602 lines
76 KiB
XML
602 lines
76 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1139.96639" ID-GBIF-Dataset="9e8ee3dd-d055-4dec-9b97-0e81afb4d22c" ID-PMC="PMC9843613" ID-Pensoft-Pub="1313-2970-1139-33" ID-Pensoft-UUID="A2C101C84CB5568A92BA5F0DB93564D2" ID-PubMed="36761277" ID-ZooBank="0A828EAE0A7E4EF690869572E8F15E21" ModsDocID="1313-2970-1139-33" checkinTime="1673408730709" checkinUser="pensoft" docAuthor="Gao, Zhizhong, Hou, Yanmeng & Zhang, Feng" docDate="2023" docId="B8502145B46C58B48237EED883C95229" docLanguage="en" docName="ZooKeys 1139: 33-69" docOrigin="ZooKeys 1139" docPubDate="2023-01-09" docSource="http://dx.doi.org/10.3897/zookeys.1139.96639" docTitle="Spelaeochthonius wulibeiensis Gao & Hou & Zhang 2023, sp. nov." docType="treatment" docUuid="296D1EC6-9D37-43A4-B971-6330674C6711" docUuidSource="ZooBank" docVersion="4" id="A2C101C84CB5568A92BA5F0DB93564D2" lastPageNumber="33" masterDocId="A2C101C84CB5568A92BA5F0DB93564D2" masterDocTitle="Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China" masterLastPageNumber="69" masterPageNumber="33" pageNumber="33" updateTime="1732483187196" updateUser="ExternalLinkService">
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<mods:mods id="F5FD8AF09F90EFB0C996B57CAFEA16CB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FC0681AF5F1E7B06E67A75665B7451A8">
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<mods:title id="1842E1864393A747B6394BC45A712108">Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China</mods:title>
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<mods:name id="8239D01FEC892EC588C35AD6644FB6DD" type="personal">
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<mods:roleTerm id="6977059A5FE63155F5C9211094A36991">Author</mods:roleTerm>
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<mods:namePart id="6833B66F432D19D7C6A1422D350264FB">Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier id="4560F90688ED8133F6BE80CB1A709AF2" type="ORCID">https://orcid.org/0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation id="22AB3792DE1A51706C74651D35D5D111">Department of Biology, Xinzhou Teachers University, Xinzhou, Shanxi 034000, China</mods:affiliation>
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<mods:roleTerm id="9EBB2B64F251AF4373BA324E56574674">Author</mods:roleTerm>
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<mods:namePart id="95A6C28E3CFF34978FC196B3E6AEECAF">Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier id="F5C58E22E0EDFC68BF6648AFFE999122" type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation id="D5816E91093090BAD1602215F227A3EB">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="53B70B559431F2C7E59AD4EB53B65945">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="57DA40C43CFC9FC87CBEDBC7149A4E51" type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="EE1746E3F15B77DD6495E52B1DFA9477">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier id="0898E9B5BCA82DADC115835191F82CAE" type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title id="2125BDA4954E7F4D358EED5399101E0F">ZooKeys</mods:title>
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<mods:date id="D9AA7A22EB86C80E4E60D4782C724F0F">2023</mods:date>
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<mods:number id="8B6E32BFE7324D1E55658C72D8B40193">2023-01-09</mods:number>
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<mods:identifier id="F8BD920E28D3311C55430C5876AE4DA5" type="DOI">http://dx.doi.org/10.3897/zookeys.1139.96639</mods:identifier>
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<subSection id="371DA9DC2E4ED137412A26A9FD300AEC" pageId="0" pageNumber="33" type="taxonomy">
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<treatment id="B8502145B46C58B48237EED883C95229" ID-GBIF-Taxon="208788639" LSID="urn:lsid:zoobank.org:act:296D1EC6-9D37-43A4-B971-6330674C6711" httpUri="http://treatment.plazi.org/id/B8502145B46C58B48237EED883C95229" lastPageNumber="33" pageId="0" pageNumber="33" scope_family="Pseudotyrannochthoniidae" scope_order="Pseudoscorpiones">
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<subSubSection id="678CD38A85804AE0F3EDA91D1834BA40" pageId="0" pageNumber="33" type="nomenclature">
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<paragraph id="576FCB9AF43D12E572C74D5286FEDF80" pageId="0" pageNumber="33">
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<taxonomicName id="A06297B3820D6E82E7443DDA92BBDB30" LSID="https://zoobank.org/296D1EC6-9D37-43A4-B971-6330674C6711" authority="Gao & Hou & Zhang, 2023" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis" status="sp. nov.">Spelaeochthonius wulibeiensis</taxonomicName>
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<taxonomicNameLabel id="C5140F5F23E54700AC634A0CDA4BEAC2" pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="66E52E1799CF23A1A6BA52C01975DD93" pageId="0" pageNumber="33" type="description">
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<paragraph id="89E4CC8165E0348D10AA6A88153322C3" pageId="0" pageNumber="33">
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<figureCitation id="0CBAB1FA0E09BE6C86D0E994A1CF6319" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">Figs 1A</figureCitation>
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<figureCitation id="6DA2393642DB99A984ABFEB697EF90E9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">, 18</figureCitation>
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<figureCitation id="9B4B1475B8D0605A03CAF00210E59F7E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">, 19</figureCitation>
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<figureCitation id="93240A023FA84A6911877E23A43A891A" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">, 20</figureCitation>
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<figureCitation id="6491F24836C147F5C1646204B512EFAA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">, 21</figureCitation>
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<figureCitation id="99D80E7EAEDB0ABCFE016610828EB40C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">, 22</figureCitation>
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<figureCitation id="D5C589791D85459DADD229D931C6BE89" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">
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, 23 Chinese name
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<normalizedToken id="E5E44DAC4B483C5EB4103F3C5CC0B969" originalValue="五里碑穴伪蝎">五里碑穴伪蝎</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="2430CA0558BAA39F1AF422192E11F574" pageId="0" pageNumber="33" type="type material">
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<paragraph id="4FDC1D3E1F69091265A2FA326E6F441E" pageId="0" pageNumber="33">Type material.</paragraph>
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<paragraph id="AC5BC6B089E6AAA0692237A2693AE1AD" pageId="0" pageNumber="33">
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<emphasis id="26ECB35F5B94676FE344E301C180BC7E" bold="true" italics="true" pageId="0" pageNumber="33">Holotype</emphasis>
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: China • ♂; Guizhou Province, Weining County, Yancang Town, Yangguan Village, Wulibei Cave;
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<geoCoordinate id="4799F26FCC291F5DBE9F55B3E64EB1D2" degrees="26" direction="north" minutes="53.82" orientation="latitude" precision="9" value="26.897">26°53.82'N</geoCoordinate>
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,
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<geoCoordinate id="DD13279FD748651327E3693680298998" degrees="104" direction="east" minutes="19.36" orientation="longitude" precision="9" value="104.32267">104°19.36'E</geoCoordinate>
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; 2425 m a.s.l.; 07 Aug. 2019; Zegang Feng, Zhaoyi Li and Chen Zhang leg.; under a stone in the deep zone; Ps.-MHBU-GZC19080701 (Figs
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<figureCitation id="779AB72C4DD1780EDBA20536765EF60E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1A</figureCitation>
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,
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<figureCitation id="2313BDBDC71592CE4D2BD29B63E41FB1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18</figureCitation>
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).
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<emphasis id="2682F4210A2C5316858BD2463745EB7C" bold="true" italics="true" pageId="0" pageNumber="33">Paratypes</emphasis>
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: • 2♂; the same data as the holotype; Ps.-MHBU-GZC19080702-GZC19080703 • 1♀; the same location as the holotype; 19 May. 2017, Zhisheng Zhang, Huiming Chen and Luyu Wang leg.; Ps.-MSWU-CZCH-17-06.
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</paragraph>
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</subSubSection>
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<subSubSection id="F5F910B10182ABA384BD7D80654259F1" pageId="0" pageNumber="33" type="diagnosis">
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<paragraph id="ECABF811932DF86902230368F69D6331" pageId="0" pageNumber="33">Diagnosis</paragraph>
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<paragraph id="CAD19A7B942B6D56DAAFF052AD8B6F5D" pageId="0" pageNumber="33">
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<emphasis id="EBA9703D2A9694DADF2B36E0039B9F57" bold="true" pageId="0" pageNumber="33">(♂♀).</emphasis>
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The new species can be recognized by the following combination of characters: surfaces mostly with fine reticulations; carapace without eyes or eyespots but eye region bulging and convex in dorsal view; anterior margin without protuberances; cheliceral palm with five setae; rallum with 11 blades (each with fine pinnate, the basal-most blade shorter than the others); coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami; pedipalps slender, femur 7.24 (♂), 6.40 (♀), chela 6.21-6.22 (♂), 5.68 (♀)
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<normalizedToken id="98E62D974C8FEACF8B92081E4C4D2AEB" originalValue="×">x</normalizedToken>
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longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 22 or 24 teeth; movable chelal finger with 16-19 teeth), slightly retrorse and pointed; chela fingers straight in dorsal view.
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</paragraph>
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</subSubSection>
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<subSubSection id="8AC9F26A5037EAF9CB6665CA8B6D0D52" pageId="0" pageNumber="33" type="etymology">
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<paragraph id="E16F800FCF8EE8FB94B298B1A4D7DEBA" pageId="0" pageNumber="33">Etymology.</paragraph>
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<paragraph id="DBE06385C3254839128F3DEF43775DE7" pageId="0" pageNumber="33">Named after the type locality, Wulibei Cave.</paragraph>
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</subSubSection>
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<subSubSection id="3D2EB266FFF2629E8D274A87BA8491CD" pageId="0" pageNumber="33" type="description">
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<paragraph id="A432C9F269C110C0DD60B09E3B079721" pageId="0" pageNumber="33">Description.</paragraph>
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<paragraph id="C89DD8E84EF7C6A6B292502E73E033B5" pageId="0" pageNumber="33">
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<emphasis id="73523F2A8F22F995424624D58D38B9FB" bold="true" pageId="0" pageNumber="33">Adult males</emphasis>
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(Figs
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<figureCitation id="29808AA97E1C0A7B1E7826F18B3857F5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18D</figureCitation>
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,
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<figureCitation id="3E2D80662BCD09DB2F47316FFE3B59FD" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19A</figureCitation>
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,
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<figureCitation id="0224EF97F6BCAB4AD50B40F6C8CABF30" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20</figureCitation>
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,
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<figureCitation id="167004391602A7DE08388C5BAE0FBA66" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A, B</figureCitation>
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,
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<figureCitation id="79EA6D8F1550A023E49849894D975FFD" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22</figureCitation>
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,
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<figureCitation id="C671F469142147766DA96D422B9ACDED" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23</figureCitation>
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).
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<emphasis id="36D40C2AF74343E0823E5763670C90D6" bold="true" italics="true" pageId="0" pageNumber="33">Color</emphasis>
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(Figs
|
||
<figureCitation id="1113A43BBC7ED3EDFCE471D6EC99C00F" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18D</figureCitation>
|
||
,
|
||
<figureCitation id="E16441307BE246245F15085A537C1F1A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19A</figureCitation>
|
||
,
|
||
<figureCitation id="099730096D8660D79F7A0E00682466ED" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20</figureCitation>
|
||
,
|
||
<figureCitation id="6ABE98773ABE48BAC928A6B78BB5D55E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A, B</figureCitation>
|
||
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
|
||
<emphasis id="4CB0D130F74D4DB4DB7468E28A6B24C9" bold="true" italics="true" pageId="0" pageNumber="33">Cephalothorax</emphasis>
|
||
(Figs
|
||
<figureCitation id="499D6144ED98BB685491E372AC3B2F4C" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20B</figureCitation>
|
||
,
|
||
<figureCitation id="4DC1ED8B3B6F95B64A6F35CC4F5E1827" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
|
||
,
|
||
<figureCitation id="2837A21C051B7CABA6E6C5A55D9984BB" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22A, C</figureCitation>
|
||
): carapace subquadrate, 1.02-1.03
|
||
<normalizedToken id="B195F19C34B2CF3800E229C7F8CD1658" originalValue="×">x</normalizedToken>
|
||
longer than broad, gently narrowed posteriorly; surface mostly with fine reticulations, without furrows but with seven or eight lyrifissures; no traces of eyes but eye region bulging and convex in dorsal view; epistome present and with some tiny spinules; with 16 setae arranged s4s: 4: 2: 2: 2, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 6-7, II 4-5, III 4, IV 4; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami (Figs
|
||
<figureCitation id="371D45F5D9916F97DAB96693745609D9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
|
||
,
|
||
<figureCitation id="258F62DB76256FF700250D6243C58D59" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22C</figureCitation>
|
||
); bisetose intercoxal tubercle present between coxae III and IV, tear drop-shaped (Fig.
|
||
<figureCitation id="CDF999E6773396A616A7ED525696C12C" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
|
||
).
