B93EF3ED767FFE1FF2CC395FEFD403B" blockId="16.[189,1398,1260,1321]" pageId="16" pageNumber="122">
Central Asia,
<collectingCountry id="F33BAFAED767FFE1FECCC395FE784014" box="[349,396,1260,1286]" name="Iran" pageId="16" pageNumber="122">Iran</collectingCountry>
(Sistan &amp; Balouchestan prov., Kuh-e Taftan;
<collectingRegion id="49E821DCD767FFE1FC31C395FBE44014" box="[928,1040,1260,1286]" country="Iran" name="Khorasan-e Shomali" pageId="16" pageNumber="122">Khorasan</collectingRegion>
and
<collectingRegion id="49E821DCD767FFE1FBD9C395FB5E4014" box="[1096,1194,1260,1286]" country="Iran" name="Markazi" pageId="16" pageNumber="122">Markazi</collectingRegion>
provinces) (
<bibRefCitation id="EFBD92CFD767FFE1FAAAC395FF08403B" author="Peck L. V." pageId="16" pageNumber="122" pagination="11 - 230" refId="ref18667" refString="Peck L. V. 1988. Family Syrphidae. In: Soos A. &amp; Papp L. (eds) Catalogue of Palaearctic Diptera. Syrphidae-Conopidae. Volume 8: 11 - 230. Akademiai Kiado, Budapest." type="journal article" year="1988">Peck 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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