treatments-xml/data/03/EA/87/03EA8785FFC6FFD8FF465EC9AA52D917.xml

<document id="10049D2B486AB95B72A7DF0C4077DD18" ID-DOI="10.11646/zootaxa.4027.2.5" ID-GBIF-Dataset="561d9c55-132f-407a-b57e-79267e72f89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="243788" ID-ZooBank="B9337E83-2035-4D74-9ADC-0B6A6F0A99CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282102617" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2015" docId="03EA8785FFC6FFD8FF465EC9AA52D917" docLanguage="en" docName="zt04027p269.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perumicrus Jałoszyński, 2015, gen. nov." docType="treatment" docVersion="9" lastPageNumber="267" masterDocId="FFD3FFFDFFCDFFD6FFD15E5EA953DE67" masterDocTitle="Revision of Neotropical species of ant-like stone beetles misplaced in Stenichnus and Scydmoraphes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="269" masterPageNumber="253" pageNumber="264" updateTime="1698614310660" updateUser="plazi">
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<mods:title id="DAA60E40BA3FEF206E2F88ED4C83C17C">Revision of Neotropical species of ant-like stone beetles misplaced in Stenichnus and Scydmoraphes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="B00D4B13C37807CB94BC528AF6602423">2015</mods:date>
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<treatment id="03EA8785FFC6FFD8FF465EC9AA52D917" ID-DOI="http://doi.org/10.5281/zenodo.6121012" ID-GBIF-Taxon="119640836" ID-Zenodo-Dep="6121012" LSID="urn:lsid:plazi:treatment:03EA8785FFC6FFD8FF465EC9AA52D917" httpUri="http://treatment.plazi.org/id/03EA8785FFC6FFD8FF465EC9AA52D917" lastPageId="14" lastPageNumber="267" pageId="11" pageNumber="264">
<subSubSection id="C3596518FFC6FFDDFF465EC9A8CADED5" box="[151,409,151,178]" pageId="11" pageNumber="264" type="nomenclature">
<paragraph id="8BFC3693FFC6FFDDFF465EC9A8CADED5" blockId="11.[151,409,151,178]" box="[151,409,151,178]" pageId="11" pageNumber="264">
<heading id="D0B481FFFFC6FFDDFF465EC9A8CADED5" bold="true" box="[151,409,151,178]" fontSize="11" level="1" pageId="11" pageNumber="264" reason="1">
<emphasis id="B937EA81FFC6FFDDFF465EC9A8CADED5" bold="true" box="[151,409,151,178]" pageId="11" pageNumber="264">
<taxonomicName id="4C434D10FFC6FFDDFF465EC9A876DED6" ID-CoL="8XNSN" box="[151,293,151,177]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC6FFDDFF465EC9A876DED6" bold="true" box="[151,293,151,177]" italics="true" pageId="11" pageNumber="264">Perumicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457FAFFC6FFDDFEFD5EC6A8CADED5" box="[300,409,152,178]" pageId="11" pageNumber="264" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3596518FFC6FFDDFF465E81AACBDE93" box="[151,920,222,245]" pageId="11" pageNumber="264" type="materials_examined">
<paragraph id="8BFC3693FFC6FFDDFF465E81AACBDE93" blockId="11.[151,920,222,245]" box="[151,920,222,245]" pageId="11" pageNumber="264">
<typeStatus id="54F88831FFC6FFDDFF465E81A99ADE93" box="[151,201,223,244]" pageId="11" pageNumber="264">Type</typeStatus>
species: 
<taxonomicName id="4C434D10FFC6FFDDFEF75E80AB88DE93" authority="Franz, 1980a: 210" authorityName="Franz" authorityPageNumber="210" authorityYear="1980" box="[294,731,222,245]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="264" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC6FFDDFEF75E80AB4DDE92" box="[294,542,222,245]" italics="true" pageId="11" pageNumber="264">Scydmoraphes peruanus</emphasis>
<bibRefCitation id="EFD24B62FFC6FFDDFDF55E81ABFFDE93" author="Franz" box="[548,684,222,244]" pageId="11" pageNumber="264" refString="Franz, H. (1980 a) Monographie der sudamerikanischen Scydmaenidae mit Einschluss einiger mittelamerikanischer Arten (Coleoptera). Folia Entomologica Hungarica, 41, 39 - 264." type="journal article" year="1980" yearSuffix="a">Franz, 1980a</bibRefCitation>
: 210
</taxonomicName>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC6FFDDFF465F7FAD43DD6A" pageId="11" pageNumber="264" type="diagnosis">
<paragraph id="8BFC3693FFC6FFDDFF465F7FAD43DD6A" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
<emphasis id="B937EA81FFC6FFDDFF465F7FA842DF5D" bold="true" box="[151,273,289,314]" pageId="11" pageNumber="264">Diagnosis.</emphasis>
The unique apomorphy of this genus is the structure of prothoracic hypomeron and lateral portion of prosternum: the notosternal suture is marked only near anterior prosternal margin and near procoxal socket, its median part is externally not visible, posterior (adcoxal) part of notosternal suture anteriorly connects with additional suture extending posterolaterally and joining hypomeral ridge, thus demarcating inner part of hypomeron from prosternum, postcoxal area of inner part of hypomeron is fused with prosternum, and hypomeral ridge in front of procoxa runs laterally to notosternal suture. Additional diagnostic characters comprise: head transverse and with tempora much shorter than eyes and without bristles, occipital constriction distinctly narrower than vertex but not narrower than frons between eyes; antenna with club composed of antennomeres IX–XI; pronotum without lateral edges, with antebasal transverse groove and one pair of lateral impressions separated from groove; sides of pronotum lacking bristles; prosternum with basisternal part slightly shorter than coxal part; prosternal process absent; mesoventral intercoxal process long, carinate but moderately elevated, extending from anterior ridge of mesoventrite to posterior margins of mesocoxae; mesothorax lacking lateral foveae; metaventral intercoxal process very narrowly separating metacoxae, with two long spines; each elytron with one small but distinct asetose basal fovea; and symmetrical aedeagus with slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC6FFDBFF165D47A8DAD992" lastPageId="13" lastPageNumber="266" pageId="11" pageNumber="264" type="description">
<paragraph id="8BFC3693FFC6FFDDFF165D47AA5CDD32" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
<emphasis id="B937EA81FFC6FFDDFF165D47A80ADD55" bold="true" box="[199,345,793,818]" pageId="11" pageNumber="264">Description.</emphasis>
Body of male moderately strongly convex, slender, with moderately long appendages, BL below 
<quantity id="4CBB9B76FFC6FFDDFF465D60A988DD32" box="[151,219,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="264" unit="mm" value="1.0">1 mm</quantity>
; cuticle glossy, brown, moderately densely setose.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF165D3FAB26DA49" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
Head (
<figureCitation id="13782A16FFC6FFDDFEC25D3FA8C6DD1D" box="[275,405,865,890]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Figs 21–22</figureCitation>
) short and approximately subtrapezoidal, with very large eyes; occipital constriction (
<figureCitation id="13782A16FFC6FFDDFABE5D3FA9E5DDF9" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 21</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFF155DD8A9B9DDFA" box="[196,234,902,925]" italics="true" pageId="11" pageNumber="264">occ</emphasis>
) in the narrowest place slightly broader than frons; 'neck' region (
<figureCitation id="13782A16FFC6FFDDFC3E5DDBAD15DDF9" box="[1007,1094,901,926]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 21</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFB855DD8AD3EDDFA" box="[1108,1133,902,925]" italics="true" pageId="11" pageNumber="264">nr</emphasis>
) much broader than long, slightly narrower than vertex; tempora shorter than eyes and strongly convergent posteromesally; vertex (
<figureCitation id="13782A16FFC6FFDDFA925DF7ACC4DDA5" box="[1347,1431,937,962]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 21</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFF465D90A9F8DD82" box="[151,171,974,997]" italics="true" pageId="11" pageNumber="264">vt</emphasis>
) strongly transverse; frons (
<figureCitation id="13782A16FFC6FFDDFE2B5D93AB01DD81" box="[506,594,973,998]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 21</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFDB35D93AB26DD82" box="[610,629,973,997]" italics="true" pageId="11" pageNumber="264">fr</emphasis>
) posteriorly confluent with vertex, subtrapezoidal, anteriorly slightly projecting at middle; frontoclypeal groove absent; antennal insertions moderately broadly separated. Tempora and posterior margin of vertex lacking bristles.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF165A67AD59DB15" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
Labrum transverse with rounded anterior margin. Each mandible subtriangular, mesal margin not visible in the studied specimen. Each maxilla with short and broad cardo (
<figureCitation id="13782A16FFC6FFDDFC835A03AAFBDA11" box="[850,936,1117,1142]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFC675A03AA82DA12" box="[950,977,1117,1141]" italics="true" pageId="11" pageNumber="264">cd</emphasis>
), subtriangular basistipes (
<figureCitation id="13782A16FFC6FFDDFAD65A03AC0FDA11" box="[1287,1372,1117,1142]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFABA5A03ACDFDA12" box="[1387,1420,1117,1141]" italics="true" pageId="11" pageNumber="264">bst</emphasis>
), short mediostipes (
<figureCitation id="13782A16FFC6FFDDFEBC5ADFA892DAFD" box="[365,449,1153,1178]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE1F5ADCA8A6DAFE" box="[462,501,1154,1177]" italics="true" pageId="11" pageNumber="264">mst</emphasis>
) and elongate galea (
<figureCitation id="13782A16FFC6FFDDFD385ADFAA6DDAFD" box="[745,830,1153,1178]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFC9A5ADFAA3CDAFE" box="[843,879,1153,1177]" italics="true" pageId="11" pageNumber="264">gal</emphasis>
) and lacinia (
<figureCitation id="13782A16FFC6FFDDFBDA5ADFAD0CDAFD" box="[1035,1119,1153,1178]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFBBD5ADFADDDDAFE" box="[1132,1166,1153,1177]" italics="true" pageId="11" pageNumber="264">lac</emphasis>
), each with dense setae along distomesal margin, maxillary palp (
<figureCitation id="13782A16FFC6FFDDFDBF5AFBAB91DAD9" box="[622,706,1189,1214]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFD1F5AF8ABAEDADA" box="[718,765,1190,1213]" italics="true" pageId="11" pageNumber="264">mxp</emphasis>
) moderately long, palpomere I minute, slightly longer than broad, palpomere II strongly elongate, nearly pipe-shaped, palpomere III relatively stout, broadest between middle and distal third, palpomere IV very slender, nearly rod-like. Labium with broad submentum (
<figureCitation id="13782A16FFC6FFDDFB035AB3AC7ADB61" box="[1234,1321,1261,1286]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFAE65AB0AC37DB62" box="[1335,1380,1262,1285]" italics="true" pageId="11" pageNumber="264">smn</emphasis>
) not demarcated laterally by sutures; subtrapezoidal mentum (
<figureCitation id="13782A16FFC6FFDDFCC75B4FAA3ADB4D" box="[790,873,1297,1322]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFCA55B4CAAC5DB4E" box="[884,918,1298,1321]" italics="true" pageId="11" pageNumber="264">mn</emphasis>
) and short prementum, ligula indistinct; labial palps (
<figureCitation id="13782A16FFC6FFDDFF355B6BA868DB29" box="[228,315,1333,1358]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE9B5B6BA833DB2A" box="[330,352,1333,1357]" italics="true" pageId="11" pageNumber="264">lp</emphasis>
) short, moderately broadly separated at bases, with palpomere II relatively long. Hypostomal ridges (
<figureCitation id="13782A16FFC6FFDDFF3A5B07A86DDB15" box="[235,318,1369,1394]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE9B5B07A836DB16" box="[330,357,1369,1393]" italics="true" pageId="11" pageNumber="264">hy</emphasis>
) distinct but posteriorly not reaching posterior tentorial pits.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF165B23ACC2DBDD" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
Gular plate (
<figureCitation id="13782A16FFC6FFDDFE8B5B23A8E3DBF1" box="[346,432,1405,1430]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE6E5B20A888DBF2" box="[447,475,1406,1429]" italics="true" pageId="11" pageNumber="264">gp</emphasis>
) broadly subtrapezoidal with rounded sides, gular sutures (
<figureCitation id="13782A16FFC6FFDDFB5C5B23ADB0DBF1" box="[1165,1251,1405,1430]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFB235B20AC58DBF2" box="[1266,1291,1406,1429]" italics="true" pageId="11" pageNumber="264">gs</emphasis>
) superficial. Posterior tentorial pits (
<figureCitation id="13782A16FFC6FFDDFE4E5BFFA8A1DBDD" box="[415,498,1441,1466]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 22</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE2F5BFCAB71DBDE" box="[510,546,1442,1465]" italics="true" pageId="11" pageNumber="264">ptp</emphasis>
) exposed, arcuate, located in transverse impression demarcating 'neck' region.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF165B98AACFDBBA" blockId="11.[151,1437,289,2006]" box="[199,924,1477,1502]" pageId="11" pageNumber="264">
Antennae (
<figureCitation id="13782A16FFC6FFDDFE905B9BA8C0DBB9" box="[321,403,1477,1502]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 21</figureCitation>
) with club composed of antennomeres IX–XI.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF165BB4AA04D8F5" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">Pronotum in dorsal view subquadrate with rounded sides, anterior margin nearly straight, anterior corners distinct, obtuse-angled, sides rounded in anterior half and distinctly narrowing posteriorly, posterior corners distinct, slightly obtuse-angled, posterior margin shallowly bisinuate. Pronotum without lateral edges, with distinct antebasal transverse groove and one pair of longitudinally elongate lateral impressions close to lateral margins and not connected with groove. Sides of pronotum lacking bristles.</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF1658C0AA50D9EA" blockId="11.[151,1437,289,2006]" pageId="11" pageNumber="264">
Prosternum (
<figureCitation id="13782A16FFC6FFDDFE8A58C3A8E2D8D1" box="[347,433,1693,1718]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 23</figureCitation>
) with basisternal part (
<figureCitation id="13782A16FFC6FFDDFD1558C3AA48D8D1" box="[708,795,1693,1718]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 23</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFCFA58C3AA1FD8D2" box="[811,844,1693,1717]" italics="true" pageId="11" pageNumber="264">bst</emphasis>
) indistinctly demarcated from coxal part and only slightly shorter than the latter; intercoxal area slightly elevated but diffused, not forming prosternal process; procoxal sockets (
<figureCitation id="13782A16FFC6FFDDFEB958BBA8EDD899" box="[360,446,1765,1790]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 23</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE1C58B8A8A1D89A" box="[461,498,1766,1789]" italics="true" pageId="11" pageNumber="264">pcs</emphasis>
) broadly closed. Notosternal sutures (
<figureCitation id="13782A16FFC6FFDDFC7C58BBAD57D899" box="[941,1028,1765,1790]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 23</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFBC358B8AD65D89A" box="[1042,1078,1766,1789]" italics="true" pageId="11" pageNumber="264">nss</emphasis>
) incomplete, externally visible only at anterior prosternal margin and in front of procoxal socket while median part is obliterated, adcoxal part of notosternal suture connected to oblique suture extending posterolaterally and connecting to hypomeral ridge. Hypomeral ridge (
<figureCitation id="13782A16FFC6FFDDFEB4590FA8EAD90D" box="[357,441,1873,1898]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 23</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFE14590FA8B9D90E" box="[453,490,1873,1897]" italics="true" pageId="11" pageNumber="264">hyr</emphasis>
) nearly complete, its precoxal part running laterally to notosternal suture. Inner part of hypomeron fused with postcoxal part of prosternum.
</paragraph>
<paragraph id="8BFC3693FFC6FFDDFF1659C7ABB4D9D5" blockId="11.[151,1437,289,2006]" box="[199,743,1945,1970]" pageId="11" pageNumber="264">Mesoscutellum in studied specimens not visible.</paragraph>
<paragraph id="8BFC3693FFC6FFDAFF1659E3AA7FDF27" blockId="11.[151,1437,289,2006]" lastBlockId="12.[151,1437,151,320]" lastPageId="12" lastPageNumber="265" pageId="11" pageNumber="264">
Mesoventrite (
<figureCitation id="13782A16FFC6FFDDFEBE59E3A896D9B1" box="[367,453,1981,2006]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 24</figureCitation>
) with narrow anterior ridge (
<figureCitation id="13782A16FFC6FFDDFCF259E3AA28D9B1" box="[803,891,1981,2006]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="11" pageNumber="264">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC6FFDDFC5D59E0AAF6D9B2" box="[908,933,1982,2005]" italics="true" pageId="11" pageNumber="264">ar</emphasis>
) and long, carinate mesoventral intercoxal process (
<figureCitation id="13782A16FFC1FFDAFF2E5EC9A804DED7" box="[255,343,151,176]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="12" pageNumber="265">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC1FFDAFEB65EC7A8F3DED7" box="[359,416,153,176]" italics="true" pageId="12" pageNumber="265">msvp</emphasis>
) which is moderately elevated (in lateral view not projecting ventrally beyond ventral margins of mesocoxae) and extending from anterior ridge to posterior margins of mesocoxae. Mesothorax lacking lateral foveae. Mesocoxal projections (
<figureCitation id="13782A16FFC1FFDAFD9B5E81ABCEDE9F" box="[586,669,223,248]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="12" pageNumber="265">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC1FFDAFD785EBFAB8BDE9F" box="[681,728,225,248]" italics="true" pageId="12" pageNumber="265">mcp</emphasis>
) prominent, with mesocoxal sockets (
<figureCitation id="13782A16FFC1FFDAFB535E81AD85DE9F" box="[1154,1238,223,248]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="12" pageNumber="265">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC1FFDAFB305EBFAC44DE9F" box="[1249,1303,225,248]" italics="true" pageId="12" pageNumber="265">mscs</emphasis>
) located on their mesoventral surface and exposed in ventral view. Mesoventrite with large asetose procoxal rests (
<figureCitation id="13782A16FFC1FFDAFA905F5AACC4DF7B" box="[1345,1431,260,285]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="12" pageNumber="265">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC1FFDAFF465F77A9EFDF27" box="[151,188,297,320]" italics="true" pageId="12" pageNumber="265">pcr</emphasis>
); area behind procoxal rests asetose and not impressed.
</paragraph>
<caption id="DF3C661BFFC1FFDAFF465884AAB8D9A0" httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="12" pageNumber="265" targetBox="[151,1434,375,1729]" targetPageId="12">
<paragraph id="8BFC3693FFC1FFDAFF465884AAB8D9A0" blockId="12.[151,1436,1754,1991]" pageId="12" pageNumber="265">
<emphasis id="B937EA81FFC1FFDAFF465884A806D888" bold="true" box="[151,341,1754,1777]" pageId="12" pageNumber="265">FIGURES 21–24.</emphasis>
Morphological structures of 
<taxonomicName id="4C434D10FFC1FFDAFD505885AAE2D897" authority="Franz" authorityName="Franz" box="[641,945,1754,1776]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="265" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC1FFDAFD505885AA0FD897" box="[641,860,1755,1776]" italics="true" pageId="12" pageNumber="265">Perumicrus peruanus</emphasis>
(Franz)
</taxonomicName>
, paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, 'neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v3, metaventrite; vlf, ventrolateral fovea; vt, vertex.
</paragraph>
</caption>
<caption id="DF3C661BFFC0FFDBFF465D01A8F1DDF3" httpUri="https://zenodo.org/record/243795/files/figure.png" pageId="13" pageNumber="266" targetBox="[186,1409,192,832]" targetPageId="13">
<paragraph id="8BFC3693FFC0FFDBFF465D01A8F1DDF3" blockId="13.[151,1435,863,916]" pageId="13" pageNumber="266">
<emphasis id="B937EA81FFC0FFDBFF465D01A806DD13" bold="true" box="[151,341,863,886]" pageId="13" pageNumber="266">FIGURES 25–27.</emphasis>
<taxonomicName id="4C434D10FFC0FFDBFE8A5D01ABFADD12" authority="Franz" authorityName="Franz" box="[347,681,863,885]" class="Insecta" family="Staphylinidae" genus="Meridaphes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="venezolanus">
<emphasis id="B937EA81FFC0FFDBFE8A5D01AB0BDD12" box="[347,600,863,885]" italics="true" pageId="13" pageNumber="266">Meridaphes venezolanus</emphasis>
(Franz)
</taxonomicName>
(25, 26) and 
<taxonomicName id="4C434D10FFC0FFDBFCE55D3FAD32DD12" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" box="[820,1121,863,886]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC0FFDBFCE55D3FAD43DD11" box="[820,1040,865,886]" italics="true" pageId="13" pageNumber="266">Perumicrus peruanus</emphasis>
(Franz)
</taxonomicName>
(27). Aedeagus in ventral (25, 27) and lateral (26) views.
</paragraph>
</caption>
<caption id="DF3C661BFFC0FFDBFF46584DAB76D82E" httpUri="https://zenodo.org/record/243796/files/figure.png" pageId="13" pageNumber="266" targetBox="[164,1418,980,1525]" targetPageId="13">
<paragraph id="8BFC3693FFC0FFDBFF46584DAB76D82E" blockId="13.[151,1436,1555,1609]" pageId="13" pageNumber="266">
<emphasis id="B937EA81FFC0FFDBFF46584DA807D84E" bold="true" box="[151,340,1555,1578]" pageId="13" pageNumber="266">FIGURES 28–30.</emphasis>
Original set of labels of holotypes. 
<taxonomicName id="4C434D10FFC0FFDBFD11584AAA88D84D" authority="Franz" authorityName="Franz" box="[704,987,1556,1578]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="andicola">
<emphasis id="B937EA81FFC0FFDBFD11584AAA78D84D" box="[704,811,1556,1578]" italics="true" pageId="13" pageNumber="266">Sciacharis</emphasis>
<emphasis id="B937EA81FFC0FFDBFCE0584AAAD9D84D" box="[817,906,1556,1578]" italics="true" pageId="13" pageNumber="266">andicola</emphasis>
(Franz)
</taxonomicName>
(28), 
<taxonomicName id="4C434D10FFC0FFDBFBC8584AAC36D84D" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" box="[1049,1381,1556,1578]" class="Insecta" family="Staphylinidae" genus="Meridaphes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="venezolanus">
<emphasis id="B937EA81FFC0FFDBFBC8584AAC47D84D" box="[1049,1300,1556,1578]" italics="true" pageId="13" pageNumber="266">Meridaphes venezolanus</emphasis>
(Franz)
</taxonomicName>
(29), and 
<taxonomicName id="4C434D10FFC0FFDBFF13586AA8BED82E" authority="(Franz)" authorityName="Franz" baseAuthorityName="Franz" box="[194,493,1587,1609]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="266" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC0FFDBFF13586AA8CFD82E" box="[194,412,1588,1609]" italics="true" pageId="13" pageNumber="266">Perumicrus peruanus</emphasis>
(Franz)
</taxonomicName>
(30).
</paragraph>
</caption>
<paragraph id="8BFC3693FFC0FFDBFF16582BADB5D89D" blockId="13.[151,1437,1653,2038]" pageId="13" pageNumber="266">
Metaventrite (
<figureCitation id="13782A16FFC0FFDBFEBF582BA899D8E9" box="[366,458,1653,1678]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="13" pageNumber="266">Fig. 24</figureCitation>
; 
<date id="FFFD1053FFC0FFDBFE0F582BA8AAD8EA" box="[478,505,1653,1677]" pageId="13" pageNumber="266">
<emphasis id="B937EA81FFC0FFDBFE0F582BA8AAD8EA" box="[478,505,1653,1677]" italics="true" pageId="13" pageNumber="266">v3</emphasis>
</date>
) subtrapezoidal, broadening posteriorly and with rounded sides; metaventral intercoxal process (
<figureCitation id="13782A16FFC0FFDBFEA558C7A89AD8D5" box="[372,457,1689,1714]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="13" pageNumber="266">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC0FFDBFE0758C4AB5FD8D6" box="[470,524,1690,1713]" italics="true" pageId="13" pageNumber="266">mtvp</emphasis>
) very narrowly separating metacoxae and with two long spines. Metanepisterna and metepimera narrow. Lateral margins of metacoxae distant from lateral margins of metaventrite. Metendosternite (metafurca) with very short stem and divergent lateral furcal arms (
<figureCitation id="13782A16FFC0FFDBFB9458BFADCBD89D" box="[1093,1176,1761,1786]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1754,1777]" captionTargetBox="[151,1434,375,1729]" captionTargetId="figure@12.[151,1436,373,1733]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Morphological structures of Perumicrus peruanus (Franz), paratype male. Head in dorsal (21) and ventral (22) views; prothorax in ventral view (23); pterothorax in ventral view (24). Abbreviations: ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fr, frons; gal, galea; gp, gular plate; gs, gular suture; h, hypomeron; hy, hypostomal ridge; hyr, hypomeral ridge; lac, lacinia; lb, labrum; lp, labial palp; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, ' neck' region; occ, occipital constriction; pf, profurca; pre, prepectus; pcc, procoxal cavity; pcs, procoxal socket; ptp, posterior tentorial pits; smn, submentum; v 3, metaventrite; vlf, ventrolateral fovea; vt, vertex." httpUri="https://zenodo.org/record/243794/files/figure.png" pageId="13" pageNumber="266">Fig. 24</figureCitation>
; 
<emphasis id="B937EA81FFC0FFDBFB7558BFAD85D89E" box="[1188,1238,1761,1785]" italics="true" pageId="13" pageNumber="266">lmfa</emphasis>
).
</paragraph>
<paragraph id="8BFC3693FFC0FFDBFF16595BAACAD979" blockId="13.[151,1437,1653,2038]" box="[199,921,1797,1822]" pageId="13" pageNumber="266">Elytra oval, each with one small but distinct asetose basal fovea.</paragraph>
<paragraph id="8BFC3693FFC0FFDBFF165974AD44D901" blockId="13.[151,1437,1653,2038]" pageId="13" pageNumber="266">Legs moderately long and slender; pro- and mesocoxae oval, metacoxae transverse; all femora weakly and gradually clavate, all tibiae slender and nearly straight; tarsi moderately slender.</paragraph>
<paragraph id="8BFC3693FFC0FFDBFF16592FAD10D9ED" blockId="13.[151,1437,1653,2038]" box="[199,1091,1905,1930]" pageId="13" pageNumber="266">Abdominal sternites unmodified, suture between two terminal sternites distinct.</paragraph>
<paragraph id="8BFC3693FFC0FFDBFF1659CBA8DAD992" blockId="13.[151,1437,1653,2038]" pageId="13" pageNumber="266">
Aedeagus (
<figureCitation id="13782A16FFC0FFDBFE9759CBA8CDD9C9" box="[326,414,1941,1966]" captionStart="FIGURES 9 – 11" captionStartId="6.[151,264,1116,1139]" captionTargetBox="[173,1415,210,1082]" captionTargetId="figure@6.[151,1436,193,1095]" captionTargetPageId="6" captionText="FIGURES 9 – 11. Meridaphes venezolanus (Franz) (9, 10) and Perumicrus peruanus (Franz) (11). Dorsal habitus of male holotype (9) and aedeagus in ventral view (10, 11)." httpUri="https://zenodo.org/record/243791/files/figure.png" pageId="13" pageNumber="266">Figs 11</figureCitation>
, 
<figureCitation id="13782A16FFC0FFDBFE7D59C8A89AD9C9" box="[428,457,1942,1966]" captionStart="FIGURES 25 – 27" captionStartId="13.[151,264,863,886]" captionTargetBox="[186,1409,192,832]" captionTargetId="figure@13.[151,1436,157,842]" captionTargetPageId="13" captionText="FIGURES 25 – 27. Meridaphes venezolanus (Franz) (25, 26) and Perumicrus peruanus (Franz) (27). Aedeagus in ventral (25, 27) and lateral (26) views." httpUri="https://zenodo.org/record/243795/files/figure.png" pageId="13" pageNumber="266">27</figureCitation>
) drop-shaped, elongate, with both median lobe and endophallus symmetrical; median lobe with subtriangular apical part; endophallus composed of paired elongate sclerites; parameres present, slender and with apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC3FFD8FF165EC9ADDCDEC8" box="[199,1167,151,176]" pageId="14" pageNumber="267" type="distribution">
<paragraph id="8BFC3693FFC3FFD8FF165EC9ADDCDEC8" blockId="14.[151,1437,151,1904]" box="[199,1167,151,176]" pageId="14" pageNumber="267">
<emphasis id="B937EA81FFC3FFD8FF165EC9AB7FDED7" bold="true" box="[199,556,151,176]" pageId="14" pageNumber="267">Distribution and composition.</emphasis>
One species of 
<taxonomicName id="4C434D10FFC3FFD8FD0F5EC6AA32DEC8" box="[734,865,152,175]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FD0F5EC6AA32DEC8" box="[734,865,152,175]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
is known to occur in 
<collectingCountry id="F3547603FFC3FFD8FB855EC6ADD8DEC8" box="[1108,1163,152,175]" name="Peru" pageId="14" pageNumber="267">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC3FFD8FF165EE2AB14DE9F" pageId="14" pageNumber="267" type="etymology">
<paragraph id="8BFC3693FFC3FFD8FF165EE2AB14DE9F" blockId="14.[151,1437,151,1904]" pageId="14" pageNumber="267">
<emphasis id="B937EA81FFC3FFD8FF165EE2A815DEB2" bold="true" box="[199,326,188,213]" pageId="14" pageNumber="267">Etymology</emphasis>
. The name 
<taxonomicName id="4C434D10FFC3FFD8FE1C5EE3AB03DEB3" box="[461,592,189,212]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FE1C5EE3AB03DEB3" box="[461,592,189,212]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
is composed of the country name and - 
<emphasis id="B937EA81FFC3FFD8FBC35EE3AD0DDEB3" box="[1042,1118,189,212]" italics="true" pageId="14" pageNumber="267">micrus</emphasis>
, referring to the small body of the 
<typeStatus id="54F88831FFC3FFD8FF0F5EBEA85DDE9F" box="[222,270,224,248]" pageId="14" pageNumber="267">type</typeStatus>
species. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC3FFD8FF165F5AAAABDCCF" pageId="14" pageNumber="267" type="discussion">
<paragraph id="8BFC3693FFC3FFD8FF165F5AAAABDCCF" blockId="14.[151,1437,151,1904]" pageId="14" pageNumber="267">
<emphasis id="B937EA81FFC3FFD8FF165F5AA868DF7A" bold="true" box="[199,315,260,285]" pageId="14" pageNumber="267">Remarks.</emphasis>
The following characters clearly indicate that 
<taxonomicName id="4C434D10FFC3FFD8FCBC5F5AADD5DF7B" box="[877,1158,260,284]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC3FFD8FCBC5F5AADD5DF7B" box="[877,1158,260,284]" italics="true" pageId="14" pageNumber="267">Scydmoraphes peruanus</emphasis>
</taxonomicName>
is not congeneric with Northern Hemisphere 
<taxonomicName id="4C434D10FFC3FFD8FE4D5F76AB6CDF27" box="[412,575,296,320]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE4D5F76AB6CDF27" box="[412,575,296,320]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
: submentum lacking lateral sutures (present in 
<taxonomicName id="4C434D10FFC3FFD8FBA95F76AC48DF27" box="[1144,1307,296,320]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FBA95F76AC48DF27" box="[1144,1307,296,320]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); posterior tentorial pits exposed (hidden in deep transverse groove demarcating 'neck region' in 
<taxonomicName id="4C434D10FFC3FFD8FB545F12AC7BDF03" box="[1157,1320,332,356]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB545F12AC7BDF03" box="[1157,1320,332,356]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); anterior margins of eyes nearly adjacent to mandibular bases (distant in 
<taxonomicName id="4C434D10FFC3FFD8FC4A5F2EAD6DDFEF" box="[923,1086,368,392]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC4A5F2EAD6DDFEF" box="[923,1086,368,392]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); antennae with distinct club composed of three terminal antennomeres (gradually thickened in 
<taxonomicName id="4C434D10FFC3FFD8FC405FCAAD67DFCB" box="[913,1076,404,428]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC405FCAAD67DFCB" box="[913,1076,404,428]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); maxillary palpomere IV very slender, nearly rod-like (subconical with broad base in 
<taxonomicName id="4C434D10FFC3FFD8FCD15FE6AAF0DFB7" box="[768,931,440,464]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FCD15FE6AAF0DFB7" box="[768,931,440,464]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); sides of pronotum without bristles (present in 
<taxonomicName id="4C434D10FFC3FFD8FF645F82A80BDF93" box="[181,344,476,500]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FF645F82A80BDF93" box="[181,344,476,500]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); prosternum in front of and behind procoxal sockets laterally broadly fused with inner parts of hypomera (not fused in 
<taxonomicName id="4C434D10FFC3FFD8FE645C5EAB0BDC7F" box="[437,600,512,536]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE645C5EAB0BDC7F" box="[437,600,512,536]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); notosternal sutures largely internalized and visible only near anterior margin of prosternum and near procoxal socket (complete in 
<taxonomicName id="4C434D10FFC3FFD8FCA45C7AAD4BDC5B" box="[885,1048,548,572]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FCA45C7AAD4BDC5B" box="[885,1048,548,572]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); mesoventral intercoxal process extending from anterior ridge of mesoventrite to posterior margins of mesocoxae (posteriorly reaching only middle of mesocoxae in 
<taxonomicName id="4C434D10FFC3FFD8FE895C32A8A8DCE3" box="[344,507,620,644]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE895C32A8A8DCE3" box="[344,507,620,644]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); anterior metaventral process absent (present in 
<taxonomicName id="4C434D10FFC3FFD8FBFA5C32AD9DDCE3" box="[1067,1230,620,644]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FBFA5C32AD9DDCE3" box="[1067,1230,620,644]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
); and metaventral intercoxal process with pair of long spines (lacking spines in 
<taxonomicName id="4C434D10FFC3FFD8FC955CCEAAB4DCCF" box="[836,999,656,680]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC955CCEAAB4DCCF" box="[836,999,656,680]" italics="true" pageId="14" pageNumber="267">Scydmoraphes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3596518FFC3FFD8FF165CEAAA52D917" pageId="14" pageNumber="267" type="reference_group">
<paragraph id="8BFC3693FFC3FFD8FF165CEAAB41DAC7" blockId="14.[151,1437,151,1904]" pageId="14" pageNumber="267">
<taxonomicName id="4C434D10FFC3FFD8FF165CEAA88ADCAB" ID-CoL="4W537" box="[199,473,692,716]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC3FFD8FF165CEAA88ADCAB" box="[199,473,692,716]" italics="true" pageId="14" pageNumber="267">Scydmoraphes peruanus</emphasis>
</taxonomicName>
shows characters resembling similarly minute species of genera 
<taxonomicName id="4C434D10FFC3FFD8FB7F5CEAA98DDC97" ID-CoL="8XMVT" authority="Franz, 1980" authorityName="Franz" authorityYear="1980" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB7F5CEAAC1DDCAB" box="[1198,1358,692,716]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
<bibRefCitation id="EFD24B62FFC3FFD8FA845CEBA98DDC97" author="Franz" pageId="14" pageNumber="267" refString="Franz, H. (1980 a) Monographie der sudamerikanischen Scydmaenidae mit Einschluss einiger mittelamerikanischer Arten (Coleoptera). Folia Entomologica Hungarica, 41, 39 - 264." type="journal article" year="1980" yearSuffix="a">Franz, 1980a</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B937EA81FFC3FFD8FF385C86A8C4DC97" box="[233,407,728,752]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
<bibRefCitation id="EFD24B62FFC3FFD8FE4C5C89AB21DC97" author="Jaloszynski" box="[413,626,727,752]" pageId="14" pageNumber="267" refString="Jaloszynski, P. (2013 b) Revision of Neotropical genera Microraphes Franz, Heteroscydmus Franz and Mimoscydmus Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3722 (2), 245 - 266. http: // dx. doi. org / 10.11646 / zootaxa. 3722.2.7" type="journal article" year="2013" yearSuffix="b">Jałoszyński, 2013b</bibRefCitation>
and 
<taxonomicName id="4C434D10FFC3FFD8FD795C86AA80DC97" ID-CoL="8XMT9" authority="Franz, 1980" authorityName="Franz" authorityYear="1980" box="[680,979,728,752]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FD795C86AA6BDC97" box="[680,824,728,752]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
<bibRefCitation id="EFD24B62FFC3FFD8FCEF5C86AA80DC97" author="Franz" box="[830,979,728,752]" pageId="14" pageNumber="267" refString="Franz, H. (1980 a) Monographie der sudamerikanischen Scydmaenidae mit Einschluss einiger mittelamerikanischer Arten (Coleoptera). Folia Entomologica Hungarica, 41, 39 - 264." type="journal article" year="1980" yearSuffix="a">Franz, 1980a</bibRefCitation>
</taxonomicName>
. However, the structure of prosternum in 
<taxonomicName id="4C434D10FFC3FFD8FF465CA2A8FCDD73" ID-CoL="4W537" box="[151,431,764,788]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B937EA81FFC3FFD8FF465CA2A8FCDD73" box="[151,431,764,788]" italics="true" pageId="14" pageNumber="267">Scydmoraphes peruanus</emphasis>
</taxonomicName>
is unique and represents an autapomorphy to define a new genus, 
<taxonomicName id="4C434D10FFC3FFD8FB335CA3AC36DD73" ID-CoL="8XNSN" box="[1250,1381,765,788]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FB335CA3AC36DD73" box="[1250,1381,765,788]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
: the notosternal suture is marked only as a short notch in anterior margin of sternum and posteriorly as a short suture running anterad from procoxal socket, the median part is internalized and externally not visible, posterior part of notosternal suture connects to an additional oblique suture extending laterocaudad to join hypomeral ridge, and these two sutures demarcate the inner (adcoxal) part of hypomeron from prosternum, while posteriorly (behind coxal socket) this part of hypomeron is fused with postcoxal part of prosternum. Moreover, the hypomeral ridge runs laterally to notosternal suture and not, as in most 
<taxonomicName id="4C434D10FFC3FFD8FCDF5D8AAAE4DD8A" ID-CoL="8VR7Z" box="[782,951,980,1005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
, connects with the suture or overlaps it in precoxal region. This relatively complex modification is not known in any other genus, although various extent of fusion of prosternum with hypomeron can be seen in some genera, especially those characterized by very small body size. The probably plesiomorphic character state in 
<taxonomicName id="4C434D10FFC3FFD8FCCB5A61AA93DA3F" ID-CoL="8VR7Z" box="[794,960,1087,1112]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
is a complete notosternal suture and inner part of hypomeron entirely demarcated from prosternum, with hypomeral ridge anteriorly confluent with, joining or overlapping notosternal suture.
</paragraph>
<paragraph id="8BFC3693FFC3FFD8FF165AF2AA52D917" blockId="14.[151,1437,151,1904]" pageId="14" pageNumber="267">
As mentioned in the previous paragraph, 
<taxonomicName id="4C434D10FFC3FFD8FD635AF3AA66DAA3" ID-CoL="8XNSN" box="[690,821,1197,1220]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FD635AF3AA66DAA3" box="[690,821,1197,1220]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
shows characters similar to those of 
<taxonomicName id="4C434D10FFC3FFD8FB245AF2A816DA8F" ID-CoL="8XMVT" authority="Amimoscydmus" authorityName="Amimoscydmus" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB245AF2ACC6DAA3" box="[1269,1429,1196,1220]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
, 
<emphasis id="B937EA81FFC3FFD8FF465A8EA816DA8F" box="[151,325,1232,1256]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C434D10FFC3FFD8FE535A8EAB41DA8F" ID-CoL="8XMT9" box="[386,530,1232,1256]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE535A8EAB41DA8F" box="[386,530,1232,1256]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
. Additionally to already listed characters, it differs from 
<taxonomicName id="4C434D10FFC3FFD8FB7E5A8EAC1CDA8F" ID-CoL="8XMVT" box="[1199,1359,1232,1256]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB7E5A8EAC1CDA8F" box="[1199,1359,1232,1256]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
</taxonomicName>
in the pronotum without lateral bristles (bristles present in 
<taxonomicName id="4C434D10FFC3FFD8FCCA5AAAAAE8DB6B" ID-CoL="8XMVT" box="[795,955,1268,1292]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FCCA5AAAAAE8DB6B" box="[795,955,1268,1292]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
</taxonomicName>
), presence of lateral pronotal antebasal impressions (lacking in 
<taxonomicName id="4C434D10FFC3FFD8FE785B46AB1ADB57" ID-CoL="8XMVT" box="[425,585,1304,1328]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE785B46AB1ADB57" box="[425,585,1304,1328]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
</taxonomicName>
), prosternal intercoxal area lacking prosternal process (carinate process in 
<taxonomicName id="4C434D10FFC3FFD8FF465B62A864DB33" ID-CoL="8XMVT" box="[151,311,1340,1364]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FF465B62A864DB33" box="[151,311,1340,1364]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
</taxonomicName>
), and lack of ventrolateral mesothoracic foveae (present in 
<taxonomicName id="4C434D10FFC3FFD8FC055B62AD27DB33" ID-CoL="8XMVT" box="[980,1140,1340,1364]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC055B62AD27DB33" box="[980,1140,1340,1364]" italics="true" pageId="14" pageNumber="267">Mimoscydmus</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C434D10FFC3FFD8FB5D5B63AC5CDB33" ID-CoL="8XNSN" box="[1164,1295,1341,1364]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FB5D5B63AC5CDB33" box="[1164,1295,1341,1364]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
differs from 
<emphasis id="B937EA81FFC3FFD8FF465B3EA816DB1F" box="[151,325,1376,1400]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
Jałoszyński in hypostomal ridges posteriorly reaching nearly to tentorial pits (ridges very short, visible only just behind cardines in 
<emphasis id="B937EA81FFC3FFD8FDF65BDAAB86DBFB" box="[551,725,1412,1436]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
), presence of lateral pronotal antebasal impressions (lacking in 
<emphasis id="B937EA81FFC3FFD8FF465BF6A816DBA7" box="[151,325,1448,1472]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
), prosternal intercoxal region lacking prosternal process (narrow carinate process present in 
<emphasis id="B937EA81FFC3FFD8FF465B92A816DB83" box="[151,325,1484,1508]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
), inner (adcoxal) part of hypomeron in front of procoxal socket demarcated by prosternum by rudiment of notosternal suture connected anteriorly to oblique suture that posteriorly connects with hypomeral ridge (inner part of hypomeron in front of procoxal socket fused with prosternum in 
<emphasis id="B937EA81FFC3FFD8FBBC584AAC48D84B" box="[1133,1307,1556,1580]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
); and each elytron with one distinct basal fovea (two rudimentary foveae in 
<emphasis id="B937EA81FFC3FFD8FC635866AD33D837" box="[946,1120,1592,1616]" italics="true" pageId="14" pageNumber="267">Amimoscydmus</emphasis>
). 
<taxonomicName id="4C434D10FFC3FFD8FBAF5866AC52D828" ID-CoL="8XNSN" box="[1150,1281,1592,1615]" class="Insecta" family="Staphylinidae" genus="Perumicrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EA81FFC3FFD8FBAF5866AC52D828" box="[1150,1281,1592,1615]" italics="true" pageId="14" pageNumber="267">Perumicrus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C434D10FFC3FFD8FF465802A83CD813" ID-CoL="8XMT9" authority="Franz" authorityName="Franz" box="[151,367,1628,1652]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FF465802A874D813" box="[151,295,1628,1652]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
Franz
</taxonomicName>
in hypostomal ridges posteriorly reaching nearly tentorial pits (ridges very short, visible only just behind cardines in 
<taxonomicName id="4C434D10FFC3FFD8FE7858DEAB6BD8FF" ID-CoL="8XMT9" box="[425,568,1664,1688]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FE7858DEAB6BD8FF" box="[425,568,1664,1688]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
), sides of pronotum without bristles (dense thick bristles in 
<taxonomicName id="4C434D10FFC3FFD8FB2D58DEACDFD8FF" ID-CoL="8XMT9" box="[1276,1420,1664,1688]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB2D58DEACDFD8FF" box="[1276,1420,1664,1688]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
), presence of lateral pronotal antebasal impressions (lacking in 
<taxonomicName id="4C434D10FFC3FFD8FC8758FAAAB5D8DB" ID-CoL="8XMT9" box="[854,998,1700,1724]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC8758FAAAB5D8DB" box="[854,998,1700,1724]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
), prosternum much broader than long, with basisternal part indistinctly demarcated from coxal part (prosternum subquadrate in 
<taxonomicName id="4C434D10FFC3FFD8FB045896AC36D887" ID-CoL="8XMT9" box="[1237,1381,1736,1760]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FB045896AC36D887" box="[1237,1381,1736,1760]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
and basisternal part demarcated by carina), inner (adcoxal) part of hypomeron in precoxal region demarcated, and in postcoxal region fused with prosternum (in 
<taxonomicName id="4C434D10FFC3FFD8FD5A594EAA48D94F" ID-CoL="8XMT9" box="[651,795,1808,1832]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FD5A594EAA48D94F" box="[651,795,1808,1832]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
fused in precoxal region while demarcated in postcoxal region), prosternal intercoxal process missing (short but distinct in 
<taxonomicName id="4C434D10FFC3FFD8FC68596AAD1AD92B" ID-CoL="8XMT9" box="[953,1097,1844,1868]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FC68596AAD1AD92B" box="[953,1097,1844,1868]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
), and metaventral intercoxal process with two long spines (lacking in 
<taxonomicName id="4C434D10FFC3FFD8FDB05906ABA3D917" ID-CoL="8XMT9" box="[609,752,1880,1904]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA81FFC3FFD8FDB05906ABA3D917" box="[609,752,1880,1904]" italics="true" pageId="14" pageNumber="267">Microraphes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>