290 lines
33 KiB
XML
290 lines
33 KiB
XML
<document id="37A5543B5CC2F91544DC22C25234F2F4" ID-DOI="10.11646/zootaxa.3872.3.5" ID-GBIF-Dataset="55ca06ce-1ac4-4f07-ac21-a75476e73536" ID-ISSN="1175-5326" ID-Zenodo-Dep="228806" ID-ZooBank="1B4717AE-9993-4166-BAA2-45DFC907080C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460927688234" checkinUser="plazi" docAuthor="Köhler, Gunther & Kieckbusch, Max" docDate="2014" docId="039D87E1FFF6467A9FDEFD95FC54FAE7" docLanguage="en" docName="zt03872p300.pdf" docOrigin="Zootaxa 3872 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atractus careolepis Köhler & Kieckbusch, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="297" masterDocId="FFA4FF99FFF2467C9F49FFB6FFA1FFEA" masterDocTitle="Two new species of Atractus from Colombia (Reptilia, Squamata, Dipsadidae)" masterLastPageNumber="300" masterPageNumber="291" pageNumber="295" updateTime="1698362779195" updateUser="plazi">
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<mods:mods id="967D100D001CEC166DAD1E2505A5515A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CD9BB855DE94262E9F8D997F830E0591">
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<mods:title id="8639D66CF91A00C6ABAA650196A91919">Two new species of Atractus from Colombia (Reptilia, Squamata, Dipsadidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A32E50FAAEA14896E590233831AFB26E" type="personal">
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<mods:roleTerm id="E9EE3E1CF1CAB12B6F1E780D757D4D42">Author</mods:roleTerm>
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<mods:namePart id="0AA207816F00955F59637F722DA673F8">Köhler, Gunther</mods:namePart>
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<mods:name id="FAEE56961BC96EB71DBFC19B2AF6C62A" type="personal">
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<mods:role id="EB6AB0720A5B1FDB9BFB60A10B0C8557">
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<mods:roleTerm id="1F6982CFA080B5003955960847BAF5CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E368D3D434DFCB0326B1552D906B233F">Kieckbusch, Max</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="09F73E56DA026C82F307064E7A817861">text</mods:typeOfResource>
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<mods:titleInfo id="4D5C0992B8455BC4AAF618CCFC09989C">
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<mods:title id="642A8E96A25912C7FF5E6A63AE6F5D9F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5DE8D4680F919B5C55CB491BB5F70B7D">
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<mods:date id="FF93E6CAA514FBB143CB916168406E7D">2014</mods:date>
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<mods:detail id="EA68AAED9AA3458C9F16E62324AE7DFF" type="volume">
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<mods:number id="8CA876B590054965F9425AD1B94EE056">3872</mods:number>
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</mods:detail>
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<mods:detail id="99AE2D1A057D9289A585677F991535B4" type="issue">
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<mods:number id="650D09CA5A99EEC765D38283714BD2AC">3</mods:number>
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<mods:end id="AD377B0A46DBF5AC6A298C038F9D15B1">300</mods:end>
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<mods:classification id="7070604A4531562A7C220B757E226DE8">journal article</mods:classification>
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<mods:identifier id="251F2DB46B169570B0BF23BD0D9745CC" type="DOI">10.11646/zootaxa.3872.3.5</mods:identifier>
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<mods:identifier id="C46B01FD107C0AD8EFE5ED5CFBC5EBDB" type="GBIF-Dataset">55ca06ce-1ac4-4f07-ac21-a75476e73536</mods:identifier>
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<mods:identifier id="7232E6562D1BAC48C27606857C38202F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DC9E9136F6818A7EC1139DE9925820F3" type="Zenodo-Dep">228806</mods:identifier>
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<mods:identifier id="FC95E36A805B70B5FAA39C15EA2E295B" type="ZooBank">1B4717AE-9993-4166-BAA2-45DFC907080C</mods:identifier>
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<treatment id="039D87E1FFF6467A9FDEFD95FC54FAE7" ID-DOI="http://doi.org/10.5281/zenodo.6124208" ID-GBIF-Taxon="119589973" ID-Zenodo-Dep="6124208" LSID="urn:lsid:plazi:treatment:039D87E1FFF6467A9FDEFD95FC54FAE7" httpUri="http://treatment.plazi.org/id/039D87E1FFF6467A9FDEFD95FC54FAE7" lastPageId="6" lastPageNumber="297" pageId="4" pageNumber="295">
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<subSubSection id="C32E657CFFF646789FDEFD95FF5EFDB5" pageId="4" pageNumber="295" type="nomenclature">
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<paragraph id="8B8B36F7FFF646789FDEFD95FE45FDD4" blockId="4.[151,484,547,607]" box="[151,484,547,574]" pageId="4" pageNumber="295">
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<heading id="D0C3819BFFF646789FDEFD95FE45FDD4" bold="true" box="[151,484,547,574]" fontSize="11" level="1" pageId="4" pageNumber="295" reason="1">
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<emphasis id="B940EAE5FFF646789FDEFD95FE45FDD4" bold="true" box="[151,484,547,574]" pageId="4" pageNumber="295">
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<taxonomicName id="4C344D74FFF646789FDEFD95FEDEFDD7" box="[151,383,547,573]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="295" phylum="Chordata" rank="species" species="careolepis" status="sp. nov.">
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<emphasis id="B940EAE5FFF646789FDEFD95FEDEFDD7" bold="true" box="[151,383,547,573]" italics="true" pageId="4" pageNumber="295">Atractus careolepis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273579EFFF646789ECFFD92FE45FDD4" box="[390,484,548,574]" pageId="4" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8B36F7FFF646789FDEFDF0FF5EFDB5" blockId="4.[151,484,547,607]" box="[151,255,582,607]" pageId="4" pageNumber="295">
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<figureCitation id="130F2A72FFF646789FDEFDF0FF5EFDB5" box="[151,255,582,607]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="4.[151,249,1553,1575]" captionStartId-1="4.[151,249,1927,1949]" captionStartId-2="5.[151,249,1520,1542]" captionTargetBox-0="[174,1428,1092,1513]" captionTargetBox-1="[151,1435,1625,1905]" captionTargetBox-2="[151,1435,193,1497]" captionTargetId-0="figure@4.[174,1459,1092,1513]" captionTargetId-1="figure@4.[151,1436,1625,1905]" captionTargetId-2="figure@5.[151,1435,193,1499]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 4. Dorsal (a) and ventral (b) views of the holotype of Atractus careolepis (SMF 68413). Scale bars equal 5.0 mm." captionText-1="FIGURE 5. Dorsal (a), lateral (b), and ventral (c) views of head of the holotype of Atractus careolepis (SMF 68413). Scale bars equal 1.0 mm." captionText-2="FIGURE 6. Dorsal view of body (a) and tail (b), lateral view body (c) and tail (d), and ventral view of body (e) and tail (f) of the holotype of Atractus careolepis (SMF 68413). The head orientation of the specimen is directed to the right in a, c, and e and to the left in b, d, and f. Scale bars equal 1.0 mm." httpUri-0="https://zenodo.org/record/228810/files/figure.png" httpUri-1="https://zenodo.org/record/228811/files/figure.png" httpUri-2="https://zenodo.org/record/228812/files/figure.png" pageId="4" pageNumber="295">Figs. 4–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E657CFFF646789FDEFD39FA20FD21" pageId="4" pageNumber="295" type="materials_examined">
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<paragraph id="8B8B36F7FFF646789FDEFD39FA20FD21" blockId="4.[151,1437,654,1039]" pageId="4" pageNumber="295">
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<emphasis id="B940EAE5FFF646789FDEFD39FEA5FD4D" bold="true" box="[151,260,655,679]" pageId="4" pageNumber="295">
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<typeStatus id="548F8855FFF646789FDEFD39FEA5FD4D" box="[151,260,655,679]" pageId="4" pageNumber="295" type="holotype">Holotype</typeStatus>
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</emphasis>
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. A male,
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<collectionCode id="ED25AE32FFF646789E3BFD39FE0BFD4D" LSID="urn:lsid:biocol.org:col:34838" box="[370,426,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="295">SMF</collectionCode>
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68413, collected on August 0 1 1970 by M. Henning and F. Klaaßen in the Punta de Betin (approximately 11.2522°, -74.2197°,
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<quantity id="4CCC9B12FFF646789D73FD04FDD2FD20" box="[570,627,690,715]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="295" unit="m" value="15.0">15 m</quantity>
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asl), municipality of Santa
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<collectingCountry id="F3237667FFF646789CE1FD05FC4FFD20" box="[936,1006,691,714]" name="Malta" pageId="4" pageNumber="295">Marta</collectingCountry>
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, province of Magdalena,
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<collectingCountry id="F3237667FFF646789A42FD04FADCFD21" box="[1291,1405,690,715]" name="Colombia" pageId="4" pageNumber="295">Colombia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E657CFFF646799F8EFD60FB45F806" lastPageId="5" lastPageNumber="296" pageId="4" pageNumber="295" type="diagnosis">
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<paragraph id="8B8B36F7FFF646789F8EFD60FEA6FBE5" blockId="4.[151,1437,654,1039]" pageId="4" pageNumber="295">
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<emphasis id="B940EAE5FFF646789F8EFD60FEE0FD05" bold="true" box="[199,321,726,751]" pageId="4" pageNumber="295">Diagnosis.</emphasis>
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A species of the genus
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<taxonomicName id="4C344D74FFF646789D13FD61FD18FD04" box="[602,697,727,750]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="295" phylum="Chordata" rank="genus">
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<emphasis id="B940EAE5FFF646789D13FD61FD18FD04" box="[602,697,727,750]" italics="true" pageId="4" pageNumber="295">Atractus</emphasis>
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</taxonomicName>
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that differs from all congeners by the following combination of characters: (1) 17 smooth dorsal scale rows; (2) preocular absent; (3) loreal absent; (4) temporals 1+2; (5) seven supralabials; (6) six or seven infralabials; (7) eight maxillary teeth; (8) three gular scale rows; (9) three preventrals; (10) 146 ventrals in single male; (11) 31/32 subcaudals in single male; (12) dorsum of body and tail reddish brown with paired, occasionally slightly alternating lateral vertical cream blotches, approximately one dorsal scale broad, not contacting in vertebral region; (13) venter cream with reddish brown blotches, forming two parallel series on anterior body and a checkered pattern on the rest of body; (14) SVL of single known male
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<quantity id="4CCC9B12FFF646789A3BFC19FF67FC00" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="295" unit="mm" value="178.0">178 mm</quantity>
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; (15) small to moderate tail length in single known male (16.3 % of SVL); (16) hemipenis morphology unknown.
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</paragraph>
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<caption id="DF4B667FFFF646789FDEF9A7FADDF9CD" box="[151,1404,1553,1575]" httpUri="https://zenodo.org/record/228810/files/figure.png" pageId="4" pageNumber="295" targetBox="[174,1428,1092,1513]" targetPageId="4">
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<paragraph id="8B8B36F7FFF646789FDEF9A7FADDF9CD" blockId="4.[151,1404,1553,1575]" box="[151,1404,1553,1575]" pageId="4" pageNumber="295">
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<emphasis id="B940EAE5FFF646789FDEF9A7FEB3F9CD" bold="true" box="[151,274,1553,1575]" pageId="4" pageNumber="295">FIGURE 4.</emphasis>
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Dorsal (a) and ventral (b) views of the holotype of
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<taxonomicName id="4C344D74FFF646789C55F9A4FC40F9CC" box="[796,993,1553,1575]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="295" phylum="Chordata" rank="species" species="careolepis">
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<emphasis id="B940EAE5FFF646789C55F9A4FC40F9CC" box="[796,993,1553,1575]" italics="true" pageId="4" pageNumber="295">Atractus careolepis</emphasis>
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</taxonomicName>
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(SMF 68413). Scale bars equal 5.0 mm.
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</paragraph>
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</caption>
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<caption id="DF4B667FFFF646789FDEF831FEF8F851" httpUri="https://zenodo.org/record/228811/files/figure.png" pageId="4" pageNumber="295" targetBox="[151,1435,1625,1905]" targetPageId="4">
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<paragraph id="8B8B36F7FFF646789FDEF831FEF8F851" blockId="4.[151,1436,1927,1979]" pageId="4" pageNumber="295">
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<emphasis id="B940EAE5FFF646789FDEF831FEB5F877" bold="true" box="[151,276,1927,1949]" pageId="4" pageNumber="295">FIGURE 5.</emphasis>
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Dorsal (a), lateral (b), and ventral (c) views of head of the holotype of
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<taxonomicName id="4C344D74FFF646789CB4F831FB65F876" box="[1021,1220,1927,1948]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="295" phylum="Chordata" rank="species" species="careolepis">
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<emphasis id="B940EAE5FFF646789CB4F831FB65F876" box="[1021,1220,1927,1948]" italics="true" pageId="4" pageNumber="295">Atractus careolepis</emphasis>
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</taxonomicName>
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(SMF 68413). Scale bars equal 1.0 mm.
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</paragraph>
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</caption>
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<caption id="DF4B667FFFF746799FDEFA46FD2BF9A9" httpUri="https://zenodo.org/record/228812/files/figure.png" pageId="5" pageNumber="296" targetBox="[151,1435,193,1497]" targetPageId="5">
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<paragraph id="8B8B36F7FFF746799FDEFA46FD2BF9A9" blockId="5.[151,1435,1520,1603]" pageId="5" pageNumber="296">
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<emphasis id="B940EAE5FFF746799FDEFA46FEB2F9EC" bold="true" box="[151,275,1520,1542]" pageId="5" pageNumber="296">FIGURE 6.</emphasis>
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Dorsal view of body (a) and tail (b), lateral view body (c) and tail (d), and ventral view of body (e) and tail (f) of the holotype of
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<taxonomicName id="4C344D74FFF746799E7CF9A6FE5BF9CE" box="[309,506,1551,1573]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">
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<emphasis id="B940EAE5FFF746799E7CF9A6FE5BF9CE" box="[309,506,1551,1573]" italics="true" pageId="5" pageNumber="296">Atractus careolepis</emphasis>
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</taxonomicName>
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(SMF 68413). The head orientation of the specimen is directed to the right in a, c, and e and to the left in b, d, and f. Scale bars equal 1.0 mm.
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</paragraph>
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</caption>
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<paragraph id="8B8B36F7FFF746799F8EF9DCFB45F806" blockId="5.[151,1436,1642,2028]" pageId="5" pageNumber="296">
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<emphasis id="B940EAE5FFF746799F8EF9DCFECDF969" bold="true" box="[199,364,1642,1667]" pageId="5" pageNumber="296">Comparisons.</emphasis>
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Among all currently recognized congeners,
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<taxonomicName id="4C344D74FFF746799C35F9DAFBF9F969" box="[892,1112,1643,1667]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">
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<emphasis id="B940EAE5FFF746799C35F9DAFBF9F969" box="[892,1112,1643,1667]" italics="true" pageId="5" pageNumber="296">Atractus careolepis</emphasis>
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</taxonomicName>
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is most similar in external morphology to
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<taxonomicName id="4C344D74FFF746799E0CF939FE18F94D" box="[325,441,1679,1703]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="lancinii">
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<emphasis id="B940EAE5FFF746799E0CF939FE18F94D" box="[325,441,1679,1703]" italics="true" pageId="5" pageNumber="296">A. lancinii</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C344D74FFF746799E80F939FDEEF94D" box="[457,591,1679,1703]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
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<emphasis id="B940EAE5FFF746799E80F939FDEEF94D" box="[457,591,1679,1703]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
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</taxonomicName>
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, and
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<emphasis id="B940EAE5FFF746799DC6F939FB9AF94D" box="[655,1083,1679,1703]" italics="true" pageId="5" pageNumber="296">
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<taxonomicName id="4C344D74FFF746799DC6F939FCF6F94D" box="[655,855,1679,1703]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="sanctaemartae">A. sanctaemartae</taxonomicName>
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.
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<taxonomicName id="4C344D74FFF746799C2BF939FB9AF94D" box="[866,1083,1679,1703]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">Atractus careolepis</taxonomicName>
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</emphasis>
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differs from all of them by the absence of the loreal scale resulting in the contact of the prefrontals with the supralabials (vs. loreal scale present in
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<taxonomicName id="4C344D74FFF746799FDEF961FEA8F905" box="[151,265,1751,1775]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="lancinii">
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<emphasis id="B940EAE5FFF746799FDEF961FEA8F905" box="[151,265,1751,1775]" italics="true" pageId="5" pageNumber="296">A. lancinii</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C344D74FFF746799E51F961FE3DF905" box="[280,412,1751,1775]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
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<emphasis id="B940EAE5FFF746799E51F961FE3DF905" box="[280,412,1751,1775]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C344D74FFF746799E90F961FD3AF905" box="[473,667,1751,1775]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="sanctaemartae">
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<emphasis id="B940EAE5FFF746799E90F961FD3AF905" box="[473,667,1751,1775]" italics="true" pageId="5" pageNumber="296">A. sanctaemartae</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C344D74FFF746799DFBF961FC28F905" box="[690,905,1751,1775]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">
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<emphasis id="B940EAE5FFF746799DFBF961FC28F905" box="[690,905,1751,1775]" italics="true" pageId="5" pageNumber="296">Atractus careolepis</emphasis>
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</taxonomicName>
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further differs from
|
||
<taxonomicName id="4C344D74FFF746799B24F961FB41F905" box="[1133,1248,1751,1775]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="lancinii">
|
||
<emphasis id="B940EAE5FFF746799B24F961FB41F905" box="[1133,1248,1751,1775]" italics="true" pageId="5" pageNumber="296">A. lancinii</emphasis>
|
||
</taxonomicName>
|
||
by having seven supralabials and seven infralabials (vs. eight supralabials and eight infralabials in
|
||
<taxonomicName id="4C344D74FFF746799B54F94AFB31F8F9" box="[1053,1168,1787,1811]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="lancinii">
|
||
<emphasis id="B940EAE5FFF746799B54F94AFB31F8F9" box="[1053,1168,1787,1811]" italics="true" pageId="5" pageNumber="296">A. lancinii</emphasis>
|
||
</taxonomicName>
|
||
) and 146 ventrals in the single male (vs. 174–186 ventrals in
|
||
<taxonomicName id="4C344D74FFF746799D71F8A9FD0CF8DD" box="[568,685,1823,1847]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="lancinii">
|
||
<emphasis id="B940EAE5FFF746799D71F8A9FD0CF8DD" box="[568,685,1823,1847]" italics="true" pageId="5" pageNumber="296">A. lancinii</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C344D74FFF746799D8FF8A9FC3FF8DD" box="[710,926,1823,1847]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">
|
||
<emphasis id="B940EAE5FFF746799D8FF8A9FC3FF8DD" box="[710,926,1823,1847]" italics="true" pageId="5" pageNumber="296">Atractus careolepis</emphasis>
|
||
</taxonomicName>
|
||
differs further from
|
||
<taxonomicName id="4C344D74FFF746799BC0F8A9FAAEF8DD" box="[1161,1295,1823,1847]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
|
||
<emphasis id="B940EAE5FFF746799BC0F8A9FAAEF8DD" box="[1161,1295,1823,1847]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
|
||
</taxonomicName>
|
||
by having a body pattern of pale transverse bands on a dark dorsum (vs. dorsum uniformly reddish brown in
|
||
<taxonomicName id="4C344D74FFF746799B81F8F2FAEDF8B1" box="[1224,1356,1859,1883]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
|
||
<emphasis id="B940EAE5FFF746799B81F8F2FAEDF8B1" box="[1224,1356,1859,1883]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
|
||
</taxonomicName>
|
||
); eight maxillary teeth (vs. nine maxillary teeth in
|
||
<taxonomicName id="4C344D74FFF746799D35F8D1FCA0F895" box="[636,769,1895,1919]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
|
||
<emphasis id="B940EAE5FFF746799D35F8D1FCA0F895" box="[636,769,1895,1919]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
|
||
</taxonomicName>
|
||
); three gular scale rows (vs. two rows in
|
||
<taxonomicName id="4C344D74FFF746799B9CF8D1FAFBF895" box="[1237,1370,1895,1919]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="macondo">
|
||
<emphasis id="B940EAE5FFF746799B9CF8D1FAFBF895" box="[1237,1370,1895,1919]" italics="true" pageId="5" pageNumber="296">A. macondo</emphasis>
|
||
</taxonomicName>
|
||
); and 31–32 subcaudals in the single known male (vs. 29 subcaudals in the single known male).
|
||
<taxonomicName id="4C344D74FFF746799B31F83AFAEFF849" box="[1144,1358,1931,1955]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="careolepis">
|
||
<emphasis id="B940EAE5FFF746799B31F83AFAEFF849" box="[1144,1358,1931,1955]" italics="true" pageId="5" pageNumber="296">Atractus careolepis</emphasis>
|
||
</taxonomicName>
|
||
differs further from
|
||
<taxonomicName id="4C344D74FFF746799E60F819FE4DF82D" box="[297,492,1967,1991]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="sanctaemartae">
|
||
<emphasis id="B940EAE5FFF746799E60F819FE4DF82D" box="[297,492,1967,1991]" italics="true" pageId="5" pageNumber="296">A. sanctaemartae</emphasis>
|
||
</taxonomicName>
|
||
by having a body pattern of pale transverse bands on a brown dorsum in the single known male (vs. black dorsum with alternate light transversal bands in males of
|
||
<taxonomicName id="4C344D74FFF746799B5BF862FB72F801" box="[1042,1235,2004,2027]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="296" phylum="Chordata" rank="species" species="sanctaemartae">
|
||
<emphasis id="B940EAE5FFF746799B5BF862FB72F801" box="[1042,1235,2004,2027]" italics="true" pageId="5" pageNumber="296">A. sanctaemartae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657CFFF4467A9F8EFF21FC30FB2F" pageId="6" pageNumber="297" type="description">
|
||
<paragraph id="8B8B36F7FFF4467A9F8EFF21FB2EFC2D" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="297">
|
||
<emphasis id="B940EAE5FFF4467A9F8EFF21FDAFFF5A" bold="true" box="[199,526,151,176]" pageId="6" pageNumber="297">
|
||
Description of the
|
||
<typeStatus id="548F8855FFF4467A9EE1FF2EFDAFFF5A" box="[424,526,152,176]" pageId="6" pageNumber="297" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Male; SVL
|
||
<quantity id="4CCC9B12FFF4467A9DECFF2EFCA7FF45" box="[677,774,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="297" unit="mm" value="178.0">178 mm</quantity>
|
||
; long TL
|
||
<quantity id="4CCC9B12FFF4467A9C30FF2EFC68FF45" box="[889,969,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="297" unit="mm" value="29.0">29 mm</quantity>
|
||
(complete; 16.3 % SVL); TTL
|
||
<quantity id="4CCC9B12FFF4467A9A7FFF2EFA36FF45" box="[1334,1431,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.07" pageId="6" pageNumber="297" unit="mm" value="207.0">207 mm</quantity>
|
||
; head not distinct from body; body cylindrical, maximum body diameter
|
||
<quantity id="4CCC9B12FFF4467A9C9DFF0AFB8DFF3E" box="[980,1068,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="6" pageNumber="297" unit="mm" value="7.1">7.1 mm</quantity>
|
||
(4 % of SVL); head rounded in dorsal view; HL
|
||
<quantity id="4CCC9B12FFF4467A9E16FF69FE66FF1D" box="[351,455,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="7.61" pageId="6" pageNumber="297" unit="mm" value="7.61">7.61 mm</quantity>
|
||
(4.3 % SVL); HW 6.0 mm (75.9 % of HL); snout rounded in dorsal view, slightly truncate in lateral view; eye diameter
|
||
<quantity id="4CCC9B12FFF4467A9D73FEB2FD32FEF6" box="[570,659,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="297" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; pupil round; interorbital distance
|
||
<quantity id="4CCC9B12FFF4467A9B5BFEB2FBC6FEF6" box="[1042,1127,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="297" unit="mm" value="3.2">3.2 mm</quantity>
|
||
(40.5 % HL); rostro-orbital distance 3.0 mm (37.9 % HL); naso-orbital distance 2.0 mm (25.3 % HL). 17/17/17 smooth dorsal scale rows, 16 dorsal scale rows above cloaca, 10 dorsal scale rows above second subcaudal; apical pits absent; 146 ventrals; 31/ 32 subcaudals; three pre-ventrals; cloacal scute entire; rostral slightly visible from above, subtriangular, wider than high (rostral height
|
||
<quantity id="4CCC9B12FFF4467A9E3FFE22FE6EFE46" box="[374,463,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="297" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, rostral width
|
||
<quantity id="4CCC9B12FFF4467A9D38FE22FD6DFE46" box="[625,716,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="297" unit="mm" value="1.2">1.2 mm</quantity>
|
||
); two subrectangular internasals, height
|
||
<quantity id="4CCC9B12FFF4467A9BDBFE22FB4AFE46" box="[1170,1259,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="297" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, width
|
||
<quantity id="4CCC9B12FFF4467A9A77FE22FA39FE46" box="[1342,1432,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="297" unit="mm" value="0.8">0.8 mm</quantity>
|
||
; internasal suture sinistral in respect to prefrontal suture; nostril in suture of prenasal and postnasal; 1/1 subrectangular postnasal, length
|
||
<quantity id="4CCC9B12FFF4467A9D5EFE6AFDD5FE1E" box="[535,628,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="297" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, postnasal height
|
||
<quantity id="4CCC9B12FFF4467A9C04FE6AFC0DFE1E" box="[845,940,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="297" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; loreal absent, prefrontals in contact with supralabials; preocular lacking; two subtriangular postoculars, upper postocular approximately equal in size to lower postocular (upper postocular length
|
||
<quantity id="4CCC9B12FFF4467A9DC4FD92FD4AFDD6" box="[653,747,548,573]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="297" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; upper postocular height
|
||
<quantity id="4CCC9B12FFF4467A9B55FD92FBDAFDD6" box="[1052,1147,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="297" unit="mm" value="0.7">0.7 mm</quantity>
|
||
); temporal formula 1+2, anterior temporal equal in size to lower posterior temporal, upper posterior temporal three times as long as lower posterior temporal (dextral) or two times as long as lower posterior temporal (sinistral) (anterior temporal: height
|
||
<quantity id="4CCC9B12FFF4467A9FDEFD26FF4EFD4D" box="[151,239,656,680]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="297" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, length
|
||
<quantity id="4CCC9B12FFF4467A9E0EFD39FE01FD4D" box="[327,416,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="297" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; upper posterior temporal: dextral: height
|
||
<quantity id="4CCC9B12FFF4467A9C31FD26FC6EFD4D" box="[888,975,656,680]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="297" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, length
|
||
<quantity id="4CCC9B12FFF4467A9B61FD26FB20FD4D" box="[1064,1153,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="297" unit="mm" value="3.4">3.4 mm</quantity>
|
||
; sinistral: height 1.0 mm, length
|
||
<quantity id="4CCC9B12FFF4467A9FAFFD02FEE2FD26" box="[230,323,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="297" unit="mm" value="2.1">2.1 mm</quantity>
|
||
); seven supralabials, first smaller than all others, third supralabial larger than second and fourth, fifth, sixth and seventh supralabial equal in size or slightly larger than third; second and third supralabial in contact with prefrontals; third and fourth supralabial in contact with the eye; six and seven infralabials, first three in contact with the chin shields; dextral third infralabial larger than all other infralabials; sinistral third and fourth larger than all other infralabials; first pair of infralabials in contact, preventing contact of mental and chin shields; gular scale rows from infralabials to preventrals three; two chin shields, length
|
||
<quantity id="4CCC9B12FFF4467A9CE3FCDEFBA4FC95" box="[938,1029,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="297" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, width
|
||
<quantity id="4CCC9B12FFF4467A9B14FCDEFB19FC95" box="[1117,1208,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="297" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; two subpentagonal prefrontals, length
|
||
<quantity id="4CCC9B12FFF4467A9E37FC3AFE41FC4E" box="[382,480,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="297" unit="mm" value="2.1">2.1 mm</quantity>
|
||
, width
|
||
<quantity id="4CCC9B12FFF4467A9D0FFC3AFD09FC4E" box="[582,680,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="297" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; subrectangular supraoculars, length
|
||
<quantity id="4CCC9B12FFF4467A9B27FC3AFB71FC4E" box="[1134,1232,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="297" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, width 1.0 mm; subtriangular frontal length
|
||
<quantity id="4CCC9B12FFF4467A9E99FC06FD89FC2D" box="[464,552,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="297" unit="mm" value="2.2">2.2 mm</quantity>
|
||
, width
|
||
<quantity id="4CCC9B12FFF4467A9D32FC06FD75FC2D" box="[635,724,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="297" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; parietal length
|
||
<quantity id="4CCC9B12FFF4467A9CCEFC06FC7EFC2D" box="[903,991,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="297" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, width
|
||
<quantity id="4CCC9B12FFF4467A9B78FC06FB2AFC2D" box="[1073,1163,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="297" unit="mm" value="2.1">2.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8B36F7FFF4467A9F8EFC65FC30FB2F" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="297">
|
||
<emphasis id="B940EAE5FFF4467A9F8EFC65FDADFC06" bold="true" box="[199,524,979,1004]" pageId="6" pageNumber="297">
|
||
Coloration of the
|
||
<typeStatus id="548F8855FFF4467A9ED7FC62FDA6FC06" box="[414,519,980,1004]" pageId="6" pageNumber="297" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
After 43 years preservation in 70% ethanol: dorsal surface of head Raw Umber (280) with Light Buff (2) blotches on most upper head scales; supralabials Light Buff (2); ventral surface of head Light Buff (2) with Raw Umber (280) speckles on anterior infralabials and chin shields; dorsal surfaces of body and tail Raw Umber (280) with paired occasionally slightly alternating lateral vertical Pale Buff (1) blotches, approximately one dorsal scale broad, not contacting in vertebral region; ventral surface Light Buff (2) with Raw Umber blotches (280), forming two parallel series on anterior body and a checkered pattern on the rest of body; dark coloration on venter more intense on posterior portion of body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657CFFF4467A9F8EFB66FC54FAE7" pageId="6" pageNumber="297" type="etymology">
|
||
<paragraph id="8B8B36F7FFF4467A9F8EFB66FC54FAE7" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="297">
|
||
<emphasis id="B940EAE5FFF4467A9F8EFB66FEE6FB02" bold="true" box="[199,327,1232,1256]" pageId="6" pageNumber="297">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName id="4C344D74FFF4467A9E86FB66FDE2FB02" box="[463,579,1232,1256]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="297" phylum="Chordata" rank="species" species="careolepis">
|
||
<emphasis id="B940EAE5FFF4467A9E86FB66FDE2FB02" box="[463,579,1232,1256]" italics="true" pageId="6" pageNumber="297">careolepis</emphasis>
|
||
</taxonomicName>
|
||
is a compound noun derived from
|
||
<emphasis id="B940EAE5FFF4467A9C9FFB67FBB4FB02" box="[982,1045,1233,1256]" italics="true" pageId="6" pageNumber="297">careo</emphasis>
|
||
(Latin for “be without”) and
|
||
<emphasis id="B940EAE5FFF4467A9A2EFB66FA3DFB02" box="[1383,1436,1232,1256]" italics="true" pageId="6" pageNumber="297">lepis</emphasis>
|
||
(Greek for “scale”) referring to the absence of the loreal scale in this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |