treatments-xml/data/03/9D/87/039D87EF6F7CFF920CC0FE85FDCFFBB7.xml

<document id="BBFA9CAB4975DE35461457E56E2B5996" ID-CLB-Dataset="21959" ID-DOI="10.5852/ejt.2016.198" ID-GBIF-Dataset="66bc6169-67f1-46ca-ba7d-08326ec09cf3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838140" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590072533401" checkinUser="valdenar" docAuthor="Constant, Jérôme &amp; Pham, Hong-Thai" docDate="2016" docId="039D87EF6F7CFF920CC0FE85FDCFFBB7" docLanguage="en" docName="ejt-2016-198.pdf.imf" docOrigin="European Journal of Taxonomy 198" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gergithus (Maculergithus) tamdao Constant &amp; Pham 2016, subgen. et sp. nov." docType="treatment" docUuid="4368D0A6-3C1D-42EB-91B2-59769EE60780" docUuidSource="ZooBank" docVersion="8" lastPageNumber="14" masterDocId="FFA4FF976F76FF9F0D17FFFFFFFEFFA2" masterDocTitle="Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="11" updateTime="1698840676411" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A7A9363CDF743DFD29C5CB20D6DDECCE">Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:namePart id="F67F87F4EB58E07C5CB7CAE57906D614">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="08D3D8DED63CF694BB83D9ED57EB1EB6">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier street 29, B- 1000 Brussels, Belgium. &amp; urn: lsid: zoobank. org: author: 6 E 6072 A 1 - 9415 - 4 C 8 D- 8 E 60 - 2504444 DB 290 &amp; Corresponding author: jerome. constant @ naturalsciences. be</mods:affiliation>
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<mods:namePart id="DAF7BF53E2E265766C02F26E0CE34617">Pham, Hong-Thai</mods:namePart>
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<mods:affiliation id="3C82B3E1936C71FA6E26320BA3A71136">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. &amp; Email: phamthai @ vnmn. vast. vn &amp; urn: lsid: zoobank. org: author: E 34 CB 863 - 7 E 3 B- 4 E 8 F- 8738 - B 41 C 07 D 9 F 5 F 9</mods:affiliation>
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<treatment id="039D87EF6F7CFF920CC0FE85FDCFFBB7" ID-DOI="http://doi.org/10.5281/zenodo.3852322" ID-GBIF-Taxon="164246218" ID-Zenodo-Dep="3852322" LSID="urn:lsid:zoobank.org:act:4368D0A6-3C1D-42EB-91B2-59769EE60780" httpUri="http://treatment.plazi.org/id/039D87EF6F7CFF920CC0FE85FDCFFBB7" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<taxonomicName id="4C344D7A6F7CFF950CC0FE85FC83FE37" ID-CoL="3FWF4" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" box="[471,893,378,405]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tamdao" status="subgen. et sp. nov." subGenus="Maculergithus">
<emphasis id="B940EAEB6F7CFF950CC0FE85FC83FE37" bold="true" box="[471,893,378,405]" italics="true" pageId="10" pageNumber="11">Gergithus (Maculergithus) tamdao</emphasis>
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<paragraph id="8B8B36F96F7CFF950FBAFE3EFC78FE7E" blockId="10.[371,1216,378,476]" box="[685,902,449,476]" pageId="10" pageNumber="11">
<figureCitation id="130F2A7C6F7CFF950FBAFE3EFD08FE79" box="[685,758,449,475]" captionStart="Fig" captionStartId="3.[189,232,1962,1988]" captionTargetBox="[190,1395,1269,1919]" captionTargetId="figure@3.[189,1398,1268,1919]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Gergithus (Maculergithus) subgen. nov. in North Vietnam and Hainan." figureDoi="http://doi.org/10.5281/zenodo.3838142" httpUri="https://zenodo.org/record/3838142/files/figure.png" pageId="10" pageNumber="11">Figs 1</figureCitation>
, 
<figureCitation id="130F2A7C6F7CFF950E13FE3EFCECFE79" box="[772,786,449,475]" captionStart="Fig" captionStartId="4.[189,232,1858,1884]" captionTargetBox="[328,1285,269,1823]" captionTargetId="figure@4.[265,1316,265,1834]" captionTargetPageId="4" captionText="Fig. 2. Gergithus (Maculergithus) subgen. nov., schema of maculation of frons and left tegmen. A–B. Gergithus (Maculergithus) luteomaculatus subgen. et sp. nov. C–D. Gergithus (Maculergithus) multipunctatus Che, Zhang &amp; Wang, 2007. E–F. Gergithus (Maculergithus) nonomaculatus Meng &amp; Wang, 2012. G–H. Gergithus (Maculergithus) tamdao subgen. et sp. nov. I–J. Gergithus (Maculergithus) sp. (drawings by J. Caudron, RBINS; C–F, I–J based on illustrations of original descriptions)." figureDoi="http://doi.org/10.5281/zenodo.3838144" httpUri="https://zenodo.org/record/3838144/files/figure.png" pageId="10" pageNumber="11">2</figureCitation>
G–H, 5–6
</paragraph>
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<paragraph id="8B8B36F96F7CFF950DAAFDFDFEC8FDBE" blockId="10.[189,310,514,540]" box="[189,310,514,540]" pageId="10" pageNumber="11">
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<emphasis id="B940EAEB6F7CFF950DAAFDFDFEC8FDBE" bold="true" box="[189,310,514,540]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAFDCEFDECFD30" blockId="10.[189,1399,561,658]" pageId="10" pageNumber="11">The species can be recognized on the following combination of characters: (1) frons with a yellow spot along dorsal margin; (2) vertex yellow; (3) 3 yellow spots on tegmina along sutural margin; (4) tegmina with 10 yellow spots in total.</paragraph>
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<emphasis id="B940EAEB6F7CFF950DAAFD46FEBAFD71" bold="true" box="[189,324,697,723]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
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The species epithet refers to the 
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location, Tam Dao National Park in North 
<collectingCountry id="F32376696F7CFF950971FD17FB33FCA0" box="[1126,1229,744,770]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
. It is used as a noun in apposition.
</paragraph>
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<heading id="D0C381956F7CFF950DAAFCB3FE92FCC4" bold="true" box="[189,364,844,870]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B940EAEB6F7CFF950DAAFCB3FE92FCC4" bold="true" box="[189,364,844,870]" pageId="10" pageNumber="11">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B5C3CA46F7CFF950DFDFC84FB7BFC79" ID-GBIF-Occurrence="2621718301" collectingDate="2011-07-25" collectingDateMax="2011-07-30" collectingDateMin="2011-07-25" collectionCode="R, RBINS" collectorName="Coll. I. &amp; Sc. N. B. &amp; Tam Dao N. P. &amp; Leg. J. Constant &amp; J. Bresseel" country="Vietnam" latitude="21.516666" location="Vietnam" longLatPrecision="1264" longitude="105.55" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548F885B6F7CFF950DFDFC84FEA2FC37" box="[234,348,891,917]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
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<collectingCountry id="F32376696F7CFF950DAAFC61FEB7FC1A" box="[189,329,926,952]" name="Vietnam" pageId="10" pageNumber="11">VIETNAM</collectingCountry>
: Ƌ (dissected, 
<figureCitation id="130F2A7C6F7CFF950CEFFC61FDBEFC1B" box="[504,576,926,953]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
): [
<collectorName id="26C1532F6F7CFF950F77FC61FD4AFC1A" box="[608,692,926,952]" pageId="10" pageNumber="11">Coll. I.</collectorName>
<collectionCode id="ED25AE3C6F7CFF950FA3FC61FD32FC1A" box="[692,716,926,952]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="11">R</collectionCode>
. 
<collectorName id="26C1532F6F7CFF950FC7FC61FCD1FC1A" box="[720,815,926,952]" pageId="10" pageNumber="11">Sc.N.B.</collectorName>
, 
<location id="8EEB60226F7CFF950E2BFC61FC5CFC1A" LSID="urn:lsid:plazi:treatment:039D87EF6F7CFF920CC0FE85FDCFFBB7:8EEB60226F7CFF950E2BFC61FC5CFC1A" box="[828,930,926,952]" country="Vietnam" latitude="21.516666" longLatPrecision="1264" longitude="105.55" name="Vietnam" pageId="10" pageNumber="11">Vietnam</location>
, 
<collectorName id="26C1532F6F7CFF950EB8FC60FBA6FC1B" box="[943,1112,927,953]" pageId="10" pageNumber="11">Tam Dao N.P.</collectorName>
, 
<geoCoordinate id="EE00503E6F7CFF950972FC61FB38FC1A" box="[1125,1222,926,952]" degrees="21" direction="north" minutes="31" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="21.516666">21°31'N</geoCoordinate>
<geoCoordinate id="EE00503E6F7CFF9509D9FC61FAC3FC1B" box="[1230,1341,926,953]" degrees="105" direction="east" minutes="33" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="105.55">105°33'E</geoCoordinate>
, 
<date id="FF8A10396F7CFF95085DFC61FEBAFC79" pageId="10" pageNumber="11" value="2011-07-25" valueMax="2011-07-30" valueMin="2011-07-25">
<collectingDate id="EFCEE9D16F7CFF95085DFC61FEBAFC79" pageId="10" pageNumber="11" value="2011-07-25" valueMax="2011-07-30" valueMin="2011-07-25">25– 30.vii.2011</collectingDate>
</date>
, day collecting, 
<collectorName id="26C1532F6F7CFF950F13FC3DFD3DFC79" box="[516,707,961,988]" pageId="10" pageNumber="11">Leg. J. Constant</collectorName>
&amp; 
<collectorName id="26C1532F6F7CFF950FFFFC3DFC96FC79" box="[744,872,961,988]" pageId="10" pageNumber="11">J. Bresseel</collectorName>
, I.G.: 31.993] (
<collectionCode id="ED25AE3C6F7CFF95090BFC3EFB87FC79" box="[1052,1145,961,987]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="10" pageNumber="11" type="Museum">RBINS</collectionCode>
).
</paragraph>
</materialsCitation>
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<emphasis id="B940EAEB6F7CFF950DFDFBFCFEA2FBBF" bold="true" box="[234,348,1027,1053]" pageId="10" pageNumber="11">
<typeStatus id="548F885B6F7CFF950DFDFBFCFEA2FBBF" box="[234,348,1027,1053]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAFBD9FCF0FBE2" blockId="10.[189,782,1027,1088]" box="[189,782,1062,1088]" pageId="10" pageNumber="11">
<collectingCountry id="F32376696F7CFF950DAAFBD9FEB7FBE2" box="[189,329,1062,1088]" name="Vietnam" pageId="10" pageNumber="11">VIETNAM</collectingCountry>
: 
<specimenCount id="9D32FD706F7CFF950C43FBD9FE7CFBE2" box="[340,386,1062,1088]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
: same data as 
<typeStatus id="548F885B6F7CFF950F3CFBD9FD6CFBE2" box="[555,658,1062,1088]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED25AE3C6F7CFF950FB6FBD9FCFFFBE2" box="[673,769,1062,1088]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="10" pageNumber="11">VNMN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65726F7CFF920DAAFB98FC43FCB1" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
<paragraph id="8B8B36F96F7CFF950DAAFB98FEB0FB23" blockId="10.[189,334,1127,1153]" box="[189,334,1127,1153]" pageId="10" pageNumber="11">
<heading id="D0C381956F7CFF950DAAFB98FEB0FB23" bold="true" box="[189,334,1127,1153]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B940EAEB6F7CFF950DAAFB98FEB0FB23" bold="true" box="[189,334,1127,1153]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAFB69FE7DFB71" blockId="10.[189,1399,1174,1235]" pageId="10" pageNumber="11">
MEASUREMENTS AND RATIOS. LT: Ƌ (n = 1): 6.0 mm; 
<specimenCount id="9D32FD706F7CFF950E10FB69FCE3FB12" box="[775,797,1174,1200]" pageId="10" pageNumber="11" type="female">♀</specimenCount>
(n = 1): 
<quantity id="4CCC9B1C6F7CFF950E97FB69FC25FB12" box="[896,987,1174,1201]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="11" unit="mm" value="6.5">6.5 mm</quantity>
. Ratio BV/LV = 4.0; LF/BF = 0.84; LTg/BTg = 1.73.
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAFB04FED6FA4A" blockId="10.[189,1399,1274,1512]" pageId="10" pageNumber="11">
HEAD. Vertex slightly excavate, bright yellow (
<figureCitation id="130F2A7C6F7CFF950FF3FB05FCC5FAB7" box="[740,827,1274,1301]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig.5A</figureCitation>
). Frons smooth and convex, largely visible from above, black-brown with broad transverse bright yellow band slightly above clypeo-frontal suture, and rounded bright yellow spot along dorsal margin, not reaching dorsolateral angles (
<figureCitation id="130F2A7C6F7CFF950986FABEFB26FAFE" box="[1169,1240,1345,1372]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
A–D). Genae black-brown, bright yellow under antennae (
<figureCitation id="130F2A7C6F7CFF950FC0FA9BFCC9FADD" box="[727,823,1380,1407]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
). Clypeus smooth, slightly carinate subapically, black-brown (
<figureCitation id="130F2A7C6F7CFF950C72FA78FE12FA00" box="[357,492,1415,1442]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5B, D</figureCitation>
). Labium dark brown, reaching median coxae, with last segment narrower and shorter than penultimate. Antennae black-brown. Scape very short; pedicel subglobulose, slightly elongate.
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAF9EFFEEAF914" blockId="10.[189,1399,1551,1719]" pageId="10" pageNumber="11">
THORAX. Pronotum minutely punctured, with anterior margin slightly carinate behind head and 2 impressed points in middle; entirely black-brown (
<figureCitation id="130F2A7C6F7CFF950E0DF9CDFC60F9EE" box="[794,926,1586,1613]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5A, C</figureCitation>
). Mesonotum minutely punctured, with transverse carina along anterior margin; black-brown with round bright yellow spot on each side before lateral angle (
<figureCitation id="130F2A7C6F7CFF950C77F986FE3EF931" box="[352,448,1656,1683]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Scutellum slightly wrinkled transversely (
<figureCitation id="130F2A7C6F7CFF950ED0F986FBD9F931" box="[967,1063,1656,1683]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Tegulae black-brown (
<figureCitation id="130F2A7C6F7CFF950850F986FEF8F914" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5A, C</figureCitation>
).
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAF921FB06F89C" blockId="10.[189,1398,1757,1854]" pageId="10" pageNumber="11">
TEGMINA. (
<figureCitation id="130F2A7C6F7CFF950C2BF922FE3DF955" box="[316,451,1757,1784]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="10" pageNumber="11">Fig. 5A, C</figureCitation>
) Black-brown, elongate, minutely punctured, with external margin and apical margins rounded. Reticulum of veins very slightly visible. Ten bright yellow spots organized, from base to apex: 1, 2, 1, 2, 1, 2, 1, with 4 spots along costal margin and 3 spots along sutural margin.
</paragraph>
<paragraph id="8B8B36F96F7CFF950DAAF899FBCFF867" blockId="10.[189,1398,1893,1990]" pageId="10" pageNumber="11">HIND WINGS. Brown, unilobed, with veins darker, slightly shorter than tegmina. Venation reticulate with main longitudinal veins distinct basally and numerous cross-veinlets. Costal margin strongly sinuate; sutural margin angularly rounded at mid-length and apical margin rounded.</paragraph>
<paragraph id="8B8B36F96F7DFF940DAAFEF3FEFFFE10" blockId="11.[189,1399,267,434]" pageId="11" pageNumber="12">
LEGS. (
<figureCitation id="130F2A7C6F7DFF940C1BFEF4FEAAFE84" box="[268,340,267,294]" captionStart="Fig" captionStartId="11.[189,232,1929,1955]" captionTargetBox="[189,1392,550,1882]" captionTargetId="figure@11.[189,1398,542,1882]" captionTargetPageId="11" captionText="Fig. 5. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, Ƌ, total length: 6.0 mm. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, frontal view. E. Label." figureDoi="http://doi.org/10.5281/zenodo.3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="11" pageNumber="12">Figs 5</figureCitation>
A–D) Elongate and slender. Pro- and mesocoxae black; metacoxae yellow-brown. Femora yellow-brown. Protibiae yellow-brown with anterodorsal and anteroventral carinae black; mesotibiae yellow-brown with externoventral carina black; metatibiae yellow-brown with 2 lateral spines near apex and 6 apical spines. Protarsi yellow-brown; mesotarsi and metatarsi yellow. Metatibotarsal formula: (2) 6/7/2.
</paragraph>
<caption id="DF4B66716F7DFF940DAAF876FED4F848" ID-DOI="http://doi.org/10.5281/zenodo.3838150" ID-Zenodo-Dep="3838150" httpUri="https://zenodo.org/record/3838150/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1929,1955]" targetBox="[189,1392,550,1882]" targetPageId="11">
<paragraph id="8B8B36F96F7DFF940DAAF876FED4F848" blockId="11.[189,1399,1929,2026]" pageId="11" pageNumber="12">
<emphasis id="B940EAEB6F7DFF940DAAF876FEEEF806" bold="true" box="[189,272,1929,1956]" pageId="11" pageNumber="12">Fig. 5.</emphasis>
<taxonomicName id="4C344D7A6F7DFF940C0AF876FD3FF801" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" box="[285,705,1929,1955]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tamdao" status="subgen. et sp. nov." subGenus="Maculergithus">
<emphasis id="B940EAEB6F7DFF940C0AF876FD3FF801" box="[285,705,1929,1955]" italics="true" pageId="11" pageNumber="12">Gergithus (Maculergithus) tamdao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357906F7DFF940FD9F876FC44F806" box="[718,954,1929,1956]" pageId="11" pageNumber="12" rank="species">subgen. et sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, total length: 6.0 mm. 
<emphasis id="B940EAEB6F7DFF940DAAF853FF2CF864" bold="true" box="[189,210,1964,1990]" pageId="11" pageNumber="12">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B940EAEB6F7DFF940CFDF852FE00F865" bold="true" box="[490,510,1965,1991]" pageId="11" pageNumber="12">B</emphasis>
. Habitus, ventral view. 
<emphasis id="B940EAEB6F7DFF940E08F853FCCAF864" bold="true" box="[799,820,1964,1990]" pageId="11" pageNumber="12">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B940EAEB6F7DFF94095AF852FB9CF865" bold="true" box="[1101,1122,1965,1991]" pageId="11" pageNumber="12">D</emphasis>
. Habitus, frontal view. 
<emphasis id="B940EAEB6F7DFF940DAAF82FFF2FF848" bold="true" box="[189,209,2000,2026]" pageId="11" pageNumber="12">E</emphasis>
. Label.
</paragraph>
</caption>
<caption id="DF4B66716F7AFF930DAAF896FBA9F84C" ID-DOI="http://doi.org/10.5281/zenodo.3838152" ID-Zenodo-Dep="3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1897,1923]" targetBox="[201,1382,268,1856]" targetPageId="12">
<paragraph id="8B8B36F96F7AFF930DAAF896FBA9F84C" blockId="12.[189,1399,1897,2030]" pageId="12" pageNumber="13">
<emphasis id="B940EAEB6F7AFF930DAAF896FEF0F821" bold="true" box="[189,270,1897,1923]" pageId="12" pageNumber="13">Fig. 6.</emphasis>
<taxonomicName id="4C344D7A6F7AFF930C0EF896FD46F821" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" box="[281,696,1897,1923]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tamdao" status="subgen. et sp. nov." subGenus="Maculergithus">
<emphasis id="B940EAEB6F7AFF930C0EF896FD46F821" box="[281,696,1897,1923]" italics="true" pageId="12" pageNumber="13">Gergithus (Maculergithus) tamdao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357906F7AFF930FD4F896FC57F826" box="[707,937,1897,1924]" pageId="12" pageNumber="13" rank="species">subgen. et sp. nov.</taxonomicNameLabel>
, holotype, male genitalia. 
<emphasis id="B940EAEB6F7AFF9309FBF896FAFFF821" bold="true" box="[1260,1281,1897,1923]" pageId="12" pageNumber="13">A</emphasis>
. Pygofer, anal tube and gonostyli, left lateral view. 
<emphasis id="B940EAEB6F7AFF930FB3F872FD46F805" bold="true" box="[676,696,1933,1959]" pageId="12" pageNumber="13">B</emphasis>
. Anal tube, dorsal view. 
<emphasis id="B940EAEB6F7AFF930ECAF873FC0CF804" bold="true" box="[989,1010,1932,1958]" pageId="12" pageNumber="13">C</emphasis>
. Left gonostylus, posterior view. 
<emphasis id="B940EAEB6F7AFF930DAAF84FFF2CF868" bold="true" box="[189,210,1968,1994]" pageId="12" pageNumber="13">D</emphasis>
. Aedeagus, postero-ventral view. 
<emphasis id="B940EAEB6F7AFF930F63F84FFD76F868" bold="true" box="[628,648,1968,1994]" pageId="12" pageNumber="13">E</emphasis>
. Aedeagus, left lateral view. 
<emphasis id="B940EAEB6F7AFF930EE3F84FFBF8F868" bold="true" box="[1012,1030,1968,1994]" pageId="12" pageNumber="13">F</emphasis>
. Aedeagus, right lateral view. Abbreviations: 
<emphasis id="B940EAEB6F7AFF930C64F82CFE6AF84F" box="[371,404,2003,2029]" italics="true" pageId="12" pageNumber="13">An</emphasis>
= anal tube; 
<emphasis id="B940EAEB6F7AFF930F3AF82BFDBCF84C" box="[557,578,2004,2030]" italics="true" pageId="12" pageNumber="13">G</emphasis>
= gonostyli; 
<emphasis id="B940EAEB6F7AFF930FC9F82BFD03F84F" box="[734,765,2004,2029]" italics="true" pageId="12" pageNumber="13">Py</emphasis>
= pygofer. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B8B36F96F7BFF920DFDFEF4FE63FE87" blockId="13.[189,1399,267,787]" box="[234,413,267,293]" pageId="13" pageNumber="14">
<emphasis id="B940EAEB6F7BFF920DFDFEF4FE63FE87" bold="true" box="[234,413,267,293]" pageId="13" pageNumber="14">Male genitalia</emphasis>
</paragraph>
<paragraph id="8B8B36F96F7BFF920DAAFED1FC43FCB1" blockId="13.[189,1399,267,787]" pageId="13" pageNumber="14">
Pygofer higher than broad and with posterior margin strongly sinuate and roundly projecting in middle in lateral view, and emarginate dorsally (
<figureCitation id="130F2A7C6F7BFF920FB0FEAEFCF6FECE" box="[679,776,337,364]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6A</figureCitation>
). Gonostyli slightly longer than high in lateral view (
<figureCitation id="130F2A7C6F7BFF920DD1FE8BFEDDFE2D" box="[198,291,372,399]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6A</figureCitation>
) with capitulum well developed; capitulum apically laminate and projecting anterointernally; lateral spine of capitulum acute, hooked and projecting lateroventrally; ventroapical margin of gonostyli rounded in lateral view (
<figureCitation id="130F2A7C6F7BFF920CFBFE44FD8AFE77" box="[492,628,443,469]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6A, C</figureCitation>
). Anal tube 1.5 times longer in median line than broad, dorsoventrally flattened and curved ventrally on posterior half; sides broadly rounded and apex truncate in middle (
<figureCitation id="130F2A7C6F7BFF920C37FDFEFE99FDBE" box="[288,359,513,540]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
A–B); posterolateral angles projecting posteroventrally (
<figureCitation id="130F2A7C6F7BFF920913FDFEFBB5FDBE" box="[1028,1099,513,540]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
A–B). Aedeagus strongly asymmetrical with ventral lobe of phallobase elongate, parallel-sided, and roundly truncate apically (
<figureCitation id="130F2A7C6F7BFF920DD1FDB7FEDBFDC1" box="[198,293,584,611]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6D</figureCitation>
). Lateroapical processes of phallobase elongate and spinose, nearly reaching apex of phallus; right process sinuate internally towards apex (
<figureCitation id="130F2A7C6F7BFF920FC4FD94FCE9FD24" box="[723,791,619,646]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
D–F). Left ventrolateral lobe of phallobase laminate and projecting laterally with 2 pointed processes projecting anterolaterally, the dorsal one much more developed, and another more posterior pointed process projecting laterally (
<figureCitation id="130F2A7C6F7BFF92095CFD4EFB6CFD6E" box="[1099,1170,689,716]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
D–E). Rather long, spinose lateral process at basal half, on each side, projecting dorsoanteriorly (
<figureCitation id="130F2A7C6F7BFF920946FD2BFB68FD4D" box="[1105,1174,724,751]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
D–F). Right lateral lobe of phallobase laminate, slightly concave internally (
<figureCitation id="130F2A7C6F7BFF920E41FD07FC4EFCB1" box="[854,944,760,787]" captionStart="Fig" captionStartId="12.[189,232,1897,1923]" captionTargetBox="[201,1382,268,1856]" captionTargetId="figure@12.[200,1383,265,1858]" captionTargetPageId="12" captionText="Fig. 6. Gergithus (Maculergithus) tamdao subgen. et sp. nov., holotype, male genitalia. A. Pygofer, anal tube and gonostyli, left lateral view. B. Anal tube, dorsal view. C. Left gonostylus, posterior view. D. Aedeagus, postero-ventral view. E. Aedeagus, left lateral view. F. Aedeagus, right lateral view. Abbreviations: An = anal tube; G = gonostyli; Py = pygofer. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838152" httpUri="https://zenodo.org/record/3838152/files/figure.png" pageId="13" pageNumber="14">Fig. 6F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65726F7BFF920DAAFCC6FB5BFC20" box="[189,1189,825,898]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B8B36F96F7BFF920DAAFCC6FEA8FCF1" blockId="13.[189,342,825,851]" box="[189,342,825,851]" pageId="13" pageNumber="14">
<heading id="D0C381956F7BFF920DAAFCC6FEA8FCF1" bold="true" box="[189,342,825,851]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B940EAEB6F7BFF920DAAFCC6FEA8FCF1" bold="true" box="[189,342,825,851]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36F96F7BFF920DAAFC97FB5BFC20" blockId="13.[189,1189,872,898]" box="[189,1189,872,898]" pageId="13" pageNumber="14">
Known from 
<collectingCountry id="F32376696F7BFF920C4CFC97FE3FFC20" box="[347,449,872,898]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
, 
<collectingRegion id="49F0F81B6F7BFF920CDBFC97FD46FC20" box="[460,696,872,898]" country="Vietnam" name="Vinh Phuc" pageId="13" pageNumber="14">Vinh Phuc province</collectingRegion>
, Tam Dao National Park (see map 
<figureCitation id="130F2A7C6F7BFF920947FC97FB6AFC20" box="[1104,1172,872,898]" captionStart="Fig" captionStartId="3.[189,232,1962,1988]" captionTargetBox="[190,1395,1269,1919]" captionTargetId="figure@3.[189,1398,1268,1919]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Gergithus (Maculergithus) subgen. nov. in North Vietnam and Hainan." figureDoi="http://doi.org/10.5281/zenodo.3838142" httpUri="https://zenodo.org/record/3838142/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65726F7BFF920DAAFC56FDCFFBB7" pageId="13" pageNumber="14" type="biology_ecology">
<paragraph id="8B8B36F96F7BFF920DAAFC56FEE2FC61" blockId="13.[189,284,937,963]" box="[189,284,937,963]" pageId="13" pageNumber="14">
<heading id="D0C381956F7BFF920DAAFC56FEE2FC61" bold="true" box="[189,284,937,963]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B940EAEB6F7BFF920DAAFC56FEE2FC61" bold="true" box="[189,284,937,963]" pageId="13" pageNumber="14">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36F96F7BFF920DAAFC27FDCFFBB7" blockId="13.[189,1398,984,1045]" pageId="13" pageNumber="14">
The specimens were collected on lower vegetation, in moist evergreen low mountain forest at an altitude around 
<quantity id="4CCC9B1C6F7BFF920C01FC04FE91FBB7" box="[278,367,1019,1045]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1000.0">1000 m</quantity>
above sea level.
</paragraph>
</subSubSection>
</treatment>
</document>