treatments-xml/data/03/D5/01/03D5014CFFF2FFF4FF1ECF9B3A61665A.xml

<document id="1383980293066340FC28CFAB984F40B8" ID-DOI="10.5281/zenodo.6391953" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391953" ID-ZooBank="B6862B03-E882-437A-B9E0-3B5FB1443602" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648504685522" checkinUser="felipe" docAuthor="Skelley, Paul E. &amp; Keller, Oliver" docDate="2022" docId="03D5014CFFF2FFF4FF1ECF9B3A61665A" docLanguage="en" docName="InsectaMundi.2022.0916.1-11.pdf" docOrigin="Insecta Mundi 2022 (916)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Haroldiellus woodruffi Skelley and Keller 2022, new species" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="FFEC7934FFF7FFF2FFDECD463F236519" masterDocTitle="A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="4" updateTime="1699308862729" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F35DC0E39ECA36548B49B708F17D81CB">A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)</mods:title>
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<mods:namePart id="0A726C0C409644DC8C7E9A96F3C5C5B6">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="F6BC3919C4A81718412829947D01EF53">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:namePart id="78C88A44477287894B6C223CA2AB90D7">Keller, Oliver</mods:namePart>
<mods:affiliation id="CD5F6FB6C7B89D5F0A67D3F347E67817">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:date id="C5E0D86ED33E576B4B0FF57EA0F74BBD">2022</mods:date>
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<subSubSection id="C366E3D1FFF2FFF7FF1ECF9B3C4A67E3" box="[192,873,733,762]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BC3B05AFFF2FFF7FF1ECF9B3C4A67E3" blockId="5.[192,873,733,796]" box="[192,873,733,762]" pageId="5" pageNumber="4">
<heading id="D08B0736FFF2FFF7FF1ECF9B3C4A67E3" box="[192,873,733,762]" fontSize="12" level="2" pageId="5" pageNumber="4" reason="2">
<emphasis id="B9086C48FFF2FFF7FF1ECF9B3C4A67E3" bold="true" box="[192,873,733,762]" pageId="5" pageNumber="4">
<taxonomicName id="4C7CCBD9FFF2FFF7FF1ECF9B3DEA67E3" authority="Skelley and Keller" authorityName="Skelley and Keller" authorityYear="2022" box="[192,713,733,762]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="B9086C48FFF2FFF7FF1ECF9B3EFE67E3" bold="true" box="[192,477,733,762]" italics="true" pageId="5" pageNumber="4">Haroldiellus woodruffi</emphasis>
Skelley and Keller
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, 
<taxonomicNameLabel id="A23BD133FFF2FFF7FD0ACF983C4A67E3" box="[724,873,733,762]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C366E3D1FFF2FFF7FF1ECE473EEB6602" box="[192,456,769,796]" pageId="5" pageNumber="4" type="description">
<paragraph id="8BC3B05AFFF2FFF7FF1ECE473EEB6602" blockId="5.[192,873,733,796]" box="[192,456,769,796]" pageId="5" pageNumber="4">
<figureCitation id="1347ACDFFFF2FFF7FF1ECE473E616605" box="[192,322,769,796]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="5" pageNumber="4">Figures 3, 5</figureCitation>
, 
<figureCitation id="1347ACDFFFF2FFF7FE93CE473E5B6602" box="[333,376,769,795]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="5" pageNumber="4">8–9</figureCitation>
, 
<figureCitation id="1347ACDFFFF2FFF7FE5DCE473EEB6602" box="[387,456,769,795]" captionStart-0="Figures 12–16" captionStart-1="Figure 17" captionStartId-0="11.[192,279,1336,1362]" captionStartId-1="12.[192,268,867,893]" captionTargetBox-0="[192,1440,213,1306]" captionTargetBox-1="[192,1440,213,837]" captionTargetId-0="figure-11@11.[192,1440,213,1306]" captionTargetId-1="figure-39@12.[192,1440,213,837]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Figures 12–16. Haroldiellus woodruffi, male holotype. 12) Lateral habitus. 13) Ventral habitus. 14) Epipharynx. 15–16) Genitalia dorsal and lateral. Scale line = 0.5 mm (figure 12 same scale bar as 13, figure 16 same scale bar as 15)." captionText-1="Figure 17. Distribution map of Haroldiellus woodruffi new species (blue) as currently known from specimen labels. Map made with SimpleMappr (Shorthouse 2010)." figureDoi-0="http://doi.org/10.5281/zenodo.6391967" figureDoi-1="http://doi.org/10.5281/zenodo.6391969" httpUri-0="https://zenodo.org/record/6391967/files/figure.png" httpUri-1="https://zenodo.org/record/6391969/files/figure.png" pageId="5" pageNumber="4">12–17</figureCitation>
</paragraph>
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<subSubSection id="C366E3D1FFF2FFF7FF1ECE6C3C3D66B3" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BC3B05AFFF2FFF7FF1ECE6C3C3D66B3" blockId="5.[191,1445,810,938]" pageId="5" pageNumber="4">
<emphasis id="B9086C48FFF2FFF7FF1ECE6C3E1F665D" bold="true" box="[192,316,810,836]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
Length 
<quantity id="4C841DBFFFF2FFF7FE47CE6C3D01665C" box="[409,546,810,837]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.8" metricValueMin="5.3" pageId="5" pageNumber="4" unit="mm" value="5.55" valueMax="5.8" valueMin="5.3">5.3–5.8 mm</quantity>
, width 
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. A member of 
<taxonomicName id="4C7CCBD9FFF2FFF7FC79CE6C3B0F665D" authorityName="Gordon and Skelley" authorityYear="2007" box="[935,1068,810,836]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9086C48FFF2FFF7FC79CE6C3B0F665D" box="[935,1068,810,836]" italics="true" pageId="5" pageNumber="4">Haroldiellus</emphasis>
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distinguished from other species by: Elytral intervals convex, striae deeply impressed; interval color pale with distinct diagonal black marking (
<figureCitation id="1347ACDFFFF2FFF7FAABCE0A3FED6691" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
). Female lacking median frontal tubercle (
<figureCitation id="1347ACDFFFF2FFF7FD6CCE283DD66691" box="[690,757,878,904]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="5" pageNumber="4">Fig. 8</figureCitation>
); male median frontal tubercle transverse and bifid (
<figureCitation id="1347ACDFFFF2FFF7FA93CE283AB76691" box="[1357,1428,878,904]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="5" pageNumber="4">Fig. 9</figureCitation>
). Protibia with dorsal surface lacking punctures (
<figureCitation id="1347ACDFFFF2FFF7FD11CEC93C3366B3" box="[719,784,911,938]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF2FFF7FF1ECEFF3E7263AF" pageId="5" pageNumber="4" type="description">
<paragraph id="8BC3B05AFFF2FFF7FF1ECEFF3BD26328" blockId="5.[191,1442,953,1718]" pageId="5" pageNumber="4">
<emphasis id="B9086C48FFF2FFF7FF1ECEFF3D2966CA" bold="true" box="[192,522,953,979]" pageId="5" pageNumber="4">
Description. Male 
<typeStatus id="54C70EF8FFF2FFF7FE42CEFF3D2666CA" box="[412,517,953,979]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
.
</emphasis>
Length 
<quantity id="4C841DBFFFF2FFF7FDB4CEFF3DE066CA" box="[618,707,953,980]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="4" unit="mm" value="5.6">5.6 mm</quantity>
, width 
<quantity id="4C841DBFFFF2FFF7FCC7CEFF3C5766CA" box="[793,884,953,979]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="4" unit="mm" value="2.6">2.6 mm</quantity>
. Body robust, not flattened. Body (
<figureCitation id="1347ACDFFFF2FFF7FADCCEFF3A6666CA" box="[1282,1349,953,979]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
, 
<figureCitation id="1347ACDFFFF2FFF7FA8CCEFF3AB466CA" box="[1362,1431,953,979]" captionStart="Figures 12–16" captionStartId="11.[192,279,1336,1362]" captionTargetBox="[192,1440,213,1306]" captionTargetId="figure-11@11.[192,1440,213,1306]" captionTargetPageId="11" captionText="Figures 12–16. Haroldiellus woodruffi, male holotype. 12) Lateral habitus. 13) Ventral habitus. 14) Epipharynx. 15–16) Genitalia dorsal and lateral. Scale line = 0.5 mm (figure 12 same scale bar as 13, figure 16 same scale bar as 15)." figureDoi="http://doi.org/10.5281/zenodo.6391967" httpUri="https://zenodo.org/record/6391967/files/figure.png" pageId="5" pageNumber="4">12–13</figureCitation>
) color dark reddish brown except head and lateral pronotum pale reddish brown; elytra yellowish brown, striae darkened over disk, intervals with an irregular diagonal dark mark notably contrasting with pale part of intervals; legs paler yellowish brown. Dorsal surface glossy; glabrous; body not fimbriate; ventral surface with moderately long, moderately dense pubescence. Anterior clypeal margin with angles rounded, margin between angles thickened, emarginate. Clypeus and epistome coarsely punctured, flat. Frontal suture trituberculate (
<figureCitation id="1347ACDFFFF2FFF7FB2DC9273A166162" box="[1267,1333,1121,1147]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="5" pageNumber="4">Fig. 9</figureCitation>
), median tubercle broad and bifid. Genae moderately developed, slightly deflexed. Pronotum shiny; with intermixed moderate and fine punctures, all punctures separated by 1–2 times their diameter on disc. Scutellar shield weakly pentagonal, not depressed. Elytra with striae deeply impressed, coarsely punctured; intervals convex, glossy, finely punctured. Ventral surface and metafemur with few coarse punctures. Male metaventrite not deeply excavated, without patch or fringe of dense setae. Protibia lacking dorsal punctures (
<figureCitation id="1347ACDFFFF2FFF7FB8AC84F3BB4603D" box="[1108,1175,1289,1316]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
), unmodified; protibial spur and protarsus unmodified. Inferior mesotibial spur straight at apex, lacking tooth. Metatrochanter with a single seta. Metafemur robust, wider than metacoxa; posterior margin of metafemur smooth, glabrous. Metatibia robust, apically flared, not compressed; apex of metatibia fringed with short, equal spinules. Metatarsus as long as metatibia, tarsomeres slender, cylindrical, with basal tarsomere as long as superior tibial spur, slightly longer than tarsomeres 2 and 3 combined. Epipharynx transverse, side rounded, anterior margin concave towards middle; epitorma conical; corypha slightly protruding, with apical clump of long, slender celtes; pedia pubescent, with several stout, scattered chaetae; chaetopariae moderately dense, moderately elongate (
<figureCitation id="1347ACDFFFF2FFF7FB49C8B23BC96317" box="[1175,1258,1524,1550]" captionStart="Figures 12–16" captionStartId="11.[192,279,1336,1362]" captionTargetBox="[192,1440,213,1306]" captionTargetId="figure-11@11.[192,1440,213,1306]" captionTargetPageId="11" captionText="Figures 12–16. Haroldiellus woodruffi, male holotype. 12) Lateral habitus. 13) Ventral habitus. 14) Epipharynx. 15–16) Genitalia dorsal and lateral. Scale line = 0.5 mm (figure 12 same scale bar as 13, figure 16 same scale bar as 15)." figureDoi="http://doi.org/10.5281/zenodo.6391967" httpUri="https://zenodo.org/record/6391967/files/figure.png" pageId="5" pageNumber="4">Fig. 14</figureCitation>
). Male genitalia unmodified; parameres slightly widening at apex, without membranous appendage (
<figureCitation id="1347ACDFFFF2FFF7FBB4CB503BC76328" box="[1130,1252,1558,1585]" captionStart="Figures 12–16" captionStartId="11.[192,279,1336,1362]" captionTargetBox="[192,1440,213,1306]" captionTargetId="figure-11@11.[192,1440,213,1306]" captionTargetPageId="11" captionText="Figures 12–16. Haroldiellus woodruffi, male holotype. 12) Lateral habitus. 13) Ventral habitus. 14) Epipharynx. 15–16) Genitalia dorsal and lateral. Scale line = 0.5 mm (figure 12 same scale bar as 13, figure 16 same scale bar as 15)." figureDoi="http://doi.org/10.5281/zenodo.6391967" httpUri="https://zenodo.org/record/6391967/files/figure.png" pageId="5" pageNumber="4">Fig. 15–16</figureCitation>
).
</paragraph>
<paragraph id="8BC3B05AFFF2FFF7FEDECB713DEA636A" blockId="5.[191,1442,953,1718]" pageId="5" pageNumber="4">
<emphasis id="B9086C48FFF2FFF7FEDECB713E9C6348" bold="true" box="[256,447,1591,1617]" pageId="5" pageNumber="4">
Female 
<typeStatus id="54C70EF8FFF2FFF7FE84CB713E996348" box="[346,442,1591,1617]" pageId="5" pageNumber="4" type="allotype">allotype</typeStatus>
.
</emphasis>
Length 
<quantity id="4C841DBFFFF2FFF7FDC4CB713D52634B" box="[538,625,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="4" unit="mm" value="5.3">5.3 mm</quantity>
, width 
<quantity id="4C841DBFFFF2FFF7FD1DCB713C38634B" box="[707,795,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="4" unit="mm" value="2.4">2.4 mm</quantity>
. Body more elongate, less robust than male; median frontal tubercle reduced to simple swelling (
<figureCitation id="1347ACDFFFF2FFF7FDA4CB1F3D98636A" box="[634,699,1625,1651]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="5" pageNumber="4">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BC3B05AFFF2FFF7FEDECB3C3E7263AF" blockId="5.[191,1442,953,1718]" pageId="5" pageNumber="4">
<emphasis id="B9086C48FFF2FFF7FEDECB3C3E55638D" bold="true" box="[256,374,1658,1684]" pageId="5" pageNumber="4">Variation.</emphasis>
<typeStatus id="54C70EF8FFF2FFF7FEA5CB3D3EC4638C" box="[379,487,1659,1685]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
show some variation in the thickness of the black elytral markings, but they are always well-defined.
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF2FFF4FF1ECB803E68679A" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BC3B05AFFF2FFF4FF1ECB803E68679A" blockId="5.[191,1441,1733,1897]" lastBlockId="6.[192,1442,213,643]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<emphasis id="B9086C48FFF2FFF7FF1ECB803EF963F9" bold="true" box="[192,474,1733,1760]" pageId="5" pageNumber="4">
Type material. 
<typeStatus id="54C70EF8FFF2FFF7FEB3CB803EF963F9" box="[365,474,1734,1760]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED6D289FFFF2FFF7FDFCCB803D4A63F9" box="[546,617,1734,1760]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
) label data: 
<collectingCountry id="F36BF0CAFFF2FFF7FD35CB803CB563F9" box="[747,918,1734,1760]" name="Guatemala" pageId="5" pageNumber="4">GUATEMALA</collectingCountry>
: El Progresso;/ Sierra de las Minas; nr.Pinalon,/ nr.Finca la Tormenta,/ nr. 
<geoCoordinate id="EE48D69DFFF2FFF7FE31CBA13D7A621B" box="[495,601,1767,1794]" degrees="15.07222" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="15.07222">15.07222</geoCoordinate>
, 
<geoCoordinate id="EE48D69DFFF2FFF7FD83CBA13DED621B" box="[605,718,1767,1794]" degrees="89.94891" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-89.94891">-89.94891</geoCoordinate>
;/ 
<date id="FFC2969AFFF2FFF7FD36CBA13CA6621B" box="[744,901,1767,1794]" pageId="5" pageNumber="4" value="2010-05-15" valueMax="2010-05-17" valueMin="2010-05-15">15-17-V-2010</date>
; ~ 
<quantity id="4C841DBFFFF2FFF7FC7BCBA13CD46218" box="[933,1015,1767,1793]" metricMagnitude="3" metricUnit="m" metricValue="2.199" pageId="5" pageNumber="4" unit="m" value="2199.0">2199m</quantity>
;/ P.Skelley, moist oak forest // [red paper] 
<typeStatus id="54C70EF8FFF2FFF7FECBCA4F3E85623A" box="[277,422,1801,1827]" pageId="5" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D7A7BD3FFF2FFF7FE6ECA4C3EE6623C" box="[432,453,1802,1829]" pageId="5" pageNumber="4" type="male">♂</specimenCount>
/ 
<taxonomicName id="4C7CCBD9FFF2FFF7FE04CA4F3D7C623A" authorityName="Gordon and Skelley" authorityYear="2007" box="[474,607,1801,1827]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9086C48FFF2FFF7FE04CA4F3D7C623A" box="[474,607,1801,1827]" italics="true" pageId="5" pageNumber="4">Haroldiellus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C7CCBD9FFF2FFF7FDADCA4F3C6A623A" authority="P. Skelley" authorityName="P. Skelley" box="[627,841,1801,1827]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="B9086C48FFF2FFF7FDADCA4F3DFE623A" box="[627,733,1801,1827]" italics="true" pageId="5" pageNumber="4">woodruffi</emphasis>
P.Skelley
</taxonomicName>
/ &amp; O.Keller. 
<emphasis id="B9086C48FFF2FFF7FC36CA4F3B6E623A" bold="true" box="[1000,1101,1801,1827]" pageId="5" pageNumber="4">
<typeStatus id="54C70EF8FFF2FFF7FC36CA4F3B6E623A" box="[1000,1101,1801,1827]" pageId="5" pageNumber="4" type="allotype">Allotype</typeStatus>
</emphasis>
female (
<collectionCode id="ED6D289FFFF2FFF7FB6ACA4F3BD8623A" box="[1204,1275,1801,1827]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
) label data: // 
<collectingCountry id="F36BF0CAFFF2FFF7FF1ECA6D3E52625C" box="[192,369,1835,1861]" name="Guatemala" pageId="5" pageNumber="4">GUATEMALA</collectingCountry>
:Guat./ Puerta Parada/ 14°33′24″ -90°27′48″/ 
<date id="FFC2969AFFF2FFF7FCB7CA6C3B38625C" box="[873,1051,1834,1861]" pageId="5" pageNumber="4" value="2005-03-26" valueMax="2005-04-06" valueMin="2005-03-26">26.III-6.IV.2005</date>
/ J.Schuster, 
<quantity id="4C841DBFFFF2FFF7FB78CA6D3BD1625C" box="[1190,1266,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="5" pageNumber="4" unit="m" value="1840.0">1840m</quantity>
// [blue paper] 
<typeStatus id="54C70EF8FFF2FFF7FF1CCA0A3E68627F" box="[194,331,1868,1894]" pageId="5" pageNumber="4" type="allotype">ALLOTYPE</typeStatus>
<specimenCount id="9D7A7BD3FFF2FFF7FE8ECA083E466270" box="[336,357,1870,1897]" pageId="5" pageNumber="4" type="female">♀</specimenCount>
/ 
<taxonomicName id="4C7CCBD9FFF2FFF7FEAACA0A3EDA627F" authorityName="Gordon and Skelley" authorityYear="2007" box="[372,505,1868,1894]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9086C48FFF2FFF7FEAACA0A3EDA627F" box="[372,505,1868,1894]" italics="true" pageId="5" pageNumber="4">Haroldiellus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C7CCBD9FFF2FFF7FDD7CA0A3DF9627F" authority="P. Skelley" authorityName="P. Skelley" box="[521,730,1868,1894]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="B9086C48FFF2FFF7FDD7CA0A3D50627F" box="[521,627,1868,1894]" italics="true" pageId="5" pageNumber="4">woodruffi</emphasis>
P.Skelley
</taxonomicName>
/ &amp; O.Keller. 
<emphasis id="B9086C48FFF2FFF7FCB4CA0A3CF2627F" bold="true" box="[874,977,1868,1894]" pageId="5" pageNumber="4">
<typeStatus id="54C70EF8FFF2FFF7FCB4CA0A3CF2627F" box="[874,977,1868,1894]" pageId="5" pageNumber="4" type="paratype">Paratype</typeStatus>
</emphasis>
(18 total) label data: 
<collectingCountry id="F36BF0CAFFF2FFF7FB62CA0A3A7D627F" box="[1212,1374,1868,1894]" name="Costa Rica" pageId="5" pageNumber="4">COSTA RICA</collectingCountry>
: Prov. of/ 
<collectingRegion id="49B87EB8FFF1FFF4FF36CD903E6965E9" box="[232,330,214,240]" country="Costa Rica" name="San Jose" pageId="6" pageNumber="5">San Jose</collectingRegion>
, San Isidro/ de Coronado, Finca, 
<collectingCountry id="F36BF0CAFFF1FFF4FD12CD903C0C65E9" box="[716,815,214,240]" name="Netherlands" pageId="6" pageNumber="5">Holanda</collectingCountry>
, 5,500 Ft. // 
<taxonomicName id="4C7CCBD9FFF1FFF4FC1DCD903B1265E9" authorityName="Hellwig" authorityYear="1798" box="[963,1073,214,240]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">Aphodius</taxonomicName>
/ near 
<taxonomicName id="4C7CCBD9FFF1FFF4FBA3CD903BCC65E9" authority="Har." authorityName="Har." box="[1149,1263,214,240]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">sallei Har.</taxonomicName>
/ # 29 [red ink] Gordon #21 [pencil]/ DET. R.E.Woodruff -72 (
<quantity id="4C841DBFFFF1FFF4FD0ECDB13DD66408" box="[720,757,247,273]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FD22CDB13C606408" box="[764,835,247,273]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="5" type="Museum">FSCA</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FC88CDB13B226408" box="[854,1025,247,273]" name="Guatemala" pageId="6" pageNumber="5">GUATEMALA</collectingCountry>
: 
<collectingCountry id="F36BF0CAFFF1FFF4FBD5CDB13BA46408" box="[1035,1159,247,273]" name="Guatemala" pageId="6" pageNumber="5">Guatemala</collectingCountry>
,/ Santa 
<taxonomicName id="4C7CCBD9FFF1FFF4FB3ECDB13AB06408" authority="Pinula" authorityName="Pinula" box="[1248,1427,247,273]" class="Insecta" family="Geometridae" genus="Catarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">Catarina Pinula</taxonomicName>
,/ Puerta Parada, Casa-Jack/ Schuster, 
<quantity id="4C841DBFFFF1FFF4FDBACC5F3D90642A" box="[612,691,281,307]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="6" pageNumber="5" unit="m" value="1860.0">1860m</quantity>
,/ N 
<geoCoordinate id="EE48D69DFFF1FFF4FD34CC5F3C57642A" box="[746,884,281,308]" degrees="14.45572" direction="west" orientation="longitude" pageId="6" pageNumber="5" precision="1" value="-14.45572">14.45572 W</geoCoordinate>
90.4653/ 
<date id="FFC2969AFFF1FFF4FC34CC5F3BAC642A" box="[1002,1167,281,307]" pageId="6" pageNumber="5" value="2009-07-04" valueMax="2009-07-11" valueMin="2009-07-04">4-11-VII-2009</date>
, J.Schuster (
<quantity id="4C841DBFFFF1FFF4FAC1CC5F3A67642A" box="[1311,1348,281,307]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FA91CC5F3AB5642A" box="[1359,1430,281,307]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="5" type="Museum">FSCA</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FF1ECC7D3E48644C" box="[192,363,315,341]" name="Guatemala" pageId="6" pageNumber="5">GUATEMALA</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FEABCC7D3ED8644C" box="[373,507,315,341]" country="Guatemala" name="San Marcos" pageId="6" pageNumber="5">San Marcos</collectingRegion>
:/ Parque Municipal Refugio/ del Quetzal 
<quantity id="4C841DBFFFF1FFF4FC14CC7D3B34644C" box="[970,1047,315,341]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="6" pageNumber="5" unit="m" value="1810.0">1810m</quantity>
<geoCoordinate id="EE48D69DFFF1FFF4FBC3CC7C3BA1644D" box="[1053,1154,314,340]" degrees="14.93916" direction="north" orientation="latitude" pageId="6" pageNumber="5" precision="1" value="14.93916">14.93916</geoCoordinate>
/ 
<geoCoordinate id="EE48D69DFFF1FFF4FB4CCC7C3A23644C" box="[1170,1280,314,341]" degrees="91.87295" direction="west" orientation="longitude" pageId="6" pageNumber="5" precision="1" value="-91.87295">-91.87295</geoCoordinate>
<date id="FFC2969AFFF1FFF4FAD9CC7C3A83644C" box="[1287,1440,314,341]" pageId="6" pageNumber="5" value="2015-06-04" valueMax="2015-06-11" valueMin="2015-06-04">4-11-VI-2015</date>
ZHFalin/ ex. flight intercept trap. cloud/ forest GUAT1F15 099 // Barcode/ SEMC1464610/ KUNHM-ENT (1f 
<collectionCode id="ED6D289FFFF1FFF4FF1CCC383E2C6481" box="[194,271,382,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="6" pageNumber="5" type="Museum">SEMC</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FEFFCC383EBB6481" box="[289,408,382,408]" name="Panama" pageId="6" pageNumber="5">PANAMA</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FE7CCC383D216481" box="[418,514,382,408]" country="Panama" name="Chiriqui" pageId="6" pageNumber="5">Chiriqui</collectingRegion>
/ Mt. Totumca Lodge/ 
<date id="FFC2969AFFF1FFF4FCDECC383C846480" box="[768,935,382,409]" pageId="6" pageNumber="5" value="2012-06-19" valueMax="2012-06-25" valueMin="2012-06-19">19-25-VI-2012</date>
/ J.B.Heppner, at light(
<quantity id="4C841DBFFFF1FFF4FB78CC383BE46481" box="[1190,1223,382,408]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FB10CC383A366481" box="[1230,1301,382,408]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="5" type="Museum">FSCA</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FAF9CC383ABD6481" box="[1319,1438,382,408]" name="Panama" pageId="6" pageNumber="5">PANAMA</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FF1FCCD93E0264A0" box="[193,289,415,441]" country="Panama" name="Chiriqui" pageId="6" pageNumber="5">Chiriqui</collectingRegion>
/ Prov. 
<quantity id="4C841DBFFFF1FFF4FEA4CCD93E8E64A0" box="[378,429,415,441]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="5" unit="km" value="2.0">2km</quantity>
W Cerro/ Punta 
<geoCoordinate id="EE48D69DFFF1FFF4FD58CCD93DF164A0" box="[646,722,415,444]" degrees="8" direction="north" minutes="51" orientation="latitude" pageId="6" pageNumber="5" precision="925" value="8.85">8°51′N</geoCoordinate>
/ 
<geoCoordinate id="EE48D69DFFF1FFF4FD39CCD93C6B64A3" box="[743,840,415,444]" degrees="82" direction="west" minutes="36" orientation="longitude" pageId="6" pageNumber="5" precision="925" value="-82.6">82°36′W</geoCoordinate>
<date id="FFC2969AFFF1FFF4FC8ACCD93C9B64A0" box="[852,952,415,441]" pageId="6" pageNumber="5" value="1977-06-03">3.vi.1977</date>
/ H.&amp; A.Howden (
<quantity id="4C841DBFFFF1FFF4FB46CCD93B9E64A0" box="[1176,1213,415,441]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FB17CCD93A2E64A0" box="[1225,1293,415,441]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="6" pageNumber="5" type="Museum">CMN</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FAFBCCE63ABF64A3" box="[1317,1436,416,442]" name="Panama" pageId="6" pageNumber="5">PANAMA</collectingCountry>
, 
<collectingRegion id="49B87EB8FFF1FFF4FF1ECC873E0364C2" box="[192,288,449,475]" country="Panama" name="Chiriqui" pageId="6" pageNumber="5">Chiriqui</collectingRegion>
/ Prov. 
<quantity id="4C841DBFFFF1FFF4FEAACC873E9D64C2" box="[372,446,449,475]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="5" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2-3km</quantity>
E Cerro/ Punta, 
<quantity id="4C841DBFFFF1FFF4FD58CC873C3664C2" box="[646,789,449,475]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="6" pageNumber="5" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000-2200m</quantity>
, 1./ VI.77 H. &amp; A. Howden // 
<taxonomicName id="4C7CCBD9FFF1FFF4FBA7CC873A2064C2" authorityName="Gordon and Skelley" authorityYear="2007" box="[1145,1283,449,475]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">Haroldiellus</taxonomicName>
/ n.sp./ det. P. Skelley/ det., 20 2012 (
<quantity id="4C841DBFFFF1FFF4FE0CCCA53ED464E4" box="[466,503,483,509]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FDDECCA53D6764E4" box="[512,580,483,509]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="6" pageNumber="5" type="Museum">CMN</collectionCode>
); Chiquihuite dint./ 
<geoCoordinate id="EE48D69DFFF1FFF4FCECCCA43CE464E4" box="[818,967,482,511]" degrees="15" direction="north" minutes="05" orientation="latitude" pageId="6" pageNumber="5" precision="1" seconds="41.0" value="15.094722">15°05′41.0″N</geoCoordinate>
/ 
<geoCoordinate id="EE48D69DFFF1FFF4FC04CCA43BA064E4" box="[986,1155,482,511]" degrees="092" direction="west" minutes="06" orientation="longitude" pageId="6" pageNumber="5" precision="1" seconds="04.3" value="-92.1012">092°06′04.3″W</geoCoordinate>
/ 
<date id="FFC2969AFFF1FFF4FB4BCCA53A3364E4" box="[1173,1296,483,509]" pageId="6" pageNumber="5" value="2007-06-23">23.VI.2007</date>
- m 2060 // 
<collectingCountry id="F36BF0CAFFF1FFF4FF1ECF423E0E6707" box="[192,301,516,542]" name="Mexico" pageId="6" pageNumber="5">MEXICO</collectingCountry>
/ Edo. 
<collectingRegion id="49B87EB8FFF1FFF4FEA5CF423EF56707" box="[379,470,516,542]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
/ Leg. Dellacasa M./ &amp; Fresi C. (
<quantity id="4C841DBFFFF1FFF4FC8ECF423C556707" box="[848,886,516,542]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
, 7f 
<collectionCode id="ED6D289FFFF1FFF4FC7ACF423CCF6707" box="[932,1004,516,542]" pageId="6" pageNumber="5">DCGI</collectionCode>
, 1f 
<collectionCode id="ED6D289FFFF1FFF4FBC6CF423B7C6707" box="[1048,1119,516,542]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="5" type="Museum">FSCA</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FBAACF423BC06707" box="[1140,1251,516,542]" name="Mexico" pageId="6" pageNumber="5">MEXICO</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FB2ECF423A686707" box="[1264,1355,516,542]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
/ Mpio: Motozintla/ Boquerón/ 
<date id="FFC2969AFFF1FFF4FE0ECF603D726759" box="[464,593,550,576]" pageId="6" pageNumber="5" value="2003-05-10">10/V/ 2003</date>
/ B. Gómez (1f 
<collectionCode id="ED6D289FFFF1FFF4FCD8CF603C6C6759" box="[774,847,550,576]" pageId="6" pageNumber="5">DCGI</collectionCode>
); 
<collectingCountry id="F36BF0CAFFF1FFF4FCBBCF603CF76759" box="[869,980,550,576]" name="Mexico" pageId="6" pageNumber="5">MEXICO</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FC3ECF603B186759" box="[992,1083,550,576]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
/ Mpio: Motozintla/ Boquerón/ 
<date id="FFC2969AFFF1FFF4FF1ECF013E166778" box="[192,309,583,609]" pageId="6" pageNumber="5" value="2004-01-29">29/I/ 2004</date>
/ B. Gómez (
<quantity id="4C841DBFFFF1FFF4FE1ACF013ECA6778" box="[452,489,583,609]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1.0">1m</quantity>
<collectionCode id="ED6D289FFFF1FFF4FE2ECF013D1A6778" box="[496,569,583,609]" pageId="6" pageNumber="5">DCGI</collectionCode>
); // 
<collectingCountry id="F36BF0CAFFF1FFF4FDB9CF013DF56778" box="[615,726,583,609]" name="Mexico" pageId="6" pageNumber="5">MEXICO</collectingCountry>
: 
<collectingRegion id="49B87EB8FFF1FFF4FD3FCF013C1F6778" box="[737,828,583,609]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
/ Mpio: Motozintla/ Boquerón/ 
<date id="FFC2969AFFF1FFF4FB7CCF013A016778" box="[1186,1314,583,609]" pageId="6" pageNumber="5" value="2003-05-11">11/V/ 2003</date>
/ B. Gómez (
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<collectionCode id="ED6D289FFFF1FFF4FF28CF2F3E1C679A" box="[246,319,617,643]" pageId="6" pageNumber="5">DCGI</collectionCode>
);
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF1FFF4FF1ECFD43EC367D6" pageId="6" pageNumber="5" type="distribution">
<paragraph id="8BC3B05AFFF1FFF4FF1ECFD43EC367D6" blockId="6.[192,1440,658,719]" pageId="6" pageNumber="5">
<emphasis id="B9086C48FFF1FFF4FF1ECFD43E4F67B5" bold="true" box="[192,364,658,684]" pageId="6" pageNumber="5">Distribution. (</emphasis>
<figureCitation id="1347ACDFFFF1FFF4FEB2CFD43E9E67B4" box="[364,445,658,685]" captionStart="Figure 17" captionStartId="12.[192,268,867,893]" captionTargetBox="[192,1440,213,837]" captionTargetId="figure-39@12.[192,1440,213,837]" captionTargetPageId="12" captionText="Figure 17. Distribution map of Haroldiellus woodruffi new species (blue) as currently known from specimen labels. Map made with SimpleMappr (Shorthouse 2010)." figureDoi="http://doi.org/10.5281/zenodo.6391969" httpUri="https://zenodo.org/record/6391969/files/figure.png" pageId="6" pageNumber="5">Fig. 17</figureCitation>
). 
<collectingCountry id="F36BF0CAFFF1FFF4FE0CCFD43D0667B5" box="[466,549,658,684]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
(
<collectingRegion id="49B87EB8FFF1FFF4FDEBCFD43DB667B4" box="[565,661,658,685]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
), 
<collectingCountry id="F36BF0CAFFF1FFF4FD74CFD43C0067B5" box="[682,803,658,684]" name="Guatemala" pageId="6" pageNumber="5">Guatemala</collectingCountry>
(
<collectingRegion id="49B87EB8FFF1FFF4FCEDCFD43C9967B4" box="[819,954,658,685]" country="Guatemala" name="El Progreso" pageId="6" pageNumber="5">El Progreso</collectingRegion>
, 
<collectingCountry id="F36BF0CAFFF1FFF4FC19CFD43B6067B5" box="[967,1091,658,684]" name="Guatemala" pageId="6" pageNumber="5">Guatemala</collectingCountry>
, 
<collectingRegion id="49B87EB8FFF1FFF4FB91CFD53BF467B4" box="[1103,1239,659,685]" country="Guatemala" name="San Marcos" pageId="6" pageNumber="5">San Marcos</collectingRegion>
), 
<collectingCountry id="F36BF0CAFFF1FFF4FB35CFD53A4667B5" box="[1259,1381,658,685]" name="Costa Rica" pageId="6" pageNumber="5">Costa Rica</collectingCountry>
(
<collectingRegion id="49B87EB8FFF1FFF4FAABCFD53FD767D7" country="Costa Rica" name="San Jose" pageId="6" pageNumber="5">San José</collectingRegion>
), 
<collectingCountry id="F36BF0CAFFF1FFF4FED8CFF33E4367D6" box="[262,352,693,719]" name="Panama" pageId="6" pageNumber="5">Panama</collectingCountry>
(
<collectingRegion id="49B87EB8FFF1FFF4FEB0CFF23EF767D6" box="[366,468,692,719]" country="Panama" name="Chiriqui" pageId="6" pageNumber="5">Chiriqui</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF1FFF4FF1ECF983C3C6603" pageId="6" pageNumber="5" type="etymology">
<paragraph id="8BC3B05AFFF1FFF4FF1ECF983C3C6603" blockId="6.[192,1440,734,794]" pageId="6" pageNumber="5">
<emphasis id="B9086C48FFF1FFF4FF1ECF983E6767E1" bold="true" box="[192,324,734,760]" pageId="6" pageNumber="5">Etymology.</emphasis>
This species is named after Robert Woodruff, friend and mentor, who first studied this species and recognized it as something different, but near 
<taxonomicName id="4C7CCBD9FFF1FFF4FD1ECE463C3A6600" box="[704,793,767,794]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">
<emphasis id="B9086C48FFF1FFF4FD1ECE463DF66600" box="[704,725,768,793]" italics="true" pageId="6" pageNumber="5">H</emphasis>
. 
<emphasis id="B9086C48FFF1FFF4FD3CCFB93C3A6600" box="[738,793,767,793]" italics="true" pageId="6" pageNumber="5">sallei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF1FFF4FF1ECE6F3A61665A" box="[192,1346,809,835]" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8BC3B05AFFF1FFF4FF1ECE6F3A61665A" blockId="6.[192,1346,809,835]" box="[192,1346,809,835]" pageId="6" pageNumber="5">
<emphasis id="B9086C48FFF1FFF4FF1ECE6F3E0C665A" bold="true" box="[192,303,809,835]" pageId="6" pageNumber="5">Remarks.</emphasis>
<taxonomicName id="4C7CCBD9FFF1FFF4FEEACE6F3D09665A" box="[308,554,809,835]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="B9086C48FFF1FFF4FEEACE6F3D09665A" box="[308,554,809,835]" italics="true" pageId="6" pageNumber="5">Haroldiellus woodruffi</emphasis>
</taxonomicName>
, like 
<taxonomicName id="4C7CCBD9FFF1FFF4FDBBCE6C3DD7665A" baseAuthorityName="Harold" baseAuthorityYear="1874" box="[613,756,809,835]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF1FFF4FDBBCE6C3D59665A" box="[613,634,810,835]" italics="true" pageId="6" pageNumber="5">H</emphasis>
. 
<emphasis id="B9086C48FFF1FFF4FD58CE6F3DD7665A" box="[646,756,809,835]" italics="true" pageId="6" pageNumber="5">lansbergei</emphasis>
</taxonomicName>
, appears to be a species occupying higher elevations.
</paragraph>
</subSubSection>
</treatment>
</document>