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<mods:title id="533806415ACCFED5E8553A55FA028C69">A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<paragraph id="0C14E4534250FFCDFF21FC7EFD831BDF" blockId="6.[151,566,963,1022]" box="[151,566,963,990]" pageId="6" pageNumber="53">
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<taxonomicName id="CBAB9FD04250FFCDFF21FC7EFD831BDF" ID-CoL="56T44" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[151,566,963,990]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF38414250FFCDFF21FC7EFD831BDF" bold="true" box="[151,566,963,990]" pageId="6" pageNumber="53">
<emphasis id="3EDF38414250FFCDFF21FC7EFE3E1BDC" bold="true" box="[151,395,964,990]" italics="true" pageId="6" pageNumber="53">Thyropygus resimus</emphasis>
<bibRefCitation id="683A99A24250FFCDFE25FC79FD831BDF" author="Attems" box="[403,566,963,989]" pageId="6" pageNumber="53" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6, 187 - 321." type="book chapter" year="1938">Attems, 1938</bibRefCitation>
</emphasis>
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(
<figureCitation id="9490F8D64250FFCDFF29FC5CFF5F1BFC" box="[159,234,998,1022]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="6" pageNumber="53">Figs. 3</figureCitation>
AE, 10AB)
</paragraph>
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<paragraph id="0C14E4534250FFCDFF21FB95FD961C46" blockId="6.[151,803,1071,1215]" box="[151,547,1071,1093]" pageId="6" pageNumber="53">
<treatmentCitationGroup id="2CBBC37D4250FFCDFF21FB95FD961C46" box="[151,547,1071,1093]" pageId="6" pageNumber="53">
<taxonomicName id="CBAB9FD04250FFCDFF21FB95FD961C46" ID-CoL="56T44" authority="Attems, 1938: 283" authorityName="Attems" authorityPageNumber="283" authorityYear="1938" box="[151,547,1071,1093]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF38414250FFCDFF21FB95FED71C47" box="[151,354,1071,1093]" italics="true" pageId="6" pageNumber="53">Thyropygus resimus</emphasis>
<treatmentCitation id="8D0AC2424250FFCDFEDEFB95FD961C46" author="Attems" box="[360,547,1071,1092]" page="283" pageId="6" pageNumber="53" year="1938">
<bibRefCitation id="683A99A24250FFCDFEDEFB95FE411C46" author="Attems" box="[360,500,1071,1092]" pageId="6" pageNumber="53" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6, 187 - 321." type="book chapter" year="1938">Attems, 1938</bibRefCitation>
: 283
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</paragraph>
<paragraph id="0C14E4534250FFCDFF21FBF7FD901C61" blockId="6.[151,803,1071,1215]" box="[151,549,1101,1123]" pageId="6" pageNumber="53">
<treatmentCitationGroup id="2CBBC37D4250FFCDFF21FBF7FD901C61" box="[151,549,1101,1123]" pageId="6" pageNumber="53">
<taxonomicName id="CBAB9FD04250FFCDFF21FBF7FD901C61" ID-CoL="56SXP" authority="Attems 1942: 80" authorityName="Attems" authorityPageNumber="80" authorityYear="1942" box="[151,549,1101,1123]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF38414250FFCDFF21FBF7FEDB1C61" box="[151,366,1101,1123]" italics="true" pageId="6" pageNumber="53">Thyropisthus resimus</emphasis>
:
<treatmentCitation id="8D0AC2424250FFCDFECAFBF4FD901C61" author="Attems" box="[380,549,1102,1123]" page="80" pageId="6" pageNumber="53" year="1942">
<bibRefCitation id="683A99A24250FFCDFECAFBF4FDB71C61" author="Attems" box="[380,514,1102,1123]" pageId="6" pageNumber="53" refString="Attems, C. (1942) Zur Kenntnis der Indischen Harpagophoridae. Annalen des Naturhistorischen Museums in Wien, 52, 66 - 105, pl. 6 - 9." type="journal article" year="1942">Attems 1942</bibRefCitation>
: 80
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</paragraph>
<paragraph id="0C14E4534250FFCDFF21FBD6FDE51CBD" blockId="6.[151,803,1071,1215]" pageId="6" pageNumber="53">
<treatmentCitationGroup id="2CBBC37D4250FFCDFF21FBD6FDE51CBD" pageId="6" pageNumber="53">
<taxonomicName id="CBAB9FD04250FFCDFF21FBD6FD2F1C83" ID-CoL="56T44" authority="Hoffman 1975: 136" authorityName="Hoffman" authorityPageNumber="136" authorityYear="1975" box="[151,666,1131,1154]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="allevatus" subSpecies="resimus">
<emphasis id="3EDF38414250FFCDFF21FBD6FE711C80" box="[151,452,1132,1154]" italics="true" pageId="6" pageNumber="53">Thyropygus allevatus resimus</emphasis>
:
<treatmentCitation id="8D0AC2424250FFCDFE67FBD1FD2F1C83" author="Hoffman" box="[465,666,1131,1153]" page="136" pageId="6" pageNumber="53" year="1975">
<bibRefCitation id="683A99A24250FFCDFE67FBD1FDDD1C83" author="Hoffman" box="[465,616,1131,1153]" pageId="6" pageNumber="53" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman 1975</bibRefCitation>
: 136
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, 137 figs 78 NOT
<taxonomicName id="CBAB9FD04250FFCDFF67FB30FD7D1CA2" ID-CoL="5F865" authority="Demange 1983" authorityName="Demange" authorityYear="1983" box="[209,712,1162,1184]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="confusus" subSpecies="tenuispinus">
<emphasis id="3EDF38414250FFCDFF67FB30FD951CA2" box="[209,544,1162,1184]" italics="true" pageId="6" pageNumber="53">Thyropygus confusus tenuispinus</emphasis>
:
<bibRefCitation id="683A99A24250FFCDFD98FB31FD7D1CA2" author="Demange" box="[558,712,1163,1184]" pageId="6" pageNumber="53" refString="Demange, J-M. (1983) Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). - Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 2 sect. A, 561 - 584." type="journal article" year="1983">Demange 1983</bibRefCitation>
</taxonomicName>
NOT
<taxonomicName id="CBAB9FD04250FFCDFF67FB13FDE51CBD" ID-CoL="56SXP" authority="Demange 1961" authorityName="Demange" authorityYear="1961" box="[209,592,1193,1215]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF38414250FFCDFF67FB13FE1D1CBD" box="[209,424,1193,1215]" italics="true" pageId="6" pageNumber="53">Thyropisthus resimus</emphasis>
:
<bibRefCitation id="683A99A24250FFCDFE03FB10FDE51CBD" author="Demange" box="[437,592,1193,1215]" pageId="6" pageNumber="53" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange 1961</bibRefCitation>
</taxonomicName>
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</paragraph>
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<emphasis id="3EDF38414250FFCDFF21FB51FE361D06" bold="true" box="[151,387,1259,1284]" pageId="6" pageNumber="53">Material examined.</emphasis>
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in same vial,
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, Savanna Khet, leg. C. Dawydoff 1933 (
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2575),
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,
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, Kalasin Province, Sahussakhan District, Phu-Kum-Khao, 16° 41ˏ 33˝ N, 103° 31ˏ 30˝ E.
<date id="7815C2934250FFCDFEE7FA8EFE591D4E" box="[337,492,1332,1357]" pageId="6" pageNumber="53" value="2007-05-18">18 May 2007</date>
. S. Satan leg., (
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).
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, Tak Province, Umphang District, Ta Kho Pi cave, 16° 3ˏ 14˝ N, 98° 49ˏ 16˝ E.
<date id="7815C2934250FFCDFD2CFAEDFC9D1D6D" box="[666,808,1367,1391]" pageId="6" pageNumber="53" value="2009-07-05">5 July 2009</date>
. Members of Animal Systematics Research Unit leg., (
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, Loei Province, Muang District, Piya cave, 17° 32ˏ 10˝ N, 101° 42ˏ 26˝ E.
<date id="7815C2934250FFCDFF55FA25FECB1DB5" box="[227,382,1439,1464]" pageId="6" pageNumber="53" value="2008-05-14">14 May 2008</date>
. R. Chanabun and members of Animal Systematics Research Unit leg., (
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).
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,
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, Mahasarakham Province, Muang District, Ban Tha-Toom, 16° 10ˏ 43˝ N, 103° 27ˏ 4˝ E.
<date id="7815C2934250FFCDFA36FA7EFEBD1DFD" pageId="6" pageNumber="53" value="2010-07-30">30 July 2010</date>
. R. Chanabun leg., (
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,
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<collectingCountry id="74BCA4C34250FFCDFC91FA52FC0E1E02" box="[807,955,1512,1536]" name="Thailand" pageId="6" pageNumber="53">THAILAND</collectingCountry>
, Kamphaengphet Province, Klonglarn District, Klong Larn waterfall National Park, 16° 7ˏ 50˝ N, 99° 16ˏ 38˝ E.
<date id="7815C2934250FFCDFC19F9B6FB8F1E26" box="[943,1082,1548,1572]" pageId="6" pageNumber="53" value="2009-06-08">8 June 2009</date>
. S. Panha and members of Animal Systematics Research Unit leg., (
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, Sakonnakorn Province, Phu-Pan District, Saerithai Cave, 17° 5ˏ 42˝ N, 103° 58ˏ 12˝ E.
<date id="7815C2934250FFCDFCB2F9EEFC511E6E" box="[772,996,1620,1644]" pageId="6" pageNumber="53" value="2009-09-10">10 September 2009</date>
. R. Chanabun and members of Animal Systematics Research Unit leg., (
<collectionCode id="6ABA7C964250FFCDFDBEF9CDFDEA1E8D" box="[520,607,1655,1679]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="6" pageNumber="53">CUMZ</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFDC7F9CDFD0A1E8D" box="[625,703,1655,1679]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
,
<specimenCount id="1AAD2FDA4250FFCDFD7DF9CDFC821E8D" box="[715,823,1655,1679]" pageId="6" pageNumber="53" type="female">5 females</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFC8BF9C2FC641E92" box="[829,977,1656,1680]" name="Thailand" pageId="6" pageNumber="53">THAILAND</collectingCountry>
, Phitsanulok Province, Wat Bod District, Kaeng Ched Kweaw entrance, 17° 8ˏ 32˝ N, 100° 26ˏ 28˝ E.
<date id="7815C2934250FFCDFCFBF926FC561EB6" box="[845,995,1692,1716]" pageId="6" pageNumber="53" value="2008-07-22">22 July 2008</date>
. R. Chanabun and members of Animal Systematics Research Unit leg., (
<collectionCode id="6ABA7C964250FFCDFDBEF905FDEA1ED5" box="[520,607,1727,1751]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="6" pageNumber="53">CUMZ</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFDC4F905FD091ED5" box="[626,700,1727,1751]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFD72F97AFCED1EDA" box="[708,856,1728,1752]" name="Thailand" pageId="6" pageNumber="53">THAILAND</collectingCountry>
, Tak Province, at Mae Usu cave, 17° 18ˏ 15˝ N, 94° 9ˏ 20˝ E.
<date id="7815C2934250FFCDFEBEF95EFE101EFE" box="[264,421,1764,1789]" pageId="6" pageNumber="53" value="2009-05-29">29 May 2009</date>
, C. Sutcharit and members of Animal Systematics Research Unit leg., (
<collectionCode id="6ABA7C964250FFCDFB54F95EFA8C1EFE" box="[1250,1337,1764,1788]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="6" pageNumber="53">CUMZ</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFAF8F95EFA2E1EFE" box="[1358,1435,1764,1788]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFF21F8B2FF651F1D" box="[151,208,1800,1823]" name="Laos" pageId="6" pageNumber="53">Laos</collectingCountry>
, Vientiane Province, Houayang National Park,
<geoCoordinate id="699F82944250FFCDFD52F8BDFCFC1F1D" box="[740,841,1799,1823]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="18.083334">18° 5ˏ N</geoCoordinate>
,
<geoCoordinate id="699F82944250FFCDFCE3F8BDFC671F1D" box="[853,978,1799,1823]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="102.666664">102° 40ˏ E</geoCoordinate>
.
<date id="7815C2934250FFCDFC6BF8BDFB141F1D" box="[989,1185,1799,1824]" pageId="6" pageNumber="53" value="2008-05-22" valueMax="2008-05-24" valueMin="2008-05-22">2224 May 2008</date>
. A. Solodovnikov and J. Pedersen leg., (
<collectionCode id="6ABA7C964250FFCDFEE9F896FE031F46" box="[351,438,1836,1860]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="53">ZMUC</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFE7CF896FD971F46" box="[458,546,1836,1860]" pageId="6" pageNumber="53" type="male">6 males</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFD9DF897FDD11F46" box="[555,612,1837,1860]" name="Laos" pageId="6" pageNumber="53">Laos</collectingCountry>
, Champasak Province, Bolaven plateau, Muang Paxong, Ban Thongvay,
<geoCoordinate id="699F82944250FFCDFF21F8F5FEBB1F65" box="[151,270,1871,1895]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="15.233334">15° 14ˏ N</geoCoordinate>
,
<geoCoordinate id="699F82944250FFCDFEAAF8F5FE2B1F65" box="[284,414,1871,1895]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="106.51667">106° 31ˏ E</geoCoordinate>
.
<date id="7815C2934250FFCDFE1DF8F5FDD41F65" box="[427,609,1871,1895]" pageId="6" pageNumber="53" value="2008-07-07" valueMax="2008-07-16" valueMin="2008-07-07">716 July 2008</date>
. A. Solodovnikov and J. Pedersen leg., (
<collectionCode id="6ABA7C964250FFCDFB8EF8F5FB3A1F65" box="[1080,1167,1871,1895]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="53">ZMUC</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFB12F8F5FB471F65" box="[1188,1266,1871,1895]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFB4AF8EAFA801F65" box="[1276,1333,1872,1895]" name="Laos" pageId="6" pageNumber="53">Laos</collectingCountry>
, Salavan Province,
<quantity id="CB5349B64250FFCDFEBEF8CEFEE61F8E" box="[264,339,1908,1932]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="53" unit="km" value="20.0">20 km</quantity>
. East of Ban Nong Bua,
<geoCoordinate id="699F82944250FFCDFDDFF8CEFD6E1F8E" box="[617,731,1908,1932]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="15.666667">15° 40ˏ N</geoCoordinate>
,
<geoCoordinate id="699F82944250FFCDFD50F8CEFCD61F8E" box="[742,867,1908,1932]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="106.55">106° 33ˏ E</geoCoordinate>
.
<date id="7815C2934250FFCDFCD9F8CEFBC41F8E" box="[879,1137,1908,1932]" pageId="6" pageNumber="53" value="2007-12-10" valueMax="2007-12-13" valueMin="2007-12-10">1013 December 2007</date>
. S. Tarasov leg., (
<collectionCode id="6ABA7C964250FFCDFA8FF8CEFA241F8E" box="[1337,1425,1908,1932]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="53">ZMUC</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFF21F82DFF511FAD" box="[151,228,1943,1967]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFF58F822FE371FB2" box="[238,386,1944,1968]" name="Thailand" pageId="6" pageNumber="53">THAILAND</collectingCountry>
, Chiang Mai Province, Fang Horticult. Exp. Station.
<date id="7815C2934250FFCDFC51F82DFB1F1FAD" box="[999,1194,1943,1967]" pageId="6" pageNumber="53" value="1981-10-21">21 October 1981</date>
. Zoological Museum Copenhagen leg., (
<collectionCode id="6ABA7C964250FFCDFED1F806FE0B1FD6" box="[359,446,1980,2004]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="53">ZMUC</collectionCode>
).
<specimenCount id="1AAD2FDA4250FFCDFE67F806FDAE1FD6" box="[465,539,1980,2004]" pageId="6" pageNumber="53" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34250FFCDFD94F806FD391FD6" box="[546,652,1980,2004]" name="Myanmar" pageId="6" pageNumber="53">BURMA</collectingCountry>
, Tenasserim, Mekane.
<date id="7815C2934250FFCDFC3CF806FBC71FD6" box="[906,1138,1980,2004]" pageId="6" pageNumber="53" value="1934-11-07" valueMax="1934-11-08" valueMin="1934-11-07">78 November 1934</date>
. R. Malaise leg., (
<collectionCode id="6ABA7C964250FFCDFA8FF806FA251FD6" box="[1337,1424,1980,2004]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="53">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="0C14E4534251FFCCFF21FF2DFE1D18D6" blockId="7.[151,1436,151,319]" pageId="7" pageNumber="54">
<specimenCount id="1AAD2FDA4251FFCCFF21FF2DFF5618AD" box="[151,227,151,175]" pageId="7" pageNumber="54" type="male">1 male</specimenCount>
<collectingCountry id="74BCA4C34251FFCCFF5DFF22FECA18B2" box="[235,383,152,176]" name="Thailand" pageId="7" pageNumber="54">THAILAND</collectingCountry>
, Loei Provine, Ban Si Than (mislabeled as Silang, Northern
<collectingCountry id="74BCA4C34251FFCCFB99FF2DFB2D18AD" box="[1071,1176,151,175]" name="Thailand" pageId="7" pageNumber="54">Thailand</collectingCountry>
).
<date id="7815C2934251FFCCFB1DFF2DFAC318AD" box="[1195,1398,151,175]" pageId="7" pageNumber="54" value="1958-12-03">3 December 1958</date>
. B. Degerbøl leg., (
<collectionCode id="6ABA7C964251FFCCFEF3FF06FE2818D6" box="[325,413,188,212]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="54">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="44B1B7D84251FFCCFF73FF65FDC3193D" pageId="7" pageNumber="54" type="diagnosis">
<paragraph id="0C14E4534251FFCCFF73FF65FDC3193D" blockId="7.[151,1436,151,319]" pageId="7" pageNumber="54">
<emphasis id="3EDF38414251FFCCFF73FF65FE8A18FA" bold="true" box="[197,319,223,248]" pageId="7" pageNumber="54">Diagnosis.</emphasis>
A species of the
<taxonomicName id="CBAB9FD04251FFCCFE48FF5AFDDC18FA" box="[510,617,224,248]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="induratus">
<emphasis id="3EDF38414251FFCCFE48FF5AFDDC18FA" box="[510,617,224,248]" italics="true" pageId="7" pageNumber="54">induratus</emphasis>
</taxonomicName>
subgroup. Differing from all other species in the subgroup by having one or more denticles in the middle of tibial spine (
<emphasis id="3EDF38414251FFCCFD0BFEBFFD79191E" box="[701,716,261,284]" italics="true" pageId="7" pageNumber="54">ti</emphasis>
) and by having anterior coxal fold (
<emphasis id="3EDF38414251FFCCFBC3FEBFFB25191E" box="[1141,1168,261,284]" italics="true" pageId="7" pageNumber="54">ac</emphasis>
) apically bent abruptly mesad with an acute narrowed, slender tip.
</paragraph>
</subSubSection>
<caption id="58D4B4DB4251FFCCFF21F905FAD81EF6" httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="7" pageNumber="54" targetBox="[240,1351,368,1706]" targetPageId="7">
<paragraph id="0C14E4534251FFCCFF21F905FAD81EF6" blockId="7.[151,1436,1727,1780]" pageId="7" pageNumber="54">
<emphasis id="3EDF38414251FFCCFF21F905FEA01ED6" bold="true" box="[151,277,1727,1750]" pageId="7" pageNumber="54">FIGURE 3.</emphasis>
<taxonomicName id="CBAB9FD04251FFCCFEA8F905FE591ED7" box="[286,492,1727,1749]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF38414251FFCCFEA8F905FE591ED7" box="[286,492,1727,1749]" italics="true" pageId="7" pageNumber="54">Thyropygus resimus</emphasis>
</taxonomicName>
(specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view.
</paragraph>
</caption>
<subSubSection id="44B1B7D84251FFC3FF73F89AFDBB1A86" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54" type="description">
<paragraph id="0C14E4534251FFCCFF73F89AFBE41FA6" blockId="7.[151,1436,1824,2029]" pageId="7" pageNumber="54">
<emphasis id="3EDF38414251FFCCFF73F89AFEE21F3B" bold="true" box="[197,343,1824,1849]" pageId="7" pageNumber="54">Description.</emphasis>
Adult males with 5771 podous rings, no apodous rings. Length
<quantity id="CB5349B64251FFCCFB8CF89AFB181F3A" box="[1082,1197,1824,1848]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="7" pageNumber="54" unit="cm" value="18.0" valueMax="21.0" valueMin="15.0">1521 cm</quantity>
, width 8.0
<quantity id="CB5349B64251FFCCFA84F89AFA2D1F3A" box="[1330,1432,1824,1848]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="7" pageNumber="54" unit="mm" value="12.9">12.9 mm</quantity>
. Adult females with 6471 podous rings, no apodous rings. Length
<quantity id="CB5349B64251FFCCFC34F8FEFC411F5E" box="[898,1012,1860,1884]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="7" pageNumber="54" unit="cm" value="16.5" valueMax="20.0" valueMin="13.0">1320 cm</quantity>
, width
<quantity id="CB5349B64251FFCCFBF0F8FEFB6A1F5E" box="[1094,1247,1860,1884]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.41" metricValueMin="0.94" pageId="7" pageNumber="54" unit="mm" value="11.75" valueMax="14.1" valueMin="9.4">9.414.1 mm</quantity>
. Overall color of living animal in some populations (
<figureCitation id="9490F8D64251FFCCFD8EF8D2FD161F82" box="[568,675,1896,1920]" captionStart="FIGURE 10. A, B" captionStartId="17.[151,250,1543,1566]" captionTargetBox="[162,1423,262,1516]" captionTargetId="figure@17.[151,1436,253,1530]" captionTargetPageId="17" captionText="FIGURE 10. A, B: Thyropygus resimus, living males (specimens from A: Phu-Kum-Khao, B: Klong Larn waterfall National Park.). C, D: Thyropygus uncinatus, living male and female (specimens from C: Roi-Et (male), D: Phukhae Botanical Garden (female)." httpUri="https://zenodo.org/record/202436/files/figure.png" pageId="7" pageNumber="54">Fig. 10A</figureCitation>
) dark brown, with prozonite brownish grey and epiproct reddish brown. In some populations, overall color of living animal (
<figureCitation id="9490F8D64251FFCCFC82F836FC2C1FA6" box="[820,921,1932,1956]" captionStart="FIGURE 10. A, B" captionStartId="17.[151,250,1543,1566]" captionTargetBox="[162,1423,262,1516]" captionTargetId="figure@17.[151,1436,253,1530]" captionTargetPageId="17" captionText="FIGURE 10. A, B: Thyropygus resimus, living males (specimens from A: Phu-Kum-Khao, B: Klong Larn waterfall National Park.). C, D: Thyropygus uncinatus, living male and female (specimens from C: Roi-Et (male), D: Phukhae Botanical Garden (female)." httpUri="https://zenodo.org/record/202436/files/figure.png" pageId="7" pageNumber="54">Fig. 10B</figureCitation>
) reddish brown.
</paragraph>
<paragraph id="0C14E4534251FFC3FF71F80AFD4F1985" blockId="7.[151,1436,1824,2029]" lastBlockId="8.[151,1437,151,824]" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
Gonopods (
<figureCitation id="9490F8D64251FFCCFEFFF80AFE201FCA" box="[329,405,1968,1992]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="7" pageNumber="54">Figs. 3</figureCitation>
AE): Anterior coxal fold (
<emphasis id="3EDF38414251FFCCFD71F80BFD541FCA" box="[711,737,1969,1992]" italics="true" pageId="7" pageNumber="54">ac</emphasis>
) (
<figureCitation id="9490F8D64251FFCCFD4CF80AFC8E1FCA" box="[762,827,1968,1992]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="7" pageNumber="54">Fig. 3</figureCitation>
A) basally slender, becoming broader towards tip, apically bent abruptly mesad, with acute, slender tip. Posterior coxal fold (
<emphasis id="3EDF38414251FFCCFC0AF86CFC631FEF" box="[956,982,2006,2029]" italics="true" pageId="7" pageNumber="54">pc</emphasis>
) (
<figureCitation id="9490F8D64251FFCCFC59F86EFB851FEE" box="[1007,1072,2004,2028]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="7" pageNumber="54">Fig. 3</figureCitation>
B) basally with moderately high lateral paracoxites (
<emphasis id="3EDF3841425EFFC3FECAFF23FE2318B2" box="[380,406,153,176]" italics="true" pageId="8" pageNumber="55">px</emphasis>
); mesal process (
<emphasis id="3EDF3841425EFFC3FDD1FF23FD2218B2" box="[615,663,153,176]" italics="true" pageId="8" pageNumber="55">pmp</emphasis>
) slightly lower than
<emphasis id="3EDF3841425EFFC3FC27FF23FC1E18B2" box="[913,939,153,176]" italics="true" pageId="8" pageNumber="55">ac</emphasis>
, forming shelf (
<emphasis id="3EDF3841425EFFC3FBDBFF22FB3318B2" box="[1133,1158,152,176]" italics="true" pageId="8" pageNumber="55">sh</emphasis>
) for accommodation of telopodite, distally directed distolaterad. Telopodite (
<figureCitation id="9490F8D6425EFFC3FD5CFF06FC8318D6" box="[746,822,188,212]" captionStart="FIGURE 3" captionStartId="7.[151,250,1727,1750]" captionTargetBox="[240,1351,368,1706]" captionTargetId="figure@7.[226,1360,361,1722]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view." httpUri="https://zenodo.org/record/202429/files/figure.png" pageId="8" pageNumber="55">Figs. 3</figureCitation>
CE) leaving coxite over shelf of posterior coxal fold; femoral spine (
<emphasis id="3EDF3841425EFFC3FEF4FF5AFEE318FA" box="[322,342,224,248]" italics="true" pageId="8" pageNumber="55">fe</emphasis>
) relatively long, curving in a horizontal plane,
<emphasis id="3EDF3841425EFFC3FCD9FF5BFC0118FA" box="[879,948,225,248]" italics="true" pageId="8" pageNumber="55">in situ</emphasis>
resting along apical part of
<emphasis id="3EDF3841425EFFC3FB43FF5BFABA18FA" box="[1269,1295,225,248]" italics="true" pageId="8" pageNumber="55">ac</emphasis>
; tibial spine (
<emphasis id="3EDF3841425EFFC3FF16FEBFFF05191E" box="[160,176,261,284]" italics="true" pageId="8" pageNumber="55">ti</emphasis>
) very long, slender, curving in an oblique-sagittal plane, in the middle sometimes with a single denticle (
<emphasis id="3EDF3841425EFFC3FACEFEBEFA26191E" box="[1400,1427,260,284]" italics="true" pageId="8" pageNumber="55">de</emphasis>
) (rarely with several denticles) directed laterally, its tip
<emphasis id="3EDF3841425EFFC3FD4DFE93FC8A1942" box="[763,831,297,320]" italics="true" pageId="8" pageNumber="55">in situ</emphasis>
close to tip of
<emphasis id="3EDF3841425EFFC3FC51FE92FC4E1942" box="[999,1019,296,320]" italics="true" pageId="8" pageNumber="55">fe</emphasis>
and
<emphasis id="3EDF3841425EFFC3FB84FE93FBF91942" box="[1074,1100,297,320]" italics="true" pageId="8" pageNumber="55">ac</emphasis>
; apical part: lamellar lobe (
<emphasis id="3EDF3841425EFFC3FA35FE92FA261942" box="[1411,1427,296,320]" italics="true" pageId="8" pageNumber="55">ll</emphasis>
) broad, rounded, transparent; palette (
<emphasis id="3EDF3841425EFFC3FD88FEF7FDEF1966" box="[574,602,333,356]" italics="true" pageId="8" pageNumber="55">pa</emphasis>
) simple, basally with an uncate subterminal lobe (
<emphasis id="3EDF3841425EFFC3FB14FEF6FB771966" box="[1186,1218,332,356]" italics="true" pageId="8" pageNumber="55">usl</emphasis>
) ventrally; distally with about six to eight brownish blepharochaetae (
<emphasis id="3EDF3841425EFFC3FD7BFECAFD5C198A" box="[717,745,368,392]" italics="true" pageId="8" pageNumber="55">bp</emphasis>
).
</paragraph>
<paragraph id="0C14E453425EFFC3FF73FE2EFDBB1A86" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="55">
<emphasis id="3EDF3841425EFFC3FF73FE2EFEBA19AE" bold="true" box="[197,271,404,428]" pageId="8" pageNumber="55">Notes.</emphasis>
Colour: some specimens are overall dark brown, some are reddish brown; specimens from
<collectingCountry id="74BCA4C3425EFFC3FAA3FE2FFAFE19AE" box="[1301,1355,405,428]" name="Laos" pageId="8" pageNumber="55">Laos</collectingCountry>
have a longitudinal brown band mid-dorsally on the body, not seen in
<collectingCountry id="74BCA4C3425EFFC3FCE5FE0DFC3319CD" box="[851,902,439,463]" name="Thailand" pageId="8" pageNumber="55">Thai</collectingCountry>
specimens. The gonopods show some variation of the tip of anterior coxal fold (
<emphasis id="3EDF3841425EFFC3FDB6FE67FDAE19F6" box="[512,539,477,500]" italics="true" pageId="8" pageNumber="55">ac</emphasis>
): some specimens with a relatively short, blunt tip, others with a narrower, relatively long, almost sharp tip. The femoral spine (
<emphasis id="3EDF3841425EFFC3FD0DFDBAFD7A1A1A" box="[699,719,512,536]" italics="true" pageId="8" pageNumber="55">fe</emphasis>
) is relatively short in some specimens, but in Tak and Kamphaengphet specimens it is much longer and is distinctly crenulated along its inner curvature. The tibial spine (
<emphasis id="3EDF3841425EFFC3FAF2FD9FFAE11A3E" box="[1348,1364,549,572]" italics="true" pageId="8" pageNumber="55">ti</emphasis>
) has a denticle (
<emphasis id="3EDF3841425EFFC3FF49FDF2FEAF1A62" box="[255,282,584,608]" italics="true" pageId="8" pageNumber="55">de</emphasis>
) (sometimes several denticles) on the outer margin at mid-length or slightly further distal in some specimens, others lack such denticles.
</paragraph>
</subSubSection>
<subSubSection id="44B1B7D8425EFFC3FF73FD35FC791B35" pageId="8" pageNumber="55" type="reference_group">
<paragraph id="0C14E453425EFFC3FF73FD35FC791B35" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="55">
Concerning
<taxonomicName id="CBAB9FD0425EFFC3FEE7FD2AFCE11AA5" ID-CoL="56SXP" authority="Demange (1961)" authorityName="Demange" authorityYear="1961" box="[337,852,655,680]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF3841425EFFC3FEE7FD2AFD3C1AAA" box="[337,649,656,680]" italics="true" pageId="8" pageNumber="55">Thyropisthus resimus sensu</emphasis>
<bibRefCitation id="683A99A2425EFFC3FD24FD2AFCE11AA5" author="Demange" box="[658,852,655,680]" pageId="8" pageNumber="55" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961)</bibRefCitation>
</taxonomicName>
, see above under
<taxonomicName id="CBAB9FD0425EFFC3FB92FD2BFB181AAA" ID-CoL="56T2J" box="[1060,1197,656,680]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="induratus">
<emphasis id="3EDF3841425EFFC3FB92FD2BFB181AAA" box="[1060,1197,656,680]" italics="true" pageId="8" pageNumber="55">T. induratus</emphasis>
</taxonomicName>
. The specimen from “Northern
<collectingCountry id="74BCA4C3425EFFC3FEA6FD0EFEC61ACE" box="[272,371,692,716]" name="Thailand" pageId="8" pageNumber="55">Thailand</collectingCountry>
Silang” recorded as
<taxonomicName id="CBAB9FD0425EFFC3FDE8FD0FFCD71ACE" ID-CoL="5F865" box="[606,866,692,716]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="confusus" subSpecies="tenuispinus">
<emphasis id="3EDF3841425EFFC3FDE8FD0FFCD71ACE" box="[606,866,692,716]" italics="true" pageId="8" pageNumber="55">T. confusus tenuispinus</emphasis>
</taxonomicName>
by
<bibRefCitation id="683A99A2425EFFC3FC3BFD0FFBFF1ACE" author="Demange" box="[909,1098,692,717]" pageId="8" pageNumber="55" refString="Demange, J-M. (1983) Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). - Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 2 sect. A, 561 - 584." type="journal article" year="1983">Demange (1983)</bibRefCitation>
is in fact a
<taxonomicName id="CBAB9FD0425EFFC3FB78FD0FFA8B1ACE" ID-CoL="56SXP" box="[1230,1342,693,716]" class="Diplopoda" family="Harpagophoridae" genus="Thyropisthus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="resimus">
<emphasis id="3EDF3841425EFFC3FB78FD0FFA8B1ACE" box="[1230,1342,693,716]" italics="true" pageId="8" pageNumber="55">T. resimus</emphasis>
</taxonomicName>
, see list of material for interpretation of Silang. A specimen from the orchid museum, Bangkok, also recorded as
<taxonomicName id="CBAB9FD0425EFFC3FAF8FD63FEE21B16" ID-CoL="5F865" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="confusus" subSpecies="tenuispinus">
<emphasis id="3EDF3841425EFFC3FAF8FD63FEE21B16" italics="true" pageId="8" pageNumber="55">T. confusus tenuispinus</emphasis>
</taxonomicName>
by
<bibRefCitation id="683A99A2425EFFC3FE37FD47FD881B16" author="Demange" box="[385,573,764,789]" pageId="8" pageNumber="55" refString="Demange, J-M. (1983) Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). - Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 2 sect. A, 561 - 584." type="journal article" year="1983">Demange (1983)</bibRefCitation>
is in fact a
<taxonomicName id="CBAB9FD0425EFFC3FD76FD47FCF51B16" ID-CoL="56SYD" box="[704,832,764,788]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="3EDF3841425EFFC3FD76FD47FCF51B16" box="[704,832,764,788]" italics="true" pageId="8" pageNumber="55">T. allevatus</emphasis>
</taxonomicName>
. These are so far the only records of
<taxonomicName id="CBAB9FD0425EFFC3FB57FD47FAEB1B16" ID-CoL="7BZC6" box="[1249,1374,764,788]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="3EDF3841425EFFC3FB57FD47FAEB1B16" box="[1249,1374,764,788]" italics="true" pageId="8" pageNumber="55">T. confusus</emphasis>
</taxonomicName>
from
<collectingCountry id="74BCA4C3425EFFC3FF21FCA5FF481B35" box="[151,253,799,823]" name="Thailand" pageId="8" pageNumber="55">Thailand</collectingCountry>
, so this species should be removed from the list of
<collectingCountry id="74BCA4C3425EFFC3FC8FFCA5FCD91B35" box="[825,876,799,823]" name="Thailand" pageId="8" pageNumber="55">Thai</collectingCountry>
species.
</paragraph>
</subSubSection>
</treatment>
</document>