treatments-xml/data/03/9D/87/039D87C9FFADFF88A0A87A9CFF1B2973.xml
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<document id="911CAB7ECCB69A5E32ADF1AD76688B52" ID-DOI="10.11646/zootaxa.5389.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10410051" ID-ZooBank="3E7237B4-85C5-4581-B469-ED1C00154413" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1703073801695" checkinUser="plazi" docAuthor="Zajonz, Uwe, Bogorodsky, Sergey V., Saeed, Fouad K. N., Aideed, Moteah S. &amp; Lavergne, Edouard" docDate="2023" docId="039D87C9FFADFF88A0A87A9CFF1B2973" docLanguage="en" docName="zootaxa.5389.3.1.pdf" docOrigin="Zootaxa 5389 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5389.3.1/52519" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scarus arabicus" docType="treatment" docVersion="3" lastPageNumber="308" masterDocId="FFA4FFB1FFAAFF8FA03F7A0BFFF82D7F" masterDocTitle="Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995" masterLastPageNumber="330" masterPageNumber="301" pageNumber="308" updateTime="1703162922346" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="640346A132E1DC53024ED5036CD5B4D4">Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995</mods:title>
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<mods:namePart id="981889C9948C1C0D193DE119BFEE1BE3">Zajonz, Uwe</mods:namePart>
<mods:affiliation id="B8AF2A2560571FF9DC9F524A879C4BE8">Senckenberg Research Institute and Museum of Nature (SMF), Marine Zoology-Senckenberganlage 25, 60325 Frankfurt am Main, Germany.</mods:affiliation>
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<mods:namePart id="1C3280937D992764A285BADBD2F238EA">Bogorodsky, Sergey V.</mods:namePart>
<mods:affiliation id="3FD0AC0E67F477319ED49DA5B74C2DDC">Senckenberg Research Institute and Museum of Nature (SMF), Marine Zoology-Senckenberganlage 25, 60325 Frankfurt am Main, Germany. &amp; Station of Naturalists, Omsk, Russia.</mods:affiliation>
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<mods:namePart id="A1F0CFBEEF6B46E0D1B492053BC56822">Saeed, Fouad K. N.</mods:namePart>
<mods:affiliation id="3E51DE7D2D709853BFE3024648060ABE">Senckenberg Research Institute and Museum of Nature (SMF), Marine Zoology-Senckenberganlage 25, 60325 Frankfurt am Main, Germany. &amp; Environmental Protection Authority, Socotra Branch, Hadibo, Yemen.</mods:affiliation>
</mods:name>
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<mods:namePart id="C69725D512FBB5026D67F40A4C7838C1">Aideed, Moteah S.</mods:namePart>
<mods:affiliation id="986AEBAAB737848563277978530F9069">Senckenberg Research Institute and Museum of Nature (SMF), Marine Zoology-Senckenberganlage 25, 60325 Frankfurt am Main, Germany. &amp; Maritime Affairs Authority, Mukalla, Yemen.</mods:affiliation>
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<mods:namePart id="7165F0DB981634FC0FA1087CEA28F16E">Lavergne, Edouard</mods:namePart>
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<treatment id="039D87C9FFADFF88A0A87A9CFF1B2973" LSID="urn:lsid:plazi:treatment:039D87C9FFADFF88A0A87A9CFF1B2973" httpUri="http://treatment.plazi.org/id/039D87C9FFADFF88A0A87A9CFF1B2973" lastPageNumber="308" pageId="7" pageNumber="308">
<subSubSection id="C32E6554FFADFF88A0A87A9CFD9A2DCE" box="[151,610,151,178]" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="8B8B36DFFFADFF88A0A87A9CFD9A2DCE" blockId="7.[151,1362,151,280]" box="[151,610,151,178]" pageId="7" pageNumber="308">
