584 lines
81 KiB
XML
584 lines
81 KiB
XML
<document id="912DA7DBB9BDEAD70D4A5FBEC84ADC02" ID-CLB-Dataset="32762" ID-DOI="10.11646/zootaxa.4286.1.2" ID-GBIF-Dataset="dddc215c-61ae-484e-bc8e-1314354620b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="828294" ID-ZooBank="3B62E1C2-79C6-4A14-9F6D-80D8872118FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108503344" checkinUser="plazi" docAuthor="Zelaya, Diego G. & Güller, Marina" docDate="2017" docId="03BCFB5FFFD0FFFAFF26FF7B31CBE880" docLanguage="en" docName="zootaxa.4286.1.2.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epitonium evanidstriatum Zelaya & Güller, 2017, new species" docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="FF858327FFDAFFF7FFB1FFEC3539EA70" masterDocTitle="Undercover speciation of wentletraps (Caenogastropoda: Epitoniidae) in the Southwestern Atlantic" masterLastPageNumber="69" masterPageNumber="41" pageNumber="51" updateTime="1698465097681" updateUser="ExternalLinkService">
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<mods:mods id="1D80C81DACBD26F3F4FE9982A3C26F02" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2352CD48145F8BACFDF8332AF59FE577">
|
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<mods:title id="0C73C90ECBAC6D90037D6656FCBE9ACD">Undercover speciation of wentletraps (Caenogastropoda: Epitoniidae) in the Southwestern Atlantic</mods:title>
|
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</mods:titleInfo>
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<mods:name id="F65363220329D02E96DC05944429498E" type="personal">
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<mods:role id="BDAE011FCCFAA22B23D4B61DDF7EFCD9">
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<mods:roleTerm id="3DB671704C3FC8868938B06FFA49E276">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ACE05B71756E5B1C4B5F2CFCD25BA0D1">Zelaya, Diego G.</mods:namePart>
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</mods:name>
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<mods:name id="01C6B357FC451D034009DA17840C0B12" type="personal">
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<mods:role id="66D1F7A1227E34A0336D739449845ACC">
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<mods:roleTerm id="27A22D2BB86608816E9C77B71067CC01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2F5F661BC6EDEF67378400699395956D">Güller, Marina</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="23641F4008EA13055B13FAD178D3BC64">text</mods:typeOfResource>
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<mods:titleInfo id="B1408D6EAC6BD2171C2F2AB597DCF1C8">
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<mods:title id="35D09E27813B81203309E404B80BBDD2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5EABF04864ADCC0323688EEE8F85EC16">
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<mods:date id="CE05F886A0BFE6F8A1D65466A447A6F8">2017</mods:date>
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<mods:detail id="EC89E50AEE21DAE2AC3C8FA6C09BAAA1" type="volume">
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<mods:number id="C013ABE5762BB938B0509842B5F90CDC">4286</mods:number>
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</mods:detail>
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<mods:detail id="D35AF1381301AC7CE9C7B0FA73E85741" type="issue">
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<mods:number id="783411ECFC154822EB0B2AB63EF76A39">1</mods:number>
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</mods:detail>
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<mods:extent id="7271AEF30B1B3BFA0BA48E4376463485" unit="page">
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<mods:start id="257BCC634C183878078C8AD1001D4851">41</mods:start>
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<mods:end id="B328F70199366275730B7A5228A0BBFB">69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="1675E9EC996B2E4AC6F188217798921E">journal article</mods:classification>
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<mods:identifier id="291EDD8B9727DB4CB134E9A4C4CF1986" type="CLB-Dataset">32762</mods:identifier>
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<mods:identifier id="AF3FFFFB4AA5121F065323C69C965316" type="DOI">10.11646/zootaxa.4286.1.2</mods:identifier>
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<mods:identifier id="F875968AE17D595928020B85D85870D0" type="GBIF-Dataset">dddc215c-61ae-484e-bc8e-1314354620b2</mods:identifier>
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<mods:identifier id="622DEF05439F2D70F5F60AEC295EEE83" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0B1796EED3D989F465EE372C361D99CA" type="Zenodo-Dep">828294</mods:identifier>
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<mods:identifier id="53ED45D7B07E9B78B034EFC2D4C61C6D" type="ZooBank">3B62E1C2-79C6-4A14-9F6D-80D8872118FA</mods:identifier>
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</mods:mods>
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<treatment id="03BCFB5FFFD0FFFAFF26FF7B31CBE880" ID-DOI="http://doi.org/10.5281/zenodo.6033221" ID-GBIF-Taxon="132548557" ID-Zenodo-Dep="6033221" LSID="urn:lsid:plazi:treatment:03BCFB5FFFD0FFFAFF26FF7B31CBE880" httpUri="http://treatment.plazi.org/id/03BCFB5FFFD0FFFAFF26FF7B31CBE880" lastPageId="13" lastPageNumber="54" pageId="10" pageNumber="51">
|
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<subSubSection id="C30F19C2FFD0FFFDFF26FF7B345BEAA3" pageId="10" pageNumber="51" type="nomenclature">
|
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<paragraph id="8BAA4A49FFD0FFFDFF26FF7B3753EAC2" blockId="10.[151,618,151,211]" box="[151,618,151,178]" pageId="10" pageNumber="51">
|
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<heading id="D0E2FD25FFD0FFFDFF26FF7B3753EAC2" bold="true" box="[151,618,151,178]" fontSize="11" level="1" pageId="10" pageNumber="51" reason="1">
|
||
<emphasis id="B961965BFFD0FFFDFF26FF7B3753EAC2" bold="true" box="[151,618,151,178]" pageId="10" pageNumber="51">
|
||
<taxonomicName id="4C1531CAFFD0FFFDFF26FF7B34EDEAC1" ID-CoL="6FXB6" box="[151,468,151,177]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="51" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD0FFFDFF26FF7B34EDEAC1" bold="true" box="[151,468,151,177]" italics="true" pageId="10" pageNumber="51">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD0FFFDFE6AFF743753EAC2" box="[475,618,152,178]" pageId="10" pageNumber="51" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF26FF56345BEAA3" blockId="10.[151,618,151,211]" box="[151,354,186,211]" pageId="10" pageNumber="51">
|
||
(
|
||
<figureCitation id="132E56CCFFD0FFFDFF2FFF563406EAA3" box="[158,319,186,211]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Figures 6A–L</figureCitation>
|
||
,
|
||
<figureCitation id="132E56CCFFD0FFFDFEFAFF573460EAA3" box="[331,345,187,211]" captionStart="FIGURE 7" captionStartId="12.[151,249,1896,1918]" captionTargetBox="[151,1436,1093,1875]" captionTargetId="figure@12.[151,1436,1093,1875]" captionTargetPageId="12" captionText="FIGURE 7. Epitonium evanidstriatum, radula and jaw. Paratype (same specimen as Fig. 6 H, I, L: MACN-In 40382). A – C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm." httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="10" pageNumber="51">7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD0FFFDFF26FEEE3437E8D7" pageId="10" pageNumber="51" type="materials_examined">
|
||
<paragraph id="8BAA4A49FFD0FFFDFF26FEEE3024EB6A" blockId="10.[151,1437,258,2011]" box="[151,1309,258,283]" pageId="10" pageNumber="51">
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||
<materialsCitation id="3B7D4014FFD0FFFDFF26FEEE3024EB6A" ID-GBIF-Occurrence="1572324892" box="[151,1309,258,283]" country="Argentina" elevation="8" latitude="-40.9215" location="Golfo San Matias" longLatPrecision="1" longitude="-65.135445" pageId="10" pageNumber="51" specimenCount="1" stateProvince="Rio Negro Province" typeStatus="holotype">
|
||
<emphasis id="B961965BFFD0FFFDFF26FEEE340CEB6B" bold="true" box="[151,309,258,283]" pageId="10" pageNumber="51">
|
||
<typeStatus id="54AEF4EBFFD0FFFDFF26FEEE35EBEB6B" box="[151,210,258,283]" pageId="10" pageNumber="51">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<geoCoordinate id="EE212C8EFFD0FFFDFE8DFEEE34EEEB6B" box="[316,471,258,283]" direction="south" orientation="latitude" pageId="10" pageNumber="51" precision="1" value="-40.9215">40°55’17.4’’S</geoCoordinate>
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||
<geoCoordinate id="EE212C8EFFD0FFFDFE6FFEEE37BDEB6B" box="[478,644,258,283]" direction="west" orientation="longitude" pageId="10" pageNumber="51" precision="1" value="-65.135445">65°08’07.6’’W</geoCoordinate>
|
||
,
|
||
<location id="8ECA1C92FFD0FFFDFD3EFEEE3661EB6B" LSID="urn:lsid:plazi:treatment:03BCFB5FFFD0FFFAFF26FF7B31CBE880:8ECA1C92FFD0FFFDFD3EFEEE3661EB6B" box="[655,856,258,283]" country="Argentina" latitude="-40.9215" longLatPrecision="1" longitude="-65.135445" name="Golfo San Matias" pageId="10" pageNumber="51" stateProvince="Rio Negro Province">Golfo San Matías</location>
|
||
,
|
||
<collectingRegion id="49D184ABFFD0FFFDFCD2FEEF317FEB6B" box="[867,1094,258,283]" country="Argentina" name="Rio Negro" pageId="10" pageNumber="51">
|
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<collectingRegion id="49D184ABFFD0FFFDFCD2FEEF36E0EB6B" box="[867,985,259,283]" country="Uruguay" name="Rio Negro" pageId="10" pageNumber="51">
|
||
Río
|
||
<collectingRegion id="49D184ABFFD0FFFDFC22FEEF36E0EB6B" box="[915,985,259,283]" inferredBySuffix="true" pageId="10" pageNumber="51">Negro</collectingRegion>
|
||
</collectingRegion>
|
||
Province
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3020AD9FFD0FFFDFBE0FEEE31FAEB6B" box="[1105,1219,258,283]" name="Argentina" pageId="10" pageNumber="51">Argentina</collectingCountry>
|
||
,
|
||
<elevation id="0038AD7AFFD0FFFDFB7EFEEF3021EB6A" box="[1231,1304,259,283]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="10" pageNumber="51" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">
|
||
<quantity id="4CEDE7ACFFD0FFFDFB7EFEEF3021EB6A" box="[1231,1304,259,283]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="10" pageNumber="51" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">7–8 m</quantity>
|
||
</elevation>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FECA36FDEB13" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76FECA3441EB4F" bold="true" box="[199,376,294,319]" pageId="10" pageNumber="51">Type material.</emphasis>
|
||
Holotype (MACN-In 40380) and 3 paratypes (MACN-In 40381) from the type locality; 2 paratypes from 40°55’29.5’’S 65°08’32.7’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFD2BFEA737D8EB12" box="[666,737,331,355]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="10" pageNumber="51" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
|
||
(MACN-In 40382).
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FE82312EE82F" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<materialsCitation id="3B7D4014FFD0FFFDFF76FE8234EDEBDB" ID-GBIF-Occurrence="1572324895" country="Argentina" latitude="-40.813" location="Rio Negro Province" longLatPrecision="1" longitude="-65.09589" pageId="10" pageNumber="51" specimenCount="1" stateProvince="Rio Negro Province">
|
||
<emphasis id="B961965BFFD0FFFDFF76FE82370FEBF7" bold="true" box="[199,566,366,391]" pageId="10" pageNumber="51">Additional material examined.</emphasis>
|
||
<collectingCountry id="F3020AD9FFD0FFFDFD8FFE82378BEBF7" box="[574,690,366,391]" name="Argentina" pageId="10" pageNumber="51">Argentina</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D184ABFFD0FFFDFD0CFE83369AEBF7" box="[701,931,366,391]" country="Argentina" name="Rio Negro" pageId="10" pageNumber="51">
|
||
<collectingRegion id="49D184ABFFD0FFFDFD0CFE83360AEBF7" box="[701,819,367,391]" country="Uruguay" name="Rio Negro" pageId="10" pageNumber="51">
|
||
Río
|
||
<collectingRegion id="49D184ABFFD0FFFDFD5CFE83360AEBF7" box="[749,819,367,391]" inferredBySuffix="true" pageId="10" pageNumber="51">Negro</collectingRegion>
|
||
</collectingRegion>
|
||
Province
|
||
</collectingRegion>
|
||
:
|
||
<geoCoordinate id="EE212C8EFFD0FFFDFC1EFE823161EBF7" box="[943,1112,366,391]" direction="south" orientation="latitude" pageId="10" pageNumber="51" precision="1" value="-40.813">40°48’46.80’’S</geoCoordinate>
|
||
<geoCoordinate id="EE212C8EFFD0FFFDFBD1FE82302DEBF7" box="[1120,1300,366,391]" direction="west" orientation="longitude" pageId="10" pageNumber="51" precision="1" value="-65.09589">65°05’45.18’’W</geoCoordinate>
|
||
, Las Grutas (MACN-In 40383: 1 sh.)
|
||
</materialsCitation>
|
||
; 40°50’12.8’’S 65°04’42.2’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFCE7FE7F36A3EBDA" box="[854,922,403,427]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="51" unit="m" value="10.0">10 m</quantity>
|
||
(MACN-In 40384: 2 sh.); 40°54’08.3’’S 65°06’28.0’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFEFFFE5B34B9EBBE" box="[334,384,439,463]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="51" unit="m" value="9.0">9 m</quantity>
|
||
(MACN-In 40385: 6 sh.); 40°54’21.7’’S 65°06’39.8’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFB9DFE5A3164EBBE" box="[1068,1117,438,463]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="51" unit="m" value="6.0">6 m</quantity>
|
||
(MACN-In 40386: 1 sh.); 40°56’27.2’S 65°07’58.2’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFE52FE37377CEB82" box="[483,581,475,499]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="10" pageNumber="51" unit="m" value="10.5" valueMax="11.0" valueMin="10.0">10–11 m</quantity>
|
||
(MACN-In 40387: 11 sh.); 41°38’06.6’’S 65°00’55.9’’W, Playas Doradas,
|
||
<quantity id="4CEDE7ACFFD0FFFDFA3FFE373594E866" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="51" unit="m" value="7.0">7 m</quantity>
|
||
(MACN-In 40388: 22 sh.); 41°39’03.1’’S 65°00’38.3’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFCE9FE133686E866" box="[856,959,511,535]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="10" pageNumber="51" unit="m" value="13.0" valueMax="14.0" valueMin="12.0">12–14 m</quantity>
|
||
(MACN-In 40389: 2 sh.); 41°39’41.3’’S 65°00’32.9’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFEFDFDCE34B3E84A" box="[332,394,546,571]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="51" unit="m" value="16.0">16 m</quantity>
|
||
(MACN-In 40390: 2 sh.); 41°40’17.8’’S 65°00’27.8’’W,
|
||
<quantity id="4CEDE7ACFFD0FFFDFB99FDCE315CE84A" box="[1064,1125,546,571]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="51" unit="m" value="16.0">16 m</quantity>
|
||
(MACN-In 40391: 2 sh.); 42°00’54.2’’S 65°03’40.9’’W, Puerto Lobos,
|
||
<quantity id="4CEDE7ACFFD0FFFDFD3FFDAB37C8E82E" box="[654,753,583,607]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="10" pageNumber="51" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10–12 m</quantity>
|
||
(MACN-In 40392: 1 sh.).
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FD863437E8D7" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<materialsCitation id="3B7D4014FFD0FFFDFF76FD863437E8D7" ID-GBIF-Occurrence="1572324901" country="Argentina" latitude="-42.0" location="Rio Negro Province" pageId="10" pageNumber="51" specimenCount="1" stateProvince="Rio Negro Province">
|
||
<emphasis id="B961965BFFD0FFFDFF76FD8634F8E8F3" bold="true" box="[199,449,618,643]" pageId="10" pageNumber="51">Known distribution.</emphasis>
|
||
Only known from Golfo San Matías (
|
||
<geoCoordinate id="EE212C8EFFD0FFFDFC10FD8736C5E8F3" box="[929,1020,619,643]" direction="south" orientation="latitude" pageId="10" pageNumber="51" precision="925" value="-40.8">40°48’S</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="EE212C8EFFD0FFFDFB9FFD8731B2E8F3" box="[1070,1163,619,643]" direction="south" orientation="latitude" pageId="10" pageNumber="51" precision="925" value="-42.0">42°00’S</geoCoordinate>
|
||
),
|
||
<collectingRegion id="49D184ABFFD0FFFDFB17FD8730A1E8F3" box="[1190,1432,618,643]" country="Argentina" name="Rio Negro" pageId="10" pageNumber="51">
|
||
<collectingRegion id="49D184ABFFD0FFFDFB17FD87301AE8F3" box="[1190,1315,619,643]" country="Uruguay" name="Rio Negro" pageId="10" pageNumber="51">
|
||
Río
|
||
<collectingRegion id="49D184ABFFD0FFFDFB6CFD87301AE8F3" box="[1245,1315,619,643]" inferredBySuffix="true" pageId="10" pageNumber="51">Negro</collectingRegion>
|
||
</collectingRegion>
|
||
Province
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3020AD9FFD0FFFDFF26FD623433E8D7" box="[151,266,654,679]" name="Argentina" pageId="10" pageNumber="51">Argentina</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD0FFFDFF76FD5E371DE89F" pageId="10" pageNumber="51" type="description">
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FD5E371DE89F" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76FD5E3464E8BB" bold="true" box="[199,349,690,715]" pageId="10" pageNumber="51">Ethymology.</emphasis>
|
||
<emphasis id="B961965BFFD0FFFDFED5FD5F3496E8BB" box="[356,431,691,715]" italics="true" pageId="10" pageNumber="51">evanid</emphasis>
|
||
- Latin for disappearing +
|
||
<emphasis id="B961965BFFD0FFFDFD6AFD5F3601E8BA" box="[731,824,691,714]" italics="true" pageId="10" pageNumber="51">striatum</emphasis>
|
||
for striae; in reference to the spiral threads that fade in the last whorls of larger specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD0FFFDFF76FD1637F5E9D3" pageId="10" pageNumber="51" type="diagnosis">
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FD1637F5E9D3" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76FD163478E963" bold="true" box="[199,321,762,787]" pageId="10" pageNumber="51">Diagnosis.</emphasis>
|
||
Shell broadly conical to narrowly elongate. Protoconch conical, of 3 to 3¼ whorls. Teleoconch with convex whorls, sculptured with lamellate or recurved axial ribs, which decrease in number with growth, and spiral threads, clearly visible in the first whorls but fading in the last whorls of larger specimens. Jaw with a row of flat, distally-blunt denticles at the anterior margin. Teeth with two secondary denticles in the central field, reduced to knobs or completely absent in the outermost teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD0FFFDFF76FC4231C1ED3B" pageId="10" pageNumber="51" type="description">
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FC423003EEEF" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76FC423460E9B7" bold="true" box="[199,345,942,967]" pageId="10" pageNumber="51">Description.</emphasis>
|
||
Shell medium sized (maximum L observed =
|
||
<quantity id="4CEDE7ACFFD0FFFDFCC0FC4236E2E9B7" box="[881,987,942,967]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="10" pageNumber="51" unit="mm" value="11.5">11.5 mm</quantity>
|
||
), broadly conical to narrowly elongate, thin, white, shiny (
|
||
<figureCitation id="132E56CCFFD0FFFDFEC3FC3E370CE99B" box="[370,565,978,1003]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6A, C, E, F</figureCitation>
|
||
). Protoconch conical, of about 300 µm in length and 310 µm in maximum diameter, composed of 3 to 3¼ whorls; whorls slightly convex in outline; usually lost in larger specimens; surface glossy, sculptured with faint axial threads (
|
||
<figureCitation id="132E56CCFFD0FFFDFDC4FBF6363DEE43" box="[629,772,1050,1075]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6D, G, I</figureCitation>
|
||
). Limit between protoconch and teleoconch clearly defined by a change in sculpture. Teleoconch with up to 7½ whorls, the first ones slightly convex, then gradually increasing in convexity (
|
||
<figureCitation id="132E56CCFFD0FFFDFE80FB8E34D0EE0B" box="[305,489,1122,1147]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6A, C, E, F</figureCitation>
|
||
); suture moderately deep (
|
||
<figureCitation id="132E56CCFFD0FFFDFCA2FB8E36BFEE0B" box="[787,902,1122,1147]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6J, K</figureCitation>
|
||
). Last whorl slightly flattened at base. Aperture subovate, expanded at the base. Outer margin thick. Umbilicus absent. Fasciole narrow (
|
||
<figureCitation id="132E56CCFFD0FFFDFBC7FB6A3014EEEF" box="[1142,1325,1158,1183]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6A, C, E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76FB46362DEC77" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
Teleoconch sculptured with prosocline axial ribs and delicate spiral threads (
|
||
<figureCitation id="132E56CCFFD0FFFDFB88FB46302AEEB3" box="[1081,1299,1194,1219]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6A, C, E, F, J</figureCitation>
|
||
). Axial ribs evenly convex along the whorl, sometimes forming a weak shoulder adapically, but without angulation at the base of last whorl; formed by fusion of several layers. Ribs on adjoining whorls discontinuous, extending well into the suture (
|
||
<figureCitation id="132E56CCFFD0FFFDFF5CFAFA345DEF5F" box="[237,356,1302,1327]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6J, K</figureCitation>
|
||
). Number of ribs consistently decreasing with shell growth: from 18–27 on the first whorls to 13–19 from the fifth whorl onwards (
|
||
<figureCitation id="132E56CCFFD0FFFDFD8AFAD637AAEF23" box="[571,659,1338,1363]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6B</figureCitation>
|
||
); ribs in the first whorl are relatively low, but gradually increasing in height in subsequent whorls, where they are either erect (lamellar) or slightly to moderately recurved (
|
||
<figureCitation id="132E56CCFFD0FFFDFA95FAB23596EFEB" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6B, J, K</figureCitation>
|
||
); in the last whorl, ribs extending to the umbilical area (
|
||
<figureCitation id="132E56CCFFD0FFFDFC94FA6E36E7EFEB" box="[805,990,1410,1435]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6A, C, E, F</figureCitation>
|
||
). Interspaces between ribs with widelyseparated spiral threads, regularly spaced along the whorl, although somewhat weaker near the suture; in number of
|
||
<quantity id="4CEDE7ACFFD0FFFDFF26FA2735E6EF93" box="[151,223,1482,1507]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="10" pageNumber="51" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">8–9 in</quantity>
|
||
the first teleoconch whorl, increasing up to 15–19, but fading from the fourth to sixth whorl onwards (
|
||
<figureCitation id="132E56CCFFD0FFFDFADEFA2635EFEC77" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6J, K</figureCitation>
|
||
). Spiral sculpture not crossing over axial sculpture.
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76F9FF3632EC03" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76F9FF3474EC5B" box="[199,333,1555,1579]" italics="true" pageId="10" pageNumber="51">Operculum:</emphasis>
|
||
Thin, subovate, paucispiral, with eccentric nucleus (
|
||
<figureCitation id="132E56CCFFD0FFFDFC0BF9FE3121EC5B" box="[954,1048,1554,1579]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6H</figureCitation>
|
||
). Adjacent whorls without raised edges. Outside surface sculptured with 34 to 36 irregular threads per
|
||
<quantity id="4CEDE7ACFFD0FFFDFC02F9DB3134EC3E" box="[947,1037,1591,1615]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="51" unit="mm" value="0.1">0.1 mm</quantity>
|
||
, obliquely oriented with respect to growth lines (
|
||
<figureCitation id="132E56CCFFD0FFFDFE80F9B634B0EC03" box="[305,393,1626,1651]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="10" pageNumber="51">Fig. 6L</figureCitation>
|
||
). Colour: light brown, translucent.
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76F993341FECCB" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76F99335CDECE6" box="[199,244,1663,1686]" italics="true" pageId="10" pageNumber="51">Jaw</emphasis>
|
||
(
|
||
<figureCitation id="132E56CCFFD0FFFDFEB5F992347FECE6" box="[260,326,1662,1687]" captionStart="FIGURE 7" captionStartId="12.[151,249,1896,1918]" captionTargetBox="[151,1436,1093,1875]" captionTargetId="figure@12.[151,1436,1093,1875]" captionTargetPageId="12" captionText="FIGURE 7. Epitonium evanidstriatum, radula and jaw. Paratype (same specimen as Fig. 6 H, I, L: MACN-In 40382). A – C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm." httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="10" pageNumber="51">Fig. 7</figureCitation>
|
||
D, E): Anterior margin with a row of flat, distally-blunt denticles, followed by a series of irregular, pitted plates.
|
||
</paragraph>
|
||
<paragraph id="8BAA4A49FFD0FFFDFF76F92A31C1ED3B" blockId="10.[151,1437,258,2011]" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76F92A3421ECAE" box="[199,280,1734,1758]" italics="true" pageId="10" pageNumber="51">Radula</emphasis>
|
||
: With numerous teeth per row, all of them showing an obscure basal denticle. Teeth of the central field short and stout, with an acute apical denticle and one or two, variably-developed secondary denticles (
|
||
<figureCitation id="132E56CCFFD0FFFDFA82F9063040ED73" box="[1331,1401,1770,1795]" captionStart="FIGURE 7" captionStartId="12.[151,249,1896,1918]" captionTargetBox="[151,1436,1093,1875]" captionTargetId="figure@12.[151,1436,1093,1875]" captionTargetPageId="12" captionText="FIGURE 7. Epitonium evanidstriatum, radula and jaw. Paratype (same specimen as Fig. 6 H, I, L: MACN-In 40382). A – C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm." httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="10" pageNumber="51">Fig. 7</figureCitation>
|
||
A). Subsequent teeth with a markedly-elongate, narrow blade, and an upturned, acute apical denticle (
|
||
<figureCitation id="132E56CCFFD0FFFDFB47F8E23002ED56" box="[1270,1339,1806,1831]" captionStart="FIGURE 7" captionStartId="12.[151,249,1896,1918]" captionTargetBox="[151,1436,1093,1875]" captionTargetId="figure@12.[151,1436,1093,1875]" captionTargetPageId="12" captionText="FIGURE 7. Epitonium evanidstriatum, radula and jaw. Paratype (same specimen as Fig. 6 H, I, L: MACN-In 40382). A – C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm." httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="10" pageNumber="51">Fig. 7</figureCitation>
|
||
B, C). A minute, knob-like secondary denticle sometimes present. Outermost teeth slightly smaller (
|
||
<figureCitation id="132E56CCFFD0FFFDFB23F8DE31EFED3B" box="[1170,1238,1842,1867]" captionStart="FIGURE 7" captionStartId="12.[151,249,1896,1918]" captionTargetBox="[151,1436,1093,1875]" captionTargetId="figure@12.[151,1436,1093,1875]" captionTargetPageId="12" captionText="FIGURE 7. Epitonium evanidstriatum, radula and jaw. Paratype (same specimen as Fig. 6 H, I, L: MACN-In 40382). A – C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm." httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="10" pageNumber="51">Fig. 7</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD0FFFBFF76F8BA31CBEB4F" lastPageId="12" lastPageNumber="53" pageId="10" pageNumber="51" type="discussion">
|
||
<paragraph id="8BAA4A49FFD0FFFCFF76F8BA37A6EA88" blockId="10.[151,1437,258,2011]" lastBlockId="11.[151,1436,151,248]" lastPageId="11" lastPageNumber="52" pageId="10" pageNumber="51">
|
||
<emphasis id="B961965BFFD0FFFDFF76F8BA3402ED1F" bold="true" box="[199,315,1878,1903]" pageId="10" pageNumber="51">Remarks.</emphasis>
|
||
The variability in shell outline described above for
|
||
<taxonomicName id="4C1531CAFFD0FFFDFC36F8BB3177ED1E" box="[903,1102,1878,1902]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="51" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD0FFFDFC36F8BB3177ED1E" box="[903,1102,1878,1902]" italics="true" pageId="10" pageNumber="51">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD0FFFDFBE6F8BA31AFED1F" box="[1111,1174,1878,1903]" pageId="10" pageNumber="51" rank="species">
|
||
<emphasis id="B961965BFFD0FFFDFBE6F8BA31AFED1F" bold="true" box="[1111,1174,1878,1903]" pageId="10" pageNumber="51">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is size-related: smaller specimens are broadly conical, and gradually become narrowly elongate with growth. This variation is associated with the change in the incremental rate of whorls, which in the first teleoconch whorl is high (with an increase in width of 50–70% from the last protoconch whorl to the first teleoconch whorl), but in subsequent whorls gradually decreases (to about 20% from the sixth to seventh whorls), showing a particularly marked decrease from the first to the second teleoconch whorl. This change in the incremental rate of whorls gives origin to a narrowly elongated aspect of the shell outline of larger specimens.
|
||
</paragraph>
|
||
<caption id="DF6A1AC1FFD1FFFCFF26F9193654EDF6" httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="11" pageNumber="52" targetBox="[151,1436,301,1760]" targetPageId="11">
|
||
<paragraph id="8BAA4A49FFD1FFFCFF26F9193654EDF6" blockId="11.[151,1436,1781,1926]" pageId="11" pageNumber="52">
|
||
<emphasis id="B961965BFFD1FFFCFF26F9193428ED7B" bold="true" box="[151,273,1781,1803]" pageId="11" pageNumber="52">FIGURE 6.</emphasis>
|
||
A–L.
|
||
<taxonomicName id="4C1531CAFFD1FFFCFEE5F91B3765ED7B" box="[340,604,1781,1804]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="52" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD1FFFCFEE5F91B3765ED7B" box="[340,604,1781,1804]" italics="true" pageId="11" pageNumber="52">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD1FFFCFDD3F91A37E5ED7B" box="[610,732,1781,1803]" pageId="11" pageNumber="52" rank="species">new species</taxonomicNameLabel>
|
||
, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40°56'27.2'S 65°07'58.2''W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of
|
||
<taxonomicName id="4C1531CAFFD1FFFCFC2CF8BE3105ED18" box="[925,1084,1874,1896]" class="Gastropoda" family="Epitoniidae" genus="Scala" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="52" phylum="Mollusca" rank="species" species="apiculata">
|
||
<emphasis id="B961965BFFD1FFFCFC2CF8BE3105ED18" box="[925,1084,1874,1896]" italics="true" pageId="11" pageNumber="52">Scala apiculata</emphasis>
|
||
</taxonomicName>
|
||
(USNM 94890). Scale bars: A–C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAA4A49FFD1FFFBFF76F85831CBEB4F" blockId="11.[151,1436,1971,2032]" lastBlockId="12.[151,1437,151,1040]" lastPageId="12" lastPageNumber="53" pageId="11" pageNumber="52">
|
||
In shell outline,
|
||
<taxonomicName id="4C1531CAFFD1FFFCFE33F858379EEDBB" box="[386,679,1971,1995]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="52" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD1FFFCFE33F858379EEDBB" box="[386,679,1971,1995]" italics="true" pageId="11" pageNumber="52">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD1FFFCFD01F85F37C8EDBC" box="[688,753,1971,1996]" pageId="11" pageNumber="52" rank="species">
|
||
<emphasis id="B961965BFFD1FFFCFD01F85F37C8EDBC" bold="true" box="[688,753,1971,1996]" pageId="11" pageNumber="52">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
closely resembles that referred to as
|
||
<taxonomicName id="4C1531CAFFD1FFFCFB2CF858345FED9F" authority="d'Orbigny, 1842" authorityName="d'Orbigny" authorityYear="1842" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="52" phylum="Mollusca" rank="species" species="candeanum">
|
||
<emphasis id="B961965BFFD1FFFCFB2CF85830A5EDBB" box="[1181,1436,1971,1995]" italics="true" pageId="11" pageNumber="52">Epitonium candeanum</emphasis>
|
||
(d’Orbigny, 1842)
|
||
</taxonomicName>
|
||
by different authors (e.g.,
|
||
<bibRefCitation id="EF8437B8FFD1FFFCFD14F83B368BED80" author="Clench" box="[677,946,2007,2032]" pageId="11" pageNumber="52" refString="Clench, W. J. & Turner, R. D. (1951) The genus Epitonium in the western Atlantic. Part 1. Johnsonia, 2, 249 - 288." type="journal article" year="1951">Clench & Turner 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8437B8FFD1FFFCFC73F83B3105ED9F" author="Rios" box="[962,1084,2007,2032]" pageId="11" pageNumber="52" refString="Rios, E. (1994) Seashells of Brazil. Editora da FURG, Rio Grande, 368 pp., 113 pls." type="book" year="1994">Rios 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8437B8FFD1FFFCFBFDF83B31D7ED9F" author="Redfern" box="[1100,1262,2007,2032]" pageId="11" pageNumber="52" refString="Redfern, C. (2000) Bahamian seashells. 1161 species from Abaco, Bahamas. Bahamianseashells. com, Inc., Florida, iii - viii + 501 pp." type="book" year="2000">Redfern 2000</bibRefCitation>
|
||
). Although the concept of
|
||
<taxonomicName id="4C1531CAFFD6FFFBFEA4FF75348FEAC0" box="[277,438,152,176]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="candeanum">
|
||
<emphasis id="B961965BFFD6FFFBFEA4FF75348FEAC0" box="[277,438,152,176]" italics="true" pageId="12" pageNumber="53">E. candeanum</emphasis>
|
||
</taxonomicName>
|
||
is variable among different authors (e.g., d’Orbigny 1842;
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFBE6FF7B3063EAC0" author="Clench" box="[1111,1370,151,176]" pageId="12" pageNumber="53" refString="Clench, W. J. & Turner, R. D. (1951) The genus Epitonium in the western Atlantic. Part 1. Johnsonia, 2, 249 - 288." type="journal article" year="1951">Clench & Turner 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFAD6FF7B3720EAA4" author="Diaz" pageId="12" pageNumber="53" refString="Diaz Merlano, J. M. & Puyana Hegedus, M. (1994) Moluscos del Caribe colombiano. Un catalogo ilustrado. Colciencias y Fundacion Natura Colombia, Santa Fe de Bogota, 291 pp., 74 pls." type="book" year="1994">Diaz Merlano & Puyana Hegedus 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFD97FF5037FAEAA4" author="Redfern" box="[550,707,188,213]" pageId="12" pageNumber="53" refString="Redfern, C. (2000) Bahamian seashells. 1161 species from Abaco, Bahamas. Bahamianseashells. com, Inc., Florida, iii - viii + 501 pp." type="book" year="2000">Redfern 2000</bibRefCitation>
|
||
), there is a general agreement on the presence of fine, secondary axial sculpture running across the spiral threads, between the strong, primary axial ribs. This condition clearly differs from that present in
|
||
<taxonomicName id="4C1531CAFFD6FFFBFE5CFEE93782EB6C" box="[493,699,260,284]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFE5CFEE93782EB6C" box="[493,699,260,284]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFD78FEE83636EB6D" box="[713,783,260,285]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFD78FEE83636EB6D" bold="true" box="[713,783,260,285]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, where this secondary sculpture is completely absent. Furthermore,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFE80FEC534C1EB30" box="[305,504,296,320]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFE80FEC534C1EB30" box="[305,504,296,320]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFE4EFECB3704EB30" box="[511,573,295,320]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFE4EFECB3704EB30" bold="true" box="[511,573,295,320]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is narrower and with less inflated whorls than
|
||
<taxonomicName id="4C1531CAFFD6FFFBFBFAFEC531D2EB30" box="[1099,1259,296,320]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="candeanum">
|
||
<emphasis id="B961965BFFD6FFFBFBFAFEC531D2EB30" box="[1099,1259,296,320]" italics="true" pageId="12" pageNumber="53">E. candeanum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD6FFFAFF76FEA1343BE868" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53" type="reference_group">
|
||
<paragraph id="8BAA4A49FFD6FFFBFF76FEA131B6EE60" blockId="12.[151,1437,151,1040]" pageId="12" pageNumber="53">
|
||
In
|
||
<taxonomicName id="4C1531CAFFD6FFFBFF5AFEA1348EEB14" ID-CoL="6FXB6" box="[235,439,332,356]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFF5AFEA1348EEB14" box="[235,439,332,356]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFE75FEA03731EB15" box="[452,520,332,357]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFE75FEA03731EB15" bold="true" box="[452,520,332,357]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the spiral sculpture fades towards the last teleoconch whorls. The vanishing condition of spirals has been previously described for a few species:
|
||
<taxonomicName id="4C1531CAFFD6FFFBFC2DFE9C3042EBF8" ID-CoL="53HFC" authority="Kiener, 1838" authorityName="Kiener" authorityYear="1838" box="[924,1403,367,392]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="costulatum">
|
||
<emphasis id="B961965BFFD6FFFBFC2DFE9C31F7EBF8" box="[924,1230,368,392]" italics="true" pageId="12" pageNumber="53">Surrepifungium costulatum</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFB6DFE83304BEBF8" author="Kiener" box="[1244,1394,367,392]" pageId="12" pageNumber="53" refString="Kiener, L. C. (1838) Species General et Iconographie des Coquilles Vivantes. Genre Scalaire. Rousseau, Paris, 22 pp., 7 pls." type="book" year="1838">Kiener, 1838</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFA36FE9C3710EBDC" ID-CoL="7B5PF" authority="Bonfitto & Sabelli, 2000" authorityName="Bonfitto & Sabelli" authorityYear="2000" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="oliverioi">
|
||
<emphasis id="B961965BFFD6FFFBFA36FE9C35CFEBDC" italics="true" pageId="12" pageNumber="53">S. oliverioi</emphasis>
|
||
(Bonfitto & Sabelli, 2000)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFD84FE7831A8EBDD" ID-CoL="9G6CJ" authority="Gittenberger & Gittenberger, 2005" authorityName="Gittenberger & Gittenberger" authorityYear="2005" box="[565,1169,404,429]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="patamakanthini">
|
||
<emphasis id="B961965BFFD6FFFBFD84FE78363BEBDC" box="[565,770,404,428]" italics="true" pageId="12" pageNumber="53">S. patamakanthini</emphasis>
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFCB8FE7831A8EBDD" author="Gittenberger" box="[777,1169,404,429]" pageId="12" pageNumber="53" refString="Gittenberger, A. & Gittenberger, E. (2005) A hitherto unnoticed adaptive radiation: epitoniid species (Gastropoda: Epitoniidae) associated with corals. Contributions to Zoology, 74, 125 - 203." type="journal article" year="2005">Gittenberger & Gittenberger, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, all from the Indo-West Pacific, and
|
||
<taxonomicName id="4C1531CAFFD6FFFBFE98FE5437F9EBA0" ID-CoL="6FXL6" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[297,704,439,464]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="apiculatum">
|
||
<emphasis id="B961965BFFD6FFFBFE98FE54371FEBA0" box="[297,550,440,464]" italics="true" pageId="12" pageNumber="53">Epitonium apiculatum</emphasis>
|
||
(Dall, 1889)
|
||
</taxonomicName>
|
||
, described from the East coast of the
|
||
<collectingCountry id="F3020AD9FFD6FFFBFB30FE5B3023EBA0" box="[1153,1306,439,464]" name="United States of America" pageId="12" pageNumber="53">United States</collectingCountry>
|
||
(
|
||
<figureCitation id="132E56CCFFD6FFFBFA9CFE5B304AEBA0" box="[1325,1395,439,464]" captionStart="FIGURE 6" captionStartId="11.[151,249,1781,1803]" captionTargetBox="[151,1436,301,1760]" captionTargetId="figure@11.[151,1436,301,1760]" captionTargetPageId="11" captionText="FIGURE 6. A – L. Epitonium evanidstriatum new species, shell and operculum. A, B. Holotype (MACN-In 40380). C, D, F, G, J. Specimens from 40 ° 56 ' 27.2 ' S 65 ° 07 ' 58.2 '' W (MACN-In 40387). E, K. Paratype (MACN-In 40381). H, I, L. Paratype (MACN-In 40382). I. Detail of protoconch sculpture. J, K. Detail of teleoconch sculpture between second and third whorls (J) and last two whorls (K); L. Detail of operculum sculpture. M. Lectotype of Scala apiculata (USNM 94890). Scale bars: A – C, E, M = 2 mm; D, G, J = 200 µm; F, H, K = 1 mm; I = 5 µm; L = 20 µm." httpUri="https://zenodo.org/record/828306/files/figure.png" pageId="12" pageNumber="53">Fig. 6</figureCitation>
|
||
M). However, in these species the spiral sculpture disappears completely at some point, whereas in
|
||
<taxonomicName id="4C1531CAFFD6FFFBFB09FE313046EB84" ID-CoL="6FXB6" box="[1208,1407,476,500]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFB09FE313046EB84" box="[1208,1407,476,500]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFA34FE303581E868" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFA34FE303581E868" bold="true" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the spirals fade in the last whorls of larger specimens but are still visible under high magnification (SEM). In addition,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFEB1FDC9371DE84C" ID-CoL="6FXB6" box="[256,548,548,572]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFEB1FDC9371DE84C" box="[256,548,548,572]" italics="true" pageId="12" pageNumber="53">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFD9AFDC83750E84D" box="[555,617,548,573]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFD9AFDC83750E84D" bold="true" box="[555,617,548,573]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
reaches larger sizes than
|
||
<taxonomicName id="4C1531CAFFD6FFFBFC36FDC9311DE84C" ID-CoL="6FXL6" box="[903,1060,548,572]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="apiculatum">
|
||
<emphasis id="B961965BFFD6FFFBFC36FDC9311DE84C" box="[903,1060,548,572]" italics="true" pageId="12" pageNumber="53">E. apiculatum</emphasis>
|
||
</taxonomicName>
|
||
, but smaller than
|
||
<emphasis id="B961965BFFD6FFFBFB5CFDC8349FE810" italics="true" pageId="12" pageNumber="53">
|
||
<taxonomicName id="4C1531CAFFD6FFFBFB5CFDC8342CE810" ID-CoL="53HFC" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="costulatum">Surrepifungium costulatum</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFE94FDA4349FE810" ID-CoL="7B5PF" box="[293,422,584,608]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="oliverioi">S. oliverioi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C1531CAFFD6FFFBFE59FDA43783E810" ID-CoL="9G6CJ" box="[488,698,584,608]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="patamakanthini">
|
||
<emphasis id="B961965BFFD6FFFBFE59FDA43783E810" box="[488,698,584,608]" italics="true" pageId="12" pageNumber="53">S. patamakanthini</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CEDE7ACFFD6FFFBFD7FFDAB3601E82F" box="[718,824,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="53" unit="mm" value="11.5">11.5 mm</quantity>
|
||
<emphasis id="B961965BFFD6FFFBFCF4FDA53665E810" box="[837,860,585,608]" italics="true" pageId="12" pageNumber="53">vs</emphasis>
|
||
.
|
||
<quantity id="4CEDE7ACFFD6FFFBFCC1FDAB36F5E82F" box="[880,972,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="53" unit="mm" value="4.5">4.5 mm</quantity>
|
||
<emphasis id="B961965BFFD6FFFBFC69FDA536D6E810" box="[984,1007,585,608]" italics="true" pageId="12" pageNumber="53">vs</emphasis>
|
||
. 44.0 mm, 20.0 mm and
|
||
<quantity id="4CEDE7ACFFD6FFFBFA9DFDA430AEE82F" box="[1324,1431,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="12" pageNumber="53" unit="mm" value="22.8">22.8 mm</quantity>
|
||
, respectively); but even when comparing similarly-sized specimens,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFC28FD81315BE8F4" ID-CoL="6FXB6" box="[921,1122,620,644]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFC28FD81315BE8F4" box="[921,1122,620,644]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFBDBFD803193E8F5" box="[1130,1194,620,645]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFBDBFD803193E8F5" bold="true" box="[1130,1194,620,645]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shows a consistently more narrowly-elongated outline than the other four species, which are widely conical all along their ontogeny. In addition,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFEB0FD5934C5E8BC" ID-CoL="6FXL6" box="[257,508,692,716]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="apiculatum">
|
||
<emphasis id="B961965BFFD6FFFBFEB0FD5934C5E8BC" box="[257,508,692,716]" italics="true" pageId="12" pageNumber="53">Epitonium apiculatum</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C1531CAFFD6FFFBFD22FD593662E8BC" ID-CoL="6FXB6" box="[659,859,692,716]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFD22FD593662E8BC" box="[659,859,692,716]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFCD2FD58369BE8BD" box="[867,930,692,717]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFCD2FD58369BE8BD" bold="true" box="[867,930,692,717]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by having a greater number of whorls at the same size (9
|
||
<emphasis id="B961965BFFD6FFFBFE9BFD34346DE880" box="[298,340,728,752]" italics="true" pageId="12" pageNumber="53">fide</emphasis>
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFEEAFD3B3763E880" author="Clench" box="[347,602,727,752]" pageId="12" pageNumber="53" refString="Clench, W. J. & Turner R. D. (1952) The genera Epitonium (Part 2), Depressiscala, Cylindriscala, Nystiella and Solutiscala in the western Atlantic. Johnsonia, 2, 289 - 356." type="journal article" year="1952">
|
||
Clench & Turner
|
||
<quantity id="4CEDE7ACFFD6FFFBFD93FD3B3763E880" box="[546,602,727,752]" metricMagnitude="1" metricUnit="m" metricValue="4.95808" pageId="12" pageNumber="53" unit="in" value="1952.0">1952</quantity>
|
||
</bibRefCitation>
|
||
in
|
||
<quantity id="4CEDE7ACFFD6FFFBFDCEFD3B37ECE89F" box="[639,725,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="53" unit="mm" value="4.5">4.5 mm</quantity>
|
||
long specimens
|
||
<emphasis id="B961965BFFD6FFFBFC23FD353689E880" box="[914,944,729,752]" italics="true" pageId="12" pageNumber="53">vs.</emphasis>
|
||
7 whorls in
|
||
<taxonomicName id="4C1531CAFFD6FFFBFB8FFD35303FE880" ID-CoL="6FXB6" box="[1086,1286,728,752]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum">
|
||
<emphasis id="B961965BFFD6FFFBFB8FFD35303FE880" box="[1086,1286,728,752]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
); fewer axial ribs on the fourth and fifth teleoconch whorl (10–11
|
||
<emphasis id="B961965BFFD6FFFBFD5DFD113633E964" box="[748,778,765,788]" italics="true" pageId="12" pageNumber="53">vs.</emphasis>
|
||
<quantity id="4CEDE7ACFFD6FFFBFCA3FD10364FE965" box="[786,886,764,789]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="5.08" metricValueMin="3.556" pageId="12" pageNumber="53" unit="in" value="17.0" valueMax="20.0" valueMin="14.0">14–20 in</quantity>
|
||
<taxonomicName id="4C1531CAFFD6FFFBFCCCFD11317CE964" ID-CoL="6FXB6" box="[893,1093,764,788]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum">
|
||
<emphasis id="B961965BFFD6FFFBFCCCFD11317CE964" box="[893,1093,764,788]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
) and the whorls only attached by axial ribs (
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFE89FCF3377DE948" author="Clench" box="[312,580,799,824]" pageId="12" pageNumber="53" refString="Clench, W. J. & Turner R. D. (1952) The genera Epitonium (Part 2), Depressiscala, Cylindriscala, Nystiella and Solutiscala in the western Atlantic. Johnsonia, 2, 289 - 356." type="journal article" year="1952">Clench & Turner 1952</bibRefCitation>
|
||
). Additional characters for distinguishing
|
||
<taxonomicName id="4C1531CAFFD6FFFBFB9BFCCC3069E948" ID-CoL="6FXB6" box="[1066,1360,800,824]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFB9BFCCC3069E948" box="[1066,1360,800,824]" italics="true" pageId="12" pageNumber="53">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFAEBFCF330A5E948" box="[1370,1436,799,824]" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFAEBFCF330A5E948" bold="true" box="[1370,1436,799,824]" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from
|
||
<emphasis id="B961965BFFD6FFFBFF65FCA837B6E92C" box="[212,655,836,860]" italics="true" pageId="12" pageNumber="53">
|
||
<taxonomicName id="4C1531CAFFD6FFFBFF65FCA83730E92C" ID-CoL="53HFC" box="[212,521,836,860]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="costulatum">Surrepifungium costulatum</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFDA5FCA837B6E92C" ID-CoL="7B5PF" box="[532,655,836,860]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="oliverioi">S. oliverioi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C1531CAFFD6FFFBFD77FCA836AAE92C" ID-CoL="9G6CJ" box="[710,915,836,860]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="patamakanthini">
|
||
<emphasis id="B961965BFFD6FFFBFD77FCA836AAE92C" box="[710,915,836,860]" italics="true" pageId="12" pageNumber="53">S. patamakanthini</emphasis>
|
||
</taxonomicName>
|
||
come from the radula and jaw: the latter three species have all radular teeth with a single denticle at the cusp (the apical denticle), and the anterior margin of the jaw with several rows of acute, slender denticles. In fact, these two characters were used by
|
||
<bibRefCitation id="EF8437B8FFD6FFFBFB5CFC60344FE9B8" author="Gittenberger" pageId="12" pageNumber="53" refString="Gittenberger, A. & Gittenberger, E. (2005) A hitherto unnoticed adaptive radiation: epitoniid species (Gastropoda: Epitoniidae) associated with corals. Contributions to Zoology, 74, 125 - 203." type="journal article" year="2005">Gittenberger & Gittenberger (2005)</bibRefCitation>
|
||
to reunite these species under the genus
|
||
<taxonomicName id="4C1531CAFFD6FFFBFC8FFC5C36D4E9B8" ID-CoL="7PHTG" box="[830,1005,944,968]" class="Gastropoda" family="Epitoniidae" genus="Surrepifungium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sorbeoconcha" pageId="12" pageNumber="53" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B961965BFFD6FFFBFC8FFC5C36D4E9B8" box="[830,1005,944,968]" italics="true" pageId="12" pageNumber="53">Surrepifungium</emphasis>
|
||
</taxonomicName>
|
||
. On the contrary,
|
||
<taxonomicName id="4C1531CAFFD6FFFBFB06FC5D3047E9B8" ID-CoL="6FXB6" box="[1207,1406,944,968]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD6FFFBFB06FC5D3047E9B8" box="[1207,1406,944,968]" italics="true" pageId="12" pageNumber="53">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD6FFFBFA34FC433581E99D" pageId="12" pageNumber="53" rank="species">
|
||
<emphasis id="B961965BFFD6FFFBFA34FC433581E99D" bold="true" pageId="12" pageNumber="53">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has the anterior margin of the jaw with a single row of blunt denticles, and the teeth of the central field of the radula with three denticles. The radula and jaw of
|
||
<taxonomicName id="4C1531CAFFD6FFFBFD7BFC1436FAEE60" ID-CoL="6FXL6" box="[714,963,1016,1040]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="apiculatum">
|
||
<emphasis id="B961965BFFD6FFFBFD7BFC1436FAEE60" box="[714,963,1016,1040]" italics="true" pageId="12" pageNumber="53">Epitonium apiculatum</emphasis>
|
||
</taxonomicName>
|
||
remain unknown.
|
||
</paragraph>
|
||
<caption id="DF6A1AC1FFD6FFFBFF26F88436E0EDCC" httpUri="https://zenodo.org/record/828308/files/figure.png" pageId="12" pageNumber="53" targetBox="[151,1436,1093,1875]" targetPageId="12">
|
||
<paragraph id="8BAA4A49FFD6FFFBFF26F88436E0EDCC" blockId="12.[151,1436,1896,1980]" pageId="12" pageNumber="53">
|
||
<emphasis id="B961965BFFD6FFFBFF26F884342CED0E" bold="true" box="[151,277,1896,1918]" pageId="12" pageNumber="53">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C1531CAFFD6FFFBFEACF885371EED0E" ID-CoL="6FXB6" box="[285,551,1896,1918]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="53" phylum="Mollusca" rank="species" species="evanidstriatum">
|
||
<emphasis id="B961965BFFD6FFFBFEACF885371EED0E" box="[285,551,1896,1918]" italics="true" pageId="12" pageNumber="53">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
, radula and jaw. Paratype (same specimen as Fig. 6H, I, L: MACN-In 40382). A–C. Radula: A. Central field. B. Outer teeth. C. Detail of cusps of intermediate teeth. D, E. Jaw. D. Anterior inner surface; E. Detail of marginal denticles. Scale bars: A = 10 µm; B = 20 µm; C, D = 5 µm; E = 1 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAA4A49FFD7FFFAFF76FF7B343BE868" blockId="13.[151,1436,151,752]" pageId="13" pageNumber="54">
|
||
Specimens of
|
||
<taxonomicName id="4C1531CAFFD7FFFAFEC5FF743799EAC0" ID-CoL="6FXB6" box="[372,672,152,176]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFEC5FF743799EAC0" box="[372,672,152,176]" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFD1FFF7B37CDEAC0" box="[686,756,151,176]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFD1FFF7B37CDEAC0" bold="true" box="[686,756,151,176]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
could be confused with the sympatric
|
||
<taxonomicName id="4C1531CAFFD7FFFAFB6BFF7530ACEADF" ID-CoL="6FXC4" box="[1242,1429,152,176]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFB6BFF7530ACEADF" box="[1242,1429,152,176]" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
, particularly if seen under low magnification. However, the former clearly differs by having spiral sculpture, and the adjacent whorls in tight contact instead appearing as “detached”. In addition, the protoconch of
|
||
<taxonomicName id="4C1531CAFFD7FFFAFA96FF0C3406EB6C" ID-CoL="6FXB6" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFA96FF0C3406EB6C" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFEF6FEE834B1EB6D" box="[327,392,260,285]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFEF6FEE834B1EB6D" bold="true" box="[327,392,260,285]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is smaller than that of
|
||
<taxonomicName id="4C1531CAFFD7FFFAFD25FEE93671EB6C" ID-CoL="6FXC4" box="[660,840,261,284]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFD25FEE93671EB6C" box="[660,840,261,284]" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
(300
|
||
<emphasis id="B961965BFFD7FFFAFC3CFEE9369DEB6C" box="[909,932,261,284]" italics="true" pageId="13" pageNumber="54">vs</emphasis>
|
||
. 500 µm long), even when having a greater number of whorls (3–3¼
|
||
<emphasis id="B961965BFFD7FFFAFE09FEC534F6EB30" box="[440,463,297,320]" italics="true" pageId="13" pageNumber="54">vs</emphasis>
|
||
. 2½); the number of axial ribs in the teleoconch of
|
||
<taxonomicName id="4C1531CAFFD7FFFAFBA8FEC531D8EB30" ID-CoL="6FXB6" box="[1049,1249,296,320]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFBA8FEC531D8EB30" box="[1049,1249,296,320]" italics="true" pageId="13" pageNumber="54">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFB58FECB3011EB30" box="[1257,1320,295,320]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFB58FECB3011EB30" bold="true" box="[1257,1320,295,320]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
decreases with growth, instead of being constant, as it is the case of
|
||
<taxonomicName id="4C1531CAFFD7FFFAFC82FEA136D1EB14" ID-CoL="6FXC4" box="[819,1000,333,356]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFC82FEA136D1EB14" box="[819,1000,333,356]" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
; and the first teleoconch whorl shows almost twice the number of ribs than those present in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFD5FFE9D369BEBF7" ID-CoL="6FXC4" box="[750,930,368,392]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFD5FFE9D369BEBF7" box="[750,930,368,392]" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
(18–27
|
||
<emphasis id="B961965BFFD7FFFAFC4EFE9D312FEBF8" box="[1023,1046,369,392]" italics="true" pageId="13" pageNumber="54">vs</emphasis>
|
||
. 12–15). Another difference arises in the morphology of the operculum, which is paucispiral in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFCF2FE79315EEBDC" ID-CoL="6FXB6" box="[835,1127,404,428]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFCF2FE79315EEBDC" box="[835,1127,404,428]" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFBDFFE783194EBDD" box="[1134,1197,404,429]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFBDFFE783194EBDD" bold="true" box="[1134,1197,404,429]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and multispiral in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFA35FE793412EBBF" ID-CoL="6FXC4" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFA35FE793412EBBF" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
. Concerning the radula,
|
||
<taxonomicName id="4C1531CAFFD7FFFAFDF9FE543654EBA0" ID-CoL="6FXB6" box="[584,877,440,464]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFDF9FE543654EBA0" box="[584,877,440,464]" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFCC7FE5B368EEBA0" box="[886,951,439,464]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFCC7FE5B368EEBA0" bold="true" box="[886,951,439,464]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shows the outermost teeth with minute or absent secondary denticles, whereas in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFDC0FE313612EB84" ID-CoL="6FXC4" box="[625,811,477,500]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="georgettinum">
|
||
<emphasis id="B961965BFFD7FFFAFDC0FE313612EB84" box="[625,811,477,500]" italics="true" pageId="13" pageNumber="54">E. georgettinum</emphasis>
|
||
</taxonomicName>
|
||
the outer teeth have two well-developed secondary denticles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F19C2FFD7FFFAFF76FDC831CBE880" pageId="13" pageNumber="54" type="description">
|
||
<paragraph id="8BAA4A49FFD7FFFAFF76FDC831CBE880" blockId="13.[151,1436,151,752]" pageId="13" pageNumber="54">
|
||
The only other species sympatric with
|
||
<taxonomicName id="4C1531CAFFD7FFFAFD3BFDC9366DE84C" box="[650,852,548,572]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFD3BFDC9366DE84C" box="[650,852,548,572]" italics="true" pageId="13" pageNumber="54">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFCEFFDC83699E84D" box="[862,928,548,573]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFCEFFDC83699E84D" bold="true" box="[862,928,548,573]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
having both axial and spiral sculpture is
|
||
<taxonomicName id="4C1531CAFFD7FFFAFA35FDC93429E810" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="striatellum">
|
||
<emphasis id="B961965BFFD7FFFAFA35FDC93429E810" italics="true" pageId="13" pageNumber="54">E. striatellum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C1531CAFFD7FFFAFE93FDA43770E810" box="[290,585,584,608]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFE93FDA43770E810" box="[290,585,584,608]" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFDE5FDAB37AFE810" box="[596,662,583,608]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFDE5FDAB37AFE810" bold="true" box="[596,662,583,608]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from this species by having a deeper suture, and the axial sculpture comprises fewer, more widely separated, higher and stronger ribs.
|
||
<taxonomicName id="4C1531CAFFD7FFFAFC48FD813027E8F4" box="[1017,1310,620,644]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFC48FD813027E8F4" box="[1017,1310,620,644]" italics="true" pageId="13" pageNumber="54">Epitonium evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFA97FD80305FE8F5" box="[1318,1382,620,645]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFA97FD80305FE8F5" bold="true" box="[1318,1382,620,645]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
also has weaker and fewer spiral threads, which fade in the last whorls, whereas in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFBBDFD7D319DE8D8" box="[1036,1188,656,680]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="striatellum">
|
||
<emphasis id="B961965BFFD7FFFAFBBDFD7D319DE8D8" box="[1036,1188,656,680]" italics="true" pageId="13" pageNumber="54">E. striatellum</emphasis>
|
||
</taxonomicName>
|
||
the spiral sculpture is regularly present along all the shell whorls. The aperture in
|
||
<taxonomicName id="4C1531CAFFD7FFFAFC8EFD5936E1E8BC" box="[831,984,692,716]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="striatellum">
|
||
<emphasis id="B961965BFFD7FFFAFC8EFD5936E1E8BC" box="[831,984,692,716]" italics="true" pageId="13" pageNumber="54">E. striatellum</emphasis>
|
||
</taxonomicName>
|
||
is larger and more markedly ovate, and this species reaches larger sizes than
|
||
<taxonomicName id="4C1531CAFFD7FFFAFD84FD3537C5E880" box="[565,764,728,752]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="evanidstriatum" status="sp. nov.">
|
||
<emphasis id="B961965BFFD7FFFAFD84FD3537C5E880" box="[565,764,728,752]" italics="true" pageId="13" pageNumber="54">E. evanidstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2522B20FFD7FFFAFCB5FD3B367BE880" box="[772,834,727,752]" pageId="13" pageNumber="54" rank="species">
|
||
<emphasis id="B961965BFFD7FFFAFCB5FD3B367BE880" bold="true" box="[772,834,727,752]" pageId="13" pageNumber="54">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(28.5
|
||
<emphasis id="B961965BFFD7FFFAFC3AFD353690E880" box="[907,937,729,752]" italics="true" pageId="13" pageNumber="54">vs.</emphasis>
|
||
<quantity id="4CEDE7ACFFD7FFFAFC01FD3B312AE89F" box="[944,1043,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="13" pageNumber="54" unit="mm" value="11.5">11.5 mm</quantity>
|
||
long, respectively).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |