269 lines
17 KiB
XML
269 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne" docDate="2023" docId="26BF2BBD0FF15342AACF7E8B6DF4E131" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila tridentata Pall-Gergely & Hunyadi 2023, sp. nov." docType="treatment" docUuid="477B871F-CCA1-478C-985B-7FE20BBBA22A" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reischuetz, Alexander</mods:namePart>
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<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
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<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart>Oerstan, Aydin</mods:namePart>
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<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:name>
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||
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>ZooKeys</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2023</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2023-02-13</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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||
<mods:number>1147</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>1</mods:start>
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||
<mods:end>177</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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||
<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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||
</mods:location>
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||
<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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||
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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||
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
|
||
</mods:mods>
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||
<subSection pageId="0" pageNumber="1" type="systematic descriptions">
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||
<treatment LSID="urn:lsid:zoobank.org:act:477B871F-CCA1-478C-985B-7FE20BBBA22A" httpUri="http://treatment.plazi.org/id/26BF2BBD0FF15342AACF7E8B6DF4E131" lastPageNumber="1" pageId="0" pageNumber="1">
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||
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="1">
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||
<taxonomicName LSID="https://zoobank.org/477B871F-CCA1-478C-985B-7FE20BBBA22A" authority="Páll-Gergely & Hunyadi" authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila tridentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tridentata" status="sp. nov.">
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Angustopila tridentata
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 95" captionStartId="F95" captionText="Figure 95. Angustopila tridentata Pall-Gergely & Hunyadi, sp. nov. (holotype, HNHM 105321). Apertural (A), apical (B), lateral (C) and ventral (D) sides of the shell; sculpture on the protoconch (E), ventral (F) and frontal (G) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure95" httpUri="https://binary.pensoft.net/fig/808904" pageId="0" pageNumber="1">Figs 95</figureCitation>
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<figureCitation captionStart="Figure 96" captionStartId="F96" captionText="Figure 96. Variability of Angustopila tridentata Pall-Gergely & Hunyadi, sp. nov. A holotype (specimen 3) B specimen 2 C specimen 1." figureDoi="10.3897/zookeys.1147.93824.figure96" httpUri="https://binary.pensoft.net/fig/808905" pageId="0" pageNumber="1">, 96</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Vietnam • 1 empty shell (H: 0.92 mm, D: 0.9 mm);
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<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
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<normalizedToken originalValue="Bình">Binh</normalizedToken>
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Province, Kim
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<normalizedToken originalValue="Bȏi">Bȏi</normalizedToken>
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District, Cao
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<normalizedToken originalValue="Dương">Dương</normalizedToken>
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, north of
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<normalizedToken originalValue="Đ">Đ</normalizedToken>
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ồng
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<normalizedToken originalValue="Phú">Phu</normalizedToken>
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, 58 km from Nho Quan towards Hanoi on the Hồ
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<normalizedToken originalValue="Chí">Chi</normalizedToken>
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Minh road (locality code: 2020/47);
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<geoCoordinate degrees="20" direction="north" minutes="42.59" orientation="latitude" precision="9" value="20.709833">20°42.59'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="39.30" orientation="longitude" precision="9" value="105.655">105°39.30'E</geoCoordinate>
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; 10 m a.s.l.; 16 Feb. 2020; A. Hunyadi leg.; HNHM 105321.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Vietnam • 2 figured shells; same data as for holotype; HNHM 105322 • 87 shells; same data as for holotype; coll. HA • 3 shells; same data as for holotype; coll. JJV.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<collectingCountry name="Vietnam">Vietnam</collectingCountry>
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• 21 j/b shells; same data as for holotype; coll. HA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A medium-sized, concave-conical
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<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species with a wide umbilicus, and three prominent apertural barriers (elevated parietal, pointed upper palatal, blunt, elongated columellar).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell small- to medium-sized for the genus, slightly higher than wide or slightly wider than high; off-white, concave-conical with regularly growing whorls except for the last one; body whorl widest from standard apertural view; protoconch consists of 1.5 whorls, with slight indication of spiral striation preceding the first teleoconch whorl; teleoconch with some fine, weak, irregular radial growth lines and much stronger, elevated, spiral striae of variable density (ca. 9-16 on body whorl in standard apertural view); whorls 4-4.5, rounded; aperture only slightly oblique to shell axis from lateral view; umbilicus wide; aperture suboval with straight parietal part; peristome expanded, not reflected; parietal callus not detached from penultimate whorl; parietal tooth elevated, mostly straight; palatal tooth situated in the middle of the palatal wall (probably homologous with the upper palatal tooth), small, denticle-like, situated close to basal lip; columellar tooth situated in the middle of columellar area, runs along peristome, lower and longer than palatal tooth.</paragraph>
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<caption doi="10.3897/zookeys.1147.93824.figure95" httpUri="https://binary.pensoft.net/fig/808904" pageId="0" pageNumber="1" start="Figure 95" startId="F95">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 95.</emphasis>
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<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tridentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tridentata">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tridentata</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi, sp. nov. (holotype, HNHM 105321). Apertural (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), apical (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), lateral (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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) and ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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) sides of the shell; sculpture on the protoconch (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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), ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) and frontal (
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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) surface of the body whorl.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1147.93824.figure96" httpUri="https://binary.pensoft.net/fig/808905" pageId="0" pageNumber="1" start="Figure 96" startId="F96">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 96.</emphasis>
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Variability of
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<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tridentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tridentata">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tridentata</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi, sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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holotype (specimen 3)
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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specimen 2
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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specimen 1.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="1">
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H = 0.76-0.99; D = 0.82-0.95, H/D*100 = 91.6-112.5 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 9), RUD = 32.9-34.1 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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See under
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<taxonomicName lsidName="A. apiaria" pageId="0" pageNumber="1" rank="species" species="apiaria">
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<emphasis italics="true" pageId="0" pageNumber="1">A. apiaria</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName lsidName="A. rara" pageId="0" pageNumber="1" rank="species" species="rara">
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<emphasis italics="true" pageId="0" pageNumber="1">A. rara</emphasis>
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</taxonomicName>
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sp. nov. Some other species that normally have two apertural barriers may have a third one, see differential diagnosis under
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<taxonomicName lsidName="A. rara" pageId="0" pageNumber="1" rank="species" species="rara">
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<emphasis italics="true" pageId="0" pageNumber="1">A. rara</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">This new species is named after its three apertural barriers.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is only known from the type locality of Cao
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<normalizedToken originalValue="Dương">Dương</normalizedToken>
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in
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<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
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<normalizedToken originalValue="Bình">Binh</normalizedToken>
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Province, Vietnam (Fig.
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure27" httpUri="https://binary.pensoft.net/fig/808836" pageId="0" pageNumber="1">27</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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||
</document> |