<heading id="D0C381B3FFADFF88A0A87A9CFD9A2DCE" bold="true" box="[151,610,151,178]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<taxonomicName id="4C344D5CFFADFF88A0A87A9CFD9A2DCE" ID-CoL="4V3FW" authority="(Steindachner, 1902)" baseAuthorityName="Steindachner" baseAuthorityYear="1902" box="[151,610,151,178]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="B940EACDFFADFF88A0A87A9CFD9A2DCE" bold="true" box="[151,610,151,178]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0A87A9CFEA22DCD" bold="true" box="[151,346,151,178]" italics="true" pageId="7" pageNumber="308">Scarus arabicus</emphasis>
(
<bibRefCitation id="EFA54B2EFFADFF88A1567A9CFDA12DCE" author="Steindachner, F." box="[361,601,151,177]" pageId="7" pageNumber="308" pagination="316 - 318" refId="ref15559" refString="Steindachner, F. (1902) Wissenschaftliche Ergebnisse der sudarabischen Expedition in den Jahren 1898 bis 1899. Fische von Sudarabien und Socotra. Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch- Naturwissenschaftliche Classe, 39 (24) (76), 316 - 318." type="journal article" year="1902">Steindachner, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0A87AB3FAAA2DAF" box="[151,1362,183,208]" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="8B8B36DFFFADFF88A0A87AB3FAAA2DAF" blockId="7.[151,1362,151,280]" box="[151,1362,183,208]" pageId="7" pageNumber="308">
<treatmentCitationGroup id="AB2411F1FFADFF88A0A87AB3FAAA2DAF" box="[151,1362,183,208]" pageId="7" pageNumber="308">
<treatmentCitation id="0A9510CEFFADFF88A0A87AB3FD7A2DAF" author="Steindachner, F." box="[151,642,184,208]" page="318" pageId="7" pageNumber="308" year="1902">
<taxonomicName id="4C344D5CFFADFF88A0A87AB3FD7A2DAF" ID-CoL="6WGJ3" authority="Steindachner, 1902: 318" authorityName="Steindachner" authorityPageNumber="318" authorityYear="1902" box="[151,642,184,208]" class="Actinopterygii" family="Scaridae" genus="Pseudoscarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="B940EACDFFADFF88A0A87AB3FE7B2DAF" box="[151,387,184,208]" italics="true" pageId="7" pageNumber="308">Pseudoscarus arabicus</emphasis>
<materialsCitation id="3B5C3C82FFADFF88A1B67AB3FD7A2DAF" box="[393,642,184,208]" collectionCode="NMW" country="Yemen" location="Gulf" municipality="Mukalla" pageId="7" pageNumber="308" specimenCode="NMW 72746" specimenCount="1" stateProvince="'Adan" typeStatus="holotype">
<bibRefCitation id="EFA54B2EFFADFF88A1B67AB3FD7A2DAF" author="Steindachner, F." box="[393,642,184,208]" pageId="7" pageNumber="308" pagination="316 - 318" refId="ref15559" refString="Steindachner, F. (1902) Wissenschaftliche Ergebnisse der sudarabischen Expedition in den Jahren 1898 bis 1899. Fische von Sudarabien und Socotra. Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch- Naturwissenschaftliche Classe, 39 (24) (76), 316 - 318." type="journal article" year="1902">Steindachner, 1902: 318</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
;
<typeStatus id="548F887DFFADFF88A2B27AB3FD122DAF" box="[653,746,184,208]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
:
<specimenCode id="DB929EA4FFADFF88A2CC7AB3FC842DB0" box="[755,892,183,207]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="308">NMW 72746</specimenCode>
; type locality:
<collectingMunicipality id="6BEFACA5FFADFF88A42E7AB3FB902DAF" box="[1041,1128,184,208]" pageId="7" pageNumber="308">Mukalla</collectingMunicipality>
,
<collectingCountry id="F323764FFFADFF88A44E7AB3FB422DB0" box="[1137,1210,184,207]" name="Yemen" pageId="7" pageNumber="308">Yemen</collectingCountry>
,
<location id="8EEB6004FFADFF88A4FA7AB3FB0B2DAF" LSID="urn:lsid:plazi:treatment:039D87C9FFADFF88A0A87A9CFF1B2973:8EEB6004FFADFF88A4FA7AB3FB0B2DAF" box="[1221,1267,184,208]" country="Yemen" municipality="Mukalla" name="Gulf" pageId="7" pageNumber="308" stateProvince="'Adan">Gulf</location>
of
<collectingRegion id="49F0F83DFFADFF88A52C7AB3FAB52DAF" box="[1299,1357,184,208]" country="Yemen" name="'Adan" pageId="7" pageNumber="308">Aden</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0A87AD1FE9E2D8B" box="[151,358,218,244]" pageId="7" pageNumber="308" type="vernacular_names">
<paragraph id="8B8B36DFFFADFF88A0A87AD1FE9E2D8B" blockId="7.[151,1362,151,280]" box="[151,358,218,244]" pageId="7" pageNumber="308">
<taxonomicName id="4C344D5CFFADFF88A0A87AD1FF092D8B" box="[151,241,218,244]" order="Arabian" pageId="7" pageNumber="325" rank="order">Arabian</taxonomicName>
Parrotfish
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0A87AF5FF0D2C67" box="[151,245,254,280]" pageId="7" pageNumber="308" type="description">
<paragraph id="8B8B36DFFFADFF88A0A87AF5FF0D2C67" blockId="7.[151,1362,151,280]" box="[151,245,254,280]" pageId="7" pageNumber="308">
<figureCitation id="130F2A5AFFADFF88A0A87AF5FF0D2C67" box="[151,245,254,280]" captionStart="FIGURE 4" captionStartId="6.[151,250,1887,1911]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-38@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURE 4. Scarus arabicus. A: initial phase, Abd al-Kuri Island, Socotra Archipelago, 7 m depth; B: SMF uncatalogued, initial phase, 26 cm SL, Hadibo fishmarket, Socotra Island; C: SMF uncatalogued, terminal male, 46 cm SL, Hadibo fishmarket, Socotra Island." figureDoi="http://doi.org/10.5281/zenodo.10410060" httpUri="https://zenodo.org/record/10410060/files/figure.png" pageId="7" pageNumber="308">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0A87B4EFD212F93" pageId="7" pageNumber="308" type="materials_examined">
<paragraph id="8B8B36DFFFADFF88A0A87B4EFC142CD7" blockId="7.[151,1437,325,1036]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0A87B4EFE7F2C20" bold="true" box="[151,391,325,351]" pageId="7" pageNumber="308">Material examined:</emphasis>
<materialsCitation id="3B5C3C82FFADFF88A1AD7B4DFCB62CFB" collectingDate="2019-04-12" collectionCode="SMF" country="Socotra Island" location="Hadibo" pageId="7" pageNumber="308" specimenCount="1">
Underwater photograph;
<collectionCode id="ED25AE1AFFADFF88A28B7B4DFD142C1F" box="[692,748,326,352]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="308">SMF</collectionCode>
uncatalogued [sample of tissue SOC19-468], initial phase,
<quantity id="4CCC9B3AFFADFF88A0A87B61FF242CFB" box="[151,220,362,388]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="308" unit="cm" value="26.0">26 cm</quantity>
SL,
<collectingCountry id="F323764FFFADFF88A1297B61FE432CFB" box="[278,443,362,388]" name="Socotra Island" pageId="7" pageNumber="308">Socotra Island</collectingCountry>
,
<location id="8EEB6004FFADFF88A1F67B61FDE32CFB" LSID="urn:lsid:plazi:treatment:039D87C9FFADFF88A0A87A9CFF1B2973:8EEB6004FFADFF88A1F67B61FDE32CFB" box="[457,539,362,388]" country="Socotra Island" name="Hadibo" pageId="7" pageNumber="308">Hadibo</location>
fishmarket,
<date id="FF8A101FFFADFF88A2937B61FCB62CFB" box="[684,846,362,388]" pageId="7" pageNumber="308" value="2019-04-12">
<collectingDate id="EFCEE9F7FFADFF88A2937B61FCB62CFB" box="[684,846,362,388]" pageId="7" pageNumber="308" value="2019-04-12">12 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B5C3C82FFADFF88A3637B61FC102CD7" collectingDate="2019-04-19" collectionCode="SMF" country="Socotra Island" location="Hadibo" pageId="7" pageNumber="308" specimenCount="1">
<collectionCode id="ED25AE1AFFADFF88A3637B61FC6C2CFB" box="[860,916,362,388]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="308">SMF</collectionCode>
uncatalogued [sample of tissue SOC19-485], terminal male,
<quantity id="4CCC9B3AFFADFF88A17F7B85FE7D2CD7" box="[320,389,398,424]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="308" unit="cm" value="46.0">46 cm</quantity>
SL,
<collectingCountry id="F323764FFFADFF88A1847B85FDA62CD7" box="[443,606,398,424]" name="Socotra Island" pageId="7" pageNumber="308">Socotra Island</collectingCountry>
,
<location id="8EEB6004FFADFF88A2557B85FD442CD7" LSID="urn:lsid:plazi:treatment:039D87C9FFADFF88A0A87A9CFF1B2973:8EEB6004FFADFF88A2557B85FD442CD7" box="[618,700,398,424]" country="Socotra Island" name="Hadibo" pageId="7" pageNumber="308">Hadibo</location>
fishmarket,
<date id="FF8A101FFFADFF88A3767B85FC102CD7" box="[841,1000,398,424]" pageId="7" pageNumber="308" value="2019-04-19">
<collectingDate id="EFCEE9F7FFADFF88A3767B85FC102CD7" box="[841,1000,398,424]" pageId="7" pageNumber="308" value="2019-04-19">19 April 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8B36DFFFADFF88A0F87BBAFB6B2F23" blockId="7.[151,1437,325,1036]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0F87BBAFE2C2CB3" bold="true" box="[199,468,433,460]" pageId="7" pageNumber="308">Distinctive characters:</emphasis>
Body depth
<quantity id="4CCC9B3AFFADFF88A25D7BB9FD2B2CB3" box="[610,723,434,460]" metricMagnitude="-2" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.858" metricValueMin="6.096" pageId="7" pageNumber="308" unit="in" value="2.55" valueMax="2.7" valueMin="2.4">2.42.7 in</quantity>
SL; dorsal profile of head of terminal males steep and distinctly convex; posterior nostril subequal to anterior nostril; teeth fully fused to form dental plates, less than one-half covered by lips; cutting edge of dental plates slightly irregular; posterior margin of upper dental plate with 4 or 5 small blunt teeth; caudal fin slightly emarginate in terminal males. Meristic values: Pectoral-fin rays 15; median predorsal scales 4, about equal in size; cheek with 2 scale rows, lower row with 45 scales.
</paragraph>
<paragraph id="8B8B36DFFFADFF88A0F8786DFD212F93" blockId="7.[151,1437,325,1036]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0F8786DFEA82FFF" box="[199,336,614,640]" italics="true" pageId="7" pageNumber="308">Colouration</emphasis>
: Initial phase gray-brown, the scale edges brown. Terminal males green often suffused with brown. Both phases with head pale blue-green ventrally, green front of snout, a broad pale blue-green band across interorbital which extends slightly behind orbit, dental plates blue-green, dorsal and anal fins colored like body with green-blue margin, and caudal fin with white posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0F878FEFDFE2E4B" pageId="7" pageNumber="308" type="distribution">
<paragraph id="8B8B36DFFFADFF88A0F878FEFDFE2E4B" blockId="7.[151,1437,325,1036]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0F878FEFE9B2E70" bold="true" box="[199,355,757,783]" pageId="7" pageNumber="308">Distribution:</emphasis>
Restricted to the
<taxonomicName id="4C344D5CFFADFF88A21178FDFD702E6F" box="[558,648,758,784]" order="Arabian" pageId="7" pageNumber="325" rank="order">Arabian</taxonomicName>
region, reported from the eastern Gulf of Aden to the Gulf of
<collectingCountry id="F323764FFFADFF88A56778FDFA642E6F" box="[1368,1436,758,784]" name="Oman" pageId="7" pageNumber="308">Oman</collectingCountry>
(Randall 1995;
<bibRefCitation id="EFA54B2EFFADFF88A17A7911FE022E4B" author="Lips, J. &amp; Lips, B. &amp; Roux, J. - M." box="[325,506,794,820]" pageId="7" pageNumber="308" refId="ref14705" refString="Lips, J., Lips, B. &amp; Roux, J. - M. (2016) Poissons de Djibouti. Universite de Djibouti, Djibouti, 216 pp." type="book" year="2016">
Lips
<emphasis id="B940EACDFFADFF88A1417910FE572E4B" box="[382,431,794,820]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFADFF88A0F87935FF1B2973" pageId="7" pageNumber="308" type="discussion">
<paragraph id="8B8B36DFFFADFF88A0F87935FF1B2973" blockId="7.[151,1437,325,1036]" pageId="7" pageNumber="308">
<emphasis id="B940EACDFFADFF88A0F87935FEC52E27" bold="true" box="[199,317,830,856]" pageId="7" pageNumber="308">Remarks:</emphasis>
Observed individuals and collected specimens match the description of
<taxonomicName id="4C344D5CFFADFF88A4437935FACB2E27" baseAuthorityName="Steindachner" baseAuthorityYear="1902" box="[1148,1331,830,856]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="B940EACDFFADFF88A4437935FACB2E27" box="[1148,1331,830,856]" italics="true" pageId="7" pageNumber="308">Scarus arabicus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA54B2EFFADFF88A57A7935FEC52E03" author="Randall, J. E. &amp; Bruce, R. W." pageId="7" pageNumber="308" pagination="1 - 39" refId="ref15077" refString="Randall, J. E. &amp; Bruce, R. W. (1983) The parrotfishes of the subfamily Scarinae of the western Indian Ocean with descriptions of three new species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 47, 1 - 39." type="journal article" year="1983">Randall &amp; Bruce 1983</bibRefCitation>
; Randall 1995), supporting the record of
<bibRefCitation id="EFA54B2EFFADFF88A3267969FBF92E03" author="Zajonz, U. &amp; Lavergne, E. &amp; Bogorodsky, S. V. &amp; Saeed, F. N. &amp; Aideed, M. S. &amp; Krupp, F." box="[793,1025,866,892]" pageId="7" pageNumber="308" pagination="1 - 108" refId="ref16035" refString="Zajonz, U., Lavergne, E., Bogorodsky, S. V., Saeed, F. N., Aideed, M. S. &amp; Krupp, F. (2019) Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa, 4636 (1), 1 - 108. https: // doi. org / 10.11646 / zootaxa. 4636.1.1" type="journal article" year="2019">
Zajonz
<emphasis id="B940EACDFFADFF88A34F7968FC5B2E03" box="[880,931,866,892]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. (2019)
</bibRefCitation>
. Individuals on
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were observed on rocky substrata at depths of
<quantity id="4CCC9B3AFFADFF88A260798DFD512EDF" box="[607,681,902,929]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="7" pageNumber="308" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">
5
<specimenCount id="9D32FD56FFADFF88A243798DFD512EDF" box="[636,681,902,929]" count="8" pageId="7" pageNumber="308" type="generic">8 m</specimenCount>
</quantity>
. Data on habitat and depth of collected specimens are not available. Previously known from
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(
<bibRefCitation id="EFA54B2EFFADFF88A26079A1FCCE2EBB" author="Zajonz, U. &amp; Lavergne, E. &amp; Bogorodsky, S. V. &amp; Saeed, F. N. &amp; Aideed, M. S. &amp; Krupp, F." box="[607,822,938,964]" pageId="7" pageNumber="308" pagination="1 - 108" refId="ref16035" refString="Zajonz, U., Lavergne, E., Bogorodsky, S. V., Saeed, F. N., Aideed, M. S. &amp; Krupp, F. (2019) Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa, 4636 (1), 1 - 108. https: // doi. org / 10.11646 / zootaxa. 4636.1.1" type="journal article" year="2019">
Zajonz
<emphasis id="B940EACDFFADFF88A28879A0FD122EBB" box="[695,746,938,964]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2019
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), the distribution range is extended to the west by the observation of the initial phase and a terminal male in a mixed coral and rock reef area of south-eastern Abd al-Kuri
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.
</paragraph>
</subSubSection>
</treatment>
</document>