|
||
<emphasis id="A0932EA4B9025C3AE5332E72C72818DD" bold="true" italics="true" pageId="0" pageNumber="33">Chelicera</emphasis>
|
||
(Figs
|
||
<figureCitation id="DFC25A094CF5AD68BCFBE8645223E4FE" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20C</figureCitation>
|
||
,
|
||
<figureCitation id="219E72DC628A86190E23902FA140163A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22B, E</figureCitation>
|
||
): large, approximately as long as carapace, 2.37-2.41
|
||
<normalizedToken id="F4DAC1CDA563ADEB3517C291E388A636" originalValue="×">x</normalizedToken>
|
||
longer than broad; five setae present on hand, movable finger with a medial seta, all setae acuminate, ventrobasal seta shorter than others; exterior condylar lyrifissure and exterior lyrifissure exist, palm with one extra (between sub-basal seta and an accessory seta). Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13-15 acute teeth, distal one largest; movable finger with 12 retrorse contiguous teeth of equal length, plus three or four round proximal teeth, 15 or 16 in total; galea represented by a very slight bump on movable finger. Serrula exterior with 21 blades and serrula interior with 17-20 blades. Rallum in two rows and composed of 11 blades with fine pinnate, of which the basal-most blade shorter than the others (Fig.
|
||
<figureCitation id="D8D1842B7AD9ADD7F983975D21BE1A0C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22E</figureCitation>
|
||
).
|
||
<emphasis id="435E2242F4058785BC74C9370A9EF8CF" bold="true" italics="true" pageId="0" pageNumber="33">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation id="74365ADEC75BB5DA92E72CDFDAA0E560" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20A</figureCitation>
|
||
,
|
||
<figureCitation id="7FD6DE9EF13227E3F2FCFDA8C9C1C383" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22D</figureCitation>
|
||
,
|
||
<figureCitation id="F042DDAD31B76E0FCC9068702935F028" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A, B</figureCitation>
|
||
): surfaces mostly with fine reticulations; long and slender, trochanter 1.78-2.00, femur 7.24, patella 2.44-2.47, chela 6.21-6.22, hand 2.26-2.36
|
||
<normalizedToken id="AE83CE8A0C287799743E33ED39AE07D7" originalValue="×">x</normalizedToken>
|
||
longer than broad; femur 2.62-2.80
|
||
<normalizedToken id="FA8F94C15BFF55201AEB3561B3B4573F" originalValue="×">x</normalizedToken>
|
||
longer than patella; movable chelal finger 1.61-1.74
|
||
<normalizedToken id="49280D73B9DD9E0B777A7F84F469642C" originalValue="×">x</normalizedToken>
|
||
longer than hand and 0.61-0.63
|
||
<normalizedToken id="1D452EA58A47F7E973FC8B94380D15F4" originalValue="×">x</normalizedToken>
|
||
longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella and femur, respectively (Fig.
|
||
<figureCitation id="1AC803F5EA0C6A249927B11A5E52B83D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22D</figureCitation>
|
||
). Chelal palm robust and slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
|
||
<emphasis id="F10F200781B33988883C1AA6E03DFF33" italics="true" pageId="0" pageNumber="33">ib</emphasis>
|
||
,
|
||
<emphasis id="F7100E3FD462197E2C94202ADE791A53" italics="true" pageId="0" pageNumber="33">isb</emphasis>
|
||
,
|
||
<emphasis id="59C07D45E8B8D4136B2013B51D210337" italics="true" pageId="0" pageNumber="33">eb</emphasis>
|
||
,
|
||
<emphasis id="E663B1E71DFEB3E9389F5FD7973D671B" italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
, and
|
||
<emphasis id="EEAD67A4C58AE6318342D3B461C35600" italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
clustered at the base of fixed finger,
|
||
<emphasis id="BA268A6B6A8103683DE6B4BC3C33F9E1" italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
slightly distal to
|
||
<emphasis id="0191CD54E0F030BEE2B3CC8D35E5CB6E" italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
,
|
||
<emphasis id="1533116B9388ED39EB424C91D1732F0C" italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
close to
|
||
<emphasis id="BB3926C16D8526E8AC802C5219E2420B" italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
than to
|
||
<emphasis id="87AEB1672C6AA5D5BCFF9AA22F63E55D" italics="true" pageId="0" pageNumber="33">eb</emphasis>
|
||
;
|
||
<emphasis id="6BB0E309DBB033758C6E350EEBD9C031" italics="true" pageId="0" pageNumber="33">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="7DD54936BA9B26C51B58FC3761780A42" italics="true" pageId="0" pageNumber="33">est</emphasis>
|
||
, situated subdistally and forming a pair;
|
||
<emphasis id="89F927AED2B62666D279670A20326AC7" italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
situated subdistally, very close to chelal teeth;
|
||
<emphasis id="CADBB52B13BE034B513789F360C61E3C" italics="true" pageId="0" pageNumber="33">dx</emphasis>
|
||
situated distal to
|
||
<emphasis id="4C49187ADFFA87E8AB1702826E8FE290" italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
, near the tip of fixed finger;
|
||
<emphasis id="D8C132F4924BF8DEF43A3B23EB89DE56" italics="true" pageId="0" pageNumber="33">sb</emphasis>
|
||
distinctly closer to
|
||
<emphasis id="01C01C8FAD5ECD9C2D3DA5D4D418D50F" italics="true" pageId="0" pageNumber="33">b</emphasis>
|
||
than to
|
||
<emphasis id="C633B52911E7D87A96FDC0067A827FB5" italics="true" pageId="0" pageNumber="33">st</emphasis>
|
||
(Fig.
|
||
<figureCitation id="96B0D5B8CFC2AF32372A0D180C35CE01" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A</figureCitation>
|
||
). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 22-24 teeth, slightly retrorse and pointed; movable chelal finger with 16-19 teeth (slightly smaller than teeth on fixed chelal finger); a small tubercle between the seventh and eighth teeth present (near trichobothrium
|
||
<emphasis id="FB747B90E457A5CADEA2B80495402A39" italics="true" pageId="0" pageNumber="33">t</emphasis>
|
||
) (Fig.
|
||
<figureCitation id="EC5DBEE27EB1175552F7383DE0A39665" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A</figureCitation>
|
||
). Chelal fingers straight in dorsal view (Fig.
|
||
<figureCitation id="FF3D8518301A091854A928DB064B9F36" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23B</figureCitation>
|
||
).
|
||
<emphasis id="ED4DA6D1C62C6DF11BEA7C87F1F4AE1D" bold="true" italics="true" pageId="0" pageNumber="33">Opisthosoma</emphasis>
|
||
: generally typical, ovate, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7-8: 8: 7: 6: 4: TT: 0, tergites VII-IX each with an unpaired median seta, one lyrifissure present on each side of tergites IV-IX. Sternal chaetotaxy III-XII: 6-9: 10-14: 13-14: 12: 12-13: 12-13: 10-11: 8-9: 0: 2, one lyrifissure present on each side of tergite III. Anterior genital operculum with 10-12 setae, genital opening pit-like, with seven marginal setae on each side, 24-26 in total, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig.
|
||
<figureCitation id="87107C07039738CADD884692D96FAD23" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21B</figureCitation>
|
||
).
|
||
<emphasis id="6B34A0C0D85881B7805485840484B1E3" bold="true" italics="true" pageId="0" pageNumber="33">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation id="E990BA1157DBAC2B1EC1DE785D9A5AEA" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23C, D</figureCitation>
|
||
): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.74
|
||
<normalizedToken id="521A1AFB1482C5D7A6270F920E2D713C" originalValue="×">x</normalizedToken>
|
||
longer than patella and with one lyrifissure at the base of femur; tarsus 2.11-2.26
|
||
<normalizedToken id="C7D1FC853E22E0D3FAE53215CB07E7BD" originalValue="×">x</normalizedToken>
|
||
longer than tibia. Femoropatella of leg IV 3.70-3.88
|
||
<normalizedToken id="343BCE87B7F74AAB9344EC3357ADA78A" originalValue="×">x</normalizedToken>
|
||
longer than deep and with one lyrifissure at the base of femur; tibia 5.92-6.00
|
||
<normalizedToken id="F86A495653982EB190E5B3340A3E14A5" originalValue="×">x</normalizedToken>
|
||
longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.00
|
||
<normalizedToken id="4867BA7B1F170C8780384EF539DE74C6" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.22-0.28), telotarsus 11.13-12.43
|
||
<normalizedToken id="A676D2B852915A96C715B51E54471A9C" originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.42-2.47
|
||
<normalizedToken id="AF9F5AB811AF4714FE88DCA8551F0AFE" originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.26-0.27). Setae of leg I (trochanter to tibia) 1: 13: 12-15: 14-17, setae of leg IV (trochanter to basitarsus) 2: 2: 6: 16: 14. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
<emphasis id="F3FD2ED2306D1F3D410E5A0CF08B7DC0" bold="true" italics="true" pageId="0" pageNumber="33">Dimensions of adult males</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm). Males: body length 2.50. Pedipalps: trochanter 0.32/0.16-0.18, femur 1.23/0.17, patella 0.44-0.47/0.18-0.19, chela 1.68-1.74/0.27-0.28, hand 0.61-0.66/0.27-0.28, movable finger length 1.06. Chelicera 0.64-0.65/0.27, movable finger length 0.34. Carapace 0.63/0.61-0.62. Leg I: trochanter 0.24-0.25/0.18, femur 0.73-0.75/0.10, patella 0.42-0.43/0.09, tibia 0.34-0.37/0.07, tarsus 0.77-0.78/0.06-0.07. Leg IV: trochanter 0.33-0.36/0.19-0.22, femoropatella 0.97-1.00/0.25-0.27, tibia 0.77-0.78/0.13, basitarsus 0.36/0.09, telotarsus 0.87-0.89/0.07-0.08.
|
||
</paragraph>
|
||
<caption id="13EC42ABD7B2F72FC5BA1943AAEAA038" doi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33" start="Figure 18" startId="F18">
|
||
<paragraph id="FAF6080B0606BD74DFD28D8696A4D0AA" pageId="0" pageNumber="33">
|
||
<emphasis id="4C99660C35C9C297538041F7455674FF" bold="true" pageId="0" pageNumber="33">Figure 18.</emphasis>
|
||
Wulibei Cave, type locality of
|
||
<taxonomicName id="9FCB60A3F2122DEDA7083438AC4D6475" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="3290E4622802A7BBC29C1974450267AD" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="C73754515680B14A567B2BF747A76ADE" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
cave location (red arrow)
|
||
<emphasis id="B34FA518657BE479282D6E5CA744BE11" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
entrance
|
||
<emphasis id="73732BF11567C486B61DD78BF340B08D" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
inside the cave entrance
|
||
<emphasis id="D10CA310150A176DCA8C14AED0608470" bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
live male of
|
||
<taxonomicName id="158EE7109A43C9CD1379865880628E9D" class="Arachnida" kingdom="Animalia" lsidName="S. wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="A1B4C279CE30B1E2157F20997ABF4F55" italics="true" pageId="0" pageNumber="33">S. wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in its natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EA59316AB629D7BBBF54831ADECAA2B2" doi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33" start="Figure 19" startId="F19">
|
||
<paragraph id="E73962C4BF2FFD34CEBDB9005614C9E1" pageId="0" pageNumber="33">
|
||
<emphasis id="187391F95702C643A195A0E27122D359" bold="true" pageId="0" pageNumber="33">Figure 19.</emphasis>
|
||
<taxonomicName id="0DD878E9CD72D3EFC8F433627FE9B8DC" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="8EA3E456C62F7933459B99E75C821799" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="19545A5ED5973428DBFBD1B60E5DE496" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view)
|
||
<emphasis id="5037873278C978927F5F5BE973206CAD" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E69616D38C33F85A4C91132861170CEC" doi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33" start="Figure 20" startId="F20">
|
||
<paragraph id="B8495C30CD9E45E65AD5E045A4BE8AB2" pageId="0" pageNumber="33">
|
||
<emphasis id="B95845304A0A4295F1646095C153C632" bold="true" pageId="0" pageNumber="33">Figure 20.</emphasis>
|
||
<taxonomicName id="7F60EAE3E3BAE24EDFF3C327A97078CA" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="E241AA4AADE0B94E2F859B28DE58B53E" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="66D7A20700416EDDB80C81793C113477" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis id="67C8C199BD960AD82DB71014897BC93B" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="B644B232606000DA97C18D1A18820C03" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
left chelicera (dorsal view). Scale bars: 0.20 mm (
|
||
<emphasis id="3D2C0C3D45AEAA3EC6A740AFD84C3AC9" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="2681072F0D577058488A4F7123CE75B7" bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3EC57750EBF8474D75C2CB1DD5B9A9C5" doi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33" start="Figure 21" startId="F21">
|
||
<paragraph id="54502CF64689C3781A9CC3948E2F88F3" pageId="0" pageNumber="33">
|
||
<emphasis id="5395C44034216BE823BA632308053A0E" bold="true" pageId="0" pageNumber="33">Figure 21.</emphasis>
|
||
<taxonomicName id="A4341C134C39E2DEBABB17B2026E9FBB" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="87252E48886F7C17625BFB43D4364A4F" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="44B5375F6FBC958AAA6D332E7637EAFC" bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
), paratype female (
|
||
<emphasis id="10129F8D91D7918ED9DFA9A5F815E6EA" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
)
|
||
<emphasis id="FE25952E047AE12A372C07AEF8FCA75C" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
coxae (ventral view)
|
||
<emphasis id="111345995848E77EDC76F58E31827BD3" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis id="EADFCAD1EA53580E0C16DCE499285E63" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.20 mm (
|
||
<emphasis id="E7DDB58CAC625DF9E698BC7D66572527" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="09108F1DFF54E3487C36A6BC08FE8554" bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0FC603194F37BECDACE791F967EAA485" doi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33" start="Figure 22" startId="F22">
|
||
<paragraph id="4DC70783DF4BCE985A41598E958689CD" pageId="0" pageNumber="33">
|
||
<emphasis id="31CBE2CD27EB659ECE4918C921236F1D" bold="true" pageId="0" pageNumber="33">Figure 22.</emphasis>
|
||
<taxonomicName id="10C652E78F15ED627B727BAE1AF89A70" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="745030C72CD92315A3DC08F483478481" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="ACEC8F974DB75111505CDF6DCF57BA6A" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
carapace (dorsal view), with a detail of anterior margin
|
||
<emphasis id="BAE3ECC986DDA1E73A806E90E66608BF" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chelicera (dorsal view), with details of teeth
|
||
<emphasis id="09AABC566C7A26B4D341DD5A0BE7EFC1" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
coxal spines on coxae I (ventral view)
|
||
<emphasis id="A4F1683A241AF14A1DBCF482D70B0661" bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis id="117AA8713F6F190B7A196105A66E6888" bold="true" pageId="0" pageNumber="33">E</emphasis>
|
||
rallum. Scale bars: 0.20 mm (
|
||
<emphasis id="30F68838A16E03DCDD43905E60F01174" bold="true" pageId="0" pageNumber="33">A, B, D</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="DD5F5AE13F7669CDC7695FDF3CCD210B" bold="true" pageId="0" pageNumber="33">C, E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="404E96F502CDB3772EF26C4CC030093D" doi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33" start="Figure 23" startId="F23">
|
||
<paragraph id="CE740229DD891362C51613905A20EFB0" pageId="0" pageNumber="33">
|
||
<emphasis id="FE49DCDE09EB87388C8A3FF71729C538" bold="true" pageId="0" pageNumber="33">Figure 23.</emphasis>
|
||
<taxonomicName id="CF9203F7B309920615B7C5A71773A12F" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="E9A4E8035EFC97BA7697AA044A1A3762" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="CC3A929415B33F5D05C5B9ECD0658CCF" bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view), with details of teeth and trichobothrial pattern
|
||
<emphasis id="D10A91E52DDF9D664F4000D45C5A2F90" bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis id="110BC3F0384ADBC54FAAD71DECF0196D" bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
leg I (lateral view)
|
||
<emphasis id="41CE09AFBE7E31FC87069189827444B0" bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6281274565389782B012ED137398A294" pageId="0" pageNumber="33">
|
||
<emphasis id="E08DBE08AAEAA4561D8E5FA1EC318AE6" bold="true" pageId="0" pageNumber="33">Adult female</emphasis>
|
||
(Figs
|
||
<figureCitation id="FC20BAFB55239BF4EA7DC17A16CD550D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19B</figureCitation>
|
||
,
|
||
<figureCitation id="12DDE904AB636C1394BF694035D8DF79" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21C</figureCitation>
|
||
). Mostly same as males; tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7: 8: 9: 7: 4: TT: 0; sternal chaetotaxy IV-XII: 10: 13: 11: 12: 12: 11: 8: 0: 2; anterior genital operculum with seven setae, posterior margin with 11 marginal setae, 18 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.60
|
||
<normalizedToken id="7B5F4E6781417FAF66E2C044A16A2BC6" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.25), telotarsus 11.63
|
||
<normalizedToken id="5CC6164BB7941139B55931A273C48951" originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.58
|
||
<normalizedToken id="A76875649C2B81E852D4B2B7383C24DE" originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.24). Body length 1.93. Pedipalps: trochanter 0.32/0.19 (1.68
|
||
<normalizedToken id="36D5494FE1E34212C3B1457270FE0B43" originalValue="×">x</normalizedToken>
|
||
), femur 1.28/0.20 (6.40
|
||
<normalizedToken id="DB8D69D3E4D4706C588E057EFF1D28B5" originalValue="×">x</normalizedToken>
|
||
), patella 0.51/0.21 (2.43
|
||
<normalizedToken id="7C9DCC005B5AB838787D6B10F5316BA0" originalValue="×">x</normalizedToken>
|
||
), chela 1.76/0.31 (5.68
|
||
<normalizedToken id="B15F8C8A7815C0F2CD1848D76B9C4DBF" originalValue="×">x</normalizedToken>
|
||
), hand 0.66/0.31 (2.13
|
||
<normalizedToken id="D305AC0C50E34058595FF27D28A5C7F7" originalValue="×">x</normalizedToken>
|
||
), movable chelal finger length 1.11. Chelicera 0.70/0.30 (2.33
|
||
<normalizedToken id="09E917571D0E40E70C6A7C7150BD0AB0" originalValue="×">x</normalizedToken>
|
||
), movable finger length 0.38. Carapace 0.68/0.68 (1.00
|
||
<normalizedToken id="A6D10D0923399827B138B0D0BDA1119E" originalValue="×">x</normalizedToken>
|
||
). Leg I: trochanter 0.24/0.19 (1.26
|
||
<normalizedToken id="2C9810F5FDC3D962353F289D0D264F38" originalValue="×">x</normalizedToken>
|
||
), femur 0.77/0.12 (6.42
|
||
<normalizedToken id="CAED652E32B24A9E1F7DF3C3F2E9395B" originalValue="×">x</normalizedToken>
|
||
), patella 0.45/0.11 (4.09
|
||
<normalizedToken id="AF246FBBFA5782F2BA285D6ADA2D75CA" originalValue="×">x</normalizedToken>
|
||
), tibia 0.36/0.08 (4.50
|
||
<normalizedToken id="7C388B3776356297E14CA5AD6F9EC290" originalValue="×">x</normalizedToken>
|
||
), tarsus 0.83/0.08 (10.38
|
||
<normalizedToken id="6F847FFB87F10643A0E5FCBE3CCA7B77" originalValue="×">x</normalizedToken>
|
||
). Leg IV: trochanter 0.35/0.22 (1.59
|
||
<normalizedToken id="33D64A80EE4730B7CD4BA6F21982CB09" originalValue="×">x</normalizedToken>
|
||
), femoropatella 1.05/0.27 (3.89
|
||
<normalizedToken id="28DCE31A4C42668C8902F45DD9712AE5" originalValue="×">x</normalizedToken>
|
||
), tibia 0.80/0.14 (5.71
|
||
<normalizedToken id="F109724F4548A1A73B3FA1E346A2D453" originalValue="×">x</normalizedToken>
|
||
), basitarsus 0.36/0.10 (3.60
|
||
<normalizedToken id="6C7BBC50E6AC12DCD2B54CE92BC82754" originalValue="×">x</normalizedToken>
|
||
), telotarsus 0.93/0.08 (11.63
|
||
<normalizedToken id="E384955E2DCFE1AE5077DE19EB66DBAF" originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4B2835672C1F79B5719567B92317AFAC" pageId="0" pageNumber="33" type="remarks">
|
||
<paragraph id="36AFBC4E7EE910690BD54BBEE6BCA262" pageId="0" pageNumber="33">Remarks.</paragraph>
|
||
<paragraph id="6736E269051A28E1E3F5BA005955D886" pageId="0" pageNumber="33">
|
||
The new species shares similar characters with most species of
|
||
<taxonomicName id="28DE60B874F2D21C832D794F16E1F440" authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9CCC97F8905B05263A262F910E4FAAF3" italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
Beier, 1931,
|
||
<taxonomicName id="B2D87E481C9EE488235C9B5643E41BE4" authorityName="Morikawa" authorityYear="1954" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="675093EB36273D80BE14B96F0C922C75" italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
and all species of "
|
||
<taxonomicName id="BB5C04A83D9FE2897142ACB17FB94941" authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BA39FB8CAEA71FA39B27496B3E62D776" italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" Beier, 1930 from the western US by the presence of only 16 setae on the carapace.
|
||
<bibRefCitation id="395F29B28B73C53D47F2BC6E141DAA4F" DOI="https://doi.org/10.1007/s12542-021-00565-8" author="Schwarze, D" journalOrPublisher="PalZ" pageId="0" pageNumber="33" pagination="11 - 27" refId="B29" refString="Schwarze, D, Harms, D, Hammel, J, Kotthoff, U, 2021. The first fossils of the most basal pseudoscorpion family (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae): Evidence for major biogeographical shifts in the European paleofauna. PalZ 96 (1): 11 - 27, DOI: https://doi.org/10.1007/s12542-021-00565-8" title="The first fossils of the most basal pseudoscorpion family (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae): Evidence for major biogeographical shifts in the European paleofauna." url="https://doi.org/10.1007/s12542-021-00565-8" volume="96" year="2021">Schwarze et al. (2021)</bibRefCitation>
|
||
emphasized the importance of the number of carapaceal setae in Holarctic pseudotyrannochthoniids, thus, it indicates that the three "
|
||
<taxonomicName id="A165F78F3DB272CE229AC1CDF4D04711" authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2BB5809CF903E96E8DC501AA0869685" italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" species in the western US were misclassified in comparison with the twelve species from Australia and the three species from Chile (including the type species, the number of carapaceal setae is more than 18). It can be said that the genus
|
||
<taxonomicName id="BAF440C99946457AACFFEEACB338DF1E" authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ABCFCB084F24E9F3652DC27AFCC57F55" italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the southern hemisphere (
|
||
<bibRefCitation id="4FAE717199497181D83FEC37E237DE37" DOI="https://doi.org/10.1636/JoA-S-21-033" author="Harvey, MS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="158 - 174" refId="B9" refString="Harvey, MS, Harms, D, 2022. The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal. The Journal of Arachnology 50 (2): 158 - 174, DOI: https://doi.org/10.1636/JoA-S-21-033" title="The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal." url="https://doi.org/10.1636/JoA-S-21-033" volume="50" year="2022">Harvey and Harms 2022</bibRefCitation>
|
||
). Thus, it is inappropriate to place this new species in
|
||
<taxonomicName id="7E14269CD2C0E8250075118C0F0802E5" authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5EF96F7C3DCBAB87553DBC4EF1625CE3" italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
, even though the shape of this new species of coxal spines is similar to that of the three "
|
||
<taxonomicName id="54FD12BB79EDDFE9553C4A7BB4F88439" authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74D378457F1EB6E4D61D108FE5CAFF43" italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" species.
|
||
</paragraph>
|
||
<paragraph id="37BC551D381D2A3A0E2963F10E634FEA" pageId="0" pageNumber="33">
|
||
The shape and number of the coxal spines are important distinguishing features between
|
||
<taxonomicName id="F92A60998910A7C2EAF9543A6EEE5D9F" authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6A1A8F659EDF1B5ED7840D391D7E7BBC" italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="213076A17F786B89176663A8800FC8A0" authorityName="Morikawa" authorityYear="1954" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1E58396A4A9B8E1A4EBABE70F6D3658C" italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="3CEB5C5F184416149240813313C15819" DOI="https://doi.org/10.1636/JoA-S-21-025" author="You, J" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="135 - 157" refId="B33" refString="You, J, Yoo, JS, Harvey, MS, Harms, D, 2022. Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. The Journal of Arachnology 50 (2): 135 - 157, DOI: https://doi.org/10.1636/JoA-S-21-025" title="Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea." url="https://doi.org/10.1636/JoA-S-21-025" volume="50" year="2022">You et al. 2022</bibRefCitation>
|
||
). In our opinion, it is appropriate to place this new species to
|
||
<taxonomicName id="DD90F6DAAF49557ABC8269F83845CF94" authorityName="Morikawa" authorityYear="1954" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ABC0EB269869893D78E5233DE2A01E6E" italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="CDCCF7EB5C93567D6AC5EE1EE0F8D8BD" authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0AFFEC26BDB385D01FDB472A5A995D48" italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the reasons are as follows: for
|
||
<taxonomicName id="93436346D62726DBB6D5E5DBB3A9906D" authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="67E59A16523F2A320D9671D85CF24DD9" italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the number of carapaceal setae is not fixed (e.g., occasionally 18 are present in
|
||
<taxonomicName id="55907C6A8855B9807A4113F5A826AB09" lsidName="C. anatonus" pageId="0" pageNumber="33" rank="species" species="anatonus">
|
||
<emphasis id="8ABE1D5A9EC5A10E98D6F80EC93F11F6" italics="true" pageId="0" pageNumber="33">C. anatonus</emphasis>
|
||
</taxonomicName>
|
||
Harvey & Harms, 2022) and only four or five coxal spines blades; for
|
||
<taxonomicName id="E9DD9D6F1F9CA30C6209CC45046E6F1E" authorityName="Morikawa" authorityYear="1954" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8B817F91252FFFB366AC03E6BBB78E09" italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the character of coxal spines is diverse (e.g., in
|
||
<taxonomicName id="755832BBC8D10255652075187C01A4EE" class="Arachnida" kingdom="Animalia" lsidName="S. undecimclavatus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="undecimclavatus">
|
||
<emphasis id="32A996129F01C30CECE8B3B9E57490BD" italics="true" pageId="0" pageNumber="33">S. undecimclavatus</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956, which is club-shaped, not distally plumose).
|
||
</paragraph>
|
||
<paragraph id="2352332AE00FB7E59A864A1AD70FEBFD" pageId="0" pageNumber="33">
|
||
<taxonomicName id="CC38578297004BCAD581DABB84F3DFA7" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="606AEA238396287EDE7E8195C754CBFF" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName id="130069093BEE5F7EAFEA59D60E8500E1" class="Arachnida" kingdom="Animalia" lsidName="S. cheonsooi" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="cheonsooi">
|
||
<emphasis id="02552264186603CCE34C3BF65D3F0B9D" italics="true" pageId="0" pageNumber="33">S. cheonsooi</emphasis>
|
||
</taxonomicName>
|
||
You, Yoo, Harvey & Harms, 2022, but differs by the number of setae on tergite I (2 vs. 4) and larger body size (body length 1.93 (♀) mm vs. 1.70 (♀) mm; chela 5.68 (♀)
|
||
<normalizedToken id="42972226778F057CB4B890895331C093" originalValue="×">x</normalizedToken>
|
||
vs. 5.32 (♀)
|
||
<normalizedToken id="834EEE59D2132014EF2619598482F562" originalValue="×">x</normalizedToken>
|
||
longer than board, length 1.76 (♀) mm vs. 1.49 (♀) mm).
|
||
</paragraph>
|
||
<paragraph id="C3A1AEEC52E6A99204D82FA6398AB068" pageId="0" pageNumber="33">
|
||
<taxonomicName id="D4EAABB09139C740554D6070736A8940" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis id="D443E6C38EB41726C78932DFE316EA81" italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="A421CC6EE65C5A53604D113B6C922D74" class="Arachnida" kingdom="Animalia" lsidName="S. seungsookae" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="seungsookae">
|
||
<emphasis id="914D9E760273E462195C3081EE60B87E" italics="true" pageId="0" pageNumber="33">S. seungsookae</emphasis>
|
||
</taxonomicName>
|
||
You, Yoo, Harvey & Harms, 2022 by the number of setae on tergite I (2 vs. 4) and smaller body size (body length 1.93 (♀) mm vs. 2.05-2.36 (♀) mm; chela length 1.68-1.74 (♂), 1.76 (♀) mm vs. 1.90 (♂), 1.92 (♀) mm); from
|
||
<taxonomicName id="94D0ED9EB8AC5558BECB16FA6BF0A464" class="Arachnida" kingdom="Animalia" lsidName="S. undecimclavatus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="undecimclavatus">
|
||
<emphasis id="163A45BB766825E5811FA8A044F65818" italics="true" pageId="0" pageNumber="33">S. undecimclavatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6A05EFF06D6EF35C888A0F734BD48A68" class="Arachnida" kingdom="Animalia" lsidName="S. dorogawaensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="dorogawaensis">
|
||
<emphasis id="CEAF75CCE6EB515367182AC3A36E3553" italics="true" pageId="0" pageNumber="33">S. dorogawaensis</emphasis>
|
||
</taxonomicName>
|
||
by the number of setae on chelicera (6 vs. 7), a slender palp (palpal femur 7.24 (♂)
|
||
<normalizedToken id="B42A923564F80E80E6063EA39678A2EC" originalValue="×">x</normalizedToken>
|
||
vs. 4.80-5.40 (♂)
|
||
<normalizedToken id="A33F5BF7FD2E095FCEE96D74DA4615C6" originalValue="×">x</normalizedToken>
|
||
longer than board; chela 6.21-6.22 (♂)
|
||
<normalizedToken id="B53068E4946886D306EDB907F3E1E959" originalValue="×">x</normalizedToken>
|
||
vs. 5.50-6.13 (♂)
|
||
<normalizedToken id="B0DB876E0B8F6696FBC0F963A7607EBC" originalValue="×">x</normalizedToken>
|
||
longer than board) and lower number of blades of coxal spines (7-8 vs. 10-11); from
|
||
<taxonomicName id="1A9910B618EF5634640586DB449C1F7A" class="Arachnida" kingdom="Animalia" lsidName="S. akiyoshiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="akiyoshiensis">
|
||
<emphasis id="5B83544CC7731215BEEEAB7B267C2D72" italics="true" pageId="0" pageNumber="33">S. akiyoshiensis</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956 and
|
||
<taxonomicName id="8C21EEF29FC5FB98BE87037EA01EB3F3" class="Arachnida" kingdom="Animalia" lsidName="S. kobayashii" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kobayashii">
|
||
<emphasis id="E97B69118597DD3AD77AACD4BC16466F" italics="true" pageId="0" pageNumber="33">S. kobayashii</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956 by the number of setae on chelicera (6 vs. 7), lower number of movable chelal finger teeth (16-19 teeth vs. min. 26 teeth) and a slender palp (palpal femur 7.24 (♂)
|
||
<normalizedToken id="206FF54EA591BBED1539A221FACCC877" originalValue="×">x</normalizedToken>
|
||
vs. 5.00-5.60 (♂)
|
||
<normalizedToken id="1E2618D41125EEDB66B8DB4AA0DE2032" originalValue="×">x</normalizedToken>
|
||
longer than board; chela 6.21-6.22 (♂)
|
||
<normalizedToken id="371F33FD5E38FF32C7F6F976D3B9C079" originalValue="×">x</normalizedToken>
|
||
vs. 5.13-5.74 (♂)
|
||
<normalizedToken id="2AF397165205AD5295092D9C009CCA1B" originalValue="×">x</normalizedToken>
|
||
longer than board); from
|
||
<taxonomicName id="AEE4DD5CA110C737753C2E6EC600FA45" class="Arachnida" kingdom="Animalia" lsidName="S. dentifer" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="dentifer">
|
||
<emphasis id="C8F220EFB5117A4C7B95E3B6CD9AAB75" italics="true" pageId="0" pageNumber="33">S. dentifer</emphasis>
|
||
</taxonomicName>
|
||
(Morikawa, 1970) by the number of setae on chelicera (6 vs. 7), lower number of movable chelal finger teeth (16-19 teeth vs. min. 36 teeth) and a shorter chela (chela 6.21-6.22 (♂), 5.68 (♀)
|
||
<normalizedToken id="B30C07FE2B242DB5BDB62830E80471E2" originalValue="×">x</normalizedToken>
|
||
vs. 6.85 (♂), 7.12 (♀)
|
||
<normalizedToken id="28E5EA450D07D5A8FC5D768A1FDA108D" originalValue="×">x</normalizedToken>
|
||
longer than board, length 1.68-1.74 (♂), 1.76 (♀) mm vs. 1.85 (♂♀) mm); from
|
||
<taxonomicName id="7AC8016EA9BE21DA6F3E992B2123BDB2" class="Arachnida" kingdom="Animalia" lsidName="S. kubotai" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kubotai">
|
||
<emphasis id="C6937D8164BF012CE3F0DA59A2E96E5D" italics="true" pageId="0" pageNumber="33">S. kubotai</emphasis>
|
||
</taxonomicName>
|
||
by the slightly smaller body size (body length 1.93 (♀) mm vs. 2.03 (♀) mm; chela 5.68 (♀)
|
||
<normalizedToken id="D6AD33B9F8AFE0C0051894AB378A5457" originalValue="×">x</normalizedToken>
|
||
vs. 5.70 (♀)
|
||
<normalizedToken id="F9F060296E9207ED3DD808E1182FC07B" originalValue="×">x</normalizedToken>
|
||
longer than board; movable chelal finger 1.68 (♀)
|
||
<normalizedToken id="044D666E9400243F189561E59D2DFC79" originalValue="×">x</normalizedToken>
|
||
vs. 1.87 (♀)
|
||
<normalizedToken id="725B27F8FC7C80C278E79B5DCBE13202" originalValue="×">x</normalizedToken>
|
||
longer than) and the number of setae of coxal spines (7 or 8 vs. 11); from
|
||
<taxonomicName id="4D47BF6F30CBAD75A8ADA0485DFF7E0D" class="Arachnida" kingdom="Animalia" lsidName="S. kishidai" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kishidai">
|
||
<emphasis id="C50771F41CA5CCEBF9E10FA4FE8E8E23" italics="true" pageId="0" pageNumber="33">S. kishidai</emphasis>
|
||
</taxonomicName>
|
||
(Morikawa, 1960) by a slender palp (palpal femur 7.24
|
||
<normalizedToken id="A532F37305DCCFF7A6913FF48342DDE2" originalValue="×">x</normalizedToken>
|
||
vs. 4.90
|
||
<normalizedToken id="87517A4D0C1AF2DD2ED79B6EF6A20547" originalValue="×">x</normalizedToken>
|
||
longer than board; movable chelal finger 0.61-0.63
|
||
<normalizedToken id="69C90820982FE091BEB17261C8DFB62C" originalValue="×">x</normalizedToken>
|
||
vs. 0.67-0.69
|
||
<normalizedToken id="72E7E24F136F06E27026007A27B516EA" originalValue="×">x</normalizedToken>
|
||
longer than board) (
|
||
<bibRefCitation id="17A56CCF04F1E3A56B3813A172E0C266" author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B24" refString="Morikawa, K, 1954. On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2(2B): 79-87." title="On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2 (2 B): 79 - 87." year="1954">Morikawa 1954</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2861F37CDC840E472EBEEB1E82077069" author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B25" refString="Morikawa, K, 1956. Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2(2B): 271-282." title="Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2 (2 B): 271 - 282." year="1956">1956</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0C0D5F89937F1FD1054A44340555A526" author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B26" refString="Morikawa, K, 1960. Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2(2B): 85-172." title="Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2 (2 B): 85 - 172." year="1960">1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7A14902DB19DFF631D86410FF0CA2C08" author="Morikawa, K" journalOrPublisher="Bulletin of the National Science Museum, Tokyo" pageId="0" pageNumber="33" pagination="141 - 148" refId="B27" refString="Morikawa, K, 1970. Results of the speleological survey in South Korea 1966. XX. New pseudoscorpions from South Korea. Bulletin of the National Science Museum, Tokyo 13: 141 - 148" title="Results of the speleological survey in South Korea 1966. XX. New pseudoscorpions from South Korea." volume="13" year="1970">1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1E2C8EC1B5F8535A0F1020E1F0208BDC" DOI="https://doi.org/10.1636/JoA-S-21-025" author="You, J" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="135 - 157" refId="B33" refString="You, J, Yoo, JS, Harvey, MS, Harms, D, 2022. Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. The Journal of Arachnology 50 (2): 135 - 157, DOI: https://doi.org/10.1636/JoA-S-21-025" title="Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea." url="https://doi.org/10.1636/JoA-S-21-025" volume="50" year="2022">You et al. 2022</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="528D779290CA48F28CCD086D5C0E64E0" pageId="0" pageNumber="33" type="distribution">
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<paragraph id="A3BEFCE12B6A2ECA7554EC6DD744B02F" pageId="0" pageNumber="33">Distribution and habitat.</paragraph>
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<paragraph id="5F153255B018429C23C4A6CF9ACCAC8F" pageId="0" pageNumber="33">
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This species is known only from the type locality, Wulibei Cave (Figs
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<figureCitation id="A16B0E66314103CA1B9FB2E80BA109D0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1A</figureCitation>
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,
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<figureCitation id="D7AA7696C65135952A70A80DCDA21140" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18A-C</figureCitation>
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), which is located ~ 1.2 km east of Yangguan Village (Weining County). This limestone cave has an elongated entrance (~ 2.5 m high and 8 m wide) with some corn stalks scattered nearby. Entrance of the cave has a large muddy cave hall, connected to a small hall through a narrow tunnel, which is a more enclosed, completely dark space, covered with gravel, with temperatures ~ 10 °C and humidity ~ 90%. The specimen was collected under a stone in a small cave hall.
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</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |