493 lines
57 KiB
XML
493 lines
57 KiB
XML
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<mods:title id="C43017ABC822117AC21F15D35CC270A4">A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:namePart id="92A5C88153A15BAC6EB9C33F812C2C2C">Pimvichai, Piyatida</mods:namePart>
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<mods:namePart id="BCCDA1C7C94E2C3A10B858CC954A06E5">Enghoff, Henrik</mods:namePart>
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<mods:namePart id="D1D31E5F605F4A21902357AA425B1183">Panha, Somsak</mods:namePart>
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<mods:date id="DB47AD100075DDD3F3B53CF4472718B9">2011</mods:date>
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<treatment id="840255454255FFCEFF21FF67FF0B19F6" ID-DOI="http://doi.org/10.5281/zenodo.5667312" ID-GBIF-Taxon="119414435" ID-Zenodo-Dep="5667312" LSID="urn:lsid:plazi:treatment:840255454255FFCEFF21FF67FF0B19F6" httpUri="http://treatment.plazi.org/id/840255454255FFCEFF21FF67FF0B19F6" lastPageId="5" lastPageNumber="52" pageId="3" pageNumber="50">
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<subSubSection id="44B1B7D84255FFC8FF21FF67FEE41915" pageId="3" pageNumber="50" type="nomenclature">
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<paragraph id="0C14E4534255FFC8FF21FF67FDF818F5" blockId="3.[151,589,221,279]" box="[151,589,221,247]" pageId="3" pageNumber="50">
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<heading id="575C533F4255FFC8FF21FF67FDF818F5" bold="true" box="[151,589,221,247]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
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<taxonomicName id="CBAB9FD04255FFC8FF21FF67FDF818F5" ID-CoL="56T2J" authority="Attems, 1936" authorityName="Attems" authorityYear="1936" box="[151,589,221,247]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="induratus">
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<emphasis id="3EDF38414255FFC8FF21FF67FDF818F5" bold="true" box="[151,589,221,247]" pageId="3" pageNumber="50">
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<emphasis id="3EDF38414255FFC8FF21FF67FE1718F5" bold="true" box="[151,418,221,247]" italics="true" pageId="3" pageNumber="50">Thyropygus induratus</emphasis>
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<bibRefCitation id="683A99A24255FFC8FE1CFF67FDF818F5" author="Attems" box="[426,589,221,247]" pageId="3" pageNumber="50" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 122 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="0C14E4534255FFC8FF21FF45FEE41915" blockId="3.[151,589,221,279]" box="[151,337,255,279]" pageId="3" pageNumber="50">
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(
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<figureCitation id="9490F8D64255FFC8FF29FF45FF521915" box="[159,231,255,279]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
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A–E, 9c)
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</paragraph>
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</subSubSection>
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<subSubSection id="44B1B7D84255FFC8FF21FEF2FDFC19F5" pageId="3" pageNumber="50" type="reference_group">
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<paragraph id="0C14E4534255FFC8FF21FEF2FD82195C" blockId="3.[151,684,328,503]" box="[151,567,328,350]" pageId="3" pageNumber="50">
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<treatmentCitationGroup id="2CBBC37D4255FFC8FF21FEF2FD82195C" box="[151,567,328,350]" pageId="3" pageNumber="50">
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<taxonomicName id="CBAB9FD04255FFC8FF21FEF2FD82195C" ID-CoL="56T2J" authority="Attems, 1936: 262" authorityName="Attems" authorityPageNumber="262" authorityYear="1936" box="[151,567,328,350]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="induratus">
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<emphasis id="3EDF38414255FFC8FF21FEF2FEC0195C" box="[151,373,328,350]" italics="true" pageId="3" pageNumber="50">Thyropygus induratus</emphasis>
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<treatmentCitation id="8D0AC2424255FFC8FECDFEF2FD82195C" author="Attems" box="[379,567,328,350]" page="262" pageId="3" pageNumber="50" year="1936">
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<bibRefCitation id="683A99A24255FFC8FECDFEF2FDB2195C" author="Attems" box="[379,519,328,350]" pageId="3" pageNumber="50" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 122 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
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: 262
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="0C14E4534255FFC8FF21FEDDFD8D1998" blockId="3.[151,684,328,503]" pageId="3" pageNumber="50">
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<treatmentCitationGroup id="2CBBC37D4255FFC8FF21FEDDFD8D1998" pageId="3" pageNumber="50">
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<taxonomicName id="CBAB9FD04255FFC8FF21FEDDFD8F197E" ID-CoL="56T3X" authority="Attems, 1938: 281" authorityName="Attems" authorityPageNumber="281" authorityYear="1938" box="[151,570,359,381]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="3EDF38414255FFC8FF21FEDDFEC3197F" box="[151,374,359,381]" italics="true" pageId="3" pageNumber="50">Thyropygus punctatus</emphasis>
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<treatmentCitation id="8D0AC2424255FFC8FECAFEDDFD8F197E" author="Attems" box="[380,570,359,380]" page="281" pageId="3" pageNumber="50" year="1938">
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<bibRefCitation id="683A99A24255FFC8FECAFEDDFDBD197E" author="Attems" box="[380,520,359,380]" pageId="3" pageNumber="50" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6, 187 - 321." type="book chapter" year="1938">Attems, 1938</bibRefCitation>
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: 281
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</treatmentCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="25EC853A4255FFC8FDF3FEDDFD3E197E" box="[581,651,359,380]" pageId="3" pageNumber="50">
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<emphasis id="3EDF38414255FFC8FDF3FEDDFD3E197E" bold="true" box="[581,651,359,380]" pageId="3" pageNumber="50">syn. n.</emphasis>
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</taxonomicNameLabel>
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<taxonomicName id="CBAB9FD04255FFC8FF21FE3FFD8D1998" ID-CoL="56SX3" authority="Attems 1942: 87" authorityName="Attems" authorityPageNumber="87" authorityYear="1942" box="[151,568,389,411]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="induratus">
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<emphasis id="3EDF38414255FFC8FF21FE3FFE341999" box="[151,385,389,411]" italics="true" pageId="3" pageNumber="50">Thyropisthus induratus</emphasis>
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:
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<treatmentCitation id="8D0AC2424255FFC8FE39FE3FFD8D1998" author="Attems" box="[399,568,389,410]" page="87" pageId="3" pageNumber="50" year="1942">
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<bibRefCitation id="683A99A24255FFC8FE39FE3FFDA01998" author="Attems" box="[399,533,389,410]" pageId="3" pageNumber="50" refString="Attems, C. (1942) Zur Kenntnis der Indischen Harpagophoridae. Annalen des Naturhistorischen Museums in Wien, 52, 66 - 105, pl. 6 - 9." type="journal article" year="1942">Attems 1942</bibRefCitation>
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: 87
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="0C14E4534255FFC8FF21FE1EFDFC19F5" blockId="3.[151,684,328,503]" pageId="3" pageNumber="50">
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<treatmentCitationGroup id="2CBBC37D4255FFC8FF21FE1EFDFC19F5" pageId="3" pageNumber="50">
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<taxonomicName id="CBAB9FD04255FFC8FF21FE1EFD1E19BB" ID-CoL="56T2J" authority="Hoffman 1975: 136" authorityName="Hoffman" authorityPageNumber="136" authorityYear="1975" box="[151,683,419,442]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="allevatus" subSpecies="induratus">
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<emphasis id="3EDF38414255FFC8FF21FE1EFE6319B8" box="[151,470,420,442]" italics="true" pageId="3" pageNumber="50">Thyropygus allevatus induratus</emphasis>
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:
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<treatmentCitation id="8D0AC2424255FFC8FE52FE19FD1E19BB" author="Hoffman" box="[484,683,419,441]" page="136" pageId="3" pageNumber="50" year="1975">
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<bibRefCitation id="683A99A24255FFC8FE52FE19FDCE19BB" author="Hoffman" box="[484,635,419,441]" pageId="3" pageNumber="50" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman 1975</bibRefCitation>
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: 136
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||
</treatmentCitation>
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||
</taxonomicName>
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||
<taxonomicName id="CBAB9FD04255FFC8FF21FE78FD1919DA" ID-CoL="56T3X" authority="Hoffman 1975: 136" authorityName="Hoffman" authorityPageNumber="136" authorityYear="1975" box="[151,684,450,472]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="allevatus" subSpecies="punctatus">
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<emphasis id="3EDF38414255FFC8FF21FE78FE6D19DA" box="[151,472,450,472]" italics="true" pageId="3" pageNumber="50">Thyropygus allevatus punctatus</emphasis>
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||
:
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<treatmentCitation id="8D0AC2424255FFC8FE53FE78FD1919DA" author="Hoffman" box="[485,684,450,472]" page="136" pageId="3" pageNumber="50" year="1975">
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<bibRefCitation id="683A99A24255FFC8FE53FE78FDC919DA" author="Hoffman" box="[485,636,450,472]" pageId="3" pageNumber="50" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman 1975</bibRefCitation>
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: 136
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</treatmentCitation>
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</taxonomicName>
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<taxonomicName id="CBAB9FD04255FFC8FF21FE5BFDFC19F5" ID-CoL="56SXP" authority="Demange 1961: 142" authorityName="Demange" authorityPageNumber="142" authorityYear="1961" box="[151,585,481,503]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="resimus">
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||
<emphasis id="3EDF38414255FFC8FF21FE5BFEDB19F5" box="[151,366,481,503]" italics="true" pageId="3" pageNumber="50">Thyropisthus resimus</emphasis>
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:
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<treatmentCitation id="8D0AC2424255FFC8FECAFE58FDFC19F5" author="Demange" box="[380,585,481,503]" page="142" pageId="3" pageNumber="50" year="1961">
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<bibRefCitation id="683A99A24255FFC8FECAFE58FDAC19F5" author="Demange" box="[380,537,481,503]" pageId="3" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange 1961</bibRefCitation>
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: 142
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||
</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="44B1B7D84255FFC8FF21FD99FC8E1CC6" pageId="3" pageNumber="50" type="materials_examined">
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<paragraph id="0C14E4534255FFC8FF21FD99FC8E1CC6" blockId="3.[151,1437,547,2012]" pageId="3" pageNumber="50">
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<emphasis id="3EDF38414255FFC8FF21FD99FE371A3E" bold="true" box="[151,386,547,572]" pageId="3" pageNumber="50">Material examined.</emphasis>
|
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<typeStatus id="D3105AF14255FFC8FE3DFD9EFDAD1A3E" box="[395,536,548,572]" pageId="3" pageNumber="50" type="syntype">SYNTYPES</typeStatus>
|
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(only gonopods)
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<collectingCountry id="74BCA4C34255FFC8FD52FD9EFCC01A3E" box="[740,885,548,572]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
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(East
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||
<collectingCountry id="74BCA4C34255FFC8FC77FD9EFC4A1A3E" box="[961,1023,548,572]" name="Thailand" pageId="3" pageNumber="50">Siam</collectingCountry>
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).
|
||
<date id="7815C2934255FFC8FBA5FD9EFB141A3E" box="[1043,1185,548,573]" pageId="3" pageNumber="50" value="1917-05-05">5 May 1917</date>
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||
. C. Boden Kloss leg., don. Calcutta Mus. (
|
||
<collectionCode id="6ABA7C964255FFC8FEC8FDFDFE721A5D" box="[382,455,583,607]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="50">NMW</collectionCode>
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2606).
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||
<specimenCount id="1AAD2FDA4255FFC8FD96FDFDFDD81A5D" box="[544,621,583,607]" pageId="3" pageNumber="50" type="male">1 male</specimenCount>
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||
<typeStatus id="D3105AF14255FFC8FDC0FDFDFCBF1A5D" box="[630,778,583,607]" pageId="3" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
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of
|
||
<taxonomicName id="CBAB9FD04255FFC8FC84FDF3FC0E1A62" box="[818,955,585,608]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414255FFC8FC84FDF3FC0E1A62" box="[818,955,585,608]" italics="true" pageId="3" pageNumber="50">T. punctatus</emphasis>
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||
</taxonomicName>
|
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<collectingCountry id="74BCA4C34255FFC8FC72FDF2FBFD1A62" box="[964,1096,584,608]" name="Vietnam" pageId="3" pageNumber="50">VIETNAM</collectingCountry>
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,
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||
<collectingCountry id="74BCA4C34255FFC8FBE2FDFDFB511A5D" box="[1108,1252,583,607]" name="Vietnam" pageId="3" pageNumber="50">Cochinchina</collectingCountry>
|
||
, Poulo-Condore
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||
<date id="7815C2934255FFC8FF21FDD6FEE71A86" box="[151,338,620,644]" pageId="3" pageNumber="50" value="1933-12">December 1933</date>
|
||
. C. Dawydoff leg. (
|
||
<collectionCode id="6ABA7C964255FFC8FDF3FDD6FD261A86" box="[581,659,620,644]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="50">NMW</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FD1DFDD6FCBF1A86" box="[683,778,620,644]" pageId="3" pageNumber="50" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FCADFDD6FC381A86" box="[795,909,620,644]" pageId="3" pageNumber="50" type="female">3 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FC2CFDD6FC561A86" box="[922,995,620,644]" name="Thailand" pageId="3" pageNumber="50">SIAM</collectingCountry>
|
||
, Angkor Vat, Coll. Brolemann, 1902 (
|
||
<collectionCode id="6ABA7C964255FFC8FF29FD35FF4E1AA5" LSID="urn:lsid:biocol.org:col:34988" box="[159,251,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="50">MNHN</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FEB9FD35FEDE1AA5" box="[271,363,655,679]" pageId="3" pageNumber="50" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FECFFD35FE521AA5" box="[377,487,655,679]" pageId="3" pageNumber="50" type="female">2 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FE47FD2AFD301AAA" box="[497,645,656,680]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Khonkean Province, Phu-Wiang District, Phu-Wiang National Park, 16° 41ˏ 59˝ N, 102° 13ˏ 41˝ E.
|
||
<date id="7815C2934255FFC8FDBAFD0EFD671ACE" box="[524,722,692,716]" pageId="3" pageNumber="50" value="2008-10-11">11 October 2008</date>
|
||
. S. Satan and P. Pimvichai leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FBE0FD0EFB181ACE" box="[1110,1197,692,716]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FB74FD0EFAAB1ACE" box="[1218,1310,692,716]" pageId="3" pageNumber="50" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FA9BFD0FFA291ACE" box="[1325,1436,692,716]" pageId="3" pageNumber="50" type="female">7 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FF21FD62FE9E1AF2" box="[151,299,728,752]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Mahasarakham Province, Muang District, Ban Tha-Toom, 16° 10ˏ 43˝ N, 103° 27ˏ 4˝ E.
|
||
<date id="7815C2934255FFC8FAF1FD6DFF661B16" pageId="3" pageNumber="50" value="2010-07-30">30 July 2010</date>
|
||
. R. Chanabun leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FE00FD46FDB81B16" box="[438,525,764,788]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FDA9FD46FD321B16" box="[543,647,764,788]" pageId="3" pageNumber="50" type="male">10 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FD24FD46FCB91B16" box="[658,780,764,788]" pageId="3" pageNumber="50" type="female">10 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FCA2FD46FC1D1B16" box="[788,936,764,788]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Srisaket Province, Kantraraluk District, 14° 38ˏ 27˝ N, 104° 38ˏ 58˝ E.
|
||
<date id="7815C2934255FFC8FE79FCA5FD271B35" box="[463,658,799,823]" pageId="3" pageNumber="50" value="2010-10-26">26 October 2010</date>
|
||
. P. Prasankok leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FCC0FCA5FC781B35" box="[886,973,799,823]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FC57FCA5FB8E1B35" box="[993,1083,799,823]" pageId="3" pageNumber="50" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FBFEFCA5FB001B35" box="[1096,1205,799,823]" pageId="3" pageNumber="50" type="female">5 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FB08FC9AFAE71B3A" box="[1214,1362,800,824]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Ubon Ratchathani Province, Nam-Yeun District, Kang Lam-duan, 14° 26ˏ 8˝ N, 105° 6ˏ 18˝ E.
|
||
<date id="7815C2934255FFC8FB36FCFEFAF51B5E" box="[1152,1344,836,860]" pageId="3" pageNumber="50" value="2010-10-10">10 October 2010</date>
|
||
. P. Pimvichai, P. Prasankok, P. Tongkerd and members of Animal Systematics Research Unit leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FB74FCDDFAAC1B7D" box="[1218,1305,871,895]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FA98FCDDFACA1B7D" box="[1326,1407,871,895]" pageId="3" pageNumber="50" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FA38FCDDFF561BA6" pageId="3" pageNumber="50" type="female">1 female</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FF5AFC36FECA1BA6" box="[236,383,908,932]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Ubon Ratchathani Province, Nam-Yeun District, Yotdome wildlife sanctuary, 14° 26ˏ 4˝ N, 105° 6ˏ 13˝ E.
|
||
<date id="7815C2934255FFC8FEF6FC15FE461BC5" box="[320,499,943,967]" pageId="3" pageNumber="50" value="2010-10-09">9 October 2010</date>
|
||
. P. Pimvichai and members of Animal Systematics Research Unit leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FA95FC15FACE1BC5" box="[1315,1403,943,967]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FA38FC15FF6E1BEE" pageId="3" pageNumber="50" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FF5EFC6EFEE01BEE" box="[232,341,980,1004]" pageId="3" pageNumber="50" type="female">6 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FEE8FC6EFE471BEE" box="[350,498,980,1004]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Udornthani Province, Non-Sa-Ard District, Nangrin waterfall, 17° 3ˏ 56˝ N, 102° 44ˏ 4˝ E.
|
||
<date id="7815C2934255FFC8FEB3FC4DFE6D1C0D" box="[261,472,1015,1039]" pageId="3" pageNumber="50" value="2009-09-08">8 September 2009</date>
|
||
. R. Chanabun and members of Animal Systematics Research Unit leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FAA3FC4DFAD91C0D" box="[1301,1388,1015,1039]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FA36FC4DFF6E1C36" pageId="3" pageNumber="50" type="male">10 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FF5EFBA6FEE31C36" box="[232,342,1052,1076]" pageId="3" pageNumber="50" type="female">5 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FEE8FBA6FE471C36" box="[350,498,1052,1076]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Surin Province, Muang District, Phanom Sawai Forest Park, 14° 15ˏ 45˝ N, 103° 22ˏ 7˝ E.
|
||
<date id="7815C2934255FFC8FEB5FB85FE101C55" box="[259,421,1087,1111]" pageId="3" pageNumber="50" value="2009-04-26">26 April 2009</date>
|
||
. S. Panha, P. Pimvichai, P. Prasankok and members of Animal Systematics Research Unit leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FF61FBDEFE9B1C7E" box="[215,302,1124,1148]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FEF5FBDEFE281C7E" box="[323,413,1124,1148]" pageId="3" pageNumber="50" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="1AAD2FDA4255FFC8FE1CFBDEFDA21C7E" box="[426,535,1124,1148]" pageId="3" pageNumber="50" type="female">3 females</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FD96FBDEFD011C7E" box="[544,692,1124,1148]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, Chaiyaphum Province, Muang District, Tard-Tone waterfall, 15° 58ˏ 52˝ N, 102° 2ˏ 8˝ E.
|
||
<date id="7815C2934255FFC8FE73FB3DFDED1C9D" box="[453,600,1159,1183]" pageId="3" pageNumber="50" value="2009-06-08">8 June 2009</date>
|
||
. R. Chanabun and members of Animal Systematics Research Unit leg., (
|
||
<collectionCode id="6ABA7C964255FFC8FF29FB16FF421CC6" box="[159,247,1196,1220]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="3" pageNumber="50">CUMZ</collectionCode>
|
||
).
|
||
<specimenCount id="1AAD2FDA4255FFC8FEBFFB16FEE11CC6" box="[265,340,1196,1220]" pageId="3" pageNumber="50" type="male">1 male</specimenCount>
|
||
<collectingCountry id="74BCA4C34255FFC8FEEDFB16FE5A1CC6" box="[347,495,1196,1220]" name="Thailand" pageId="3" pageNumber="50">THAILAND</collectingCountry>
|
||
, bought in pet shop (
|
||
<collectionCode id="6ABA7C964255FFC8FD6EFB16FC9A1CC6" box="[728,815,1196,1220]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="50">ZMUC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="44B1B7D84255FFC8FF73FB75FDE11D56" pageId="3" pageNumber="50" type="diagnosis">
|
||
<paragraph id="0C14E4534255FFC8FF73FB75FDE11D56" blockId="3.[151,1437,547,2012]" pageId="3" pageNumber="50">
|
||
<emphasis id="3EDF38414255FFC8FF73FB75FE8A1CEA" bold="true" box="[197,319,1231,1256]" pageId="3" pageNumber="50">Diagnosis.</emphasis>
|
||
A species of the
|
||
<taxonomicName id="CBAB9FD04255FFC8FDB6FB6AFDDE1CEA" box="[512,619,1232,1256]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414255FFC8FDB6FB6AFDDE1CEA" box="[512,619,1232,1256]" italics="true" pageId="3" pageNumber="50">induratus</emphasis>
|
||
</taxonomicName>
|
||
subgroup. Differing from all other species (except
|
||
<taxonomicName id="CBAB9FD04255FFC8FB1FFB6BFAA01CEA" box="[1193,1301,1233,1256]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414255FFC8FB1FFB6BFAA01CEA" box="[1193,1301,1233,1256]" italics="true" pageId="3" pageNumber="50">T. quietus</emphasis>
|
||
</taxonomicName>
|
||
) in the subgroup by having two processes of anterior coxal fold (
|
||
<emphasis id="3EDF38414255FFC8FCACFB4FFC801D0E" box="[794,821,1269,1292]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
), the outer forming a slender, sharp spine, the inner broadly rounded and flexible, and by a very thick
|
||
<emphasis id="3EDF38414255FFC8FD7FFAA3FD511D32" box="[713,740,1305,1328]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
laterally. Differs from
|
||
<taxonomicName id="CBAB9FD04255FFC8FC50FAA3FBE61D32" box="[998,1107,1305,1328]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414255FFC8FC50FAA3FBE61D32" box="[998,1107,1305,1328]" italics="true" pageId="3" pageNumber="50">T. quietus</emphasis>
|
||
</taxonomicName>
|
||
by being much larger and by having extremely high paracoxites (
|
||
<emphasis id="3EDF38414255FFC8FD9CFA87FDF11D56" box="[554,580,1341,1364]" italics="true" pageId="3" pageNumber="50">px</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="44B1B7D84255FFCFFF73FAE5FC90191E" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="description">
|
||
<paragraph id="0C14E4534255FFC8FF73FAE5FEEF1DE6" blockId="3.[151,1437,547,2012]" pageId="3" pageNumber="50">
|
||
<emphasis id="3EDF38414255FFC8FF73FAE5FEE21D7A" bold="true" box="[197,343,1375,1400]" pageId="3" pageNumber="50">Description.</emphasis>
|
||
Adult males with 61–66 podous rings, no apodous rings. Length
|
||
<quantity id="CB5349B64255FFC8FB86FAE5FB141D7A" box="[1072,1185,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="3" pageNumber="50" unit="cm" value="19.0" valueMax="21.0" valueMin="17.0">17–21 cm</quantity>
|
||
, width
|
||
<quantity id="CB5349B64255FFC8FB45FAE5FA2D1D7A" box="[1267,1432,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.43" metricValueMin="1.02" pageId="3" pageNumber="50" unit="mm" value="12.25" valueMax="14.3" valueMin="10.2">10.2–14.3 mm</quantity>
|
||
. Adult females with 59–67 podous rings, no apodous rings. Length
|
||
<quantity id="CB5349B64255FFC8FC3BFA3EFBB51D9E" box="[909,1024,1412,1436]" metricMagnitude="-1" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" pageNumber="50" unit="cm" value="20.5" valueMax="22.0" valueMin="19.0">19–22 cm</quantity>
|
||
, width
|
||
<quantity id="CB5349B64255FFC8FBE2FA3EFB4E1D9E" box="[1108,1275,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" metricValueMax="1.51" metricValueMin="1.05" pageId="3" pageNumber="50" unit="mm" value="12.8" valueMax="15.1" valueMin="10.5">10.5–15.1 mm</quantity>
|
||
. Overall color of living animal (
|
||
<figureCitation id="9490F8D64255FFC8FEEEFA1DFE2F1DBD" box="[344,410,1447,1471]" captionStart="FIGURE 9. A" captionStartId="16.[151,250,1491,1514]" captionTargetBox="[162,1422,201,1456]" captionTargetId="figure@16.[151,1436,193,1470]" captionTargetPageId="16" captionText="FIGURE 9. A: Thyropygus siamensis (specimen from Sam Lan waterfall National Park), living male. B: Thyropygus macrosiamensis, living male (paratype). C: Thyropygus induratus (specimen from Yotdome), living male. D: Thyropygus laterolobatus, living male (paratype)." httpUri="https://zenodo.org/record/202435/files/figure.png" pageId="3" pageNumber="50">Fig. 9</figureCitation>
|
||
C) dark brown. Head, anterior body rings, epiproct and paraprocts reddish brown. Prozonite yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="0C14E4534255FFC8FF73FA55FE921F72" blockId="3.[151,1437,547,2012]" pageId="3" pageNumber="50">
|
||
Gonopods (
|
||
<figureCitation id="9490F8D64255FFC8FEFEFA55FE241E05" box="[328,401,1519,1543]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
|
||
A–E): Anterior coxal fold (
|
||
<emphasis id="3EDF38414255FFC8FD7FFA4BFD511E0A" box="[713,740,1521,1544]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
) (
|
||
<figureCitation id="9490F8D64255FFC8FD4AFA55FCF01E05" box="[764,837,1519,1543]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
|
||
A–B) basally slender, becoming broader towards tip, distally with two processes: the outer forming a slender, sharp spine, curving mesad; the inner broadly rounded, flattened, somewhat transparent, flexible;
|
||
<emphasis id="3EDF38414255FFC8FDCEF983FD261E52" box="[632,659,1593,1616]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
in posterior view very thickened laterally. Posterior coxal fold (
|
||
<emphasis id="3EDF38414255FFC8FACEF983FA261E52" box="[1400,1427,1593,1616]" italics="true" pageId="3" pageNumber="50">pc</emphasis>
|
||
) (
|
||
<figureCitation id="9490F8D64255FFC8FF29F9E6FF571E76" box="[159,226,1628,1652]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Fig. 1</figureCitation>
|
||
C) basally with extremely high lateral paracoxites (
|
||
<emphasis id="3EDF38414255FFC8FC9CF9E7FCF11E76" box="[810,836,1629,1652]" italics="true" pageId="3" pageNumber="50">px</emphasis>
|
||
); mesal process (
|
||
<emphasis id="3EDF38414255FFC8FBBCF9E7FB8F1E76" box="[1034,1082,1629,1652]" italics="true" pageId="3" pageNumber="50">pmp</emphasis>
|
||
) slightly shorter than
|
||
<emphasis id="3EDF38414255FFC8FA85F9E7FAFB1E76" box="[1331,1358,1629,1652]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
, forming shelf (
|
||
<emphasis id="3EDF38414255FFC8FEBAF93AFE901E9A" box="[268,293,1664,1688]" italics="true" pageId="3" pageNumber="50">sh</emphasis>
|
||
) for accommodation of telopodite, distally directed anteriad;
|
||
<emphasis id="3EDF38414255FFC8FC53F93BFBB51E9A" box="[997,1024,1665,1688]" italics="true" pageId="3" pageNumber="50">pc</emphasis>
|
||
fused with
|
||
<emphasis id="3EDF38414255FFC8FB3FF93BFB161E9A" box="[1161,1187,1665,1688]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
at base of the latter’s outer process. Telopodite (
|
||
<figureCitation id="9490F8D64255FFC8FE70F91EFDA01EBE" box="[454,533,1700,1724]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
|
||
D–E) leaving coxite over shelf of posterior coxal fold; femoral spine (
|
||
<emphasis id="3EDF38414255FFC8FAF4F91EFAE31EBE" box="[1346,1366,1700,1724]" italics="true" pageId="3" pageNumber="50">fe</emphasis>
|
||
) relatively long, apically curved,
|
||
<emphasis id="3EDF38414255FFC8FE6DF973FD971EE2" box="[475,546,1737,1760]" italics="true" pageId="3" pageNumber="50">in situ</emphasis>
|
||
resting behind (sometime in front of) inner process of
|
||
<emphasis id="3EDF38414255FFC8FB27F973FB1E1EE2" box="[1169,1195,1737,1760]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
; tibial spine (
|
||
<emphasis id="3EDF38414255FFC8FAFFF973FAED1EE2" box="[1353,1368,1737,1760]" italics="true" pageId="3" pageNumber="50">ti</emphasis>
|
||
) very long, slender, curving in an oblique-sagittal plane, its tip
|
||
<emphasis id="3EDF38414255FFC8FCA1F957FCE81F06" box="[791,861,1773,1796]" italics="true" pageId="3" pageNumber="50">in situ</emphasis>
|
||
resting close to distal-lateral part of
|
||
<emphasis id="3EDF38414255FFC8FB4CF957FA9F1F06" box="[1274,1322,1773,1796]" italics="true" pageId="3" pageNumber="50">pmp</emphasis>
|
||
or resting between tips of
|
||
<emphasis id="3EDF38414255FFC8FEE7F8ABFEDE1F2A" box="[337,363,1809,1832]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
; apical part: lamellar lobe (
|
||
<emphasis id="3EDF38414255FFC8FD19F8AAFD0A1F2A" box="[687,703,1808,1832]" italics="true" pageId="3" pageNumber="50">ll</emphasis>
|
||
) relatively broad, rounded, bent down, transparent; palette (
|
||
<emphasis id="3EDF38414255FFC8FAC0F8ABFA271F2A" box="[1398,1426,1809,1832]" italics="true" pageId="3" pageNumber="50">pa</emphasis>
|
||
) simple, basally with an uncate subterminal lobe (
|
||
<emphasis id="3EDF38414255FFC8FD09F88EFD551F4E" box="[703,736,1844,1868]" italics="true" pageId="3" pageNumber="50">usl</emphasis>
|
||
) ventrally; distally with about six to eight brownish blepharochaetae (
|
||
<emphasis id="3EDF38414255FFC8FF4DF8E2FEA21F72" box="[251,279,1880,1904]" italics="true" pageId="3" pageNumber="50">bp</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0C14E4534255FFCFFF71F8C6FC90191E" blockId="3.[151,1437,547,2012]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
|
||
<emphasis id="3EDF38414255FFC8FF71F8C6FEA41F96" bold="true" box="[199,273,1916,1940]" pageId="3" pageNumber="50">Notes.</emphasis>
|
||
Anterior coxal fold (
|
||
<emphasis id="3EDF38414255FFC8FE48F8C7FDAC1F96" box="[510,537,1917,1940]" italics="true" pageId="3" pageNumber="50">ac</emphasis>
|
||
) with some variation (
|
||
<figureCitation id="9490F8D64255FFC8FCA0F8C6FCD61F96" box="[790,867,1916,1940]" captionStart="FIGURE 1" captionStartId="4.[151,250,1658,1681]" captionTargetBox="[166,1423,346,1612]" captionTargetId="figure@4.[151,1436,337,1637]" captionTargetPageId="4" captionText="FIGURE 1. Thyropygus induratus (specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
|
||
A–B): in some individuals the inner process originates lower than the base of the outer process (does not fuse directly with curve at the basal part of outer process) and the outer process fuses directly with the distal part of
|
||
<emphasis id="3EDF38414255FFC8FCACF87FFCFF1FDE" box="[794,842,1989,2012]" italics="true" pageId="3" pageNumber="50">pmp</emphasis>
|
||
. In some individuals the outer and inner processes of
|
||
<emphasis id="3EDF38414252FFCFFF21FF23FF0718B2" box="[151,178,153,176]" italics="true" pageId="4" pageNumber="51">ac</emphasis>
|
||
are shorter and smaller than in others. The femoral spine (
|
||
<emphasis id="3EDF38414252FFCFFCFFFF22FCE818B2" box="[841,861,152,176]" italics="true" pageId="4" pageNumber="51">fe</emphasis>
|
||
) of specimens from Phu-Wiang, Nangrin waterfall and Tard Tone waterfall is relatively longer than in other specimens. All variable characters mainly separate northern (Phu-Wiang, Nangrin waterfall, Tard Tone waterfall and Ban Tha Toom) and southern populations (Kantraraluk District, Kang Lam-duan and Phanom Sawai Forest Park).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="58D4B4DB4252FFCFFF21F9C0FCB41ECF" httpUri="https://zenodo.org/record/202426/files/figure.png" pageId="4" pageNumber="51" targetBox="[166,1423,346,1612]" targetPageId="4">
|
||
<paragraph id="0C14E4534252FFCFFF21F9C0FCB41ECF" blockId="4.[151,1437,1658,1741]" pageId="4" pageNumber="51">
|
||
<emphasis id="3EDF38414252FFCFFF21F9C0FEA71E92" bold="true" box="[151,274,1658,1681]" pageId="4" pageNumber="51">FIGURE 1.</emphasis>
|
||
<taxonomicName id="CBAB9FD04252FFCFFEAFF9C0FE431E92" box="[281,502,1658,1680]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414252FFCFFEAFF9C0FE431E92" box="[281,502,1658,1680]" italics="true" pageId="4" pageNumber="51">Thyropygus induratus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Phu-wiang, except B), gonopods. A: anterior view, left telopodite removed. B: anterior view, anterior coxal fold (specimen from Yotdome). C: posterior view, left telopodite removed. D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="44B1B7D84252FFCEFF71F94FFF0B19F6" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51" type="discussion">
|
||
<paragraph id="0C14E4534252FFCEFF71F94FFAAC18F5" blockId="4.[151,1437,1781,2022]" lastBlockId="5.[151,1437,151,500]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">
|
||
<emphasis id="3EDF38414252FFCFFF71F94FFEF91F0C" bold="true" box="[199,332,1781,1806]" pageId="4" pageNumber="51">Discussion.</emphasis>
|
||
We found the gonopods of a
|
||
<taxonomicName id="CBAB9FD04252FFCFFD2FF94CFC941F0F" box="[665,801,1781,1805]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414252FFCFFD2FF94CFC941F0F" box="[665,801,1781,1805]" italics="true" pageId="4" pageNumber="51">T. induratus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="D3105AF14252FFCFFC9EF94FFC031F0F" box="[808,950,1781,1805]" pageId="4" pageNumber="51" type="syntype">SYNTYPES</typeStatus>
|
||
and those of the
|
||
<taxonomicName id="CBAB9FD04252FFCFFBCEF94CFAB41F0F" box="[1144,1281,1782,1805]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414252FFCFFBCEF94CFAB41F0F" box="[1144,1281,1782,1805]" italics="true" pageId="4" pageNumber="51">T. punctatus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="D3105AF14252FFCFFABEF94FFA291F0F" box="[1288,1436,1781,1805]" pageId="4" pageNumber="51" type="holotype">HOLOTYPE</typeStatus>
|
||
to be identical. These nominal species are also similar in diameter (
|
||
<quantity id="CB5349B64252FFCFFC39F8A3FC411F33" box="[911,1012,1817,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="51" unit="mm" value="11.5">11.5 mm</quantity>
|
||
and 11.0 mm respectively); the number of podous body rings in
|
||
<taxonomicName id="CBAB9FD04252FFCFFE56F887FDF91F57" box="[480,588,1853,1877]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414252FFCFFE56F887FDF91F57" box="[480,588,1853,1877]" italics="true" pageId="4" pageNumber="51">induratus</emphasis>
|
||
</taxonomicName>
|
||
is ca. 61–66 whereas in
|
||
<taxonomicName id="CBAB9FD04252FFCFFCDBF884FC6F1F57" box="[877,986,1854,1877]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414252FFCFFCDBF884FC6F1F57" box="[877,986,1854,1877]" italics="true" pageId="4" pageNumber="51">punctatus</emphasis>
|
||
</taxonomicName>
|
||
it is 56 (information on
|
||
<taxonomicName id="CBAB9FD04252FFCFFB4CF887FAD01F57" box="[1274,1381,1853,1877]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414252FFCFFB4CF887FAD01F57" box="[1274,1381,1853,1877]" italics="true" pageId="4" pageNumber="51">induratus</emphasis>
|
||
</taxonomicName>
|
||
also from
|
||
<bibRefCitation id="683A99A24252FFCFFF63F8DBFEC41F7B" author="Attems" box="[213,369,1889,1913]" pageId="4" pageNumber="51" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 122 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="683A99A24252FFCFFE33F8D8FDC81F7B" author="Demange" box="[389,637,1889,1914]" pageId="4" pageNumber="51" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961: 146)</bibRefCitation>
|
||
noticed the strong similarity between
|
||
<taxonomicName id="CBAB9FD04252FFCFFB99F8D8FB351F7B" box="[1071,1152,1890,1913]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414252FFCFFB99F8D8FB351F7B" box="[1071,1152,1890,1913]" italics="true" pageId="4" pageNumber="51">quietus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CBAB9FD04252FFCFFB0FF8D8FA931F7B" box="[1209,1318,1890,1913]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414252FFCFFB0FF8D8FA931F7B" box="[1209,1318,1890,1913]" italics="true" pageId="4" pageNumber="51">punctatus</emphasis>
|
||
</taxonomicName>
|
||
and noted only a very small difference between these two species.
|
||
<bibRefCitation id="683A99A24252FFCFFCB9F83FFC701F9F" author="Hoffman" box="[783,965,1925,1949]" pageId="4" pageNumber="51" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman (1975)</bibRefCitation>
|
||
mentioned the similarities between
|
||
<taxonomicName id="CBAB9FD04252FFCFFAEAF83CFF781FC3" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414252FFCFFAEAF83CFF781FC3" italics="true" pageId="4" pageNumber="51">punctatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="CBAB9FD04252FFCFFF6AF810FE841FC3" box="[220,305,1962,1985]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="resimus">
|
||
<emphasis id="3EDF38414252FFCFFF6AF810FE841FC3" box="[220,305,1962,1985]" italics="true" pageId="4" pageNumber="51">resimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CBAB9FD04252FFCFFEDFF810FE0F1FC3" box="[361,442,1962,1985]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414252FFCFFEDFF810FE0F1FC3" box="[361,442,1962,1985]" italics="true" pageId="4" pageNumber="51">quietus</emphasis>
|
||
</taxonomicName>
|
||
, noticing that Attems’ gonopod drawings are so similar that they might have been made from the same gonopods in different positions.
|
||
<bibRefCitation id="683A99A24252FFCFFD03F874FCEC1FE7" author="Attems" box="[693,857,1997,2022]" pageId="4" pageNumber="51" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 122 - 323." type="journal article" year="1936">Attems (1936)</bibRefCitation>
|
||
distinguished these three species by three variable characters: size, segmental sculpture and form of the epiproct. Moreover,
|
||
<bibRefCitation id="683A99A24253FFCEFC6AFF2DFB2118AD" author="Hoffman" box="[988,1172,151,175]" pageId="5" pageNumber="52" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman (1975)</bibRefCitation>
|
||
discussed the gonopod structure of
|
||
<taxonomicName id="CBAB9FD04253FFCEFEA9FF07FEC118D6" box="[287,372,189,212]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="resimus">
|
||
<emphasis id="3EDF38414253FFCEFEA9FF07FEC118D6" box="[287,372,189,212]" italics="true" pageId="5" pageNumber="52">resimus</emphasis>
|
||
</taxonomicName>
|
||
as described by
|
||
<bibRefCitation id="683A99A24253FFCEFD85FF07FC2E18D6" author="Demange" box="[563,923,188,213]" pageId="5" pageNumber="52" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961: figs. 184, 185)</bibRefCitation>
|
||
which he found not typical of
|
||
<taxonomicName id="CBAB9FD04253FFCEFB4FFF07FAFB18D6" box="[1273,1358,189,212]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="resimus">
|
||
<emphasis id="3EDF38414253FFCEFB4FFF07FAFB18D6" box="[1273,1358,189,212]" italics="true" pageId="5" pageNumber="52">resimus</emphasis>
|
||
</taxonomicName>
|
||
; Hoffman preferred to regard this as an intermediate between
|
||
<taxonomicName id="CBAB9FD04253FFCEFCBBFF5AFCCD18FA" box="[781,888,224,248]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFCBBFF5AFCCD18FA" box="[781,888,224,248]" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
and some more southern population.
|
||
</paragraph>
|
||
<paragraph id="0C14E4534253FFCEFF71FEBEFF0B19F6" blockId="5.[151,1437,151,500]" pageId="5" pageNumber="52">
|
||
On the basis of our material we propose to treat
|
||
<taxonomicName id="CBAB9FD04253FFCEFD55FEBFFCE5191E" box="[739,848,261,284]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414253FFCEFD55FEBFFCE5191E" box="[739,848,261,284]" italics="true" pageId="5" pageNumber="52">punctatus</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName id="CBAB9FD04253FFCEFBA3FEBEFB34191E" box="[1045,1153,260,284]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFBA3FEBEFB34191E" box="[1045,1153,260,284]" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
despite the smaller number of podous rings in the
|
||
<taxonomicName id="CBAB9FD04253FFCEFE0FFE93FD931942" box="[441,550,297,320]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="3EDF38414253FFCEFE0FFE93FD931942" box="[441,550,297,320]" italics="true" pageId="5" pageNumber="52">punctatus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="D3105AF14253FFCEFD9AFE9DFD27193D" box="[556,658,295,319]" pageId="5" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
. For
|
||
<taxonomicName id="CBAB9FD04253FFCEFD7FFE93FCAC1942" box="[713,793,297,320]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414253FFCEFD7FFE93FCAC1942" box="[713,793,297,320]" italics="true" pageId="5" pageNumber="52">quietus</emphasis>
|
||
</taxonomicName>
|
||
, we fully agree that the gonopods are identical with those of
|
||
<taxonomicName id="CBAB9FD04253FFCEFF00FEF6FE941966" box="[182,289,332,356]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFF00FEF6FE941966" box="[182,289,332,356]" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
, but the body size is significantly different (
|
||
<figureCitation id="9490F8D64253FFCEFCA1FEF6FCDE1966" box="[791,875,332,356]" captionStart="FIGURE 12" captionStartId="19.[151,250,1954,1977]" captionTargetBox="[219,1372,1357,1909]" captionTargetId="figure@19.[151,1436,1313,1951]" captionTargetPageId="19" captionText="FIGURE 12. Relationship between number of podous rings and body diameter in adult males of Thyropygus induratus and Thyropygus quietus." httpUri="https://zenodo.org/record/202438/files/figure.png" pageId="5" pageNumber="52">Fig. 12</figureCitation>
|
||
). Furthermore, genetic distances between
|
||
<taxonomicName id="CBAB9FD04253FFCEFAFEFEF6FF0D198A" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFAFEFEF6FF0D198A" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CBAB9FD04253FFCEFF58FECBFE8A198A" box="[238,319,369,392]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414253FFCEFF58FECBFE8A198A" box="[238,319,369,392]" italics="true" pageId="5" pageNumber="52">quietus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="4129D1E84253FFCEFEFBFED5FE141985" box="[333,417,367,391]" captionStart="TABLE 1" captionStartId="19.[151,235,152,173]" captionTargetBox="[159,1233,267,1194]" captionTargetPageId="19" captionText="TABLE 1. Pairwise comparisons of closely related species in the Thyropygus induratus subgroup; Thyropygus induratus and Thyropygus quietus (1 – 6); Thyropygus siamensis and Thyropygus macrosiamensis (7); compared with conspecific comparisons (8 – 22)." httpUri="http://table.plazi.org/id/58D4B4DB4245FFD8FF21FF22FF5418E9" pageId="5" pageNumber="52" tableUuid="58D4B4DB4245FFD8FF21FF22FF5418E9">Table 1</tableCitation>
|
||
) are consistently higher than conspecific distances between
|
||
<taxonomicName id="CBAB9FD04253FFCEFBFFFECAFB01198A" box="[1097,1204,368,392]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFBFFFECAFB01198A" box="[1097,1204,368,392]" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CBAB9FD04253FFCEFB5DFECBFA89198A" box="[1259,1340,369,392]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414253FFCEFB5DFECBFA89198A" box="[1259,1340,369,392]" italics="true" pageId="5" pageNumber="52">quietus</emphasis>
|
||
</taxonomicName>
|
||
individuals, see also discussion at end of paper. Altogether, we prefer to treat
|
||
<taxonomicName id="CBAB9FD04253FFCEFC05FE2EFBAA19AE" box="[947,1055,404,428]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFC05FE2EFBAA19AE" box="[947,1055,404,428]" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CBAB9FD04253FFCEFBE3FE2FFB1319AE" box="[1109,1190,405,428]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="quietus">
|
||
<emphasis id="3EDF38414253FFCEFBE3FE2FFB1319AE" box="[1109,1190,405,428]" italics="true" pageId="5" pageNumber="52">quietus</emphasis>
|
||
</taxonomicName>
|
||
as a separate species. On the other hand, we found the gonopod configuration of
|
||
<taxonomicName id="CBAB9FD04253FFCEFC80FE03FC3F19D2" box="[822,906,441,464]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="resimus">
|
||
<emphasis id="3EDF38414253FFCEFC80FE03FC3F19D2" box="[822,906,441,464]" italics="true" pageId="5" pageNumber="52">resimus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9490F8D64253FFCEFC2AFE0DFC5519CD" box="[924,992,439,463]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="5" pageNumber="52">Fig. 3</figureCitation>
|
||
) to be distinctly different from
|
||
<taxonomicName id="CBAB9FD04253FFCEFAFEFE02FF0D19F6" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="induratus">
|
||
<emphasis id="3EDF38414253FFCEFAFEFE02FF0D19F6" italics="true" pageId="5" pageNumber="52">induratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |